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<document id="7955B399FB9FB4304EEB7D342D0BB0D7" ID-DOI="10.11646/zootaxa.4743.2.13" ID-GBIF-Dataset="1ff2d63e-6658-4645-8583-c901b133ddff" ID-ISSN="1175-5326" ID-Zenodo-Dep="3687738" ID-ZooBank="795EF592-C7FB-4D80-B599-0E196B770E78" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582709255748" checkinUser="plazi" docAuthor="Jiang, Ri-Xin &amp; Wang, Shuo" docDate="2020" docId="3F3E8788D209FF9B5596F89CDC39FD33" docLanguage="en" docName="zootaxa.4743.2.13.pdf" docOrigin="Zootaxa 4743 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Termitodiellus hainanensis Jiang &amp; Wang 2020, sp. nov." docType="treatment" docUuid="1855B904-86B3-4179-9679-C07B2D9CF85D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="292" masterDocId="C307FFF0D209FF985501FF95D865FFB5" masterDocTitle="Discovery of the genus Termitodiellus Nakane, 1961 in China, with description of Termitodiellus hainanensis Jiang &amp; Wang, new species (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="294" masterPageNumber="289" pageNumber="289" updateTime="1698721529990" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6DA9A1FC3B20323A98E87FF036B505E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E1317FCD0102D543FD4EF99E33FF993E">
<mods:title id="14FCAC4E16FF8C6A0C0C5BE81E5BFB62">Discovery of the genus Termitodiellus Nakane, 1961 in China, with description of Termitodiellus hainanensis Jiang &amp; Wang, new species (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
</mods:titleInfo>
<mods:name id="89EEA7E252AF2BF9F765A1DFE1A59D2A" type="personal">
<mods:role id="8D79D03E6C7CBA1B5D1CAC37E0B8E12C">
<mods:roleTerm id="29710C55930E397BB6BC671554CB25B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="30A4315786055F08DB1D770F03E4959F">Jiang, Ri-Xin</mods:namePart>
</mods:name>
<mods:name id="ED26F46FDFEB54403ADC99033BF5464A" type="personal">
<mods:role id="E8714F227C9DA3BB6866D6DB598FAA19">
<mods:roleTerm id="60CF566696A1B1EB0BC2317A6232A5F4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="42ACA6DF4B2EB03F994D6651639FA978">Wang, Shuo</mods:namePart>
</mods:name>
<mods:typeOfResource id="E7C6B2E4B1D2CD1C020105625AC5014B">text</mods:typeOfResource>
<mods:relatedItem id="7158711F99F47F5D1B0722B15E213B63" type="host">
<mods:titleInfo id="4DB624FB4DB8304FA9A48C66A8A5DD63">
<mods:title id="5CFE3C3F6EED3357D7C59522EB92DD4A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="33C5876D200AEEB954BC3048FEC94519">
<mods:date id="2DF464D43072552A8CF5543BA07B2A35">2020</mods:date>
<mods:detail id="5F34D06BF074542C32A554D87650A99E" type="pubDate">
<mods:number id="F197ACE8A6D0A755C8CA2EA7EB5BEA4F">2020-02-25</mods:number>
</mods:detail>
<mods:detail id="2E7004A45EA0950200FD898BD7783F4B" type="volume">
<mods:number id="4075732FEA02752183E68FF115EE633E">4743</mods:number>
</mods:detail>
<mods:detail id="B99F62C982D987042B5395BC526E6FB2" type="issue">
<mods:number id="0A734B0B687C876FCC47F0785CC5A234">2</mods:number>
</mods:detail>
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<mods:start id="EDDCF68B7404280A365B617F5A2BB8CB">289</mods:start>
<mods:end id="E04691490F83198454D0756DE9A79A70">294</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BE73033801A8DC7D09548665416825A6">journal article</mods:classification>
<mods:identifier id="F5A9F1E2615DFF491FA6E6400EAE5280" type="DOI">10.11646/zootaxa.4743.2.13</mods:identifier>
<mods:identifier id="21688AD689C3C16A5BA78C06A755E1D1" type="GBIF-Dataset">1ff2d63e-6658-4645-8583-c901b133ddff</mods:identifier>
<mods:identifier id="7C130E9DC762E1BBF41D0C312F6AA9C1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="634ED9003515AF6B94C3EF35D53FCBE8" type="Zenodo-Dep">3687738</mods:identifier>
<mods:identifier id="62FA102B0E94C5305F2248238FA5016A" type="ZooBank">795EF592-C7FB-4D80-B599-0E196B770E78</mods:identifier>
</mods:mods>
<treatment id="3F3E8788D209FF9B5596F89CDC39FD33" ID-DOI="http://doi.org/10.5281/zenodo.3687982" ID-GBIF-Taxon="162267531" ID-Zenodo-Dep="3687982" LSID="urn:lsid:zoobank.org:act:1855B904-86B3-4179-9679-C07B2D9CF85D" httpUri="http://treatment.plazi.org/id/3F3E8788D209FF9B5596F89CDC39FD33" lastPageId="3" lastPageNumber="292" pageId="0" pageNumber="289">
<subSubSection id="FF8D6515D209FF985596F89CDC04F896" box="[151,1121,1801,1828]" pageId="0" pageNumber="289" type="nomenclature">
<paragraph id="B728369ED209FF985596F89CDC04F896" blockId="0.[151,1436,1801,2043]" box="[151,1121,1801,1828]" pageId="0" pageNumber="289">
<heading id="EC6081F2D209FF985596F89CDC04F896" bold="true" box="[151,1121,1801,1828]" fontSize="11" level="1" pageId="0" pageNumber="289" reason="1">
<emphasis id="85E3EA8CD209FF985596F89CDC04F896" bold="true" box="[151,1121,1801,1828]" pageId="0" pageNumber="289">
<taxonomicName id="70974D1DD209FF985596F89CD9BAF891" authority="Jiang &amp; Wang, 2020" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[151,479,1801,1828]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="289" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="85E3EA8CD209FF985596F89CD9BAF891" bold="true" box="[151,479,1801,1828]" italics="true" pageId="0" pageNumber="289">Termitodiellus hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9ED057F7D209FF9854E7F89FDA26F891" box="[486,579,1802,1828]" pageId="0" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
Jiang &amp; Wang, new species (
<figureCitation id="2FAC2A1BD209FF9856AFF89CDC3DF896" box="[942,1112,1801,1827]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,244,1463,1485]" captionStartId-1="2.[151,244,1920,1942]" captionStartId-2="3.[151,244,1689,1711]" captionTargetBox-0="[166,1419,188,1427]" captionTargetBox-1="[168,1425,500,1879]" captionTargetBox-2="[158,1431,676,1654]" captionTargetId-0="figure@1.[151,1436,182,1436]" captionTargetId-1="figure@2.[151,1436,489,1893]" captionTargetId-2="figure@3.[151,1436,669,1663]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Habitus of Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype, male. A, dorsal view; B, ventral view." captionText-1="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." captionText-2="FIGURE 3. Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype (A, C) and Termitodiellus besucheti (Paulian, 1983) (B, D). A, B, pronotum, dorsal view; C, D, apex of elytra. Figs. B, D, were taken by Mr. Showtaro Kakizoe, specimen from Malaysia: Selangor, deposited in Showtaro Kakizoe private collection (Fukuoka, Japan)." figureDoi-0="http://doi.org/10.5281/zenodo.3687740" figureDoi-1="http://doi.org/10.5281/zenodo.3687742" figureDoi-2="http://doi.org/10.5281/zenodo.3687744" httpUri-0="https://zenodo.org/record/3687740/files/figure.png" httpUri-1="https://zenodo.org/record/3687742/files/figure.png" httpUri-2="https://zenodo.org/record/3687744/files/figure.png" pageId="0" pageNumber="289">Figs. 1–3A, C</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D209FF985596F8BAD946F806" pageId="0" pageNumber="289" type="materials_examined">
<paragraph id="B728369ED209FF985596F8BAD946F806" blockId="0.[151,1436,1801,2043]" pageId="0" pageNumber="289">
<emphasis id="85E3EA8CD209FF985596F8BAD952F8F2" bold="true" box="[151,311,1839,1863]" pageId="0" pageNumber="289">Type material</emphasis>
(
<specimenCount id="A191FD17D209FF985449F8BAD9EDF8F2" box="[328,392,1839,1863]" pageId="0" pageNumber="289" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="A191FD17D209FF985494F8BAD9B7F8F2" box="[405,466,1839,1863]" pageId="0" pageNumber="289" type="female">2 ♀♀</specimenCount>
).
<typeStatus id="682C883CD209FF9854F3F8BADA3DF8F2" box="[498,600,1839,1863]" pageId="0" pageNumber="289" type="holotype">Holotype</typeStatus>
,
<materialsCitation id="07FF3CC3D209FF985765F8A5DA03F83A" ID-GBIF-Occurrence="2571176116" collectingDate="2018-04-20" collectingDateMax="2018-04-21" collectingDateMin="2018-04-20" collectionCode="QUST" collectorName="Jiang Ri-Xin" country="China" county="Ledong County" elevation="1002" latitude="18.745277" location="Jianfengling National Forest" longLatPrecision="21" longitude="108.83889" municipality="Jianfengling National Forest Park" pageId="0" pageNumber="289" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<specimenCount id="A191FD17D209FF985765F8A5DA1FF8F2" box="[612,634,1840,1863]" pageId="0" pageNumber="289" type="male">♂</specimenCount>
, labeled “
<collectingCountry id="CF80760ED209FF9857E6F8BADB48F8F2" box="[743,813,1839,1863]" name="China" pageId="0" pageNumber="289">China</collectingCountry>
:
<collectingRegion id="7553F87CD209FF985639F8BADBEDF8F2" box="[824,904,1839,1863]" country="China" name="Hainan" pageId="0" pageNumber="289">Hainan</collectingRegion>
,
<collectingCounty id="5E494E12D209FF985692F8BADC59F8F2" box="[915,1084,1839,1863]" pageId="0" pageNumber="289">Ledong County</collectingCounty>
(
<emphasis id="85E3EA8CD209FF98514CF8B8DCF8F8F6" bold="true" box="[1101,1181,1837,1859]" pageId="0" pageNumber="289">mṛu</emphasis>
),
<collectingMunicipality id="574CACE4D209FF9851B5F8BAD977F8DE" pageId="0" pageNumber="289">
<location id="B2486045D209FF9851B5F8BAD8BFF8DE" LSID="urn:lsid:plazi:treatment:3F3E8788D209FF9B5596F89CDC39FD33:B2486045D209FF9851B5F8BAD8BFF8DE" country="China" county="Ledong County" latitude="18.745277" longLatPrecision="21" longitude="108.83889" municipality="Jianfengling National Forest Park" name="Jianfengling National Forest" pageId="0" pageNumber="289" stateProvince="Hainan">Jianfengling National Forest</location>
Park
</collectingMunicipality>
(
<emphasis id="85E3EA8CD209FF985423F8C4DA77F8D2" bold="true" box="[290,530,1873,1895]" pageId="0" pageNumber="289">ṪṞKOiẎŵẇẌẘ</emphasis>
),
<location id="B2486045D209FF985728F8C6DAC8F8DE" LSID="urn:lsid:plazi:treatment:3F3E8788D209FF9B5596F89CDC39FD33:B2486045D209FF985728F8C6DAC8F8DE" box="[553,685,1875,1899]" country="China" county="Ledong County" latitude="18.745277" longLatPrecision="21" longitude="108.83889" municipality="Jianfengling National Forest Park" name="Mingfenggu" pageId="0" pageNumber="289" stateProvince="Hainan">Mingfenggu</location>
(
<emphasis id="85E3EA8CD209FF9857BCF8C4DB68F8D2" bold="true" box="[701,781,1873,1895]" pageId="0" pageNumber="289">
<location id="B2486045D209FF9857BCF8C4DB68F8D2" LSID="urn:lsid:plazi:treatment:3F3E8788D209FF9B5596F89CDC39FD33:B2486045D209FF9857BCF8C4DB68F8D2" box="[701,781,1873,1895]" country="China" county="Ledong County" latitude="18.745277" longLatPrecision="21" longitude="108.83889" municipality="Jianfengling National Forest Park" name="Bmo" pageId="0" pageNumber="289" stateProvince="Hainan">Ḃṃỡ</location>
</emphasis>
),
<geoCoordinate id="D2A35059D209FF985625F8C7DBC3F8D9" box="[804,934,1874,1900]" degrees="18" direction="north" minutes="44" orientation="latitude" pageId="0" pageNumber="289" precision="15" seconds="43" value="18.745277">18°44′43″N</geoCoordinate>
,
<geoCoordinate id="D2A35059D209FF9856B0F8C7DC59F8D9" box="[945,1084,1874,1900]" degrees="108" direction="east" minutes="50" orientation="longitude" pageId="0" pageNumber="289" precision="15" seconds="20" value="108.83889">108°50′20″E</geoCoordinate>
, H:
<quantity id="706F9B7BD209FF98516BF8C6DC89F8DE" box="[1130,1260,1875,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.002" metricValueMax="1.048" metricValueMin="0.9559999999999998" pageId="0" pageNumber="289" unit="m" value="1002.0" valueMax="1048.0" valueMin="956.0">
<elevation id="3CBAD1ADD209FF98516BF8C6DC89F8DE" box="[1130,1260,1875,1899]" metricMagnitude="3" metricUnit="m" metricValue="1.002" metricValueMax="1.048" metricValueMin="0.9559999999999998" pageId="0" pageNumber="289" unit="m" value="1002.0" valueMax="1048.0" valueMin="956.0">956–1048m</elevation>
</quantity>
,
<date id="C329105ED209FF9851F9F8C6DDFDF8DE" box="[1272,1432,1875,1899]" pageId="0" pageNumber="289" value="2018-04-20" valueMax="2018-04-21" valueMin="2018-04-20">
<collectingDate id="D36DE9B6D209FF9851F9F8C6DDFDF8DE" box="[1272,1432,1875,1899]" pageId="0" pageNumber="289" value="2018-04-20" valueMax="2018-04-21" valueMin="2018-04-20">20–21.IV.2018</collectingDate>
</date>
, in rotten wood,
<collectorName id="1A625348D209FF985440F8E2D9AFF83A" box="[321,458,1911,1935]" pageId="0" pageNumber="289">Jiang Ri-Xin</collectorName>
leg.” (
<collectionCode id="D186AE5BD209FF985712F8E2DA3BF83A" box="[531,606,1911,1935]" pageId="0" pageNumber="289">QUST</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="07FF3CC3D209FF985771F8EDD945F806" ID-GBIF-Occurrence="2571176115" collectingDate="2018-04-20" collectingDateMax="2018-04-21" collectingDateMin="2018-04-20" collectionCode="MYNU, QUST" collectorName="Jiang Ri-Xin" country="China" county="Ledong County" elevation="1002" latitude="18.745277" location="Jianfengling National Forest" longLatPrecision="21" longitude="108.83889" municipality="Jianfengling National Forest Park" pageId="0" pageNumber="289" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<typeStatus id="682C883CD209FF985771F8EDDABFF825" box="[624,730,1912,1936]" pageId="0" pageNumber="289" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="A191FD17D209FF9857E4F8E2DB75F83A" box="[741,784,1911,1935]" pageId="0" pageNumber="289" type="male">1 ♂</specimenCount>
,
<specimenCount id="A191FD17D209FF98561AF8E2DB3CF83A" box="[795,857,1911,1935]" pageId="0" pageNumber="289" type="female">2 ♀♀</specimenCount>
, same data as for the holotype (
<specimenCount id="A191FD17D209FF9851B1F8E2DCBDF83A" box="[1200,1240,1911,1935]" pageId="0" pageNumber="289" type="male">1 ♂</specimenCount>
,
<specimenCount id="A191FD17D209FF9851E5F8E2DD69F83A" box="[1252,1292,1911,1935]" pageId="0" pageNumber="289" type="female">1 ♀</specimenCount>
in
<collectionCode id="D186AE5BD209FF98502EF8EDDDE1F83A" box="[1327,1412,1912,1935]" pageId="0" pageNumber="289">MYNU</collectionCode>
;
<specimenCount id="A191FD17D209FF98508EF8E2D8CEF806" pageId="0" pageNumber="289" type="female">1 ♀</specimenCount>
in
<collectionCode id="D186AE5BD209FF9855CCF80ED972F806" box="[205,279,1947,1971]" pageId="0" pageNumber="289">QUST</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D209FF9B55C6F82ADA7DFED0" lastPageId="3" lastPageNumber="292" pageId="0" pageNumber="289" type="description">
<paragraph id="B728369ED209FF9855C6F82AD8BDF84E" blockId="0.[151,1436,1801,2043]" pageId="0" pageNumber="289">
<emphasis id="85E3EA8CD209FF9855C6F82AD9B2F862" bold="true" box="[199,471,1983,2007]" pageId="0" pageNumber="289">
Description of
<typeStatus id="682C883CD209FF98546EF82AD9B7F862" box="[367,466,1983,2007]" pageId="0" pageNumber="289" type="holotype">holotype</typeStatus>
,
</emphasis>
male (
<figureCitation id="2FAC2A1BD209FF985721F82ADAFAF862" box="[544,671,1983,2007]" captionStart="FIGURE 1" captionStartId="1.[151,244,1463,1485]" captionTargetBox="[166,1419,188,1427]" captionTargetId="figure@1.[151,1436,182,1436]" captionTargetPageId="1" captionText="FIGURE 1. Habitus of Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype, male. A, dorsal view; B, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3687740" httpUri="https://zenodo.org/record/3687740/files/figure.png" pageId="0" pageNumber="289">Figs. 1A, B</figureCitation>
;
<figureCitation id="2FAC2A1BD209FF9857ABF82ADAD2F862" box="[682,695,1983,2007]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="0" pageNumber="289">2</figureCitation>
A–C, E–H; 3A, C). Small, dark brown, dorsal surface obviously carinated.
</paragraph>
<caption id="E3E86616D208FF995596FA22DD0CFA78" ID-DOI="http://doi.org/10.5281/zenodo.3687740" ID-Zenodo-Dep="3687740" box="[151,1385,1463,1485]" httpUri="https://zenodo.org/record/3687740/files/figure.png" pageId="1" pageNumber="290" startId="1.[151,244,1463,1485]" targetBox="[166,1419,188,1427]" targetPageId="1">
<paragraph id="B728369ED208FF995596FA22DD0CFA78" blockId="1.[151,1385,1463,1485]" box="[151,1385,1463,1485]" pageId="1" pageNumber="290">
<emphasis id="85E3EA8CD208FF995596FA22D968FA79" bold="true" box="[151,269,1463,1485]" pageId="1" pageNumber="290">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="70974D1DD208FF99547CFA22DB74FA79" authority="Jiang &amp; Wang" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[381,785,1463,1485]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="290" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="85E3EA8CD208FF99547CFA22DAE5FA79" box="[381,640,1463,1484]" italics="true" pageId="1" pageNumber="290">Termitodiellus hainanensis</emphasis>
Jiang &amp; Wang
</taxonomicName>
,
<emphasis id="85E3EA8CD208FF99561BFA22DBF5FA78" bold="true" box="[794,912,1463,1485]" pageId="1" pageNumber="290">
<taxonomicNameLabel id="9ED057F7D208FF99561BFA22DBF5FA78" box="[794,912,1463,1485]" pageId="1" pageNumber="290" rank="species">new species</taxonomicNameLabel>
</emphasis>
, holotype, male.
<emphasis id="85E3EA8CD208FF995136FA22DC2BFA79" bold="true" box="[1079,1102,1463,1484]" pageId="1" pageNumber="290">A,</emphasis>
dorsal view;
<emphasis id="85E3EA8CD208FF9951D1FA22DC83FA79" bold="true" box="[1232,1254,1463,1484]" pageId="1" pageNumber="290">B,</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="B728369ED208FF9955C6FA68DA4FF958" blockId="1.[151,1437,1533,2026]" pageId="1" pageNumber="290">Head traverse, about twice as wide as long, widest near middle. Dorsal surface crude, sparsely punctuated, most punctures with minute setae. Paramedian costae and discolateral costae longitudinal and interrupted, submarginal costae long and uninterrupted. Compound eyes not visible from above, anterior margin obviously sinuated on each side. Antenna of nine antennomeres, club trimerous. Antennomere I elongated and expanded, about twice as long as II; antennomere II expanded and short; antennomere III–VI similar, much short and traverse; antennomere VII–IX strongly expanded, antennomere VI and VIII traverse and near trapezoidal, antennomere IX with round apex. Frontodiscal costae long and interrupted, frontolateral costae short.</paragraph>
<paragraph id="B728369ED208FF9955C6F96FDA99F814" blockId="1.[151,1437,1533,2026]" pageId="1" pageNumber="290">
Pronotum (
<figureCitation id="2FAC2A1BD208FF995443F96CD9F2F8A4" box="[322,407,1785,1809]" captionStart="FIGURE 3" captionStartId="3.[151,244,1689,1711]" captionTargetBox="[158,1431,676,1654]" captionTargetId="figure@3.[151,1436,669,1663]" captionTargetPageId="3" captionText="FIGURE 3. Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype (A, C) and Termitodiellus besucheti (Paulian, 1983) (B, D). A, B, pronotum, dorsal view; C, D, apex of elytra. Figs. B, D, were taken by Mr. Showtaro Kakizoe, specimen from Malaysia: Selangor, deposited in Showtaro Kakizoe private collection (Fukuoka, Japan)." figureDoi="http://doi.org/10.5281/zenodo.3687744" httpUri="https://zenodo.org/record/3687744/files/figure.png" pageId="1" pageNumber="290">Fig. 3A</figureCitation>
) wider than long, with six longitudinal costae, all costae interrupted by transverse depression across middle area. Anterior part of costae shorter than posterior part, strongly elevated. All costae with small punctures at both sides, the area between the inner two costae with strong and big punctures. All punctures on pronotum with minute setae. Lateral margin with three lobes, anterior lobe big and strong, intermediate lobe bisected to two small lobes, and located at middle of pronotum, all lobes with round apex.
</paragraph>
<paragraph id="B728369ED208FF9A55C6F838DB2DFF4C" blockId="1.[151,1437,1533,2026]" lastBlockId="2.[151,1437,154,466]" lastPageId="2" lastPageNumber="291" pageId="1" pageNumber="290">
Elytra (
<figureCitation id="2FAC2A1BD208FF995416F838D90DF870" box="[279,360,1965,1989]" captionStart="FIGURE 3" captionStartId="3.[151,244,1689,1711]" captionTargetBox="[158,1431,676,1654]" captionTargetId="figure@3.[151,1436,669,1663]" captionTargetPageId="3" captionText="FIGURE 3. Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype (A, C) and Termitodiellus besucheti (Paulian, 1983) (B, D). A, B, pronotum, dorsal view; C, D, apex of elytra. Figs. B, D, were taken by Mr. Showtaro Kakizoe, specimen from Malaysia: Selangor, deposited in Showtaro Kakizoe private collection (Fukuoka, Japan)." figureDoi="http://doi.org/10.5281/zenodo.3687744" httpUri="https://zenodo.org/record/3687744/files/figure.png" pageId="1" pageNumber="290">Fig. 3C</figureCitation>
) widest at basal 1/3, gradually attenuated apically. Each elytron with three strongly elevated, narrow longitudinal costae, all three costae with double row of fine setae along summit, postdiscal bulbs strong and rounded, apex covered with long setae. Each posterior portion of humeral and marginal costae with single row of setae. Area between costae with two rows of big punctures. Caudal bulb with three protrusions, internal and external protrusion small, median protrusion much bigger, all protrusions covered with short setae.
</paragraph>
<paragraph id="B728369ED20BFF9A55C6FE93DC2FFE18" blockId="2.[151,1437,154,466]" pageId="2" pageNumber="291">
All legs sturdy. Profemora (
<figureCitation id="2FAC2A1BD20BFF9A54EAFE93DA59FEAB" box="[491,572,262,286]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="2" pageNumber="291">Fig. 2B</figureCitation>
) strongly expanded, widest near basal 1/3, ventral surface with a tuft of long setae at about basal 1/3. Apex of protibiae (
<figureCitation id="2FAC2A1BD20BFF9A572FFEBFDAE4FEF7" box="[558,641,298,322]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="2" pageNumber="291">Fig. 2B</figureCitation>
) with three small teeth, the lateral one larger than other two. Mesofemora and mesotibia (
<figureCitation id="2FAC2A1BD20BFF9A543EFEDBD9F6FED3" box="[319,403,334,358]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="2" pageNumber="291">Fig. 2C</figureCitation>
) expanded, apex of mesotibia with a large projection and a small tooth. Metafemora (
<figureCitation id="2FAC2A1BD20BFF9A5043FEDBDDF1FED3" box="[1346,1428,334,358]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="2" pageNumber="291">Fig. 2E</figureCitation>
) slender than mesofemora, metatibia expanded and with a small tooth at apex. Surface of mesotibia, metatibia and apex of mesofemora and metafemora covered with small punctuations and triangular short setae.
</paragraph>
<paragraph id="B728369ED20BFF9A55C6FE2FD9B9FE67" blockId="2.[151,1437,154,466]" box="[199,476,442,466]" pageId="2" pageNumber="291">Scutellum weakly visible.</paragraph>
<caption id="E3E86616D20BFF9A5596F815DB13F86B" ID-DOI="http://doi.org/10.5281/zenodo.3687742" ID-Zenodo-Dep="3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="2" pageNumber="291" startId="2.[151,244,1920,1942]" targetBox="[168,1425,500,1879]" targetPageId="2">
<paragraph id="B728369ED20BFF9A5596F815DB13F86B" blockId="2.[151,1437,1920,2014]" pageId="2" pageNumber="291">
<emphasis id="85E3EA8CD20BFF9A5596F815D975F820" bold="true" box="[151,272,1920,1942]" pageId="2" pageNumber="291">FIGURE 2.</emphasis>
Diagnostic features of
<taxonomicName id="70974D1DD20BFF9A54FAF815DBFCF820" authority="Jiang &amp; Wang" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[507,921,1920,1942]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="291" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="85E3EA8CD20BFF9A54FAF815DB65F820" box="[507,768,1920,1941]" italics="true" pageId="2" pageNumber="291">Termitodiellus hainanensis</emphasis>
Jiang &amp; Wang
</taxonomicName>
,
<emphasis id="85E3EA8CD20BFF9A56A4F815DC7BF823" bold="true" box="[933,1054,1920,1942]" pageId="2" pageNumber="291">
<taxonomicNameLabel id="9ED057F7D20BFF9A56A4F815DC7BF823" box="[933,1054,1920,1942]" pageId="2" pageNumber="291" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
<emphasis id="85E3EA8CD20BFF9A512CF815DC21F820" bold="true" box="[1069,1092,1920,1941]" pageId="2" pageNumber="291">A,</emphasis>
habitus, lateral view;
<emphasis id="85E3EA8CD20BFF9A5025F815DD5FF820" bold="true" box="[1316,1338,1920,1941]" pageId="2" pageNumber="291">B,</emphasis>
profemur and protibia, male;
<emphasis id="85E3EA8CD20BFF9A5456F831D90BF80F" bold="true" box="[343,366,1956,1978]" pageId="2" pageNumber="291">C,</emphasis>
mesofemur and mesotibia male;
<emphasis id="85E3EA8CD20BFF9A57B7F831DAA8F80C" bold="true" box="[694,717,1956,1977]" pageId="2" pageNumber="291">D,</emphasis>
mesofemur and mesotibia, female;
<emphasis id="85E3EA8CD20BFF9A512FF831DC21F80C" bold="true" box="[1070,1092,1956,1977]" pageId="2" pageNumber="291">E,</emphasis>
metafemur and metatibia, male;
<emphasis id="85E3EA8CD20BFF9A508BF831DDF8F80C" bold="true" box="[1418,1437,1956,1977]" pageId="2" pageNumber="291">F,</emphasis>
aedeagus, ventral view;
<emphasis id="85E3EA8CD20BFF9A547EF85DD9FDF868" bold="true" box="[383,408,1992,2014]" pageId="2" pageNumber="291">G,</emphasis>
aedeagus, lateral view;
<emphasis id="85E3EA8CD20BFF9A577EF85DDAFDF868" bold="true" box="[639,664,1992,2013]" pageId="2" pageNumber="291">H,</emphasis>
aedeagus, dorsal view.
</paragraph>
</caption>
<paragraph id="B728369ED20AFF9B55C6FF0FDD3AFF07" blockId="3.[151,1437,154,646]" box="[199,1375,154,178]" pageId="3" pageNumber="292">
Aedeagus (
<figureCitation id="2FAC2A1BD20AFF9B543FFF0FD9E4FF07" box="[318,385,154,178]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="3" pageNumber="292">Figs 2</figureCitation>
F–H), slender and simple, finely curved at lateral view, parameres is shorter than phallobase.
</paragraph>
<paragraph id="B728369ED20AFF9B55C6FF2BDB7EFF4C" blockId="3.[151,1437,154,646]" pageId="3" pageNumber="292">
Measurements: BL:
<quantity id="706F9B7BD20AFF9B54A7FF2BDA23FF60" box="[422,582,190,214]" metricMagnitude="-3" metricUnit="m" metricValue="3.63" metricValueMax="3.64" metricValueMin="3.62" pageId="3" pageNumber="292" unit="mm" value="3.63" valueMax="3.64" valueMin="3.62">3.62–3.64 mm</quantity>
; HL:
<quantity id="706F9B7BD20AFF9B5786FF2BDB42FF60" box="[647,807,190,214]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.699999999999999" metricValueMin="4.6000000000000005" pageId="3" pageNumber="292" unit="mm" value="0.46499999999999997" valueMax="0.47" valueMin="0.46">0.46–0.47 mm</quantity>
, HW:
<quantity id="706F9B7BD20AFF9B5671FF2BDC74FF60" box="[880,1041,190,214]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" metricValueMax="1.08" metricValueMin="1.06" pageId="3" pageNumber="292" unit="mm" value="1.07" valueMax="1.08" valueMin="1.06">1.06–1.08 mm</quantity>
; PL:
<quantity id="706F9B7BD20AFF9B514CFF2BDC88FF60" box="[1101,1261,190,214]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="8.7" metricValueMin="8.6" pageId="3" pageNumber="292" unit="mm" value="0.865" valueMax="0.87" valueMin="0.86">0.86–0.87 mm</quantity>
, PW:
<quantity id="706F9B7BD20AFF9B5033FF2BD8A1FF4C" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.41" metricValueMin="1.4" pageId="3" pageNumber="292" unit="mm" value="1.4049999999999998" valueMax="1.41" valueMin="1.4">1.40–1.41 mm</quantity>
; EL:
<quantity id="706F9B7BD20AFF9B55FCFF77D9FEFF4C" box="[253,411,226,250]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.31" metricValueMin="2.29" pageId="3" pageNumber="292" unit="mm" value="2.3" valueMax="2.31" valueMin="2.29">2.29–2.31 mm</quantity>
, EW:
<quantity id="706F9B7BD20AFF9B54DDFF77DA1FFF4C" box="[476,634,226,250]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" metricValueMax="1.39" metricValueMin="1.37" pageId="3" pageNumber="292" unit="mm" value="1.38" valueMax="1.39" valueMin="1.37">1.37–1.39 mm</quantity>
; AL:
<quantity id="706F9B7BD20AFF9B57B4FF74DB72FF4C" box="[693,791,225,249]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="292" unit="mm" value="0.75">0.75 mm</quantity>
.
</paragraph>
<paragraph id="B728369ED20AFF9B55C6FE93DA7DFED0" blockId="3.[151,1437,154,646]" pageId="3" pageNumber="292">
Female (
<figureCitation id="2FAC2A1BD20AFF9B5426FE93D9E5FEAB" box="[295,384,262,286]" captionStart="FIGURE 2" captionStartId="2.[151,244,1920,1942]" captionTargetBox="[168,1425,500,1879]" captionTargetId="figure@2.[151,1436,489,1893]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, habitus, lateral view; B, profemur and protibia, male; C, mesofemur and mesotibia male; D, mesofemur and mesotibia, female; E, metafemur and metatibia, male; F, aedeagus, ventral view; G, aedeagus, lateral view; H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3687742" httpUri="https://zenodo.org/record/3687742/files/figure.png" pageId="3" pageNumber="292">Fig. 2D</figureCitation>
): similar to the male, the projection of apex of mesotibia much weaker than that of male. Measurements: BL:
<quantity id="706F9B7BD20AFF9B5444FEBFD980FEF4" box="[325,485,298,322]" metricMagnitude="-3" metricUnit="m" metricValue="3.62" metricValueMax="3.63" metricValueMin="3.61" pageId="3" pageNumber="292" unit="mm" value="3.62" valueMax="3.63" valueMin="3.61">3.61–3.63 mm</quantity>
; HL:
<quantity id="706F9B7BD20AFF9B5725FEBCDAA6FEF4" box="[548,707,297,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" metricValueMax="5.1" metricValueMin="4.699999999999999" pageId="3" pageNumber="292" unit="mm" value="0.49" valueMax="0.51" valueMin="0.47">0.47–0.51 mm</quantity>
, HW:
<quantity id="706F9B7BD20AFF9B560AFEBFDBCEFEF4" box="[779,939,298,322]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.08" metricValueMin="1.07" pageId="3" pageNumber="292" unit="mm" value="1.075" valueMax="1.08" valueMin="1.07">1.07–1.08 mm</quantity>
; PL:
<quantity id="706F9B7BD20AFF9B56E7FEBFDCE0FEF4" box="[998,1157,298,322]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.6" metricValueMin="7.7" pageId="3" pageNumber="292" unit="mm" value="0.815" valueMax="0.86" valueMin="0.77">0.77–0.86 mm</quantity>
, PW:
<quantity id="706F9B7BD20AFF9B51C9FEBFDD0DFEF4" box="[1224,1384,298,322]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="8.6" metricValueMin="8.4" pageId="3" pageNumber="292" unit="mm" value="0.85" valueMax="0.86" valueMin="0.84">0.84–0.86 mm</quantity>
; EL:
<quantity id="706F9B7BD20AFF9B5596FEDBD950FED0" box="[151,309,334,358]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" metricValueMax="2.3" metricValueMin="2.26" pageId="3" pageNumber="292" unit="mm" value="2.28" valueMax="2.3" valueMin="2.26">2.26–2.30 mm</quantity>
, EW:
<quantity id="706F9B7BD20AFF9B5477FEDBDA71FED0" box="[374,532,334,358]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" metricValueMax="1.38" metricValueMin="1.36" pageId="3" pageNumber="292" unit="mm" value="1.37" valueMax="1.38" valueMin="1.36">1.36–1.38 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D20AFF9B55C6FEE4DBC3FE40" pageId="3" pageNumber="292" type="diagnosis">
<paragraph id="B728369ED20AFF9B55C6FEE4DBC3FE40" blockId="3.[151,1437,154,646]" pageId="3" pageNumber="292">
<emphasis id="85E3EA8CD20AFF9B55C6FEE4D9A7FE3C" bold="true" box="[199,450,369,393]" pageId="3" pageNumber="292">Differential diagnosis.</emphasis>
<taxonomicName id="70974D1DD20AFF9B54C9FEE7DA82FE3F" authority="Jiang &amp; Wang, 2020" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[456,743,370,394]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="85E3EA8CD20AFF9B54C9FEE7DA82FE3F" box="[456,743,370,394]" italics="true" pageId="3" pageNumber="292">Termitodiellus hainanensis</emphasis>
</taxonomicName>
<emphasis id="85E3EA8CD20AFF9B57EFFEE7DB17FE3C" bold="true" box="[750,882,369,394]" pageId="3" pageNumber="292">
<taxonomicNameLabel id="9ED057F7D20AFF9B57EFFEE7DB17FE3C" box="[750,882,369,394]" pageId="3" pageNumber="292" rank="species">new species</taxonomicNameLabel>
</emphasis>
can be readily distinguished to other
<taxonomicName id="70974D1DD20AFF9B5002FEE7DDF9FE3F" authorityName="Nakane" authorityYear="1961" box="[1283,1436,370,394]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="85E3EA8CD20AFF9B5002FEE7DDF9FE3F" box="[1283,1436,370,394]" italics="true" pageId="3" pageNumber="292">Termitodiellus</emphasis>
</taxonomicName>
species by the intermediate lobe bisected to two similar small lobes. However, the new species highly resembles
<taxonomicName id="70974D1DD20AFF9B5057FE03DA6CFE67" authority="(Paulian, 1983)" baseAuthorityName="Paulian" baseAuthorityYear="1983" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="85E3EA8CD20AFF9B5057FE03D93AFE67" italics="true" pageId="3" pageNumber="292">Termitodiellus besucheti</emphasis>
(
<bibRefCitation id="D3064B6FD20AFF9B546DFE2FDA67FE67" author="Paulian, R." box="[364,514,442,466]" pageId="3" pageNumber="292" pagination="615 - 622" refId="ref2339" refString="Paulian, R. (1983) Sur quelques Coleopteres Scarabaeoidea de la region Orientale. Revue Suisse de Zoologie, 90, 615 - 622. https: // doi. org / 10.5962 / bhl. part. 82000" type="journal article" year="1983">Paulian, 1983</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="2FAC2A1BD20AFF9B5716FE2FDAF1FE67" box="[535,660,442,466]" captionStart="FIGURE 3" captionStartId="3.[151,244,1689,1711]" captionTargetBox="[158,1431,676,1654]" captionTargetId="figure@3.[151,1436,669,1663]" captionTargetPageId="3" captionText="FIGURE 3. Termitodiellus hainanensis Jiang &amp; Wang, new species, holotype (A, C) and Termitodiellus besucheti (Paulian, 1983) (B, D). A, B, pronotum, dorsal view; C, D, apex of elytra. Figs. B, D, were taken by Mr. Showtaro Kakizoe, specimen from Malaysia: Selangor, deposited in Showtaro Kakizoe private collection (Fukuoka, Japan)." figureDoi="http://doi.org/10.5281/zenodo.3687744" httpUri="https://zenodo.org/record/3687744/files/figure.png" pageId="3" pageNumber="292">Figs. 3B, D</figureCitation>
) in general appearance, but the latter species with a strong posterior lobe on pronotum, and also different in the form of protrusions on elytra apex.
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D20AFF9B55C6FD94DA67FDAF" box="[199,514,513,538]" pageId="3" pageNumber="292" type="distribution">
<paragraph id="B728369ED20AFF9B55C6FD94DA67FDAF" blockId="3.[151,1437,154,646]" box="[199,514,513,538]" pageId="3" pageNumber="292">
<emphasis id="85E3EA8CD20AFF9B55C6FD94D93FFDAC" bold="true" box="[199,346,513,537]" pageId="3" pageNumber="292">Distribution.</emphasis>
<collectingCountry id="CF80760ED20AFF9B5461FD97D9C1FDAF" box="[352,420,514,538]" name="China" pageId="3" pageNumber="292">China</collectingCountry>
:
<collectingRegion id="7553F87CD20AFF9B54AFFD97D99AFDAF" box="[430,511,514,538]" country="China" name="Hainan" pageId="3" pageNumber="292">Hainan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D20AFF9B55C6FDB0DCC3FDD7" pageId="3" pageNumber="292" type="biology_ecology">
<paragraph id="B728369ED20AFF9B55C6FDB0DCC3FDD7" blockId="3.[151,1437,154,646]" pageId="3" pageNumber="292">
<emphasis id="85E3EA8CD20AFF9B55C6FDB0D946FD88" bold="true" box="[199,291,549,573]" pageId="3" pageNumber="292">Biology.</emphasis>
All adults were collected from rotten wood (
<figureCitation id="2FAC2A1BD20AFF9B5604FDB3DB2CFD8B" box="[773,841,550,574]" captionStart="FIGURE 4" captionStartId="4.[151,244,1610,1632]" captionTargetBox="[151,1437,309,1590]" captionTargetId="figure@4.[151,1437,309,1590]" captionTargetPageId="4" captionText="FIGURE 4. Habitat of Termitodiellus hainanensis Jiang &amp; Wang, new species. A, type locality: Jianfengling National Forest Park (Hainan, China); B, the environment of the forest; C, microbiotop of the type locality (rotten wood in the forest)." figureDoi="http://doi.org/10.5281/zenodo.3687746" httpUri="https://zenodo.org/record/3687746/files/figure.png" pageId="3" pageNumber="292">Figs 4</figureCitation>
A–C), and living in the internal gap which may caused by termites, ants or larvae of other beetles, but no termites or ants living in the same rotten wood.
</paragraph>
</subSubSection>
<subSubSection id="FF8D6515D20AFF9B55C6FDFBDC39FD33" box="[199,1116,622,646]" pageId="3" pageNumber="292" type="etymology">
<paragraph id="B728369ED20AFF9B55C6FDFBDC39FD33" blockId="3.[151,1437,154,646]" box="[199,1116,622,646]" pageId="3" pageNumber="292">
<emphasis id="85E3EA8CD20AFF9B55C6FDFBD92DFD33" bold="true" box="[199,328,622,646]" pageId="3" pageNumber="292">Etymology.</emphasis>
The new species is named after its
<typeStatus id="682C883CD20AFF9B57C3FDFBDA95FD33" box="[706,752,622,646]" pageId="3" pageNumber="292">type</typeStatus>
locality:
<collectingRegion id="7553F87CD20AFF9B5654FDFBDC6FFD33" box="[853,1034,622,646]" country="China" name="Hainan" pageId="3" pageNumber="292">Hainan Province</collectingRegion>
,
<collectingCountry id="CF80760ED20AFF9B5114FDFBDC3DFD33" box="[1045,1112,622,646]" name="China" pageId="3" pageNumber="292">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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