103 lines
10 KiB
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103 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.238.3624" ID-PMC="PMC3496945" ID-Pensoft-Pub="1313-2970-238-57" ID-Pensoft-UUID="FFCDE657FFBEFFAEFF94FFFDA240FFB3" ID-PubMed="23226708" ID-Zenodo-Dep="577750" ID-ZooBank="928C2017C15A4F67956EB5747F3BF6D8" ModsDocID="1313-2970-238-57" checkinTime="1451248584044" checkinUser="pensoft" docAuthor="Zhang, Junfeng" docDate="2012" docId="3FB92F07289E5929BE576D5F9B2E8E8E" docLanguage="en" docName="ZooKeys 238: 57-76" docOrigin="ZooKeys 238" docPubDate="2012-11-06" docSource="http://dx.doi.org/10.3897/zookeys.238.3624" docTitle="Orientisargidae Zhang 2012, fam. n." docType="treatment" docUuid="6E5DC307-FFD3-48A6-895E-12047C8A9001" docUuidSource="ZooBank" docVersion="4" id="FFCDE657FFBEFFAEFF94FFFDA240FFB3" lastPageNumber="58" masterDocId="FFCDE657FFBEFFAEFF94FFFDA240FFB3" masterDocTitle="Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China" masterLastPageNumber="76" masterPageNumber="57" pageNumber="58" updateTime="1668154673256" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Orientisargidae fam. n., a new Jurassic family of Archisargoidea (Diptera, Brachycera), with review of Archisargidae from China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Junfeng</mods:namePart>
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<mods:affiliation>College of Palaeontology, Shenyang Normal University, Shenyang, 110034, P. R. China & Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, P. R. China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2012-11-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>238</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>57</mods:start>
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<mods:end>76</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.238.3624</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.238.3624</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-238-57</mods:identifier>
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<mods:identifier type="ZooBank">928C2017C15A4F67956EB5747F3BF6D8</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFCDE657FFBEFFAEFF94FFFDA240FFB3</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577750</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182235766" LSID="urn:lsid:zoobank.org:act:6E5DC307-FFD3-48A6-895E-12047C8A9001" httpUri="http://treatment.plazi.org/id/3FB92F07289E5929BE576D5F9B2E8E8E" lastPageNumber="58" pageId="1" pageNumber="58">
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<subSubSection pageId="1" pageNumber="58" type="nomenclature">
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<paragraph pageId="1" pageNumber="58">
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Family
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<taxonomicName LSID="urn:lsid:zoobank.org:act:6E5DC307-FFD3-48A6-895E-12047C8A9001" authority="Zhang, 2012" authorityName="Zhang" authorityYear="2012" class="Insecta" family="Orientisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family" status="fam. n.">Orientisargidae</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="58">fam. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="58" type="type genus">
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<paragraph pageId="1" pageNumber="58">Type genus.</paragraph>
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<paragraph pageId="1" pageNumber="58">
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<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Archisargidae" genus="Orientisargus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="1" pageNumber="58">Orientisargus</emphasis>
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</taxonomicName>
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gen. n.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="58" type="included genus">
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<paragraph pageId="1" pageNumber="58">Included genus.</paragraph>
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<paragraph pageId="1" pageNumber="58">The type genus only.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="58" type="diagnosis">
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<paragraph pageId="1" pageNumber="58">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="58">Large sized (more than 12 mm) flies. Legs and abdomen slender, strongly pubescent but devoid of bristles; antennal pedicel longest, first flagellomere neither segmented nor arista or style on its tip; wing narrow and long, subpetiolate, alula absent; venation not costalized; R pectinate; C running around entire wing margin although thinned beyond wing tip; C, Sc, R and CuA strong; Sc, R1 and R2+3 long; R4+5 simple (not bifurcated); origin of Rs proximal, nearly at level of M fork; discoidal cell slightly shifted distally; crossvein m-cu absent; M3+4 stem strongly flexed, and touching CuA; cell m3 closed; hind legs stout and long, tibial spurs well developed, empodium wanting; female cerci foliaceous.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="58" type="remarks">
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<paragraph pageId="1" pageNumber="58">Remarks.</paragraph>
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<paragraph pageId="1" pageNumber="58">
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This new family demonstrates similar body structures and wing venation to the family
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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Rohdendorf, 1962 based on the following characters: large sized flies with hind legs and abdomen stout and long, body strongly pubescent but devoid of bristles; wing narrow and long, subpetiolate, alula absent; venation: C running around wing margin although thinned beyond wing tip; long and strong Sc and R1 (R1 more than four-fifths of wing length, and clearly stouter than M); and the position of r-m, which meets R4+5 and fore margin of d, respectively; as well as the position of disciodal cell, which is more or less shifted distally. However, from all known representatives of
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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it differs by the simple R4+5, which is not bifurcated, the origin of Rs which is clearly proximal, the strongly flexed M3+4 stem, of which bM3+4 section becomes short, crossvein-like, and touches CuA instead of m-cu; Furthermore, considering the characteristic features that R4+5 is simple, R furcated pectinately and the origin of Rs is proximal,
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<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Orientisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Orientisargidae</taxonomicName>
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fam. n. is similar to the family
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<taxonomicName authorityName="Mostovski" authorityYear="1997" class="Insecta" family="Kovalevisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Kovalevisargidae</taxonomicName>
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Mostovski, 1997, the sister group of
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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within
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<taxonomicName authorityName="Rohdendorf" authorityYear="1962" genus="Archisargoidea" lsidName="" pageId="1" pageNumber="58" rank="genus">Archisargoidea</taxonomicName>
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. It is distinct from all the kovalevisargids in having longer Sc, R1 and R2+3, the closed cell m3, and the absence of m-cu. It is interesting that the long pedicel, the absence of arista or style on the tip of antenna, the absence of empodium may be the unique features of
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<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Orientisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Orientisargidae</taxonomicName>
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and are found neither in
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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nor in
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<taxonomicName authorityName="Mostovski" authorityYear="1997" class="Insecta" family="Kovalevisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Kovalevisargidae</taxonomicName>
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. It is also interesting that the new family has a pair of foliaceous cerci on the female terminalia, which is only present in
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<taxonomicName lsidName="" pageId="1" pageNumber="58" rank="subFamily" subFamily="Uranorhagioninae">Uranorhagioninae</taxonomicName>
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(=
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<taxonomicName authorityName="Zhang" authorityYear="2012" lsidName="" pageId="1" pageNumber="58" rank="subFamily" subFamily="Mostovskisarginae">Mostovskisarginae</taxonomicName>
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) within
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<taxonomicName class="Insecta" family="Archisargidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Diptera" pageId="1" pageNumber="58" phylum="Arthropoda" rank="family">Archisargidae</taxonomicName>
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(JF Zhang, 2010a).
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</paragraph>
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<paragraph pageId="1" pageNumber="58">
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As for m-cu is concerned, an alternative explanation is possible that a very short, but thick, rudimentary m-cu connecting CuA and flex point of M3+4 is present (see
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Orientisargus illecebrosus gen et sp. n., NIGP DHG 901 a, NIGP DHG 901 b, part and counterpart, holotype, photographs, female, dorsal view. A B body C antenna D left wing E right wing F part enlarged of left wing G part enlarged of right wing H tarsus of hind leg I abdominal apex of female. Scale bars represent 1 mm except for C and I for which scale bars represent 0.1 mm." httpUri="https://binary.pensoft.net/fig/23013" pageId="1" pageNumber="58">Figure 1F, G</figureCitation>
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). In such case, however, it becomes too short to measure.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |