335 lines
46 KiB
XML
335 lines
46 KiB
XML
<document id="AD80B4BE364CB43C7A585B51A46FF9F7" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. & Herbert, D. G." docDate="2013" docId="03BC1F3CE53ED90D07A5FF5D47E6B325" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Ampelita beanka Griffiths & Herbert 2013, sp. n." docType="treatment" docUuid="F9496435-F4B7-45C6-8B3C-25D8895AC9B4" docUuidSource="ZooBank" docVersion="8" lastPageNumber="26" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="23" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B7D2A6D96001075544776B7F2410DDD1">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:name id="601E5152EEEC0000D4E2061A2B5AB394" type="personal">
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<mods:roleTerm id="C209F433B293567C097A73AFB6ED1EFA">Author</mods:roleTerm>
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<mods:namePart id="5B2E0206DB70000E7790CF3AE1535839">Griffiths, O. L.</mods:namePart>
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<mods:affiliation id="A6FC434E7A7B17BD33972D54DBFBA067">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
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<mods:nameIdentifier id="6B1D9E03113D616BD492AD176C3B3EED" type="email">owen@bcm.intnet.mu</mods:nameIdentifier>
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<mods:namePart id="6AD6AB4580DD8578B8248529C173D664">Herbert, D. G.</mods:namePart>
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<mods:affiliation id="7340BC9C3369A1804A26A7F58080B5AD">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
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<mods:nameIdentifier id="9F95898183BF5403396CFCC60D1C5BD2" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
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<mods:title id="159DA8D08A0710DD10115000C6FFE14E">African Invertebrates</mods:title>
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<mods:part id="2741AE04681129644EC268C495074F2A">
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<mods:date id="689E086E034B894CE1E8812D9458B9DA">2013</mods:date>
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<mods:number id="04E5C25EBDCECD0C2D27B34D76370A38">2013-02-13</mods:number>
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<mods:number id="9CD21DD3EF847F311A1BC294EE1CB819">54</mods:number>
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<treatment id="03BC1F3CE53ED90D07A5FF5D47E6B325" ID-DOI="http://doi.org/10.5281/zenodo.7670267" ID-GBIF-Taxon="209341533" ID-Zenodo-Dep="7670267" LSID="urn:lsid:zoobank.org:act:F9496435-F4B7-45C6-8B3C-25D8895AC9B4" httpUri="http://treatment.plazi.org/id/03BC1F3CE53ED90D07A5FF5D47E6B325" lastPageId="26" lastPageNumber="26" pageId="23" pageNumber="23">
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<subSubSection id="C30FFDA1E53ED90007A5FF5D44D6B7A5" box="[488,759,203,230]" pageId="23" pageNumber="23" type="nomenclature">
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<paragraph id="8BAAAE2AE53ED90007A5FF5D44D6B7A5" blockId="23.[144,1105,203,1038]" box="[488,759,203,230]" pageId="23" pageNumber="23">
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<heading id="D0E21946E53ED90007A5FF5D44D6B7A5" box="[488,759,203,230]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="23" reason="2">
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<taxonomicName id="4C15D5A9E53ED90007A5FF5D4495B7A5" ID-CoL="CY75" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[488,692,203,230]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="23" phylum="Mollusca" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9617238E53ED90007A5FF5D4495B7A5" bold="true" box="[488,692,203,230]" pageId="23" pageNumber="23">Ampelita beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252CF43E53ED90004F6FF5A44D6B7A5" box="[699,759,204,230]" pageId="23" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FFDA1E53ED90D07B9FF6547E6B325" lastPageId="26" lastPageNumber="26" pageId="23" pageNumber="23" type="description">
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<paragraph id="8BAAAE2AE53ED90007B9FF6544CDB64E" blockId="23.[144,1105,203,1038]" box="[500,748,243,269]" pageId="23" pageNumber="23">
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<figureCitation id="132EB2AFE53ED90007B9FF654468B64E" box="[500,585,243,269]" captionStart="Fig" captionStartId="23.[144,177,1558,1579]" captionTargetBox="[151,1096,1100,1536]" captionTargetId="figure-274@23.[143,1103,1094,1539]" captionTargetPageId="23" captionText="Fig. 15. Ampelita beanka sp. n.: (A–C) holotype, max. diameter 17.7 mm, height 8.0 mm,AMS C.474166; (D) specimen with less strongly shouldered whorls, max. diameter 19.1 mm, st’n 07/09, paratype, NMSA L8501/T2981; (E–F) specimens of the same size and collected from the same station R01/09, showing the differing extentto which the suture descends below the shoulder, max.diameter 17.1mm, paratypes, NMSA L8507/T2982." figureDoi="http://doi.org/10.5281/zenodo.7670134" httpUri="https://zenodo.org/record/7670134/files/figure.png" pageId="23" pageNumber="23">Figs 15</figureCitation>
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,
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<figureCitation id="132EB2AFE53ED9000418FF654452B64E" box="[597,627,243,269]" captionStart="Fig" captionStartId="24.[181,214,837,858]" captionTargetBox="[185,1136,209,816]" captionTargetId="figure-254@24.[181,1141,204,834]" captionTargetPageId="24" captionText="Fig. 16. Ampelita beanka sp. n., genitalia, paratype, NMSA L8504/T2983: (A) entire reproductive tract; (B) external view of penial complex; (C) penis opened longitudinally to show internal folds and raised papilla at insertion of epiphallus. Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, pp – penis papilla, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670138" httpUri="https://zenodo.org/record/7670138/files/figure.png" pageId="23" pageNumber="23">16</figureCitation>
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,
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<figureCitation id="132EB2AFE53ED9000432FF6544CDB64E" box="[639,748,243,269]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="23" pageNumber="23">17A, 17B</figureCitation>
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</paragraph>
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<paragraph id="8BAAAE2AE53ED90006DDFE8D453DB676" blockId="23.[144,1105,203,1038]" box="[144,796,283,310]" pageId="23" pageNumber="23">
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Etymology: Named after the
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<typeStatus id="54AE1088E53ED900079BFE8A4427B675" box="[470,518,284,310]" pageId="23" pageNumber="23">type</typeStatus>
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locality,
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<taxonomicName id="4C15D5A9E53ED9000420FE8D4539B676" box="[621,792,283,309]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="23" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BAAAE2AE53ED90006DDFED545D6B6DE" blockId="23.[144,1105,203,1038]" pageId="23" pageNumber="23">Diagnosis: Profile distinctive due to low spire, strongly shouldered whorls and descendant suture; umbilicus wide; protoconch with close-set axial riblets; teleoconch with close-set lamellate axial pliculae; lustreless, more or less uniformly mid to dark brown.</paragraph>
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<paragraph id="8BAAAE2AE53ED90006DDFE3D473AB686" blockId="23.[144,1105,203,1038]" box="[144,283,427,453]" pageId="23" pageNumber="23">Description:</paragraph>
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<paragraph id="8BAAAE2AE53ED90006DDFE40442AB4CD" blockId="23.[144,1105,203,1038]" pageId="23" pageNumber="23">
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<emphasis id="B9617238E53ED90006DDFE4046E9B6AD" box="[144,200,470,494]" italics="true" pageId="23" pageNumber="23">Shell</emphasis>
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: Relatively small with sharply angled periphery situated well above mid-whorl; spire very low, frequently almost flat; apical surface of whorls weakly convex, becoming slightly concave toward periphery; suture shallowly indented, but progressively descending below peripheral keel on last adult whorl giving shell a stepped profile; umbilicus wide, its rim roundly angular. Protoconch of 1¼–1½ whorls, max. diameter variable
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<quantity id="4CED03CFE53ED90006DDFDE54738B5CD" box="[144,281,627,654]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="23" pageNumber="23" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 mm</quantity>
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; initially smooth, rather flat and somewhat sunken, thereafter more convex, bearing numerous, close-set, curved, axial riblets; riblets occasionally bifurcating or anastomosing (
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<figureCitation id="132EB2AFE53ED9000777FD254785B58E" box="[314,420,691,717]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="23" pageNumber="23">Fig. 17A</figureCitation>
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). Teleoconch of a further 2–2¼ whorls; sculptured by closeset, axial pliculae, each with a thin periostracal crest in living specimens, rendering the pliculae somewhat lamellate (
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<figureCitation id="132EB2AFE53ED9000796FD654461B44E" box="[475,576,755,781]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="23" pageNumber="23">Fig. 17B</figureCitation>
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); growth irregularities frequent; basal sculpture the same, continuing into umbilicus. Aperture rounded basally, somewhat angled at periphery and flattened above this; strongly oblique to vertical axis of shell; peristome incomplete, interrupted in parietal region; rim of peristome slightly thickened and flaring in adult, more strongly so basally.
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</paragraph>
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<paragraph id="8BAAAE2AE53ED90006E7FC02446AB34D" blockId="23.[144,1105,203,1038]" pageId="23" pageNumber="23">Shell lustreless with a well-developed, predominantly mid to dark brown periostracum when fresh; slight radial unevenness in intensity; apical whorls often slightly paler; umbilicus commonly darker internally, its margin sometimes delineated by a paler band, but this rarely evident in fresh material.</paragraph>
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<caption id="DF6AFEA2E53ED90006DDF98047DAB1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670134" ID-Zenodo-Dep="7670134" httpUri="https://zenodo.org/record/7670134/files/figure.png" pageId="23" pageNumber="23" startId="23.[144,177,1558,1579]" targetBox="[151,1096,1100,1536]" targetPageId="23">
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<paragraph id="8BAAAE2AE53ED90006DDF98047DAB1C8" blockId="23.[144,1105,1558,1675]" pageId="23" pageNumber="23">
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Fig. 15.
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<taxonomicName id="4C15D5A9E53ED9000697F98E4753B168" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[218,370,1560,1579]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="23" phylum="Mollusca" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9617238E53ED9000697F98E4753B168" box="[218,370,1560,1579]" italics="true" pageId="23" pageNumber="23">Ampelita beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252CF43E53ED9000735F980478BB168" box="[376,426,1558,1579]" pageId="23" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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: (A–C) holotype, max. diameter 17.7 mm, height 8.0 mm, AMS C.474166; (D) specimen with less strongly shouldered whorls, max. diameter 19.1 mm, st’n 07/09, paratype, NMSA L8501/T2981; (E–F) specimens of the same size and collected from the same station R01/09, showing the differing extent to which the suture descends below the shoulder, max. diameter 17.1 mm, paratypes, NMSA L8507/T2982.
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</paragraph>
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</caption>
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<caption id="DF6AFEA2E531D90F06F8FCD34425B491" ID-DOI="http://doi.org/10.5281/zenodo.7670138" ID-Zenodo-Dep="7670138" httpUri="https://zenodo.org/record/7670138/files/figure.png" pageId="24" pageNumber="24" startId="24.[181,214,837,858]" targetBox="[185,1136,209,816]" targetPageId="24">
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<paragraph id="8BAAAE2AE531D90F06F8FCD34425B491" blockId="24.[181,1142,837,978]" pageId="24" pageNumber="24">
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Fig. 16.
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<taxonomicName id="4C15D5A9E531D90F0749FCD147BFB419" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[260,414,839,858]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="24" pageNumber="24" phylum="Mollusca" rank="species" species="beanka" status="sp. nov.">
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<emphasis id="B9617238E531D90F0749FCD147BFB419" box="[260,414,839,858]" italics="true" pageId="24" pageNumber="24">Ampelita beanka</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252CF43E531D90F07EBFCD347F8B419" box="[422,473,837,858]" pageId="24" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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||
, genitalia, paratype, NMSA L8504/T2983: (A) entire reproductive tract; (B) external view of penial complex; (C) penis opened longitudinally to show internal folds and raised papilla at insertion of epiphallus. Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, pp – penis papilla, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens.
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</paragraph>
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</caption>
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<paragraph id="8BAAAE2AE531D90F069DFB9547E9B37D" blockId="24.[181,1142,1027,1678]" pageId="24" pageNumber="24">
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Dimensions:
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<typeStatus id="54AE1088E531D90F072BFB9547F0B35E" box="[358,465,1027,1053]" pageId="24" pageNumber="24" type="holotype">Holotype</typeStatus>
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, max. diameter
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<quantity id="4CED03CFE531D90F04C5FB9544CEB35D" box="[648,751,1027,1054]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="24" pageNumber="24" unit="mm" value="17.7">17.7 mm</quantity>
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, height 8.0 mm; largest specimen, max. diameter
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<quantity id="4CED03CFE531D90F0710FBB547E5B37D" box="[349,452,1059,1086]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="24" pageNumber="24" unit="mm" value="19.1">19.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BAAAE2AE531D90F06F8FBD844ABB3A6" blockId="24.[181,1142,1027,1678]" pageId="24" pageNumber="24">
|
||
<emphasis id="B9617238E531D90F06F8FBD84754B325" box="[181,373,1102,1126]" italics="true" pageId="24" pageNumber="24">External features</emphasis>
|
||
: Data on coloration of living animals not available; in preserved specimens head-foot mostly dark brown to blackish, but with evidence of a paler longitudinal band extending down neck from each optic tentacle; skin texture uniformly granular; lining of mantle cavity heavily pigmented with black and patterned with pale circular blotches, mostly on the right (upper) side.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE531D90F06F8FB604460B1CE" blockId="24.[181,1142,1027,1678]" pageId="24" pageNumber="24">
|
||
<emphasis id="B9617238E531D90F06F8FB604743B24D" box="[181,354,1270,1294]" italics="true" pageId="24" pageNumber="24">Distal genitalia</emphasis>
|
||
(
|
||
<figureCitation id="132EB2AFE531D90F0722FB6547E0B24E" box="[367,449,1267,1293]" captionStart="Fig" captionStartId="24.[181,214,837,858]" captionTargetBox="[185,1136,209,816]" captionTargetId="figure-254@24.[181,1141,204,834]" captionTargetPageId="24" captionText="Fig. 16. Ampelita beanka sp. n., genitalia, paratype, NMSA L8504/T2983: (A) entire reproductive tract; (B) external view of penial complex; (C) penis opened longitudinally to show internal folds and raised papilla at insertion of epiphallus. Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, ga – genital atrium, hd – hermaphrodite duct, p – penis, pf – folds lining penis lumen, pp – penis papilla, prm – penial retractor muscle, sod – spermoviduct, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670138" httpUri="https://zenodo.org/record/7670138/files/figure.png" pageId="24" pageNumber="24">Fig. 16</figureCitation>
|
||
): Relatively simple and largely typical of the genus. Penis short to moderate in length (4.0–5.0 mm) and rather stout; vas deferens running beside and loosely attached to its basal portion, but becoming more closely attached near penis apex; at apex of penis vas deferens turns sharply back on itself and broadens noticeably before inserting some distance below penis apex; this latter portion, perhaps constituting an epiphallus, has a thicker, pale flesh-coloured wall and is fused to outer wall of penis. Penis retractor muscle attaches apically, enveloping recurved portion of vas deferens and penis apex. Interior of penis with a Y-shaped longitudinal fold in the centre of which is a distinct, rounded papilla at insertion of epiphallus; this primary fold bordered on each side by a further longitudinal fold; remainder of lumen largely smooth, but with evidence of microscopic papillation in apical region. Genital atrium and vagina, simple and short; bursa copulatrix duct long, basal portion somewhat broader; bursa itself of moderate size, ovate to subcircular.
|
||
</paragraph>
|
||
<caption id="DF6AFEA2E530D90E06DDFB3D4733B27B" ID-DOI="http://doi.org/10.5281/zenodo.7670142" ID-Zenodo-Dep="7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="25" pageNumber="25" startId="25.[144,177,1195,1216]" targetBox="[154,1091,206,1175]" targetPageId="25">
|
||
<paragraph id="8BAAAE2AE530D90E06DDFB3D4733B27B" blockId="25.[144,1104,1195,1336]" pageId="25" pageNumber="25">
|
||
Fig. 17.
|
||
<taxonomicName id="4C15D5A9E530D90E0696FB3B4752B383" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[219,371,1197,1216]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="beanka" status="sp. nov.">
|
||
<emphasis id="B9617238E530D90E0696FB3B4752B383" box="[219,371,1197,1216]" italics="true" pageId="25" pageNumber="25">Ampelita beanka</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252CF43E530D90E0734FB3D478AB383" box="[377,427,1195,1216]" pageId="25" pageNumber="25" rank="species">sp. n.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C15D5A9E530D90E07F9FB3B44E2B383" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[436,707,1195,1216]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E530D90E07F9FB3B4429B383" box="[436,520,1197,1216]" italics="true" pageId="25" pageNumber="25">A. milloti</emphasis>
|
||
Fischer-Piette, 1952
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E530D90E04A2FB3B426DB383" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[751,1100,1195,1216]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E530D90E04A2FB3B45AEB383" box="[751,911,1197,1216]" italics="true" pageId="25" pageNumber="25">A. namerokoensis</emphasis>
|
||
Fischer-Piette, 1952
|
||
</taxonomicName>
|
||
, protoconch morphology and teleoconch microsculpture: (A, B)
|
||
<taxonomicName id="4C15D5A9E530D90E056BFB5345A2B39B" authorityName="Griffiths & Herbert" authorityYear="2013" box="[806,899,1221,1240]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E530D90E056BFB5345A2B39B" box="[806,899,1221,1240]" italics="true" pageId="25" pageNumber="25">A. beanka</emphasis>
|
||
</taxonomicName>
|
||
, protoconch diameter 4.1 mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D)
|
||
<taxonomicName id="4C15D5A9E530D90E0743FB634743B24B" authorityName=": Fischer-Piette" authorityYear="1952" box="[270,354,1269,1288]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E530D90E0743FB634743B24B" box="[270,354,1269,1288]" italics="true" pageId="25" pageNumber="25">A. milloti</emphasis>
|
||
</taxonomicName>
|
||
, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F)
|
||
<taxonomicName id="4C15D5A9E530D90E0598FB6346DCB263" authorityName=": Fischer-Piette" authorityYear="1952" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E530D90E0598FB6346DCB263" italics="true" pageId="25" pageNumber="25">A. namerokoensis</emphasis>
|
||
</taxonomicName>
|
||
, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAAAE2AE530D90E06DDFAD84533B2D0" blockId="25.[144,1104,1358,1676]" pageId="25" pageNumber="25">
|
||
<materialsCitation id="3B7DA477E530D90E06DDFAD84533B2D0" collectingDate="2009-04" collectionCode="AMS" collectorName="Central W Madagascar & R. Randalana" country="Madagascar" elevation="380" latitude="-18.05028" location="Andohanandranogedro" longLatPrecision="1" longitude="44.53786" pageId="25" pageNumber="25" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54AE1088E530D90E06DDFAD846C9B220" box="[144,232,1358,1379]" pageId="25" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F302EEBAE530D90E06BCFAD847B7B220" box="[241,406,1358,1379]" name="Madagascar" pageId="25" pageNumber="25">MADAGASCAR</collectingCountry>
|
||
:
|
||
<collectorName id="26E0CBFCE530D90E07EDFAD84454B220" box="[416,629,1358,1379]" pageId="25" pageNumber="25">Central W Madagascar</collectorName>
|
||
,
|
||
<emphasis id="B9617238E530D90E0432FAC644B5B220" box="[639,660,1360,1379]" italics="true" pageId="25" pageNumber="25">ca</emphasis>
|
||
<locationDeviation id="9A18B708E530D90E04D1FAD84555B220" box="[668,884,1358,1379]" location="Maintirano" pageId="25" pageNumber="25">
|
||
<quantity id="4CED03CFE530D90E04D1FAD844F6B220" box="[668,727,1358,1379]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="25" unit="km" value="60.0">60 km</quantity>
|
||
E of Maintirano
|
||
</locationDeviation>
|
||
, in tall dense dry forest growing above cliffs on E side of
|
||
<taxonomicName id="4C15D5A9E530D90E07F4FAF0441FB238" box="[441,574,1382,1403]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="25" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
at
|
||
<location id="8ECAF8F1E530D90E041BFAF04509B238" LSID="urn:lsid:plazi:treatment:03BC1F3CE53ED90D07A5FF5D47E6B325:8ECAF8F1E530D90E041BFAF04509B238" box="[598,808,1382,1403]" country="Madagascar" latitude="-18.05028" longLatPrecision="1" longitude="44.53786" name="Andohanandranogedro" pageId="25" pageNumber="25">Andohanandranogedro</location>
|
||
,
|
||
<geoCoordinate id="EE21C8EDE530D90E0562FAF045BBB238" box="[815,922,1382,1403]" degrees="18.05028" direction="south" orientation="latitude" pageId="25" pageNumber="25" precision="1" value="-18.05028">18.05028°S</geoCoordinate>
|
||
<geoCoordinate id="EE21C8EDE530D90E05D3FAF0422DB238" box="[926,1036,1382,1403]" degrees="44.53786" direction="east" orientation="longitude" pageId="25" pageNumber="25" precision="1" value="44.53786">44.53786°E</geoCoordinate>
|
||
,
|
||
<quantity id="4CED03CFE530D90E0259FAF0426DB238" box="[1044,1100,1382,1403]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="25" unit="m" value="380.0">
|
||
<elevation id="00384919E530D90E0259FAF0426DB238" box="[1044,1100,1382,1403]" metricMagnitude="2" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="25" unit="m" value="380.0">380m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFAB88EAE530D90E06DDFAE846F4B2D0" box="[144,213,1406,1427]" pageId="25" pageNumber="25" value="2009-04">
|
||
<collectingDate id="EFEF7102E530D90E06DDFAE846F4B2D0" box="[144,213,1406,1427]" pageId="25" pageNumber="25" value="2009-04">iv.2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26E0CBFCE530D90E0693FAE8477AB2D0" box="[222,347,1406,1427]" pageId="25" pageNumber="25">R. Randalana</collectorName>
|
||
, st’n R04/09 (
|
||
<collectionCode id="ED0436EFE530D90E0797FAE8442DB2D0" box="[474,524,1406,1427]" pageId="25" pageNumber="25">AMS</collectionCode>
|
||
C.474166, body in alcohol).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE530D90E06DDFA084574B2B8" blockId="25.[144,1104,1358,1676]" pageId="25" pageNumber="25">
|
||
<materialsCitation id="3B7DA477E530D90E06DDFA084488B288" collectingDate="2009-04" collectionCode="NMSA" collectorName="Central W Madagascar & R. Randalana" country="Madagascar" elevation="380" latitude="-18.05028" location="Andohanandranogedro" longLatPrecision="1" longitude="44.53786" pageId="25" pageNumber="25" specimenCode="L8504, NHMUK 20120014, L8525" specimenCount="16" typeStatus="paratype">
|
||
<typeStatus id="54AE1088E530D90E06DDFA0846CBB2F0" box="[144,234,1438,1459]" pageId="25" pageNumber="25" type="paratype">Paratypes</typeStatus>
|
||
: Same data as holotype (
|
||
<collectionCode id="ED0436EFE530D90E07F1FA0847DCB2F0" box="[444,509,1438,1459]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E530D90E044CFA08441DB2F0" box="[513,572,1438,1459]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">L8504</specimenCode>
|
||
/T2983,
|
||
<specimenCount id="9D1365A3E530D90E04C8FA0844D2B2F0" box="[645,755,1438,1459]" pageId="25" pageNumber="25" type="generic">9 specimens</specimenCount>
|
||
, bodies in alcohol;
|
||
<specimenCode id="DBB30651E530D90E05D5FA08426CB2F0" box="[920,1101,1438,1459]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="25" pageNumber="25" type="Museum">NHMUK 20120014</specimenCode>
|
||
,
|
||
<specimenCount id="9D1365A3E530D90E06DDFA2046D9B288" box="[144,248,1462,1483]" pageId="25" pageNumber="25" type="generic">1 specimen</specimenCount>
|
||
); st’n 07/10 (
|
||
<collectionCode id="ED0436EFE530D90E0725FA204788B288" box="[360,425,1462,1483]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E530D90E07E0FA2047C9B288" box="[429,488,1462,1483]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">L8525</specimenCode>
|
||
/T2980,
|
||
<specimenCount id="9D1365A3E530D90E047FFA204483B288" box="[562,674,1462,1483]" pageId="25" pageNumber="25" type="generic">6 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7DA477E530D90E04FDFA2044F7B2A0" collectionCode="NMSA" country="Madagascar" pageId="25" pageNumber="25" specimenCode="L8501, L8507" specimenCount="6" typeStatus="paratype">
|
||
st’n 07/09 (
|
||
<collectionCode id="ED0436EFE530D90E055EFA204575B288" box="[787,852,1462,1483]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E530D90E0515FA2045B2B288" box="[856,915,1462,1483]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">L8501</specimenCode>
|
||
/T2981,
|
||
<specimenCount id="9D1365A3E530D90E0590FA20426DB288" box="[989,1100,1462,1483]" pageId="25" pageNumber="25" type="generic">4 specimens</specimenCount>
|
||
, one in alcohol); st’n R01/09 (
|
||
<collectionCode id="ED0436EFE530D90E07DFFA5847F5B2A0" box="[402,468,1486,1507]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E530D90E0795FA584432B2A0" box="[472,531,1486,1507]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="25" pageNumber="25" type="Museum">L8507</specimenCode>
|
||
/T2982,
|
||
<specimenCount id="9D1365A3E530D90E0413FA5844F1B2A0" box="[606,720,1486,1507]" pageId="25" pageNumber="25" type="generic">2 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7DA477E530D90E0493FA584570B2B8" collectionCode="AMS, TMAM, MNHN" country="Madagascar" pageId="25" pageNumber="25" specimenCode="T165" specimenCount="6" typeStatus="paratype">
|
||
st’n 01/10 (
|
||
<collectionCode id="ED0436EFE530D90E050FFA584555B2A0" box="[834,884,1486,1507]" pageId="25" pageNumber="25">AMS</collectionCode>
|
||
C.469583,
|
||
<specimenCount id="9D1365A3E530D90E0596FA58426DB2A0" box="[987,1100,1486,1507]" pageId="25" pageNumber="25" type="generic">4 specimens</specimenCount>
|
||
;
|
||
<collectionCode id="ED0436EFE530D90E06DDFA7046F6B2B8" box="[144,215,1510,1531]" pageId="25" pageNumber="25">TMAM</collectionCode>
|
||
<specimenCode id="DBB30651E530D90E0696FA70472CB2B8" box="[219,269,1510,1531]" collectionCode="TMAM" pageId="25" pageNumber="25">T165</specimenCode>
|
||
,
|
||
<specimenCount id="9D1365A3E530D90E075AFA7047A3B2B8" box="[279,386,1510,1531]" pageId="25" pageNumber="25" type="generic">1 specimen</specimenCount>
|
||
); st’n 09/09 (
|
||
<collectionCode id="ED0436EFE530D90E07B5FA70441EB2B8" box="[504,575,1510,1531]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="25">MNHN</collectionCode>
|
||
IM-2010-20071,
|
||
<specimenCount id="9D1365A3E530D90E04ADFA70456AB2B8" box="[736,843,1510,1531]" pageId="25" pageNumber="25" type="generic">1 specimen</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE530D90E06DDF99044C8B158" blockId="25.[144,1104,1358,1676]" box="[144,745,1542,1563]" pageId="25" pageNumber="25">
|
||
Additional locality data:
|
||
<taxonomicName id="4C15D5A9E530D90E073FF99E47D7B158" box="[370,502,1544,1563]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="25" pageNumber="25" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E530D90E073FF99E47D7B158" box="[370,502,1544,1563]" italics="true" pageId="25" pageNumber="25">Tsingy Beanka</emphasis>
|
||
</taxonomicName>
|
||
: st’ns 11/06, 09/09, 01/10.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE530D90E06DDF9BC421AB107" blockId="25.[144,1104,1358,1676]" box="[144,1083,1578,1605]" pageId="25" pageNumber="25">
|
||
Distribution: A narrow-range endemic currently recorded only from
|
||
<taxonomicName id="4C15D5A9E530D90E05C1F9BC4216B107" box="[908,1079,1578,1604]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="25" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE530D90E06DDF9C44271B1CF" blockId="25.[144,1104,1358,1676]" pageId="25" pageNumber="25">Habitat. Fresh dead shells found most commonly in and amongst limestone rocks predominantly in the taller moister forests. Living individuals usually found in leaf-litter.</paragraph>
|
||
<paragraph id="8BAAAE2AE533D90D06F8FF5D47B4B6E6" blockId="26.[181,1142,203,1126]" pageId="26" pageNumber="26">
|
||
Remarks: In terms of its shape and coloration,
|
||
<taxonomicName id="4C15D5A9E533D90D0487FF58451AB7A5" box="[714,827,206,230]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E533D90D0487FF58451AB7A5" box="[714,827,206,230]" italics="true" pageId="26" pageNumber="26">A. beanka</emphasis>
|
||
</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="4C15D5A9E533D90D05F6FF584204B7A5" authorityName=": Fischer-Piette" authorityYear="1952" box="[955,1061,206,230]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E533D90D05F6FF584204B7A5" box="[955,1061,206,230]" italics="true" pageId="26" pageNumber="26">A. milloti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E533D90D0210FF58477DB645" authorityName=": Fischer-Piette" authorityYear="1952" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E533D90D0210FF58477DB645" italics="true" pageId="26" pageNumber="26">A. namerokoensis</emphasis>
|
||
</taxonomicName>
|
||
(below), both of which occur in the Bemaraha-Beanka region.
|
||
<taxonomicName id="4C15D5A9E533D90D025FFF78473EB665" authorityName=": Fischer-Piette" authorityYear="1952" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E533D90D025FFF78473EB665" italics="true" pageId="26" pageNumber="26">A. namerokoensis</emphasis>
|
||
</taxonomicName>
|
||
differs in lacking significant axial sculpture on the protoconch and in having distinct spiral microsculpture on the teleoconch (
|
||
<figureCitation id="132EB2AFE533D90D04AFFEBD4549B605" box="[738,872,299,326]" captionStart="Fig" captionStartId="25.[144,177,1195,1216]" captionTargetBox="[154,1091,206,1175]" captionTargetId="figure-149@25.[149,1095,202,1179]" captionTargetPageId="25" captionText="Fig. 17. Ampelita beanka sp. n., A. milloti Fischer-Piette, 1952 and A. namerokoensis Fischer-Piette, 1952, protoconch morphology and teleoconch microsculpture: (A, B) A. beanka, protoconch diameter 4.1mm, st’n R04/09, respectively paratype, NMSA L8504/T2983, and holotype, AMS C.474166; (C, D) A. milloti, protoconch diameter 4.4 mm, both st’n 07/09, NMSA L8502; (E, F) A. namerokoensis, protoconch diameter 3.5 mm, respectively st’n 18/06, NMSA L8439, and st’n 03/06, NMSA L7038." figureDoi="http://doi.org/10.5281/zenodo.7670142" httpUri="https://zenodo.org/record/7670142/files/figure.png" pageId="26" pageNumber="26">Fig. 17E, F</figureCitation>
|
||
). It also attains a larger size, has the peripheral keel closer to mid-whorl, has a somewhat narrower umbilicus, and the suture, if descendant at all, is at most weakly so and then only near the end of the last adult whorl.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE533D90D069DFE3845BBB5A6" blockId="26.[181,1142,203,1126]" pageId="26" pageNumber="26">
|
||
<taxonomicName id="4C15D5A9E533D90D069DFE3847A2B685" authorityName=": Fischer-Piette" authorityYear="1952" box="[208,387,430,454]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E533D90D069DFE3847A2B685" box="[208,387,430,454]" italics="true" pageId="26" pageNumber="26">Ampelita milloti</emphasis>
|
||
</taxonomicName>
|
||
is perhaps more similar in having axial riblets on the protoconch and lacking spiral microsculpture, but its teleoconch sculpture is finer, the axial elements more irregular and fragmented, appearing granular rather than lamellate (particularly evident on the base). Furthermore, as in
|
||
<taxonomicName id="4C15D5A9E533D90D04CEFD98456BB565" authorityName=": Fischer-Piette" authorityYear="1952" box="[643,842,526,550]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E533D90D04CEFD98456BB565" box="[643,842,526,550]" italics="true" pageId="26" pageNumber="26">A. namerokoensis</emphasis>
|
||
</taxonomicName>
|
||
, the suture of
|
||
<taxonomicName id="4C15D5A9E533D90D05BCFD98427BB565" authorityName=": Fischer-Piette" authorityYear="1952" box="[1009,1114,526,550]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E533D90D05BCFD98427BB565" box="[1009,1114,526,550]" italics="true" pageId="26" pageNumber="26">A. milloti</emphasis>
|
||
</taxonomicName>
|
||
is never strongly descendant. The smaller size, shouldered profile, strongly descendant suture and lamellate axial sculpture of
|
||
<taxonomicName id="4C15D5A9E533D90D0417FDD844E8B525" box="[602,713,590,614]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E533D90D0417FDD844E8B525" box="[602,713,590,614]" italics="true" pageId="26" pageNumber="26">A. beanka</emphasis>
|
||
</taxonomicName>
|
||
render it distinctive. The descent of the suture below the peripheral keel, though characteristic of the species and well developed in the
|
||
<typeStatus id="54AE1088E533D90D06B6FD1D4741B5E6" box="[251,352,651,677]" pageId="26" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
, is somewhat variable in its extent, but usually commences at least one half whorl prior to the aperture. The position of the peripheral keel is also somewhat variable, lying closer to mid-whorl in some specimens (
|
||
<figureCitation id="132EB2AFE533D90D0568FD5D45ACB5A6" box="[805,909,715,741]" captionStart="Fig" captionStartId="23.[144,177,1558,1579]" captionTargetBox="[151,1096,1100,1536]" captionTargetId="figure-274@23.[143,1103,1094,1539]" captionTargetPageId="23" captionText="Fig. 15. Ampelita beanka sp. n.: (A–C) holotype, max. diameter 17.7 mm, height 8.0 mm,AMS C.474166; (D) specimen with less strongly shouldered whorls, max. diameter 19.1 mm, st’n 07/09, paratype, NMSA L8501/T2981; (E–F) specimens of the same size and collected from the same station R01/09, showing the differing extentto which the suture descends below the shoulder, max.diameter 17.1mm, paratypes, NMSA L8507/T2982." figureDoi="http://doi.org/10.5281/zenodo.7670134" httpUri="https://zenodo.org/record/7670134/files/figure.png" pageId="26" pageNumber="26">Fig. 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE533D90D069DFD7A47E6B325" blockId="26.[181,1142,203,1126]" pageId="26" pageNumber="26">
|
||
We have considered the possibility that these differences may reflect ecological factors and that the stepped profile may result from damage to the mantle edge at the suture, as is evident in some specimens of both
|
||
<taxonomicName id="4C15D5A9E533D90D043FFCB844F8B405" authorityName=": Fischer-Piette" authorityYear="1952" box="[626,729,814,838]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E533D90D043FFCB844F8B405" box="[626,729,814,838]" italics="true" pageId="26" pageNumber="26">A. milloti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E533D90D0543FCB845F2B405" authorityName=": Fischer-Piette" authorityYear="1952" box="[782,979,814,838]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E533D90D0543FCB845F2B405" box="[782,979,814,838]" italics="true" pageId="26" pageNumber="26">A. namerokoensis</emphasis>
|
||
</taxonomicName>
|
||
. However, the stepped profile of
|
||
<taxonomicName id="4C15D5A9E533D90D0736FCD847C8B425" box="[379,489,846,870]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E533D90D0736FCD847C8B425" box="[379,489,846,870]" italics="true" pageId="26" pageNumber="26">A. beanka</emphasis>
|
||
</taxonomicName>
|
||
is present in specimens showing no growth scars indicative of earlier damage to the outer lip, and we can consistently discriminate between
|
||
<taxonomicName id="4C15D5A9E533D90D0210FCF84727B4E5" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E533D90D0210FCF84727B4E5" italics="true" pageId="26" pageNumber="26">A. beanka</emphasis>
|
||
</taxonomicName>
|
||
and the other two species in terms of the microsculptural differences detailed above. There are also significant differences evident in the morphology of the penis (see below). All three species occur in the
|
||
<taxonomicName id="4C15D5A9E533D90D04D9FC5A451DB4A5" box="[660,828,972,998]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="26" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
and have been found to cooccur at a number of sampling sites. Emberton (1999) described a similar lamellar axial microsculpture in
|
||
<taxonomicName id="4C15D5A9E533D90D07CDFB9844C3B365" authority="Emberton, 1999" authorityName="Emberton" authorityYear="1999" box="[384,738,1036,1062]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="26" phylum="Mollusca" rank="species" species="analamerae">
|
||
<emphasis id="B9617238E533D90D07CDFB984404B365" box="[384,549,1038,1062]" italics="true" pageId="26" pageNumber="26">A. analamerae</emphasis>
|
||
Emberton, 1999
|
||
</taxonomicName>
|
||
, from far north-eastern
|
||
<collectingCountry id="F302EEBAE533D90D05A5FB9A4250B365" box="[1000,1137,1036,1062]" name="Madagascar" pageId="26" pageNumber="26">Madagascar</collectingCountry>
|
||
, but that species is larger, is keeled at mid-whorl and has a much more prominent spire and narrower umbilicus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |