358 lines
42 KiB
XML
358 lines
42 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E781FFFDFFF013FD2AE5FE815" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Colossobolus semicyclus Wesener & Enghoff & Sierwald 2009, sp. n." docType="treatment" docUuid="8205B095-B802-46B7-8BA7-BEF078E551A8" docUuidSource="ZooBank" docVersion="11" lastPageNumber="26" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="25" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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||
</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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||
<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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||
</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791395" ID-GBIF-Taxon="163747654" ID-Zenodo-Dep="3791395" LSID="urn:lsid:zoobank.org:act:8205B095-B802-46B7-8BA7-BEF078E551A8" httpUri="http://treatment.plazi.org/id/1772122E781FFFDFFF013FD2AE5FE815" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<subSubSection box="[140,603,166,193]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph blockId="24.[140,956,166,229]" box="[140,603,166,193]" pageId="24" pageNumber="25">
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<heading box="[140,603,166,193]" fontSize="11" level="2" pageId="24" pageNumber="25" reason="3">
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<emphasis bold="true" box="[140,603,166,193]" pageId="24" pageNumber="25">
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<taxonomicName authority="Wesener" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,526,166,193]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" status="sp. n.">
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<emphasis bold="true" box="[140,418,166,193]" italics="true" pageId="24" pageNumber="25">Colossobolus semicyclus</emphasis>
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Wesener
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</taxonomicName>
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,
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<taxonomicNameLabel box="[538,603,167,193]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,956,202,229]" pageId="24" pageNumber="25" type="description">
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<paragraph blockId="24.[140,956,166,229]" box="[140,956,202,229]" pageId="24" pageNumber="25">
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<uri box="[140,956,202,229]" pageId="24" pageNumber="25">
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urn:lsid:zoobank.org:act:
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<uuid box="[420,956,202,229]" pageId="24" pageNumber="25">8205B095-B802-46B7-8BA7-BEF078E551A8</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="materials_examined">
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<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
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<emphasis bold="true" box="[140,375,272,298]" pageId="24" pageNumber="25">
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<emphasis box="[140,241,272,298]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Material</emphasis>
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examined:
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608818300" collectingDate="2003-12-12" collectionCode="CAS, BLF" collectorName="B. L. Fisher" country="Madagascar" elevation="550" latitude="-13.211666" location="Foret d'Antsahabe" longLatPrecision="21" longitude="49.556667" municipality="Province" pageId="24" pageNumber="25" specimenCount="3" specimenCount-female="2" stateProvince="Antsiranana" typeStatus="holotype">
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2 ♂,
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<specimenCount box="[453,500,272,299]" pageId="24" pageNumber="25" type="female">2 ♀</specimenCount>
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,
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<specimenCount box="[519,607,272,298]" pageId="24" pageNumber="25" type="generic">1 imm.</specimenCount>
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<emphasis box="[618,721,272,298]" italics="true" pageId="24" pageNumber="25">
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<typeStatus box="[618,715,272,298]" pageId="24" pageNumber="25">Holotype</typeStatus>
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:
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</emphasis>
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1 ♂ (
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<quantity box="[801,903,272,299]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="mm" value="120.0">120 mm</quantity>
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long),
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<collectionCode box="[993,1046,272,298]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/BLF10114" pageId="24" pageNumber="25">
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<collectionCode box="[1058,1108,273,299]" pageId="24" pageNumber="25">BLF</collectionCode>
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10114
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</accessionNumber>
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a.
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<collectingCountry box="[248,382,307,333]" name="Madagascar" pageId="24" pageNumber="25">Madagascar</collectingCountry>
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,
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<collectingMunicipality box="[401,499,307,333]" pageId="24" pageNumber="25">Province</collectingMunicipality>
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<collectingRegion box="[514,653,307,333]" country="Madagascar" name="Antsiranana" pageId="24" pageNumber="25">Antsiranana</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:1772122E781FFFDFFF013FD2AE5FE815:9A04F5E3781FFFDEFD2D3E40AC21EB1C" box="[672,888,307,334]" country="Madagascar" latitude="-13.211666" longLatPrecision="21" longitude="49.556667" municipality="Province" name="Foret d'Antsahabe" pageId="24" pageNumber="25" stateProvince="Antsiranana">Forêt d’Antsahabe</location>
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,
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<locationDeviation location="Daraina" pageId="24" pageNumber="25">
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<quantity box="[907,1011,307,334]" metricMagnitude="4" metricUnit="m" metricValue="1.14" pageId="24" pageNumber="25" unit="km" value="11.4">11.4 km</quantity>
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275°W Daraina
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</locationDeviation>
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, tropical dry forest,
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<quantity box="[470,549,342,369]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="m" value="550.0">
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<elevation box="[470,549,342,369]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="m" value="550.0">550 m</elevation>
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</quantity>
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,
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<geoCoordinate box="[563,705,343,369]" degrees="13" direction="south" minutes="12" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="42" value="-13.211666">13°12’42” S</geoCoordinate>
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,
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<geoCoordinate box="[719,879,343,369]" degrees="049" direction="east" minutes="33" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="24" value="49.556667">049°33’24” E</geoCoordinate>
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, leg.
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<collectorName box="[942,1077,343,369]" pageId="24" pageNumber="25">B. L. Fisher</collectorName>
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et al.,
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<date box="[183,329,378,404]" pageId="24" pageNumber="25" value="2003-12-12">
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<collectingDate box="[183,329,378,404]" pageId="24" pageNumber="25" value="2003-12-12">12.XII.2003</collectingDate>
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</date>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2608818213" box="[342,1103,378,404]" collectingDate="2003-12-12" collectionCode="CAS, BLF" collectorName="B. L. Fisher" country="Madagascar" elevation="550" latitude="-13.211666" location="Foret d'Antsahabe" longLatPrecision="21" longitude="49.556667" municipality="Province" pageId="24" pageNumber="25" specimenCount="2" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
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<emphasis box="[342,448,379,404]" italics="true" pageId="24" pageNumber="25">
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<typeStatus box="[342,444,379,404]" pageId="24" pageNumber="25">Paratypes</typeStatus>
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:
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</emphasis>
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<specimenCount box="[458,504,378,404]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
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,
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<specimenCount box="[521,609,378,404]" pageId="24" pageNumber="25" type="generic">1 imm.</specimenCount>
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,
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<collectionCode box="[623,676,378,404]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
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<accessionNumber box="[686,820,378,404]" httpUri="http://www.ncbi.nlm.nih.gov/protein/BLF10114" pageId="24" pageNumber="25">
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<collectionCode box="[686,736,378,404]" pageId="24" pageNumber="25">BLF</collectionCode>
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10114
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</accessionNumber>
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a, same data as holotype
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2608818216" collectingDate="2001-11-03" collectingDateMax="2001-11-10" collectingDateMin="2001-11-03" collectionCode="FMMC" collectorName="S. Goodman" country="Madagascar" elevation="463" latitude="-13.255" location="Analamozava River" longLatPrecision="129" longitude="49.616665" municipality="Province" pageId="24" pageNumber="25" specimenCode="FMMC 5488" specimenCount="1" specimenCount-female="1" stateProvince="Antsiranana" typeStatus="paratype">
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1 ♂,
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<specimenCount box="[199,243,413,440]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
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,
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<specimenCode box="[258,418,413,440]" collectionCode="FMMC" pageId="24" pageNumber="25">FMMC 5488</specimenCode>
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,
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<collectingMunicipality box="[430,528,413,439]" pageId="24" pageNumber="25">Province</collectingMunicipality>
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<collectingRegion box="[536,675,413,439]" country="Madagascar" name="Antsiranana" pageId="24" pageNumber="25">Antsiranana</collectingRegion>
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, near
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<location LSID="urn:lsid:plazi:treatment:1772122E781FFFDFFF013FD2AE5FE815:9A04F5E3781FFFDEFD653EE9AC9AEBE6" box="[744,963,413,439]" country="Madagascar" latitude="-13.255" longLatPrecision="129" longitude="49.616665" municipality="Province" name="Analamozava River" pageId="24" pageNumber="25" stateProvince="Antsiranana">Analamozava River</location>
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,
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<locationDeviation location="Daraina" pageId="24" pageNumber="25">
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<quantity box="[975,1057,413,440]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="24" pageNumber="25" unit="km" value="7.5">7.5 km</quantity>
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SW Daraina
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</locationDeviation>
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, undisturbed humid lowland forest,
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<quantity box="[650,786,448,475]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="6.0" metricValueMin="3.25" pageId="24" pageNumber="25" unit="m" value="462.5" valueMax="600.0" valueMin="325.0">
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<elevation box="[650,786,448,475]" metricMagnitude="2" metricUnit="m" metricValue="4.625" metricValueMax="6.0" metricValueMin="3.25" pageId="24" pageNumber="25" unit="m" value="462.5" valueMax="600.0" valueMin="325.0">325–600 m</elevation>
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</quantity>
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,
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<geoCoordinate box="[798,919,448,475]" degrees="13" direction="south" minutes="15.3" orientation="latitude" pageId="24" pageNumber="25" precision="92" value="-13.255">13°15.3’ S</geoCoordinate>
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,
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<geoCoordinate box="[932,1056,448,475]" degrees="49" direction="east" minutes="37.0" orientation="longitude" pageId="24" pageNumber="25" precision="92" value="49.616665">49°37.0’ E</geoCoordinate>
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, leg.
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<collectorName box="[140,288,483,510]" pageId="24" pageNumber="25">S. Goodman</collectorName>
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,
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<date box="[299,463,484,510]" pageId="24" pageNumber="25" value="2001-11-03" valueMax="2001-11-10" valueMin="2001-11-03">
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<collectingDate box="[299,463,484,510]" pageId="24" pageNumber="25" value="2001-11-03" valueMax="2001-11-10" valueMin="2001-11-03">3–10.XI.2001</collectingDate>
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</date>
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,
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<collectingMethod box="[474,597,483,510]" pageId="24" pageNumber="25">pitfall trap</collectingMethod>
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</materialsCitation>
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.
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</paragraph>
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<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
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<emphasis bold="true" box="[185,493,518,544]" pageId="24" pageNumber="25">
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<emphasis box="[185,257,518,544]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Other</emphasis>
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material examined:
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</emphasis>
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2 ♂,
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<specimenCount box="[559,603,519,545]" pageId="24" pageNumber="25" type="female">1 ♀</specimenCount>
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||
,
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<collectionCode box="[618,671,519,545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
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<accessionNumber box="[678,811,519,545]" httpUri="http://www.ncbi.nlm.nih.gov/protein/BLF10877" pageId="24" pageNumber="25">
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||
<collectionCode box="[678,728,519,545]" pageId="24" pageNumber="25">BLF</collectionCode>
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||
10877
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||
</accessionNumber>
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a, Forêt de Binara,
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<quantity box="[1026,1107,519,545]" metricMagnitude="3" metricUnit="m" metricValue="9.1" pageId="24" pageNumber="25" unit="km" value="9.1">9.1 km</quantity>
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SW Daraina, rainforest,
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||
<quantity box="[426,565,554,581]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="8.0" metricValueMin="6.5" pageId="24" pageNumber="25" unit="m" value="725.0" valueMax="800.0" valueMin="650.0">650–800 m</quantity>
|
||
,
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||
<geoCoordinate box="[580,723,554,581]" degrees="13" direction="south" minutes="15" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="48" value="-13.263333">13°15’48” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[739,899,554,580]" degrees="049" direction="east" minutes="36" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="12" value="49.603333">049°36’12” E</geoCoordinate>
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||
, leg. B. L. Fisher,
|
||
<date box="[140,275,589,615]" pageId="24" pageNumber="25" value="2004-11-19">19.XI.2004</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[185,455,624,650]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,324,624,650]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Differential</emphasis>
|
||
diagnosis:
|
||
</emphasis>
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[467,718,624,650]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[467,718,624,650]" italics="true" pageId="24" pageNumber="25">Colossobolus semicyclus</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel box="[730,795,625,651]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
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cannot be separated from other sympatric
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||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[327,459,659,685]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[327,459,659,685]" italics="true" pageId="24" pageNumber="25">Colossobolus</emphasis>
|
||
</taxonomicName>
|
||
species by external features alone (
|
||
<figureCitation box="[859,943,659,686]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="24" pageNumber="25">Fig. 10</figureCitation>
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||
). Differs from all other species in the unique shape of the posterior gonopod telopodite (
|
||
<figureCitation box="[988,1057,695,721]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9</figureCitation>
|
||
F–I) and in the presence of a well-rounded coxite process on the anterior gonopod. Shares with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[197,372,765,792]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[197,372,765,792]" italics="true" pageId="24" pageNumber="25">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[378,438,766,792]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[493,625,766,792]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[493,625,766,792]" italics="true" pageId="24" pageNumber="25">C. giganteus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[632,692,766,792]" pageId="24" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
|
||
a finger-shaped coxite process on the posterior gonopods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||
<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[185,335,835,861]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,330,835,861]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[345,497,836,861]" italics="true" pageId="24" pageNumber="25">Measurements</emphasis>
|
||
: males with 49–52 body rings, circa
|
||
<quantity box="[939,1039,836,862]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="mm" value="120.0">120 mm</quantity>
|
||
long,
|
||
<quantity box="[140,298,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.14" metricValueMin="0.97" pageId="24" pageNumber="25" unit="mm" value="10.55" valueMax="11.4" valueMin="9.7">9.7–11.4 mm</quantity>
|
||
wide. Females with 49–52 body rings,
|
||
<quantity box="[756,914,870,897]" metricMagnitude="-1" metricUnit="m" metricValue="1.125" metricValueMax="1.25" metricValueMin="1.0" pageId="24" pageNumber="25" unit="mm" value="112.5" valueMax="125.0" valueMin="100.0">100–125 mm</quantity>
|
||
long, 10.5–11.0 mm wide.
|
||
</paragraph>
|
||
<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,303,941,967]" italics="true" pageId="24" pageNumber="25">Coloration</emphasis>
|
||
affected by alcohol. Head, legs, antennae and telson red. Meso- and metazonites of body rings reddish-brown, posterior margin with a thin, dark brown line (
|
||
<figureCitation box="[198,269,1011,1038]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9</figureCitation>
|
||
A–C).
|
||
<emphasis box="[349,454,1012,1038]" italics="true" pageId="24" pageNumber="25">Antennae</emphasis>
|
||
protruding back to ring 3 (
|
||
<figureCitation box="[776,868,1011,1038]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9A</figureCitation>
|
||
).
|
||
<emphasis box="[888,1107,1011,1037]" italics="true" pageId="24" pageNumber="25">Male coxal processes</emphasis>
|
||
present on coxae 3 and 4, well-developed. Coxae 5–7 with visible, but short processes (
|
||
<figureCitation box="[209,299,1082,1108]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[140,1108,272,1672]" box="[185,797,1117,1144]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,346,1117,1143]" italics="true" pageId="24" pageNumber="25">Preanal process</emphasis>
|
||
well-rounded, not projecting (
|
||
<figureCitation box="[694,783,1117,1143]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,373,1152,1179]" italics="true" pageId="24" pageNumber="25">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, broadly rounded lobe (
|
||
<figureCitation box="[1000,1094,1152,1178]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9D</figureCitation>
|
||
). Mesal process of coxite weakly developed, protruding into a short wide lobe which slightly extends beyond sternite (
|
||
<figureCitation box="[513,606,1223,1249]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9D</figureCitation>
|
||
). Telopodite on posterior side basally with a circular groove (
|
||
<figureCitation box="[322,409,1258,1284]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9E</figureCitation>
|
||
). Telopodite process long and relatively slender. Mesal margin laterally sharp-edged, but not projecting (
|
||
<figureCitation box="[609,697,1293,1319]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Fig. 9E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[140,1108,272,1672]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,383,1328,1354]" italics="true" pageId="24" pageNumber="25">Posterior gonopods</emphasis>
|
||
unique (
|
||
<figureCitation box="[482,549,1328,1354]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9</figureCitation>
|
||
F–I). Telopodites positioned face-to face with one another. Coxite branch relatively short, wide and stout (
|
||
<figureCitation box="[777,846,1363,1390]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9</figureCitation>
|
||
F–I). Lateral branch of telopodite curved, forming a ‘C’ together with coxite branch. Central membranous area protruding (
|
||
<figureCitation box="[332,456,1434,1460]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9F, H</figureCitation>
|
||
). Mesal main branch well-developed, apical part overlapping lateral branch (
|
||
<figureCitation box="[381,507,1469,1496]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9G, I</figureCitation>
|
||
). Mesal margin of main branch with two overlapping membranous folds. Membranous folds large, well-rounded, forming a half-circle (
|
||
<figureCitation box="[147,266,1539,1566]" captionStart="Figure 9" captionStartId="17.[140,212,1462,1484]" captionTargetBox="[140,1108,165,1395]" captionTargetId="figure@17.[140,1108,732,1395]" captionTargetPageId="17" captionText="Figure 9. Colossobolus semicyclus sp. n., male holotype. A head B legs 1–7, ventral view C telson D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view H posterior gonopod, anterior view I posterior gonopod, posterior view. av = anal valves; Co = collum; Cx = coxite; Cx-br = coxite branch; Gn = gnathochilarium; ma = membranous area; Md = mandible; ms = marginal setae; Pre = preanal ring; sf = setiferous foveolae; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767294" httpUri="https://zenodo.org/record/3767294/files/figure.png" pageId="24" pageNumber="25">Figs 9G, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[140,1108,272,1672]" lastBlockId="25.[140,1108,167,580]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[185,461,1574,1600]" pageId="24" pageNumber="25">
|
||
<emphasis box="[185,333,1574,1600]" inLineHeading="true" pageId="24" pageNumber="25" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
animals from the
|
||
<typeStatus box="[692,740,1575,1601]" pageId="24" pageNumber="25">type</typeStatus>
|
||
series (Forêt Antsahabe) differ greatly in size from those collected only some kilometers away in an area close to the Analamozava River. Specimens from the
|
||
<typeStatus box="[616,664,1646,1672]" pageId="24" pageNumber="25">type</typeStatus>
|
||
series have 52 body rings while those from the Analamozava River have only 49. Specimens from Antsahabe are approximately
|
||
<quantity box="[226,312,202,228]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="10.0">10 mm</quantity>
|
||
longer but
|
||
<quantity box="[454,525,202,228]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="25" pageNumber="26" unit="mm" value="1.0">1 mm</quantity>
|
||
slimmer than those from the Analamozava River. Males from both localities show identical gonopods. Those differences can probably be accounted for by the fact that the habitat on the Antsahabe site is much drier than the Analamozava River site. However, more specimens from both sites are necessary to further refine this intraspecific variation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="26" type="distribution">
|
||
<paragraph blockId="25.[140,1108,167,580]" pageId="25" pageNumber="26">
|
||
<emphasis bold="true" box="[185,494,377,404]" pageId="25" pageNumber="26">
|
||
<emphasis box="[185,336,377,403]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Distribution</emphasis>
|
||
and ecology
|
||
</emphasis>
|
||
: this species has hitherto been recorded only in dry and subhumid forests at mid-altitudes (
|
||
<quantity box="[606,745,413,440]" metricMagnitude="2" metricUnit="m" metricValue="5.625" metricValueMax="8.0" metricValueMin="3.25" pageId="25" pageNumber="26" unit="m" value="562.5" valueMax="800.0" valueMin="325.0">325–800 m</quantity>
|
||
) around Daraina (
|
||
<figureCitation box="[968,1056,413,439]" captionStart="Figure 10" captionStartId="18.[140,212,1654,1676]" captionTargetBox="[160,1106,1078,1634]" captionTargetId="figure@18.[140,1109,1071,1635]" captionTargetPageId="18" captionText="Figure 10. Distribution of the genus Colossobolus gen. n." figureDoi="http://doi.org/10.5281/zenodo.3820794" httpUri="https://zenodo.org/record/3820794/files/figure.png" pageId="25" pageNumber="26">Fig. 10</figureCitation>
|
||
).
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis italics="true" pageId="25" pageNumber="26">C. semicyclus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[256,318,449,475]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
occurs at Antsahabe sympatrically with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[779,956,448,475]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[779,956,448,475]" italics="true" pageId="25" pageNumber="26">C. oblongopedus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[964,1028,449,475]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
, at the Analamozava river site sympatrically with
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[615,746,483,510]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" status="sp. n.">
|
||
<emphasis box="[615,746,483,510]" italics="true" pageId="25" pageNumber="26">C. aculeatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[754,815,484,510]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph blockId="25.[140,1108,167,580]" pageId="25" pageNumber="26">
|
||
<emphasis bold="true" box="[185,323,519,545]" pageId="25" pageNumber="26">
|
||
<emphasis box="[185,318,519,545]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Etymology</emphasis>
|
||
:
|
||
</emphasis>
|
||
<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[330,437,519,545]" class="Diplopoda" family="Pachybolidae" genus="Colossobolus" kingdom="Animalia" order="Spirobolida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[330,437,519,545]" italics="true" pageId="25" pageNumber="26">semicyclus</emphasis>
|
||
</taxonomicName>
|
||
, adjective, refers to the uniquely shaped posterior gonopod telopodite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |