treatments-xml/data/03/F7/96/03F7965AFF8FFFE457C1EB85D3C6E99D.xml
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<document id="C1404DE803AA3DC1F5E1F3DE10241163" ID-CLB-Dataset="46913" ID-DOI="10.5281/zenodo.200069" ID-GBIF-Dataset="97be6d67-b44e-467c-9356-7118502955c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="200069" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460352717287" checkinUser="plazi" docAuthor="Rowley, Jodi J. L., Duong, Le Thi Thuy, Dao, Tran Thi Anh, Stuart, Bryan L. &amp; Huy, Hoang Duc" docDate="2010" docId="03F7965AFF8FFFE457C1EB85D3C6E99D" docLanguage="en" docName="zt02727p055.pdf" docOrigin="Zootaxa 2727" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhacophorus vampyrus Rowley, Duong, Dao, Stuart &amp; Huy, 2010, sp. nov." docType="treatment" docVersion="14" lastPageNumber="53" masterDocId="FFCEEE22FF8EFFED5756EF40D00DE949" masterDocTitle="A new tree frog of the genus Rhacophorus (Anura: Rhacophoridae) from southern Vietnam" masterLastPageNumber="55" masterPageNumber="45" pageNumber="46" updateTime="1698293720631" updateUser="ExternalLinkService">
<mods:mods id="4E1E640DC16A1B33FE1A3DBD8D0A0C15" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0A8CE5AB554A45B68E942B1665577D7C">
<mods:title id="E83316F68F8C1F716CF13E6DDA90D6E3">A new tree frog of the genus Rhacophorus (Anura: Rhacophoridae) from southern Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="8FEEEB0AD170AD66E4D1B27B66C4C94A" type="personal">
<mods:role id="F8704470BEB2EC9436B0250766084374">
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</mods:role>
<mods:namePart id="3DA8152105C8652366C5A32D02A1CB8A">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="B34B27035B9559B58838EF1916AAFBC2">Duong, Le Thi Thuy</mods:namePart>
</mods:name>
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<mods:namePart id="37BEB8D3E708EF459CEC9823A3DC5FF7">Dao, Tran Thi Anh</mods:namePart>
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<mods:role id="18309801A6A99FD4BD7B685D00FDDE54">
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<mods:namePart id="795CD09F75F20CE537E566F14665525A">Stuart, Bryan L.</mods:namePart>
</mods:name>
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<mods:roleTerm id="DA60EF59761B41B9395A74AFC1786442">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F96659B27C393353882B14CF7FFFBA3">Huy, Hoang Duc</mods:namePart>
</mods:name>
<mods:typeOfResource id="C993FDB30E924F9E232211FC342D1A40">text</mods:typeOfResource>
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<mods:title id="994CC87584785A0C5477B5946B7448D5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F42757A8E0FAF5C6B8C0AB0538C811D3">
<mods:date id="1521B916FEB4D99B603B294A60628495">2010</mods:date>
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<mods:identifier id="4D0E2F35497BF15C649067669BD0CA09" type="DOI">10.5281/zenodo.200069</mods:identifier>
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<treatment id="03F7965AFF8FFFE457C1EB85D3C6E99D" ID-DOI="http://doi.org/10.5281/zenodo.5691173" ID-GBIF-Taxon="119408206" ID-Zenodo-Dep="5691173" LSID="urn:lsid:plazi:treatment:03F7965AFF8FFFE457C1EB85D3C6E99D" httpUri="http://treatment.plazi.org/id/03F7965AFF8FFFE457C1EB85D3C6E99D" lastPageId="9" lastPageNumber="53" pageId="1" pageNumber="46">
<subSubSection id="C34474C7FF8FFFEC57C1EB85D216ED96" box="[151,539,1221,1247]" pageId="1" pageNumber="46" type="nomenclature">
<paragraph id="8BE1274CFF8FFFEC57C1EB85D216ED96" blockId="1.[151,539,1221,1247]" box="[151,539,1221,1247]" pageId="1" pageNumber="46">
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<emphasis id="B92AFB5EFF8FFFEC57C1EB85D216ED96" bold="true" box="[151,539,1221,1247]" pageId="1" pageNumber="46">
<taxonomicName id="4C5E5CCFFF8FFFEC57C1EB85D1BBED96" box="[151,438,1221,1247]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="46" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF8FFFEC57C1EB85D1BBED96" bold="true" box="[151,438,1221,1247]" italics="true" pageId="1" pageNumber="46">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194625FF8FFFEC56EBEB85D216ED96" box="[445,539,1221,1247]" pageId="1" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34474C7FF8FFFEC57C1EA4CD17EEEBD" pageId="1" pageNumber="46" type="materials_examined">
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<emphasis id="B92AFB5EFF8FFFEC57C1EA4CD106EC6D" bold="true" box="[151,267,1292,1316]" pageId="1" pageNumber="46">
<typeStatus id="54E599EEFF8FFFEC57C1EA4CD10BEC6D" box="[151,262,1292,1316]" pageId="1" pageNumber="46" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED4FBF89FF8FFFEC5642EA4CD15CEC6D" box="[276,337,1292,1316]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173127, adult male, on a tree branch in montane evergreen forest in Bidoup-Nui Ba National Park, Lac Duong District, Lam Dong Province,
<collectingCountry id="F34967DCFF8FFFEC55E5EA6FD31FEC0E" box="[691,786,1327,1351]" name="Vietnam" pageId="1" pageNumber="46">Vietnam</collectingCountry>
(
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,
<geoCoordinate id="EE6A418BFF8FFFEC54E7EA6FD432EC0E" box="[945,1087,1327,1351]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.7151">108.7151 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC531DEA6FD4A9EC0E" box="[1099,1188,1327,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="1" pageNumber="46" unit="m" value="1625.0">1625 m</quantity>
). Collected at 21:00 h on
<date id="FFE0018CFF8FFFEC57ECEA14D146EC25" box="[186,331,1364,1388]" pageId="1" pageNumber="46" value="2010-07-26">26 July 2010</date>
by Le T. T. D.
</paragraph>
<paragraph id="8BE1274CFF8FFFEC5793EA38D2D8EE2D" blockId="1.[151,1437,1292,2036]" pageId="1" pageNumber="46">
<emphasis id="B92AFB5EFF8FFFEC5793EA38D149ECD9" bold="true" box="[197,324,1400,1424]" pageId="1" pageNumber="46">
<typeStatus id="54E599EEFF8FFFEC5793EA38D132ECD9" box="[197,319,1400,1424]" pageId="1" pageNumber="46" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED4FBF89FF8FFFEC561AEA37D184ECC6" box="[332,393,1399,1423]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173126, female, on a tree branch approximately
<quantity id="4CA68AA9FF8FFFEC5484EA37D3F3ECC6" box="[978,1022,1399,1423]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="46" unit="m" value="2.0">2 m</quantity>
above ground in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC57B3EADCD164ECFD" box="[229,361,1436,1460]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1865">12.1865 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC5622EADCD20CECFD" box="[372,513,1436,1460]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.715">108.7150 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC555BEADCD269ECFD" box="[525,612,1436,1460]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="1" pageNumber="46" unit="m" value="1625.0">1625 m</quantity>
), collected at 22:30 h on
<date id="FFE0018CFF8FFFEC5428EADCD402ECFD" box="[894,1039,1436,1460]" pageId="1" pageNumber="46" value="2010-07-26">26 July 2010</date>
by Le T. T. D.
<collectionCode id="ED4FBF89FF8FFFEC53EFEADCD4FCECFD" box="[1209,1265,1436,1460]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="1" pageNumber="46">UNS</collectionCode>
00103/
<collectionCode id="ED4FBF89FF8FFFEC5210EADCD58FECFD" box="[1350,1410,1436,1460]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173128 male, on a leaf approximately
<quantity id="4CA68AA9FF8FFFEC5506EAFFD270EC9E" box="[592,637,1471,1495]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="46" unit="m" value="2.0">2 m</quantity>
above ground in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC5320EAFFD4F1EC9E" box="[1142,1276,1471,1495]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1736">12.1736 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC525FEAFFD59AEC9E" box="[1289,1431,1471,1495]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.6987">108.6987 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC57C1EAA4D0E1ECB5" box="[151,236,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="46" unit="m" value="1470.0">1470 m</quantity>
) collected at 22:05 h on
<date id="FFE0018CFF8FFFEC56A9EAA4D2A5ECB5" box="[511,680,1508,1532]" pageId="1" pageNumber="46" value="2009-03-31">31 March 2009</date>
by Le T. T. D.
<collectionCode id="ED4FBF89FF8FFFEC5407EAA4D384ECB5" box="[849,905,1508,1532]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="1" pageNumber="46">UNS</collectionCode>
00104/
<collectionCode id="ED4FBF89FF8FFFEC5488EAA4D414ECB5" box="[990,1049,1508,1532]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173129, male, on a leaf approximately
<quantity id="4CA68AA9FF8FFFEC57A7E947D13CEF56" box="[241,305,1543,1567]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="46" unit="m" value="1.5">1.5 m</quantity>
above ground,
<quantity id="4CA68AA9FF8FFFEC568BE947D210EF56" box="[477,541,1543,1567]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="46" unit="m" value="0.5">0.5 m</quantity>
from rocky stream in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC5368E947D4CCEF56" box="[1086,1217,1543,1567]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1865">12.1865 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC539AE947D554EF56" box="[1228,1369,1543,1567]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.7151">108.7151 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC5232E947D0A3EF0D" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="1" pageNumber="46" unit="m" value="1625.0">1625 m</quantity>
) collected at 20:30 h on
<date id="FFE0018CFF8FFFEC5699E96CD273EF0D" box="[463,638,1580,1604]" pageId="1" pageNumber="46" value="2009-03-30">30 March 2009</date>
by Le T. T. D.
<collectionCode id="ED4FBF89FF8FFFEC5463E96CD37FEF0D" box="[821,882,1580,1604]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173507, female, on a leaf approximately
<quantity id="4CA68AA9FF8FFFEC5239E96CD591EF0D" box="[1391,1436,1580,1604]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="46" unit="m" value="1.0">1 m</quantity>
above ground in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC55D2E90FD304EF2E" box="[644,777,1615,1639]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1736">12.1736 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC5443E90FD3AEEF2E" box="[789,931,1615,1639]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.6987">108.6987 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC54F9E90FD405EF2E" box="[943,1032,1615,1639]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="46" unit="m" value="1470.0">1470 m</quantity>
), collected at 21:50 h on
<date id="FFE0018CFF8FFFEC5271E90FD0C2EFC5" pageId="1" pageNumber="46" value="2009-08-18">18 August 2009</date>
by Le T. T. D.
<collectionCode id="ED4FBF89FF8FFFEC5629E934D1C3EFC5" box="[383,462,1652,1676]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="1" pageNumber="46">NCSM</collectionCode>
77318, adult male, on a trunk of tree away from water,
<quantity id="4CA68AA9FF8FFFEC531FE934D478EFC5" box="[1097,1141,1652,1676]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="46" unit="m" value="1.0">1 m</quantity>
above ground in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC560CE9D7D1D2EFE6" box="[346,479,1687,1711]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1864">12.1864 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC56BDE9D7D27AEFE6" box="[491,631,1687,1711]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.7149">108.7149 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC55D2E9D7D2D1EFE6" box="[644,732,1687,1711]" metricMagnitude="3" metricUnit="m" metricValue="1.627" pageId="1" pageNumber="46" unit="m" value="1627.0">1627 m</quantity>
), collected at 21:00 h on
<date id="FFE0018CFF8FFFEC54AEE9D7D49BEFE6" box="[1016,1174,1687,1711]" pageId="1" pageNumber="46" value="2008-03-06">6 March 2008</date>
by B. L. Stuart, J. J. L. Rowley, Le T. T. D., Tran T. A. D., and Hoang D. H.
<collectionCode id="ED4FBF89FF8FFFEC544AE9FCD359EF9D" box="[796,852,1724,1748]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="1" pageNumber="46">UNS</collectionCode>
0 0 105, adult female, in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC57C8E99FD12EEFBE" box="[158,291,1759,1783]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1736">12.1736 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC5679E99FD1B1EFBE" box="[303,444,1759,1783]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.6987">108.6987 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC569FE99FD22CEFBE" box="[457,545,1759,1783]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="46" unit="m" value="1470.0">1470 m</quantity>
), collected on
<date id="FFE0018CFF8FFFEC5590E99FD37EEFBE" box="[710,883,1759,1783]" pageId="1" pageNumber="46" value="2010-03-18">18 March 2010</date>
by Tran T. A. D.
<collectionCode id="ED4FBF89FF8FFFEC536BE9A0D486EFBE" LSID="urn:lsid:biocol.org:col:34613" box="[1085,1163,1760,1783]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="1" pageNumber="46">ZFMK</collectionCode>
91076, adult male, on a tree
<quantity id="4CA68AA9FF8FFFEC579FE844D0F8EE55" box="[201,245,1796,1820]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="46" unit="m" value="1.0">1 m</quantity>
above ground near 1
<quantity id="4CA68AA9FF8FFFEC56A5E844D212EE55" box="[499,543,1796,1820]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="46" unit="m" value="2.0">2 m</quantity>
wide, dry stream in montane evergreen forest (
<geoCoordinate id="EE6A418BFF8FFFEC536EE844D4B0EE55" box="[1080,1213,1796,1820]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.0096">12.0096 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC539FE844D55AEE55" box="[1225,1367,1796,1820]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.6607">108.6607 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC5232E844D0BDEE76" metricMagnitude="3" metricUnit="m" metricValue="2.004" pageId="1" pageNumber="46" unit="m" value="2004.0">2004 m</quantity>
), collected on
<date id="FFE0018CFF8FFFEC5600E867D209EE76" box="[342,516,1831,1855]" pageId="1" pageNumber="46" value="2010-03-23">23 March 2010</date>
by Tran T. A. D. All specimens were collected in Bidoup-Nui Ba National Park, Lac Duong District, Lam Dong Province,
<collectingCountry id="F34967DCFF8FFFEC5538E80CD2DCEE2D" box="[622,721,1868,1892]" name="Vietnam" pageId="1" pageNumber="46">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BE1274CFF8FFFEC5793E82FD17EEEBD" blockId="1.[151,1437,1292,2036]" pageId="1" pageNumber="46">
<emphasis id="B92AFB5EFF8FFFEC5793E82FD18DEEC1" bold="true" box="[197,384,1903,1928]" pageId="1" pageNumber="46">Other material.</emphasis>
<collectionCode id="ED4FBF89FF8FFFEC56D1E82FD1CEEECE" box="[391,451,1903,1927]" pageId="1" pageNumber="46">AMS</collectionCode>
R 173132, six tadpoles, inside small pool of water in an approximately
<quantity id="4CA68AA9FF8FFFEC53BBE82FD53FEEC1" box="[1261,1330,1903,1928]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="46" unit="cm" value="10.0">10 cm</quantity>
diameter tree-hole, approximately
<quantity id="4CA68AA9FF8FFFEC56E4E8D4D1D2EEE5" box="[434,479,1940,1964]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="46" unit="m" value="1.0">1 m</quantity>
above the ground in montane evergreen forest in Bidoup-Nui Ba National Park, Lac Duong District, Lam Dong Province,
<collectingCountry id="F34967DCFF8FFFEC556BE8F8D291EE99" box="[573,668,1976,2000]" name="Vietnam" pageId="1" pageNumber="46">Vietnam</collectingCountry>
(
<geoCoordinate id="EE6A418BFF8FFFEC55FCE8F8D322EE99" box="[682,815,1976,2000]" direction="north" orientation="latitude" pageId="1" pageNumber="46" precision="5" value="12.1736">12.1736 N</geoCoordinate>
,
<geoCoordinate id="EE6A418BFF8FFFEC546CE8F8D3CBEE99" box="[826,966,1976,2000]" direction="east" orientation="longitude" pageId="1" pageNumber="46" precision="5" value="108.6987">108.6987 E</geoCoordinate>
,
<quantity id="4CA68AA9FF8FFFEC5484E8F8D426EE99" box="[978,1067,1976,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="46" unit="m" value="1470.0">1470 m</quantity>
). Collected at 10:30 h on
<date id="FFE0018CFF8FFFEC521CE8F8D0C2EEBD" pageId="1" pageNumber="46" value="2010-07-28">28 July 2010</date>
by Le T. T. D.
</paragraph>
</subSubSection>
<caption id="DF2177C4FF8CFFEF57C1EBD0D319EDEF" box="[151,788,1168,1191]" httpUri="https://zenodo.org/record/200070/files/figure.png" pageId="2" pageNumber="47" targetBox="[151,1436,195,1144]" targetPageId="2">
<paragraph id="8BE1274CFF8CFFEF57C1EBD0D319EDEF" blockId="2.[151,788,1168,1191]" box="[151,788,1168,1191]" pageId="2" pageNumber="47">
<emphasis id="B92AFB5EFF8CFFEF57C1EBD0D11EEDEF" bold="true" box="[151,275,1168,1191]" pageId="2" pageNumber="47">FIGURE 1.</emphasis>
Collection site of
<taxonomicName id="4C5E5CCFFF8CFFEF569BEBD1D2B0EDEE" box="[461,701,1169,1191]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF8CFFEF569BEBD1D2B0EDEE" box="[461,701,1169,1191]" italics="true" pageId="2" pageNumber="47">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194625FF8CFFEF5592EBD1D319EDEF" box="[708,788,1169,1190]" pageId="2" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C34474C7FF8CFFEF5793EB94D4C9EC7A" pageId="2" pageNumber="47" type="etymology">
<paragraph id="8BE1274CFF8CFFEF5793EB94D162EC46" blockId="2.[151,1437,1235,2016]" pageId="2" pageNumber="47">
<emphasis id="B92AFB5EFF8CFFEF5793EB94D146EDA5" bold="true" box="[197,331,1236,1260]" pageId="2" pageNumber="47">Etymology.</emphasis>
Specific epithet in reference to the unusual tadpole of the new species (see below), applied as a noun in apposition.
</paragraph>
<paragraph id="8BE1274CFF8CFFEF5793EA5BD4C9EC7A" blockId="2.[151,1437,1235,2016]" box="[197,1220,1301,1332]" pageId="2" pageNumber="47">
<emphasis id="B92AFB5EFF8CFFEF5793EA5BD1F4EC7D" bold="true" box="[197,505,1307,1332]" pageId="2" pageNumber="47">Suggested common name.</emphasis>
Vampire tree frog (English). Ế ch cây ma cà rông (Vietnamese).
</paragraph>
</subSubSection>
<subSubSection id="C34474C7FF8CFFEF5793EA7FD17DEF1A" pageId="2" pageNumber="47" type="diagnosis">
<paragraph id="8BE1274CFF8CFFEF5793EA7FD17DEF1A" blockId="2.[151,1437,1235,2016]" pageId="2" pageNumber="47">
<emphasis id="B92AFB5EFF8CFFEF5793EA7FD132EC11" bold="true" box="[197,319,1343,1368]" pageId="2" pageNumber="47">Diagnosis.</emphasis>
The new species is assigned to the genus
<taxonomicName id="4C5E5CCFFF8CFFEF5461EA00D3C0EC11" box="[823,973,1344,1368]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B92AFB5EFF8CFFEF5461EA00D3C0EC11" box="[823,973,1344,1368]" italics="true" pageId="2" pageNumber="47">Rhacophorus</emphasis>
</taxonomicName>
by the presence of intercalary cartilage between the terminal and penultimate phalanges of digits, Y-shaped distal end of terminal phalanx, tips of digits expanded into large disks bearing circummarginal grooves, webbed fingers, a supracloacal dermal ridge, vomerine teeth, and horizontal pupil (
<bibRefCitation id="EFCF5ABDFF8CFFEF5698EAEBD2C0EC8A" author="Brown" box="[462,717,1451,1475]" pageId="2" pageNumber="47" refString="Brown, W. C. &amp; Alcala, A. C. (1994) Philippine frogs of the family Rhacophoridae. Proceedings of the Californian Academy of Sciences 48, 185 - 220." type="journal article" year="1994">Brown &amp; Alcala 1994</bibRefCitation>
;
<bibRefCitation id="EFCF5ABDFF8CFFEF558CEAEBD3FFEC8A" author="Duellman" box="[730,1010,1451,1475]" pageId="2" pageNumber="47" refString="Duellman, W. E. &amp; Trueb, L. (1986) Biology of Amphibians. The John Hopkins University Press, Baltimore and London. 670 pp." type="book" year="1986">Duellman &amp; Trueb 1986</bibRefCitation>
;
<bibRefCitation id="EFCF5ABDFF8CFFEF54A9EAEBD473EC8A" author="Liem" box="[1023,1150,1451,1475]" pageId="2" pageNumber="47" refString="Liem, S. S. (1970) The morphology, systematic, and evolution of the old world treefrogs (Rhacophoridae and Hyperoliidae). Fieldiana Zoology, 57, 1 - 145." type="journal article" year="1970">Liem 1970</bibRefCitation>
).
<taxonomicName id="4C5E5CCFFF8CFFEF53C4EAECD591EC8D" box="[1170,1436,1452,1476]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="47" phylum="Arthropoda" rank="species" species="vampyrus">
<emphasis id="B92AFB5EFF8CFFEF53C4EAECD591EC8D" box="[1170,1436,1452,1476]" italics="true" pageId="2" pageNumber="47">Rhacophorus vampyrus</emphasis>
</taxonomicName>
is distinguished from all other
<taxonomicName id="4C5E5CCFFF8CFFEF56A5EA90D285ECA1" box="[499,648,1488,1512]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B92AFB5EFF8CFFEF56A5EA90D285ECA1" box="[499,648,1488,1512]" italics="true" pageId="2" pageNumber="47">Rhacophorus</emphasis>
</taxonomicName>
in mainland Southeast Asia by a combination of (1) pale tan to brick red dorsum, (2) white throat, chest and belly, (3) mostly black flanks and anterior and posterior surface of thighs, (4) grey to black webbing between fingers and toes, (5) reduced finger webbing, and (6) pointed projection at tibiotarsal articulation.
</paragraph>
</subSubSection>
<subSubSection id="C34474C7FF8CFFE45793E91FD3C6E99D" lastPageId="9" lastPageNumber="54" pageId="2" pageNumber="47" type="description">
<paragraph id="8BE1274CFF8CFFEE5793E91FD4F2EB54" blockId="2.[151,1437,1235,2016]" lastBlockId="3.[151,1436,151,541]" lastPageId="3" lastPageNumber="48" pageId="2" pageNumber="47">
<emphasis id="B92AFB5EFF8CFFEF5793E91FD1E9EF31" bold="true" box="[197,484,1631,1656]" pageId="2" pageNumber="47">
Description of
<typeStatus id="54E599EEFF8CFFEF5620E920D1D2EF31" box="[374,479,1632,1656]" pageId="2" pageNumber="47" type="holotype">holotype</typeStatus>
.
</emphasis>
Body dorsoventrally compressed; head length 90% of head width; snout truncate in dorsal view, bluntly truncate in profile, projecting slightly beyond margin of the lower jaw, with slight point on tip of snout visible in ventral view; canthus rostralis distinct, bluntly angular; loreal region sloping, slightly concave; lips slightly flared; interorbital region slightly convex; nostrils oval, slightly protuberant, without flap of skin laterally, much closer to tip of snout than eye; pupil horizontal, tympanum barely visible externally, tympanic rim slightly elevated relative to skin of temporal region, 41% of eye diameter; pineal ocellus absent; skin not co-ossified to forehead; vomerine teeth present in oblique groups, separated by a distance about as long as each group, closer to choanae than wide; choanae oval, at margins of roof of mouth; tongue attached anteriorly, deeply notched posteriorly; tooth-like projections on lower jaw absent; pair of distinct, oval vocal sac openings at base of jaw; external paired subgular vocal sacs; weak supratympanic fold extending to just beyond level of axilla. Forelimbs relatively robust, relative length of fingers I &lt;II &lt;IV &lt;III; tips of all fingers with well-developed disks with distinct circummarginal grooves, disks relatively wide compared to finger width (third finger disk 170% third finger width), third finger disk width greater (163%) than tympanum diameter; webbing formula
<date id="FFE0018CFF8DFFEE53DCEF80D4C3E99E" box="[1162,1230,191,215]" pageId="3" pageNumber="48">I 2 2</date>
+
<date id="FFE0018CFF8DFFEE538CEF80D50FE99E" box="[1242,1282,191,215]" pageId="3" pageNumber="48">II 1</date>
+ 2+
<date id="FFE0018CFF8DFFEE5269EF80D57DE99E" box="[1343,1392,191,215]" pageId="3" pageNumber="48">III 2</date>
-
<date id="FFE0018CFF8DFFEE52D8EFFFD0BBE9B3" pageId="3" pageNumber="48">1 IV</date>
; subarticular tubercles prominent, rounded, formula 1, 1, 2, 2; palmar tubercle absent; accessory palmar tubercles indistinct; thenar tubercle absent; prepollex enlarged, with low, relatively indistinct tubercle; nuptial pads or nuptial excrescences absent. Relative length of toes I &lt;II &lt;III &lt;V &lt;IV; tips of toes with well-developed disks with distinct circummarginal grooves; disks smaller than those of fingers; webbing formula
<date id="FFE0018CFF8DFFEE53CCEE12D59EE820" box="[1178,1427,337,361]" pageId="3" pageNumber="48">I 1 2 II 1 2 III 1 2</date>
+
<date id="FFE0018CFF8DFFEE57C1EE36D0F9E8C4" box="[151,244,373,397]" pageId="3" pageNumber="48">IV 2 1</date>
V; subarticular tubercles rounded, distal subarticular tubercles distinct, inner less distinct, formula 1, 1, 2, 3, 2; inner metatarsal tubercle low, oval; outer metatarsal tubercle and supernumary tubercles absent. Dorsal skin smooth, ventral surface of thighs and belly coarsely granular, chest and throat smooth. Loose skin on either side of the throat over vocal sacs. Tarsal fold absent. Outer margin of forearm and foot with low dermal ridges; low supracloacal dermal ridge; pointed projection at tibiotarsal articulation present, approximately
<quantity id="4CA68AA9FF8DFFEE5328ED45D4B2EB54" box="[1150,1215,517,541]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="48" unit="mm" value="1.0">1 mm</quantity>
long.
</paragraph>
<caption id="DF2177C4FF8DFFEE57C1E908D1D7EFF3" httpUri="https://zenodo.org/record/200071/files/figure.png" pageId="3" pageNumber="48" targetBox="[151,1436,603,1586]" targetPageId="3">
<paragraph id="8BE1274CFF8DFFEE57C1E908D1D7EFF3" blockId="3.[151,1436,1608,1722]" pageId="3" pageNumber="48">
<emphasis id="B92AFB5EFF8DFFEE57C1E908D119EF14" bold="true" box="[151,276,1608,1631]" pageId="3" pageNumber="48">FIGURE 2.</emphasis>
Dorsolateral view of
<taxonomicName id="4C5E5CCFFF8DFFEE56AEE908D2E7EF17" box="[504,746,1608,1630]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF8DFFEE56AEE908D2E7EF17" box="[504,746,1608,1630]" italics="true" pageId="3" pageNumber="48">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194625FF8DFFEE55A7E909D34EEF17" box="[753,835,1609,1630]" pageId="3" pageNumber="48" rank="species">
<emphasis id="B92AFB5EFF8DFFEE55A7E909D34EEF17" bold="true" box="[753,835,1609,1630]" pageId="3" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
in life (A) adult male holotype (AMS R 173127), diurnal colouration, (B) female paratype (AMS R 173126), diurnal colouration, (C) adult male, non-vouchered specimen,
<emphasis id="B92AFB5EFF8DFFEE527EE928D565EF34" box="[1320,1384,1640,1661]" italics="true" pageId="3" pageNumber="48">in situ</emphasis>
with nocturnal colouration, and (D) adult male paratype NCSM 77318, diurnal colouration (note modified skin under angles of jaw over paired subgular vocal sac).
</paragraph>
</caption>
<paragraph id="8BE1274CFF8DFFE95793E9A6D23AE855" blockId="3.[151,1436,1766,2007]" lastBlockId="4.[151,1436,151,391]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
<emphasis id="B92AFB5EFF8DFFEE5793E9A6D20DEFB6" bold="true" box="[197,512,1766,1791]" pageId="3" pageNumber="48">
Colour of
<typeStatus id="54E599EEFF8DFFEE5615E9A7D1A7EFB6" box="[323,426,1767,1791]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsal surface pale copper-brown with faint darker brown mottling along back; dorsolateral surfaces with very small, sparse, white and darker brown flecks; dorsal surface of lower arms, thigh and tibiotarsus copper-brown with diffuse darker brown barring; dorsal surface of hands and feet copper-brown proximally, fading distally to pinkish-cream on fingers and toes I II, and grey on fingers III IV and toes III V; dark grey to black webbing dorsally. Black flanks, upper arms, ventral surface of lower arms, anterior and posterior surface of thighs, and ventral surface of crus, with small, irregular white spots within the black on the flanks and upper arms. Ventral surface of throat, chest and belly immaculate except for black mottling extending slightly onto lateral margins of chest at axilla; ventral surfaces of toes and fingers pale grey; ventral surface of webbing grey with dark grey/black margins. Iris pale yellowish gold with a network of fine dark gold reticulations concentrated around the pupil; iris periphery black; posterior periphery of eye blue. Dorsal colouration varied from pale tan (diurnally) to brick red (nocturnally).
</paragraph>
<paragraph id="8BE1274CFF8AFFE95793EE67D466E876" blockId="4.[151,1436,151,391]" box="[197,1131,295,320]" pageId="4" pageNumber="49">
<emphasis id="B92AFB5EFF8AFFE95793EE67D268E809" bold="true" box="[197,613,295,320]" pageId="4" pageNumber="49">
Colour of
<typeStatus id="54E599EEFF8AFFE95668EE68D1A8E809" box="[318,421,296,320]" pageId="4" pageNumber="49" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
<collectingCountry id="F34967DCFF8AFFE9553DEE68D287E809" box="[619,650,296,320]" name="American Samoa" pageId="4" pageNumber="49">As</collectingCountry>
in life, but with dorsum fading to pale tan.
</paragraph>
<paragraph id="8BE1274CFF8AFFE95793EE0CD359E8CE" blockId="4.[151,1436,151,391]" pageId="4" pageNumber="49">
<emphasis id="B92AFB5EFF8AFFE95793EE0CD176E82D" bold="true" box="[197,379,332,356]" pageId="4" pageNumber="49">Measurements.</emphasis>
<typeStatus id="54E599EEFF8AFFE956D4EE0CD1E3E82D" box="[386,494,332,356]" pageId="4" pageNumber="49" type="holotype">Holotype</typeStatus>
: SVL 43.6, HDL 14.6, HDW 16.3, SNT 5.5, EYE 4.6, IOD 4.8, TMP 1.9, TEY 1.6, IN 3.8, NS 1.2, EN 3.6, TIB 20.7, ML 13.2, PL 19.1, IML 2.4.
</paragraph>
<caption id="DF2177C4FF8AFFE957C1EB02D150ED3E" httpUri="https://zenodo.org/record/200072/files/figure.png" pageId="4" pageNumber="49" targetBox="[155,1428,440,1065]" targetPageId="4">
<paragraph id="8BE1274CFF8AFFE957C1EB02D150ED3E" blockId="4.[151,1436,1090,1143]" pageId="4" pageNumber="49">
<emphasis id="B92AFB5EFF8AFFE957C1EB02D11FED11" bold="true" box="[151,274,1090,1113]" pageId="4" pageNumber="49">FIGURE 3.</emphasis>
Dorsal (A) and ventral (B) views of preserved male holotype of
<taxonomicName id="4C5E5CCFFF8AFFE954CDEB03D487ED10" box="[923,1162,1091,1113]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF8AFFE954CDEB03D487ED10" box="[923,1162,1091,1113]" italics="true" pageId="4" pageNumber="49">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<emphasis id="B92AFB5EFF8AFFE953C6EB03D4EDED11" bold="true" box="[1168,1248,1091,1112]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="A2194625FF8AFFE953C6EB03D4EDED11" box="[1168,1248,1091,1112]" pageId="4" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(AMS R 173127). Scale bar = 10 mm.
</paragraph>
</caption>
<paragraph id="8BE1274CFF8AFFE85793EBDED253E99D" blockId="4.[151,1437,1182,2034]" lastBlockId="5.[151,1437,151,212]" lastPageId="5" lastPageNumber="50" pageId="4" pageNumber="49">
<emphasis id="B92AFB5EFF8AFFE95793EBDED134EDFE" bold="true" box="[197,313,1182,1207]" pageId="4" pageNumber="49">Variation.</emphasis>
Measurements of the
<typeStatus id="54E599EEFF8AFFE95565EBE0D26EEDF1" box="[563,611,1184,1208]" pageId="4" pageNumber="49">type</typeStatus>
series are shown in
<tableCitation id="C6DC12F7FF8AFFE95412EBDFD39AEDFE" box="[836,919,1183,1207]" captionStart="TABLE 1" captionStartId="5.[151,235,255,276]" captionTargetBox="[151,1421,304,1172]" captionTargetPageId="5" captionText="TABLE 1. Measurements (mm) of Rhacophorus vampyrus sp. nov. Abbreviations defined in text." httpUri="http://table.plazi.org/id/DF2177C4FF8BFFE857C1EFBFD477E85D" pageId="4" pageNumber="49" tableUuid="DF2177C4FF8BFFE857C1EFBFD477E85D">Table 1</tableCitation>
. AMS R 173126 has dark brownish-grey marbling on the dorsal surface running from the interorbital region down the back; pinkish coloured webbing, particularly ventrally, between toes III IV; more distinct barring on limbs in preserve; and less dense black colouration on the ventral surfaces of the thighs. AMS R 173507 has a uniformly coloured dorsum, with no black or white spotting or dark patterns; indistinct barring on dorsal surface of tibiotarsus; and grey webbing on hands and fingers. AMS R 173129 has an indistinct, slightly darker brown, irregular pattern running along dorsum; two,
<quantity id="4CA68AA9FF8AFFE953FBEA12D4E3EC23" box="[1197,1262,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="49" unit="mm" value="1.0">1 mm</quantity>
diameter white spots on the back posteriorly; and completely black posterior and anterior surfaces of thighs, without white spotting. AMS R 173128 is strongly patterned with a darker brown, irregular pattern and scattered darker brown specks on the entire dorsum; very distinct barring on limbs; and less black colouration on ventral surfaces of thighs. NCSM 77318 has pale brown or cream mottling on the dorsal surface of head, back and limbs in life; and a dark brown dorsum with pale dusting only visible in between barring on the dorsal surface of the legs in preserve. ZFMK 91076 is almost uniformly pale tan in life and in preserve; has faint barring on arms and legs; slightly darker blotches in the interorbital region and along back; less extensive black colouration marbled with white on flanks, anterior and posterior surfaces of upper arms, anterior surface of lower arms, and anterior and posterior surface of thighs; and only faint black speckling on ventral surfaces of fingers II IV and finger webbing, ventral surface of crus, and anterior half of pes, including ventral surface of toes I III and toe-webbing. UNS 0 0 105 has an almost uniformly pale tan dorsum in life and in preserve; has faint barring on arms and legs; and the most extensive black colouration, without white spotting on flanks, anterior and posterior surfaces of upper arms, anterior surface of lower arms, ventral surfaces of fingers I IV and finger-webbing, anterior and posterior surface of thighs, ventral surface of crus, and anterior half of pes, including ventral surface of toes I III and toe-webbing. UNS 0 0 105 also has fine black specks concentrated on ventral surface of chest, but also sparsely distributed on the belly and outer margins of the throat. Males vary in the extent and colouration of loose skin in the gular region; UNS 00103/AMS R 173128 has only slightly loose skin in the gular region, only weakly coloured grey; UNS00104/AMS R 173129 has loose skin in the gular region, coloured greyish black; NCSM 77318 and ZFMK 91076 have the most pronounced loose skin in the gular region, with dark grey colouration also extending ventrally along margins of throat. All males lack nuptial pads or asperities.
</paragraph>
<caption id="DF2177C4FF8BFFE857C1EFBFD477E85D" ID-Table-UUID="DF2177C4FF8BFFE857C1EFBFD477E85D" box="[151,1146,254,277]" httpUri="http://table.plazi.org/id/DF2177C4FF8BFFE857C1EFBFD477E85D" pageId="5" pageNumber="50" targetBox="[151,1421,304,1172]" targetIsTable="true" targetPageId="5">
<paragraph id="8BE1274CFF8BFFE857C1EFBFD477E85D" blockId="5.[151,1146,254,277]" box="[151,1146,254,277]" pageId="5" pageNumber="50">
<emphasis id="B92AFB5EFF8BFFE857C1EFBFD109E85D" bold="true" box="[151,260,255,276]" pageId="5" pageNumber="50">TABLE 1.</emphasis>
Measurements (mm) of
<taxonomicName id="4C5E5CCFFF8BFFE856A8EFBFD2E3E85C" box="[510,750,255,277]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF8BFFE856A8EFBFD2E3E85C" box="[510,750,255,277]" italics="true" pageId="5" pageNumber="50">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194625FF8BFFE855A2EFBFD348E85D" box="[756,837,255,276]" pageId="5" pageNumber="50" rank="species">
<emphasis id="B92AFB5EFF8BFFE855A2EFBFD332E85D" bold="true" box="[756,831,255,276]" pageId="5" pageNumber="50">sp. nov</emphasis>
.
</taxonomicNameLabel>
Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="8BE1274CFF8BFFE85678EE70D54CEDDD" pageId="5" pageNumber="50">
<table id="F95ED5ECFF8B001257C1EE70D580EDDD" box="[151,1421,304,1172]" gridcols="7" gridrows="8" pageId="5" pageNumber="50">
<tr id="356E250EFF8B001257C1EE70D580E8CB" box="[151,1421,304,386]" gridrow="0" pageId="5" pageNumber="50" rowspan-0="1">
<th id="76BF4C72FF8B00125678EE70D18BE8CB" box="[302,390,304,386]" gridcol="1" gridrow="0" pageId="5" pageNumber="50">AMS R 173127*</th>
<th id="76BF4C72FF8B001256EAEE70D32FE8CB" box="[444,802,304,386]" gridcol="2" gridrow="0" pageId="5" pageNumber="50">UNS 00103/ UNS 00104/ NCSM AMS R AMS R 77318 173128 173129</th>
<th id="76BF4C72FF8B0012543FEE70D3A2E8CB" box="[873,943,304,386]" gridcol="3" gridrow="0" pageId="5" pageNumber="50">ZFMK 91076</th>
<th id="76BF4C72FF8B001254A1EE70D44BE8CB" box="[1015,1094,304,386]" gridcol="4" gridrow="0" pageId="5" pageNumber="50">AMS R 173126</th>
<th id="76BF4C72FF8B001253D0EE70D4D9E8CB" box="[1158,1236,304,386]" gridcol="5" gridrow="0" pageId="5" pageNumber="50">AMS R 173507</th>
<th id="76BF4C72FF8B00125243EE70D580E8CB" box="[1301,1421,304,386]" gridcol="6" gridrow="0" pageId="5" pageNumber="50">UNS 0 0 105</th>
</tr>
<tr id="356E250EFF8B001257C1EED6D580E8E5" box="[151,1421,406,428]" gridrow="1" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1EED6D0F2E8E5" box="[151,255,406,428]" gridcol="0" gridrow="1" pageId="5" pageNumber="50">Sex</th>
<td id="76BF4C72FF8B00125678EED6D18BE8E5" box="[302,390,406,428]" gridcol="1" gridrow="1" pageId="5" pageNumber="50">Male</td>
<td id="76BF4C72FF8B001256EAEED6D32FE8E5" box="[444,802,406,428]" gridcol="2" gridrow="1" pageId="5" pageNumber="50">Male Male Male</td>
<td id="76BF4C72FF8B0012543FEED6D3A2E8E5" box="[873,943,406,428]" gridcol="3" gridrow="1" pageId="5" pageNumber="50">Male</td>
<td id="76BF4C72FF8B001254A1EED6D44BE8E5" box="[1015,1094,406,428]" gridcol="4" gridrow="1" pageId="5" pageNumber="50">Female</td>
<td id="76BF4C72FF8B001253D0EED6D4D9E8E5" box="[1158,1236,406,428]" gridcol="5" gridrow="1" pageId="5" pageNumber="50">Female</td>
<td id="76BF4C72FF8B00125243EED6D580E8E5" box="[1301,1421,406,428]" gridcol="6" gridrow="1" pageId="5" pageNumber="50">Female</td>
</tr>
<tr id="356E250EFF8B001257C1EE80D580EB18" box="[151,1421,448,593]" gridrow="2" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1EE80D0F2EB18" box="[151,255,448,593]" gridcol="0" gridrow="2" pageId="5" pageNumber="50">SVL HDL HDW SNT</th>
<td id="76BF4C72FF8B00125678EE80D18BEB18" box="[302,390,448,593]" gridcol="1" gridrow="2" pageId="5" pageNumber="50">43.6 14.6 16.3 5.5</td>
<td id="76BF4C72FF8B001256EAEE80D32FEB18" box="[444,802,448,593]" gridcol="2" gridrow="2" pageId="5" pageNumber="50">42.5 44.8 44.2 14.9 15.2 15.3 17.0 17.3 17.0 5.8 6.2 6.3</td>
<td id="76BF4C72FF8B0012543FEE80D3A2EB18" box="[873,943,448,593]" gridcol="3" gridrow="2" pageId="5" pageNumber="50">43.3 14.4 17.1 5.8</td>
<td id="76BF4C72FF8B001254A1EE80D44BEB18" box="[1015,1094,448,593]" gridcol="4" gridrow="2" pageId="5" pageNumber="50">38.9 15.0 16.8 5.5</td>
<td id="76BF4C72FF8B001253D0EE80D4D9EB18" box="[1158,1236,448,593]" gridcol="5" gridrow="2" pageId="5" pageNumber="50">41.4 15.3 16.6 6.1</td>
<td id="76BF4C72FF8B00125243EE80D580EB18" box="[1301,1421,448,593]" gridcol="6" gridrow="2" pageId="5" pageNumber="50">53.4 18.0 20.8 7.3</td>
</tr>
<tr id="356E250EFF8B001257C1ED25D580EBED" box="[151,1421,613,676]" gridrow="3" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1ED25D0F2EBED" box="[151,255,613,676]" gridcol="0" gridrow="3" pageId="5" pageNumber="50">EYE IOD</th>
<td id="76BF4C72FF8B00125678ED25D18BEBED" box="[302,390,613,676]" gridcol="1" gridrow="3" pageId="5" pageNumber="50">4.6 4.8</td>
<td id="76BF4C72FF8B001256EAED25D32FEBED" box="[444,802,613,676]" gridcol="2" gridrow="3" pageId="5" pageNumber="50">4.6 4.5 4.7 6.1 5.6 4.7</td>
<td id="76BF4C72FF8B0012543FED25D3A2EBED" box="[873,943,613,676]" gridcol="3" gridrow="3" pageId="5" pageNumber="50">4.5 5.5</td>
<td id="76BF4C72FF8B001254A1ED25D44BEBED" box="[1015,1094,613,676]" gridcol="4" gridrow="3" pageId="5" pageNumber="50">4.2 4.9</td>
<td id="76BF4C72FF8B001253D0ED25D4D9EBED" box="[1158,1236,613,676]" gridcol="5" gridrow="3" pageId="5" pageNumber="50">4.7 5.2</td>
<td id="76BF4C72FF8B00125243ED25D580EBED" box="[1301,1421,613,676]" gridcol="6" gridrow="3" pageId="5" pageNumber="50">5.3 5.1</td>
</tr>
<tr id="356E250EFF8B001257C1EDF8D580EA69" box="[151,1421,696,800]" gridrow="4" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1EDF8D0F2EA69" box="[151,255,696,800]" gridcol="0" gridrow="4" pageId="5" pageNumber="50">TMP TEY IN</th>
<td id="76BF4C72FF8B00125678EDF8D18BEA69" box="[302,390,696,800]" gridcol="1" gridrow="4" pageId="5" pageNumber="50">1.9 1.6 3.8</td>
<td id="76BF4C72FF8B001256EAEDF8D32FEA69" box="[444,802,696,800]" gridcol="2" gridrow="4" pageId="5" pageNumber="50">2.4 2.0 2.2 1.2 1.6 1.6 4.1 3.6 4.2</td>
<td id="76BF4C72FF8B0012543FEDF8D3A2EA69" box="[873,943,696,800]" gridcol="3" gridrow="4" pageId="5" pageNumber="50">2.7 1.3 4.5</td>
<td id="76BF4C72FF8B001254A1EDF8D44BEA69" box="[1015,1094,696,800]" gridcol="4" gridrow="4" pageId="5" pageNumber="50">2.6 1.5 3.8</td>
<td id="76BF4C72FF8B001253D0EDF8D4D9EA69" box="[1158,1236,696,800]" gridcol="5" gridrow="4" pageId="5" pageNumber="50">2.5 2.0 3.9</td>
<td id="76BF4C72FF8B00125243EDF8D580EA69" box="[1301,1421,696,800]" gridcol="6" gridrow="4" pageId="5" pageNumber="50">2.7 2.0 4.8</td>
</tr>
<tr id="356E250EFF8B001257C1EC74D580EAD5" box="[151,1421,820,924]" gridrow="5" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1EC74D0F2EAD5" box="[151,255,820,924]" gridcol="0" gridrow="5" pageId="5" pageNumber="50">NS EN TIB</th>
<td id="76BF4C72FF8B00125678EC74D18BEAD5" box="[302,390,820,924]" gridcol="1" gridrow="5" pageId="5" pageNumber="50">1.2 3.6 20.7</td>
<td id="76BF4C72FF8B001256EAEC74D32FEAD5" box="[444,802,820,924]" gridcol="2" gridrow="5" pageId="5" pageNumber="50">2.1 2.4 2.7 3.9 3.9 3.6 20.7 21.3 20.7</td>
<td id="76BF4C72FF8B0012543FEC74D3A2EAD5" box="[873,943,820,924]" gridcol="3" gridrow="5" pageId="5" pageNumber="50">2.3 3.5 20.5</td>
<td id="76BF4C72FF8B001254A1EC74D44BEAD5" box="[1015,1094,820,924]" gridcol="4" gridrow="5" pageId="5" pageNumber="50">1.7 3.6 20.1</td>
<td id="76BF4C72FF8B001253D0EC74D4D9EAD5" box="[1158,1236,820,924]" gridcol="5" gridrow="5" pageId="5" pageNumber="50">2.0 3.7 21.1</td>
<td id="76BF4C72FF8B00125243EC74D580EAD5" box="[1301,1421,820,924]" gridcol="6" gridrow="5" pageId="5" pageNumber="50">3.0 4.4 26.4</td>
</tr>
<tr id="356E250EFF8B001257C1ECF0D580ED0B" box="[151,1421,944,1090]" gridrow="6" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1ECF0D0F2ED0B" box="[151,255,944,1090]" gridcol="0" gridrow="6" pageId="5" pageNumber="50">ML PL IML HW:HL</th>
<td id="76BF4C72FF8B00125678ECF0D18BED0B" box="[302,390,944,1090]" gridcol="1" gridrow="6" pageId="5" pageNumber="50">13.2 19.1 2.4 1.12</td>
<td id="76BF4C72FF8B001256EAECF0D32FED0B" box="[444,802,944,1090]" gridcol="2" gridrow="6" pageId="5" pageNumber="50">12.5 13.1 14.4 17.4 17.4 19.3 1.9 1.9 2.1 1.14 1.14 1.11</td>
<td id="76BF4C72FF8B0012543FECF0D3A2ED0B" box="[873,943,944,1090]" gridcol="3" gridrow="6" pageId="5" pageNumber="50">13.8 18.0 1.6 1.19</td>
<td id="76BF4C72FF8B001254A1ECF0D44BED0B" box="[1015,1094,944,1090]" gridcol="4" gridrow="6" pageId="5" pageNumber="50">12.2 17.6 2.0 1.12</td>
<td id="76BF4C72FF8B001253D0ECF0D4D9ED0B" box="[1158,1236,944,1090]" gridcol="5" gridrow="6" pageId="5" pageNumber="50">13.8 18.2 1.6 1.0 8</td>
<td id="76BF4C72FF8B00125243ECF0D580ED0B" box="[1301,1421,944,1090]" gridcol="6" gridrow="6" pageId="5" pageNumber="50">16.8 23.1 2.4 1.16</td>
</tr>
<tr id="356E250EFF8B001257C1EB16D580EDDD" box="[151,1421,1110,1172]" gridrow="7" pageId="5" pageNumber="50">
<th id="76BF4C72FF8B001257C1EB16D0F2EDDD" box="[151,255,1110,1172]" gridcol="0" gridrow="7" pageId="5" pageNumber="50">HL:HW TIB:SVL</th>
<td id="76BF4C72FF8B00125678EB16D18BEDDD" box="[302,390,1110,1172]" gridcol="1" gridrow="7" pageId="5" pageNumber="50">0.9 0.47</td>
<td id="76BF4C72FF8B001256EAEB16D32FEDDD" box="[444,802,1110,1172]" gridcol="2" gridrow="7" pageId="5" pageNumber="50">0.88 0.87 0.9 0.49 0.47 0.47</td>
<td id="76BF4C72FF8B0012543FEB16D3A2EDDD" box="[873,943,1110,1172]" gridcol="3" gridrow="7" pageId="5" pageNumber="50">0.84 0.47</td>
<td id="76BF4C72FF8B001254A1EB16D44BEDDD" box="[1015,1094,1110,1172]" gridcol="4" gridrow="7" pageId="5" pageNumber="50">0.89 0.52</td>
<td id="76BF4C72FF8B001253D0EB16D4D9EDDD" box="[1158,1236,1110,1172]" gridcol="5" gridrow="7" pageId="5" pageNumber="50">0.92 0.51</td>
<td id="76BF4C72FF8B00125243EB16D580EDDD" box="[1301,1421,1110,1172]" gridcol="6" gridrow="7" pageId="5" pageNumber="50">0.87 0.49</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE1274CFF8BFFE857C9EBE8D510EC59" blockId="5.[159,1346,1192,1296]" pageId="5" pageNumber="50">HDL:SVL 0.33 0.35 0.34 0.35 0.33 0.38 0.37 0.34 TMP:EYE 0.41 0.53 0.45 0.47 0.60 0.61 0.54 0.51 Weight (g) 3.9 - - - - 3.5 4.0 -</paragraph>
<paragraph id="8BE1274CFF8BFFE857C1EA71D0F1EC0E" blockId="5.[151,252,1329,1351]" box="[151,252,1329,1351]" pageId="5" pageNumber="50">
*
<typeStatus id="54E599EEFF8BFFE857F6EA71D0F1EC0E" box="[160,252,1329,1351]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="8BE1274CFF8BFFE85793EA34D222EF76" blockId="5.[151,1436,1396,2033]" pageId="5" pageNumber="50">
<emphasis id="B92AFB5EFF8BFFE85793EA34D125ECC5" bold="true" box="[197,296,1396,1420]" pageId="5" pageNumber="50">Tadpole.</emphasis>
Tadpoles were assigned to the new species based upon the extremely low sequence divergences between the tadpole and adults frogs (0.0 0.6% at the 16S rRNA gene). The tadpole is elongate, with a tail length about 3 times body length, and the tail is about as tall as the body. The tadpole has a greatly reduced oral disc, only an upper jaw sheath, and most notably, a pair of keratinized hooks on the edge of the lower labium that face away from the mouth. Colour in life dark brownish grey, slightly paler in preservative. A detailed description of the new tadpole will be published separately.
</paragraph>
<paragraph id="8BE1274CFF8BFFE85793E90CD47DEF86" blockId="5.[151,1436,1396,2033]" pageId="5" pageNumber="50">
<emphasis id="B92AFB5EFF8BFFE85793E90CD124EF2D" bold="true" box="[197,297,1612,1636]" pageId="5" pageNumber="50">Ecology.</emphasis>
The species is a phytotelm breeder, depositing foam nests in small tree-holes, away from streams or ponds. Between May-July 2010, we observed eight foam nests and/or groups of tadpoles consistent with the new species, all in water-filled tree-holes, 0.3
<quantity id="4CA68AA9FF8BFFE8553BE9D4D2A3EFE5" box="[621,686,1684,1708]" metricMagnitude="0" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="50" unit="m" value="1.2">1.2 m</quantity>
above the ground. Adults of the new species were also adjacent to these tree-holes on several occasions. The advertisement call of the species is unknown.
</paragraph>
<paragraph id="8BE1274CFF8BFFE85793E99BD508EECD" blockId="5.[151,1436,1396,2033]" pageId="5" pageNumber="50">
<emphasis id="B92AFB5EFF8BFFE85793E99BD1B6EFBD" bold="true" box="[197,443,1755,1780]" pageId="5" pageNumber="50">Conservation status.</emphasis>
<taxonomicName id="4C5E5CCFFF8BFFE85692E99CD2C2EFBD" box="[452,719,1756,1780]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="vampyrus">
<emphasis id="B92AFB5EFF8BFFE85692E99CD2C2EFBD" box="[452,719,1756,1780]" italics="true" pageId="5" pageNumber="50">Rhacophorus vampyrus</emphasis>
</taxonomicName>
is known only from Bidoup-Nui Ba National Park. The actual distribution of the new species is unknown but probably extends into montane evergreen forest in the Langbian Plateau, including Chu Yang
<collectingCountry id="F34967DCFF8BFFE8568BE864D20FEE75" box="[477,514,1828,1852]" name="China" pageId="5" pageNumber="50">Sin</collectingCountry>
National Park in Dak Lak Province, and Phuoc Binh National Park in Ninh Thuan Province, which are continuous with Bidoup-Nui Ba National Park. Given the available information, we suggest the species should be considered Data Deficient following IUCNs Red List categories (IUCN 2001).
</paragraph>
<paragraph id="8BE1274CFF8BFFE85793E8CFD235EEB9" blockId="5.[151,1436,1396,2033]" pageId="5" pageNumber="50">
<emphasis id="B92AFB5EFF8BFFE85793E8CFD166EEEE" bold="true" box="[197,363,1935,1960]" pageId="5" pageNumber="50">Comparisons.</emphasis>
The pale tan to brick red dorsum, white venter, and mostly black flanks, ventral surface of upper arms and crus, and anterior and posterior surface of thighs distinguishes the new species from all mainland Southeast Asian congeners (Table 2).
</paragraph>
<paragraph id="8BE1274CFF88FFEB5721ED2CD558EB36" blockId="6.[119,1662,620,1569]" box="[119,1365,620,639]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB5721ED2CD0DFEB36" bold="true" box="[119,210,620,639]" pageId="6" pageNumber="51">TABLE 2</emphasis>
.
<taxonomicName id="4C5E5CCFFF88FFEB5788ED2CD159EB36" box="[222,340,620,639]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B92AFB5EFF88FFEB5788ED2CD159EB36" box="[222,340,620,639]" italics="true" pageId="6" pageNumber="51">Rhacophorus</emphasis>
</taxonomicName>
species known from mainland Southeast Asia. Fields that differ to those of the new species are highlighted in grey.
</paragraph>
<paragraph id="8BE1274CFF88FFEB521BEDC6D5E7EBD1" blockId="6.[119,1662,620,1569]" box="[1357,1514,646,664]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB521BEDC6D5E7EBD1" bold="true" box="[1357,1514,646,664]" pageId="6" pageNumber="51">Pointed projection</emphasis>
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EDDDD673EBBB" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EDDDD64EEBE6" bold="true" box="[129,1603,669,687]" pageId="6" pageNumber="51">Species Dorsum colour Ventral colour Pattern on sides of body Dorsal colour of webbing Finger webbing on heel Source</emphasis>
<bibRefCitation id="EFCF5ABDFF88FFEB515FEDF4D665EB8C" author="Smith" box="[1545,1640,692,709]" pageId="6" pageNumber="51" refString="Smith, M. A. (1924) New tree-frogs from Indo-China and the Malay Peninsula. Proceedings of the Zoological Society of London 1924, 225 - 234." type="proceedings paper" year="1924">Smith 1924</bibRefCitation>
; variable, white and/or yellow; Inger
<emphasis id="B92AFB5EFF88FFEB516EED8CD651EB95" box="[1592,1628,715,732]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
. brown with dark scattered black/grey/brown variable; dark greyish 1999; JR pers.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EDB9D621EA43" blockId="6.[119,1662,620,1569]" box="[129,1580,761,778]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7EDB9D0D2EA40" box="[129,223,761,777]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="annamensis">
<emphasis id="B92AFB5EFF88FFEB57D7EDB9D0D2EA40" box="[129,223,761,777]" italics="true" pageId="6" pageNumber="51">annamensis</emphasis>
</taxonomicName>
markings white spots brown/black complete absent obs.
</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EC50D659EA71" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
greyish brown with
<bibRefCitation id="EFCF5ABDFF88FFEB515FEC50D677EA68" author="Gunther" box="[1545,1658,784,801]" pageId="6" pageNumber="51" refString="Gunther, A. C. L. G. (1858) Neue Batrachier in der Sammlung des britischen Museums. Archiv fur Naturgeschichte, 24, 319 - 328." type="journal article" year="1858">Günther 1858</bibRefCitation>
; dark obscure cream coloured; no absent (narrow flap Brown &amp;
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EC7DD670EA35" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EC7DD0F6EA07" box="[129,251,829,846]" italics="true" pageId="6" pageNumber="51">appendiculatus</emphasis>
mottling spots reduced only)
<bibRefCitation id="EFCF5ABDFF88FFEB515FEC7ED667EA06" author="Alcala" box="[1545,1642,830,847]" pageId="6" pageNumber="51" refString="Brown, W. C. &amp; Alcala, A. C. (1994) Philippine frogs of the family Rhacophoridae. Proceedings of the Californian Academy of Sciences 48, 185 - 220." type="journal article" year="1994">Alcala 1994</bibRefCitation>
<bibRefCitation id="EFCF5ABDFF88FFEB515FEC15D656EA2F" author="Ahl" box="[1545,1627,853,870]" pageId="6" pageNumber="51" refString="Ahl, E. (1927) Zur Systematik der asiatischen Arten der Froschgattung Rhacophorus. Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1927, 35 - 47." type="journal article" year="1927">Ahl, 1927</bibRefCitation>
; Bordoloi
<emphasis id="B92AFB5EFF88FFEB5103EC2CD675EA35" box="[1621,1656,875,892]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7ECC2D63CEADD" blockId="6.[119,1662,620,1569]" box="[129,1585,897,916]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7ECC2D0D3EADA" box="[129,222,898,915]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B92AFB5EFF88FFEB57D7ECC2D0D3EADA" box="[129,222,898,915]" italics="true" pageId="6" pageNumber="51">bipunctatus</emphasis>
</taxonomicName>
green yellow 13 black spots red reduced present 2007
</paragraph>
<paragraph id="8BE1274CFF88FFEB5685ECD9D272EAE3" blockId="6.[119,1662,620,1569]" box="[467,639,921,938]" pageId="6" pageNumber="51">chocolate brown with</paragraph>
<paragraph id="8BE1274CFF88FFEB5685ECF0D664EA91" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">white markings Andersson anteriorly, whitish 1939; Ohler</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EC9ED63CEAA6" blockId="6.[119,1662,620,1569]" box="[129,1585,988,1007]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EC9ED0DCEAA6" box="[129,209,990,1007]" italics="true" pageId="6" pageNumber="51">burmanus</emphasis>
green posteriorly* reduced absent 2009
</paragraph>
<paragraph id="8BE1274CFF88FFEB5648ECB5D1A9ED4C" blockId="6.[119,1662,620,1569]" box="[286,420,1013,1030]" pageId="6" pageNumber="51">olive/pale green;</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EB4CD396ED54" blockId="6.[119,1662,620,1569]" box="[286,923,1036,1053]" pageId="6" pageNumber="51">small pale yellow yellow, 2 black spots (one side</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EB63D66BED7D" blockId="6.[119,1662,620,1569]" box="[129,1638,1058,1076]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7EB63D0DCED7D" box="[129,209,1059,1076]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="calcaneus">
<emphasis id="B92AFB5EFF88FFEB57D7EB63D0DCED7D" box="[129,209,1059,1076]" italics="true" pageId="6" pageNumber="51">calcaneus</emphasis>
</taxonomicName>
dots pale yellow only) yellow/olive, dark spots reduced present
<bibRefCitation id="EFCF5ABDFF88FFEB515FEB63D66BED7D" author="Smith" box="[1545,1638,1059,1076]" pageId="6" pageNumber="51" refString="Smith, M. A. (1924) New tree-frogs from Indo-China and the Malay Peninsula. Proceedings of the Zoological Society of London 1924, 225 - 234." type="proceedings paper" year="1924">Smith 1924</bibRefCitation>
</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EB7AD185ED03" blockId="6.[119,1662,620,1569]" box="[286,392,1082,1099]" pageId="6" pageNumber="51">dark green or</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EB11D668ED28" blockId="6.[119,1662,620,1569]" box="[286,1637,1105,1122]" pageId="6" pageNumber="51">
brown; small pale bright yellow with 2 black Orlov
<emphasis id="B92AFB5EFF88FFEB516BEB11D66DED2B" box="[1597,1632,1105,1122]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EB27D63CED30" blockId="6.[119,1662,620,1569]" box="[129,1585,1127,1145]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7EB27D109ED31" box="[129,260,1127,1144]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="chuyangsinensis">
<emphasis id="B92AFB5EFF88FFEB57D7EB27D109ED31" box="[129,260,1127,1144]" italics="true" pageId="6" pageNumber="51">chuyangsinensis</emphasis>
</taxonomicName>
spots bright yellow spots bright yellow reduced present 2008
</paragraph>
<paragraph id="8BE1274CFF88FFEB5685EB3ED28FEDC6" blockId="6.[119,1662,620,1569]" box="[467,642,1150,1167]" pageId="6" pageNumber="51">throat and chest white</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EBD5D36CEDEC" blockId="6.[119,1662,620,1569]" box="[286,865,1173,1190]" pageId="6" pageNumber="51">sand-coloured with with brown speckles; posterior dark brown to</paragraph>
<paragraph id="8BE1274CFF88FFEB5648EBECD669EDF4" blockId="6.[119,1662,620,1569]" box="[286,1636,1196,1213]" pageId="6" pageNumber="51">cinnamon coloured belly light yellow to blackish with iridescent blue Manthey &amp;</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EB84D670EC51" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7EB84D0F3ED9D" box="[129,254,1220,1236]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="cyanopunctatus">
<emphasis id="B92AFB5EFF88FFEB57D7EB84D0F3ED9D" box="[129,254,1220,1236]" italics="true" pageId="6" pageNumber="51">cyanopunctatus</emphasis>
</taxonomicName>
cross bars dark down the flanks spots reduced absent Seiof 1998
<bibRefCitation id="EFCF5ABDFF88FFEB515FEB9AD63EEC4B" author="Blanford" box="[1545,1616,1242,1282]" pageId="6" pageNumber="51" refString="Blanford, W. T. (1881) On a collection of reptiles and frogs chiefly from Singapore. Proceedings of the Zoological Society of London, 1881, 215 - 226" type="proceedings paper" year="1881">Blanford 1881</bibRefCitation>
; dirty white, mottled Bordoloi
<emphasis id="B92AFB5EFF88FFEB5103EA48D675EC50" box="[1621,1656,1288,1305]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EA5FD63CEC79" blockId="6.[119,1662,620,1569]" box="[129,1585,1311,1328]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EA5FD0B3EC79" box="[129,190,1311,1328]" italics="true" pageId="6" pageNumber="51">dennysi</emphasis>
green with dusky none whitish reduced absent 2007
</paragraph>
<paragraph id="8BE1274CFF88FFEB55F3EA76D362EC0E" blockId="6.[119,1662,620,1569]" box="[677,879,1334,1351]" pageId="6" pageNumber="51">white with variable black</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EA0CD669ECC9" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EA0CD0ECEC14" box="[129,225,1356,1373]" italics="true" pageId="6" pageNumber="51">dorsoviridis</emphasis>
green white spotting reduced absent
<bibRefCitation id="EFCF5ABDFF88FFEB515FEA0DD67EEC14" author="Bourret" box="[1545,1651,1356,1373]" pageId="6" pageNumber="51" refString="Bourret, R. (1937) Notes herpetologiques sur lIndochine francaise. XIV. Les batraciens de la collection du Laboratoire des Sciences Naturelles de lUniversite. Descriptions de quinze especes ou varietes nouvelles. Annexe au Bulletin General de l'Instruction Publique. Hanoi, 1937, 5 - 56." type="journal article" year="1937">Bourret1937</bibRefCitation>
fleshy with brown lower flanks blackish with Ohler
<emphasis id="B92AFB5EFF88FFEB516AEA30D652ECC8" box="[1596,1631,1392,1409]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EAC7D66DEC8C" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EAC7D0B1ECD1" box="[129,188,1415,1432]" italics="true" pageId="6" pageNumber="51">duboisi</emphasis>
dark olive green spots white spots marbled black and white reduced absent 2000 green, with round
<bibRefCitation id="EFCF5ABDFF88FFEB515FEADED66AECE6" author="David" box="[1545,1639,1438,1455]" pageId="6" pageNumber="51" refString="David, A. (1872 &quot; 1871 &quot;) Rapport adresse a MM. les Professeurs-Administrateurs du Museum d'histoire naturelle. Nouvelles Archives du Museum d'Histoire Naturelle, 7, 75 - 100." type="journal article" year="1872">David 1872</bibRefCitation>
spots of a golden cream yellow, spotted &quot;1871; Liu
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7EA8BD673EF40" blockId="6.[119,1662,620,1569]" pageId="6" pageNumber="51">
<emphasis id="B92AFB5EFF88FFEB57D7EA8BD0CDEC95" box="[129,192,1483,1500]" italics="true" pageId="6" pageNumber="51">dugritei</emphasis>
metallic ash with dark grey marbled with cream yellow reduced absent 1950 mottled grey, Inger
<emphasis id="B92AFB5EFF88FFEB516EEAA3D651ECBA" box="[1592,1628,1506,1523]" italics="true" pageId="6" pageNumber="51">et al</emphasis>
. brown, and olive white anteriorly; marbled back and white/pale absent (large 1999; JR pers.
</paragraph>
<paragraph id="8BE1274CFF88FFEB57D7E950D621EF68" blockId="6.[119,1662,620,1569]" box="[129,1580,1551,1569]" pageId="6" pageNumber="51">
<taxonomicName id="4C5E5CCFFF88FFEB57D7E950D0EAEF68" box="[129,231,1552,1569]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="51" phylum="Arthropoda" rank="species" species="exechopygus">
<emphasis id="B92AFB5EFF88FFEB57D7E950D0EAEF68" box="[129,231,1552,1569]" italics="true" pageId="6" pageNumber="51">exechopygus</emphasis>
</taxonomicName>
green yellow posteriorly blue as dorsum complete tubercle only) obs.
</paragraph>
<caption id="DF2177C4FF86FFE557C1EBB3D214EC60" httpUri="https://zenodo.org/record/200073/files/figure.png" pageId="8" pageNumber="53" targetBox="[151,1436,193,1246]" targetPageId="8">
<paragraph id="8BE1274CFF86FFE557C1EBB3D214EC60" blockId="8.[151,1435,1267,1321]" pageId="8" pageNumber="53">
<emphasis id="B92AFB5EFF86FFE557C1EBB3D118EC40" bold="true" box="[151,277,1267,1290]" pageId="8" pageNumber="53">FIGURE 4.</emphasis>
Ventral surfaces of (A) right hand and (B) right foot of the male holotype of
<taxonomicName id="4C5E5CCFFF86FFE5531BEBB4D532EC43" box="[1101,1343,1268,1290]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="vampyrus" status="sp. nov.">
<emphasis id="B92AFB5EFF86FFE5531BEBB4D532EC43" box="[1101,1343,1268,1290]" italics="true" pageId="8" pageNumber="53">Rhacophorus vampyrus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2194625FF86FFE5521EEBB4D596EC40" box="[1352,1435,1268,1289]" pageId="8" pageNumber="53" rank="species">
<emphasis id="B92AFB5EFF86FFE5521EEBB4D596EC40" bold="true" box="[1352,1435,1268,1289]" pageId="8" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
(AMS R 173127). Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8BE1274CFF86FFE45793EA15D3C6E99D" blockId="8.[151,1436,1365,2038]" lastBlockId="9.[151,1436,151,212]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
The presence of a distinct pointed projection at the tibiotarsal articulation further distinguishes the new species from all mainland Southeast Asian congeners with the exception of
<emphasis id="B92AFB5EFF86FFE554FBEA39D3E7ECFC" italics="true" pageId="8" pageNumber="53">
<taxonomicName id="4C5E5CCFFF86FFE554FBEA39D4DEECD8" box="[941,1235,1401,1425]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="bipunctatus">Rhacophorus bipunctatus</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE553B6EA3AD578ECD8" box="[1248,1397,1401,1425]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="calcaneus">R. calcaneus</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE552D2EA3AD15FECFC" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="chuyangsinensis">R. chuyangsinensis</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE5560AEADED228ECFC" box="[348,549,1437,1461]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="hoanglienensis">R. hoanglienensis</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE55566EADED279ECFC" box="[560,628,1437,1461]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="kio">R. kio</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE55529EADED355ECFC" box="[639,856,1437,1461]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="marmoridorsum">R. marmoridorsum</taxonomicName>
,
<taxonomicName id="4C5E5CCFFF86FFE55434EADED3E7ECFC" box="[866,1002,1437,1461]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="rhodopus">R. rhodopus</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C5E5CCFFF86FFE55370EADED4A8ECFC" box="[1062,1189,1437,1461]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="B92AFB5EFF86FFE55370EADED4A8ECFC" box="[1062,1189,1437,1461]" italics="true" pageId="8" pageNumber="53">R. spelaeus</emphasis>
</taxonomicName>
(Table 2). From these species,
<taxonomicName id="4C5E5CCFFF86FFE557AEEA82D18AEC90" box="[248,391,1474,1497]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="vampyrus">
<emphasis id="B92AFB5EFF86FFE557AEEA82D18AEC90" box="[248,391,1474,1497]" italics="true" pageId="8" pageNumber="53">R. vampyrus</emphasis>
</taxonomicName>
can be distinguished further on the basis of colouration and extent of hand webbing.
<taxonomicName id="4C5E5CCFFF86FFE55233EA81D17EECB4" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="vampyrus">
<emphasis id="B92AFB5EFF86FFE55233EA81D17EECB4" italics="true" pageId="8" pageNumber="53">Rhacophorus vampyrus</emphasis>
</taxonomicName>
differs from:
<taxonomicName id="4C5E5CCFFF86FFE5555BEAA6D2A2ECB4" box="[525,687,1509,1533]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B92AFB5EFF86FFE5555BEAA6D2A2ECB4" box="[525,687,1509,1533]" italics="true" pageId="8" pageNumber="53">R. bipunctatus</emphasis>
</taxonomicName>
in having a pale tan to brick red dorsum (versus green), white ventral surface with black margins (versus yellow), black flanks (versus 1 3 black spots), and grey/black webbing on dorsal surface (versus red); from
<taxonomicName id="4C5E5CCFFF86FFE55544E96ED2AEEF0C" box="[530,675,1581,1605]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="calcaneus">
<emphasis id="B92AFB5EFF86FFE55544E96ED2AEEF0C" box="[530,675,1581,1605]" italics="true" pageId="8" pageNumber="53">R. calcaneus</emphasis>
</taxonomicName>
in having a pale tan to brick red dorsum (versus olive/pale green), white ventral surface with black margins (versus pale yellow), black flanks (versus yellow with black spots), and grey/black webbing on dorsal surface (versus yellow/olive with dark spots); from
<taxonomicName id="4C5E5CCFFF86FFE5536AE936D519EFC4" box="[1084,1300,1653,1677]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="chuyangsinensis">
<emphasis id="B92AFB5EFF86FFE5536AE936D519EFC4" box="[1084,1300,1653,1677]" italics="true" pageId="8" pageNumber="53">R. chuyangsinensis</emphasis>
</taxonomicName>
in having a pale tan to brick red dorsum (versus dark green or brown with small pale spots), white ventral surface with black margins (versus bright yellow), black flanks (versus bright yellow with black spots), and grey/black webbing on dorsal surface (versus bright yellow); from
<taxonomicName id="4C5E5CCFFF86FFE555D1E9A2D2C7EFB0" box="[647,714,1761,1785]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="kio">
<emphasis id="B92AFB5EFF86FFE555D1E9A2D2C7EFB0" box="[647,714,1761,1785]" italics="true" pageId="8" pageNumber="53">R. kio</emphasis>
</taxonomicName>
in having incomplete finger webbing (versus complete), a pale tan to brick red dorsum (versus green), white ventral surface with black margins (versus lemon yellow), black flanks (versus with one black spot), and grey/black webbing on dorsal surface (versus orange and black); from
<taxonomicName id="4C5E5CCFFF86FFE552D2E86AD140EE2C" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="marmoridorsum">
<emphasis id="B92AFB5EFF86FFE552D2E86AD140EE2C" italics="true" pageId="8" pageNumber="53">R. marmoridorsum</emphasis>
</taxonomicName>
in having a pale cream to brick red dorsum (versus flesh coloured with marble chocolate pattern), white ventral surface with black margins (versus light cream), and black flanks (versus irregular spots and lines of chocolate colour on cream background); from
<taxonomicName id="4C5E5CCFFF86FFE555FAE8D6D33BEEE4" box="[684,822,1941,1965]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="rhodopus">
<emphasis id="B92AFB5EFF86FFE555FAE8D6D33BEEE4" box="[684,822,1941,1965]" italics="true" pageId="8" pageNumber="53">R. rhodopus</emphasis>
</taxonomicName>
in having a white ventral surface with black margins (versus yellow), black flanks (versus 1 black spot), and grey/black webbing on dorsal surface (versus red); and from
<taxonomicName id="4C5E5CCFFF86FFE55783E89FD15BEEBF" box="[213,342,2014,2038]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="53" phylum="Arthropoda" rank="species" species="spelaeus">
<emphasis id="B92AFB5EFF86FFE55783E89FD15BEEBF" box="[213,342,2014,2038]" italics="true" pageId="8" pageNumber="53">R. spelaeus</emphasis>
</taxonomicName>
in having a pale tan to brick red dorsum (versus greyish brown with dark, irregular spots), white ventral surface with black margins (versus light grey with dark specks), black flanks (versus light grey), and grey/ black webbing on dorsal surface (versus grey-brown with dark speckles).
</paragraph>
</subSubSection>
</treatment>
</document>