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Mansibang, Aumentado &amp; Y.P.Ang
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§ 
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&amp; 
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</paragraph>
</subSubSection>
<subSubSection id="448C656D0B6BFF905BEB830EFEF5E14F" pageId="5" pageNumber="258" type="diagnosis">
<paragraph id="0C2936E60B6BFF905BEB830EFEF5E14F" blockId="5.[136,1452,1701,2088]" pageId="5" pageNumber="258">
<emphasis id="3EE2EAF40B6BFF905BEB830EFEEAEE4C" bold="true" box="[189,342,1809,1835]" pageId="5" pageNumber="258">Diagnosis:—</emphasis>
<taxonomicName id="CB964D650B6BFF905A00830DFE44EE4C" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[342,504,1809,1835]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="258" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B6BFF905A00830DFE44EE4C" box="[342,504,1809,1835]" italics="true" pageId="5" pageNumber="258">Begonia akaw</emphasis>
</taxonomicName>
belongs to 
<emphasis id="3EE2EAF40B6BFF9059D2830DFD29EE4C" box="[644,661,1810,1835]" italics="true" pageId="5" pageNumber="258">B</emphasis>
. sect. 
<taxonomicName id="CB964D650B6BFF9059B5830DFCC8EE4C" box="[739,884,1810,1835]" pageId="5" pageNumber="264" rank="section" section="Petermannia">
<emphasis id="3EE2EAF40B6BFF9059B5830DFCC8EE4C" box="[739,884,1810,1835]" italics="true" pageId="5" pageNumber="258">Petermannia</emphasis>
</taxonomicName>
(
<bibRefCitation id="68074B170B6BFF9058D3830EFBD0EE4B" author="Klotzsch, J. F." box="[901,1132,1809,1836]" pageId="5" pageNumber="258" pagination="119 - 128" refId="ref6458" refString="Klotzsch, J. F. (1854) Begoniaceen-Gattungen und Arten. Monatsberichte der Ko ¨ niglich preussischen Akademie der Wissenschaften zu Berlin 1854: 119 - 128." type="journal article" year="1854">Klotzsch 1854: 124</bibRefCitation>
; de 
<bibRefCitation id="68074B170B6BFF905FF6830EFA37EE4B" author="Candolle, A. P. de" box="[1184,1419,1809,1836]" pageId="5" pageNumber="258" pagination="128 - 134" refId="ref5924" refString="Candolle, A. P. de (1859) Meimoire sur la famille des Beigoniaceies. Annales des Sciences Naturelles, Ortanique 11 (4): 128 - 134." type="journal article" year="1859">Candolle (1859:128)</bibRefCitation>
in having an erect habit and protogynous inflorescence (
<bibRefCitation id="68074B170B6BFF9059B8832AFC48EE37" author="Doorenbos, J. &amp; Sosef, M. S. M. &amp; De Wilde, J. J. F. E." box="[750,1012,1845,1872]" pageId="5" pageNumber="258" pagination="1 - 266" refId="ref6120" refString="Doorenbos, J., Sosef, M. S. M. &amp; De Wilde, J. J. F. E. (1998) The sections of Begonia including descriptions, keys and species lists (studies in Begoniaceae VI). Wageningen Agricultural University Papers 98 (2): 1 - 266." type="journal article" year="1998">
Doorenbos 
<emphasis id="3EE2EAF40B6BFF9058228329FC14EE28" box="[884,936,1845,1871]" italics="true" pageId="5" pageNumber="258">et al</emphasis>
. 1998
</bibRefCitation>
). It resembles 
<taxonomicName id="CB964D650B6BFF905FCB8329FF07EE13" authority="Merr. (1911: 388)" authorityName="Merr." authorityPageNumber="388" authorityYear="1911" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="258" phylum="Tracheophyta" rank="species" species="ramosii">
<emphasis id="3EE2EAF40B6BFF905FCB8329FAA8EE28" box="[1181,1300,1846,1871]" italics="true" pageId="5" pageNumber="258">B. ramosii</emphasis>
Merr. (1911: 388)
</taxonomicName>
in having erect, rarely branched stems, few glabrous, long-ovate leaves, and a clustered terminal inflorescence of staminate flowers with two tepals. 
<taxonomicName id="CB964D650B6BFF9059658361FD68EEF0" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[563,724,1917,1943]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="258" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B6BFF9059658361FD68EEF0" box="[563,724,1917,1943]" italics="true" pageId="5" pageNumber="258">Begonia akaw</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="CB964D650B6BFF90583D8361FC5DEEF0" box="[875,993,1918,1943]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="258" phylum="Tracheophyta" rank="species" species="ramosii">
<emphasis id="3EE2EAF40B6BFF90583D8361FC5DEEF0" box="[875,993,1918,1943]" italics="true" pageId="5" pageNumber="258">B. ramosii</emphasis>
</taxonomicName>
by having larger stipules [(10–)15–21 × 
<quantity id="CB6E9B030B6BFF905BDE83BDFF4EEEDB" box="[136,242,1954,1980]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="258" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
(vs. 10–15 × 
<emphasis id="3EE2EAF40B6BFF905ADE83BDFE15EEDC" box="[392,425,1954,1979]" italics="true" pageId="5" pageNumber="258">ca.</emphasis>
<quantity id="CB6E9B030B6BFF905AF983BEFE49EEDB" box="[431,501,1953,1980]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="258" unit="mm" value="5.0">5 mm</quantity>
)], longer inflorescences (
<quantity id="CB6E9B030B6BFF90585F83BEFCC3EEDB" box="[777,895,1953,1980]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="12.0" metricValueMin="4.5" pageId="5" pageNumber="258" unit="cm" value="8.25" valueMax="12.0" valueMin="4.5">4.5–12 cm</quantity>
vs. &lt;
<quantity id="CB6E9B030B6BFF90589683BDFC46EEDB" box="[960,1018,1954,1980]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="258" unit="cm" value="4.0">4 cm</quantity>
), obliquely ovate bracts (vs. orbicular), larger tepals on staminate flowers (12–17 × 
<quantity id="CB6E9B030B6BFF90593A83D9FD58EE87" box="[620,740,1990,2016]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="5" pageNumber="258" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
vs. 
<emphasis id="3EE2EAF40B6BFF90585883D9FC8CEEB8" box="[782,816,1990,2015]" italics="true" pageId="5" pageNumber="258">ca.</emphasis>
8 × 
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), a solitary pistillate flower (vs. in pairs), larger tepals on pistillate flowers (14–15 × 
<quantity id="CB6E9B030B6BFF90597583F5FD31E163" box="[547,653,2026,2052]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="5" pageNumber="258" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
vs. 9–10 × 
<emphasis id="3EE2EAF40B6BFF90584683F5FC8EE164" box="[784,818,2026,2051]" italics="true" pageId="5" pageNumber="258">ca.</emphasis>
<quantity id="CB6E9B030B6BFF90586F83F6FCC2E163" box="[825,894,2025,2052]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="258" unit="mm" value="6.0">6 mm</quantity>
), and a larger ovary (
<emphasis id="3EE2EAF40B6BFF905F3D83F5FB31E164" box="[1131,1165,2026,2051]" italics="true" pageId="5" pageNumber="258">ca.</emphasis>
18.5 × 
<quantity id="CB6E9B030B6BFF905FB483F5FA9FE163" box="[1250,1315,2026,2052]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="258" unit="mm" value="8.0">8 mm</quantity>
vs. 
<emphasis id="3EE2EAF40B6BFF905E0683F5FACEE164" box="[1360,1394,2026,2051]" italics="true" pageId="5" pageNumber="258">ca.</emphasis>
10 × 
<quantity id="CB6E9B030B6BFF905BDE8C11FF76E14F" box="[136,202,2062,2088]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="258" unit="mm" value="4.0">4 mm</quantity>
) (
<emphasis id="3EE2EAF40B6BFF905BB48C11FEFEE140" bold="true" box="[226,322,2061,2088]" pageId="5" pageNumber="258">
<tableCitation id="4114035D0B6BFF905BB48C11FE86E140" box="[226,314,2061,2088]" captionStart="TABLE 2" captionStartId="7.[136,224,772,796]" captionTargetPageId="7" captionText="TABLE 2. Morphological differences between Begonia akaw and B. ramosii" httpUri="http://table.plazi.org/id/58E9666E0B69FF925BDE871BFC01EA7B" pageId="5" pageNumber="258" tableUuid="58E9666E0B69FF925BDE871BFC01EA7B">Table 2</tableCitation>
)
</emphasis>
.
</paragraph>
</subSubSection>
<caption id="58E9666E0B68FF935BDE824FFC68EFEE" ID-DOI="http://doi.org/10.5281/zenodo.8297091" ID-Zenodo-Dep="8297091" httpUri="https://zenodo.org/record/8297091/files/figure.png" pageId="6" pageNumber="259" startId="6.[136,229,1616,1638]" targetBox="[173,1432,193,1589]" targetPageId="6" targetType="figure">
<paragraph id="0C2936E60B68FF935BDE824FFC68EFEE" blockId="6.[136,1451,1616,1674]" pageId="6" pageNumber="259">
<emphasis id="3EE2EAF40B68FF935BDE824FFE2EEF01" bold="true" box="[136,402,1616,1638]" pageId="6" pageNumber="259">
FIGURE 4. 
<taxonomicName id="CB964D650B68FF935A52824FFE2EEF01" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[260,402,1616,1638]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="6" pageNumber="259" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B68FF935A52824FFE2EEF01" bold="true" box="[260,402,1616,1638]" italics="true" pageId="6" pageNumber="259">Begonia akaw</emphasis>
</taxonomicName>
</emphasis>
. 
<emphasis id="3EE2EAF40B68FF935AC8824FFE13EF02" bold="true" box="[414,431,1616,1637]" pageId="6" pageNumber="259">A</emphasis>
. Habit; 
<emphasis id="3EE2EAF40B68FF935AA8824FFDB2EF02" bold="true" box="[510,526,1616,1637]" pageId="6" pageNumber="259">B</emphasis>
. Stipule; 
<emphasis id="3EE2EAF40B68FF93593A824FFDC1EF01" bold="true" box="[620,637,1616,1638]" pageId="6" pageNumber="259">C</emphasis>
. Bract; 
<emphasis id="3EE2EAF40B68FF93599D824FFD60EF02" bold="true" box="[715,732,1616,1637]" pageId="6" pageNumber="259">D</emphasis>
. Inflorescence; 
<emphasis id="3EE2EAF40B68FF935821824FFC3BEF02" bold="true" box="[887,903,1616,1637]" pageId="6" pageNumber="259">E</emphasis>
. Androecium; 
<emphasis id="3EE2EAF40B68FF935F41824FFB9AEF02" bold="true" box="[1047,1062,1616,1637]" pageId="6" pageNumber="259">F</emphasis>
. Pistillate flower lateral view; 
<emphasis id="3EE2EAF40B68FF935E06824FFADFEF01" bold="true" box="[1360,1379,1616,1638]" pageId="6" pageNumber="259">G</emphasis>
. Ovary cross section. Illustrated by J. Mansibang based on AAMNP173 (PNH, BRIT, CAHUP).
</paragraph>
</caption>
<subSubSection id="448C656D0B68FF925BEB82A7FE0EEBB2" lastPageId="7" lastPageNumber="260" pageId="6" pageNumber="259" type="description">
<paragraph id="0C2936E60B68FF925BEB82A7FE0EEBB2" blockId="6.[136,1452,1720,2071]" lastBlockId="7.[136,1452,158,725]" lastPageId="7" lastPageNumber="260" pageId="6" pageNumber="259">
<emphasis id="3EE2EAF40B68FF935BEB82A7FED1EFB5" bold="true" box="[189,365,1720,1746]" pageId="6" pageNumber="259">Description:—</emphasis>
Lithophytic herb, monoecious, erect, lower parts of the stems prostrate or ascending, 
<emphasis id="3EE2EAF40B68FF935E5082A6FA94EFB5" box="[1286,1320,1721,1746]" italics="true" pageId="6" pageNumber="259">ca.</emphasis>
<quantity id="CB6E9B030B68FF935E7A82A6FAD2EFB4" box="[1324,1390,1721,1747]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="259" unit="cm" value="30.0">30 cm</quantity>
high. 
<emphasis id="3EE2EAF40B68FF935BDE82C2FF02EF90" box="[136,190,1757,1783]" italics="true" pageId="6" pageNumber="259">Stem</emphasis>
terete, unbranched, with occasional shallow and longitudinal groove, pale green, 
<quantity id="CB6E9B030B68FF935F0682C3FB11EF90" box="[1104,1197,1756,1783]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="6" pageNumber="259" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
in diam., subglabrous, glabrescent, with inconspicuous microscopic indumentum, internodes (1.5–)25–60(–85) mm long. 
<emphasis id="3EE2EAF40B68FF935F8C831FFA8FEE7D" box="[1242,1331,1792,1818]" italics="true" pageId="6" pageNumber="259">Stipules</emphasis>
persistent, pale green, hyaline, ovate to oblong-ovate, obliquely inequilateral, (10–)15–21 × 
<quantity id="CB6E9B030B68FF935F7E833AFB2BEE58" box="[1064,1175,1829,1855]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="6" pageNumber="259" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
, abaxially and adaxially glabrous, shallowly keeled, margin entire, apex cuspidate, cusp 
<emphasis id="3EE2EAF40B68FF9358268356FC2EEE05" box="[880,914,1865,1890]" italics="true" pageId="6" pageNumber="259">ca.</emphasis>
<quantity id="CB6E9B030B68FF9358CB8357FC4AEE04" box="[925,1014,1864,1891]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="259" unit="mm" value="1.5">1.5 mm</quantity>
long. 
<emphasis id="3EE2EAF40B68FF935F128356FB2EEE05" box="[1092,1170,1865,1890]" italics="true" pageId="6" pageNumber="259">Leaves</emphasis>
alternate; petiole terete, pale green, succulent, 4.0–9.0 × 
<quantity id="CB6E9B030B68FF9359518372FD28EEE0" box="[519,660,1901,1927]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="6" pageNumber="259" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
, glabrous; lamina asymmetric, basifixed, oblong-ovate, 13.3–17.2 × 4.3–8.0(–10.0) cm, glabrous, adaxially olive green, abaxially pale green, veins raised prominently; base obliquely and prominently cordate on one side of lamina, both lobes rounded, unequal in length, margin very shallowly serrate, apex acuminate; venation basally 
<emphasis id="3EE2EAF40B68FF935A9D83C6FE50EE95" box="[459,492,2009,2034]" italics="true" pageId="6" pageNumber="259">ca.</emphasis>
8-palmate, branching dichotomously or nearly so, veins sunken adaxially and raised abaxially. 
<emphasis id="3EE2EAF40B68FF935BAC83E3FE2CE171" box="[250,400,2044,2070]" italics="true" pageId="6" pageNumber="259">Inflorescence</emphasis>
terminal, bisexual, protogynous, panicle 
<quantity id="CB6E9B030B68FF93580A83E3FC6DE170" box="[860,977,2044,2071]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="12.0" metricValueMin="4.5" pageId="6" pageNumber="259" unit="cm" value="8.25" valueMax="12.0" valueMin="4.5">4.5–12 cm</quantity>
long, staminate flowers initially clustered, maturing into a solitary monochasial raceme, peduncle 
<quantity id="CB6E9B030B69FF9259B08481FCE1E9DE" box="[742,861,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="5.0" metricValueMin="2.4" pageId="7" pageNumber="260" unit="mm" value="37.0" valueMax="50.0" valueMin="24.0">24–50 mm</quantity>
long, glabrous, green, reddish pink at nodes, pistillate flower solitary, axillary on terminal leaf node, basal to staminate flowers along the same panicle. 
<emphasis id="3EE2EAF40B69FF925FEC84DCFABFE9BB" box="[1210,1283,195,220]" italics="true" pageId="7" pageNumber="260">Bracts</emphasis>
sub-persistent, hyaline, white, obliquely ovate, consistent in size throughout inflorescence, (12–)18–22 × (12–) 
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, glabrous, margin entire, apex mucronate. 
<emphasis id="3EE2EAF40B69FF925AA58514FCB5E843" box="[499,777,266,293]" italics="true" pageId="7" pageNumber="260">Staminate inflorescence:</emphasis>
pedicel 
<quantity id="CB6E9B030B69FF92583A8515FC5AE842" box="[876,998,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.1" metricValueMin="1.1" pageId="7" pageNumber="260" unit="mm" value="16.0" valueMax="21.0" valueMin="11.0">11–21 mm</quantity>
long, glabrous, tepals 2, predominantly white suffused with pink at base, venation distinct, pinkish, oblong-ovate to elliptic,12–17 × 
<quantity id="CB6E9B030B69FF925FEE8531FA8AE82E" box="[1208,1334,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="7" pageNumber="260" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
, glabrous, margin entire, apex rounded, 
<emphasis id="3EE2EAF40B69FF925A8E854DFDE1E80B" box="[472,605,338,364]" italics="true" pageId="7" pageNumber="260">androecium</emphasis>
zygomorphic, filament 
<quantity id="CB6E9B030B69FF925826854DFC5FE80A" box="[880,995,338,365]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="7" pageNumber="260" unit="mm" value="0.65" valueMax="1.0" valueMin="0.3">0.3–1 mm</quantity>
long, fused at base; stamens yellow, 18 to 20, oblong in outline, apex rounded, 1.5–2.0 mm long, locules 2. 
<emphasis id="3EE2EAF40B69FF9258DE8569FB2EE8F7" box="[904,1170,374,400]" italics="true" pageId="7" pageNumber="260">Pistillate inflorescence:</emphasis>
pedicel 
<quantity id="CB6E9B030B69FF925FA48569FAD7E8F6" box="[1266,1387,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="7" pageNumber="260" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
long, glabrous; tepals 5, white to pale pink with distinct pinkish venation, subequal, glabrous, margin entire, inner tepals orbicular-obovate, 
<emphasis id="3EE2EAF40B69FF925A0985A0FE3DE8BF" box="[351,385,447,472]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
<quantity id="CB6E9B030B69FF925ADC85A1FE67E8BE" box="[394,475,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="260" unit="mm" value="15.0">15 mm</quantity>
× 
<quantity id="CB6E9B030B69FF925AAB85A1FDD0E8BE" box="[509,620,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="7" pageNumber="260" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
, apex rounded, outer tepals somewhat broader, orbicular-ovate, 14–15 × 
<quantity id="CB6E9B030B69FF925BDE85FDFEBEE89A" box="[136,258,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="7" pageNumber="260" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
, apex rounded; styles yellow, forming 3 divided arms, each arm forked at distal half, 
<emphasis id="3EE2EAF40B69FF925FF985FCFB6CE89B" box="[1199,1232,483,508]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
<quantity id="CB6E9B030B69FF925F8085FDFAAAE89A" box="[1238,1302,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="260" unit="mm" value="8.0">8 mm</quantity>
long, twisted and papillose all around; ovary trigonous-ellipsoid, 
<emphasis id="3EE2EAF40B69FF92598A8618FD42EB47" box="[732,766,519,544]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
18.5 × 
<quantity id="CB6E9B030B69FF92580A8619FC1FEB46" box="[860,931,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="260" unit="mm" value="8.0">8 mm</quantity>
; locules 3, placenta bifid, wings 3, subequal, dorsal wing oblong-lunate, larger than 2 lateral wings, rounded proximally, truncate to subtruncate distally, 17–20 × 
<emphasis id="3EE2EAF40B69FF925BDE8650FF16EB0F" box="[136,170,591,616]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
<quantity id="CB6E9B030B69FF925BE78651FF49EB0E" box="[177,245,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="260" unit="mm" value="9.0">9 mm</quantity>
, lateral wings triangular, acute to obtuse proximally, truncate distally, 
<emphasis id="3EE2EAF40B69FF925F468650FB8DEB0F" box="[1040,1073,591,616]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
<quantity id="CB6E9B030B69FF925F6F8651FB35EB0E" box="[1081,1161,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="7" pageNumber="260" unit="mm" value="17.0">17 mm</quantity>
× 
<quantity id="CB6E9B030B69FF925FFE8651FB50EB0E" box="[1192,1260,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="260" unit="mm" value="8.0">8 mm</quantity>
, base rounded to subacute. 
<emphasis id="3EE2EAF40B69FF925BAF866DFEE9EBEB" box="[249,341,626,652]" italics="true" pageId="7" pageNumber="260">Capsule</emphasis>
recurved on stiff pedicel, pedicel 
<quantity id="CB6E9B030B69FF925980866DFCECEBEA" box="[726,848,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="7" pageNumber="260" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
long, dorsal wing distinctly larger than lateral wings, nodding when mature, 2 lateral wings cucullate to form a splash cup, dehiscing along ovary wing attachments when dried, 16–20 × 
<emphasis id="3EE2EAF40B69FF925A6586A4FEE9EBB3" box="[307,341,699,724]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
<quantity id="CB6E9B030B69FF925A0A86A5FE12EBB2" box="[348,430,698,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="260" unit="mm" value="18.0">18 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="58E9666E0B69FF925BDE871BFC01EA7B" ID-Table-UUID="58E9666E0B69FF925BDE871BFC01EA7B" box="[136,957,772,796]" httpUri="http://table.plazi.org/id/58E9666E0B69FF925BDE871BFC01EA7B" pageId="7" pageNumber="260" startId="7.[136,224,772,796]" targetBox="[148,1201,813,1653]" targetIsTable="true" targetPageId="7" targetType="table">
<paragraph id="0C2936E60B69FF925BDE871BFC01EA7B" blockId="7.[136,957,772,796]" box="[136,957,772,796]" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BDE871BFF47EA7B" bold="true" box="[136,251,772,796]" pageId="7" pageNumber="260">TABLE 2.</emphasis>
Morphological differences between 
<taxonomicName id="CB964D650B69FF9259D5871AFCA7EA7B" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[643,795,772,796]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B69FF9259D5871AFCA7EA7B" box="[643,795,772,796]" italics="true" pageId="7" pageNumber="260">Begonia akaw</emphasis>
</taxonomicName>
and 
<taxonomicName id="CB964D650B69FF925819871AFC01EA7B" box="[847,957,773,796]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="ramosii">B. ramosii</taxonomicName>
</paragraph>
</caption>
<paragraph id="0C2936E60B69FF9259108732FB3BEF12" pageId="7" pageNumber="260">
<table id="7E96C4460B69006A5BC28732FB0DEF12" box="[148,1201,813,1653]" gridcols="3" gridrows="20" pageId="7" pageNumber="260">
<tr id="B2A634A40B69006A5BC28732FB0DEA24" box="[148,1201,813,835]" gridrow="0" pageId="7" pageNumber="260" rowspan-0="1">
<th id="F1775DD80B69006A59108732FCE8EA24" box="[582,852,813,835]" gridcol="1" gridrow="0" pageId="7" pageNumber="260">
<taxonomicName id="CB964D650B69FF9259108732FD6EEA24" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[582,722,813,835]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B69FF9259108732FD6EEA24" bold="true" box="[582,722,813,835]" italics="true" pageId="7" pageNumber="260">Begonia akaw</emphasis>
</taxonomicName>
</th>
<th id="F1775DD80B69006A5F588732FB0DEA24" box="[1038,1201,813,835]" gridcol="2" gridrow="0" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925F588732FB0DEA25" bold="true" box="[1038,1201,813,834]" italics="true" pageId="7" pageNumber="260">Begonia ramosii</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC2879FFB0DEAF1" box="[148,1201,896,918]" gridrow="1" pageId="7" pageNumber="260" rowspan-1="1" rowspan-2="1">
<th id="F1775DD80B69006A5BC2879FFE6CEAF1" box="[148,464,896,918]" gridcol="0" gridrow="1" pageId="7" pageNumber="260">Stipule</th>
</tr>
<tr id="B2A634A40B69006A5BC287B7FB0DEAD9" box="[148,1201,936,958]" gridrow="2" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC287B7FE6CEAD9" box="[148,464,936,958]" gridcol="0" gridrow="2" pageId="7" pageNumber="260">size (mm)</th>
<td id="F1775DD80B69006A591087B7FCE8EAD9" box="[582,852,936,958]" gridcol="1" gridrow="2" pageId="7" pageNumber="260">(10–)15–21 × 8–10</td>
<td id="F1775DD80B69006A5F5887B7FB0DEAD9" box="[1038,1201,936,958]" gridcol="2" gridrow="2" pageId="7" pageNumber="260">
10–15 × 
<emphasis id="3EE2EAF40B69FF925F3587B6FB3DEAD9" box="[1123,1153,937,958]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
5
</td>
</tr>
<tr id="B2A634A40B69006A5BC287CEFB0DEA80" box="[148,1201,977,999]" gridrow="3" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC287CEFB0DEA80" box="[148,1201,977,999]" colspan="3" colspanRight="2" gridcol="0" gridrow="3" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BC287CEFEA0EA80" bold="true" box="[148,284,977,999]" pageId="7" pageNumber="260">Inflorescence</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC287E5FB0DED77" box="[148,1201,1018,1040]" gridrow="4" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC287E5FE6CED77" box="[148,464,1018,1040]" gridcol="0" gridrow="4" pageId="7" pageNumber="260">length (cm)</th>
<td id="F1775DD80B69006A591087E5FCE8ED77" box="[582,852,1018,1040]" gridcol="1" gridrow="4" pageId="7" pageNumber="260">4.5–12</td>
<td id="F1775DD80B69006A5F5887E5FB0DED77" box="[1038,1201,1018,1040]" gridcol="2" gridrow="4" pageId="7" pageNumber="260">&lt;4</td>
</tr>
<tr id="B2A634A40B69006A5BC2803CFB0DED5F" box="[148,1201,1059,1080]" gridrow="5" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC2803CFB0DED5F" box="[148,1201,1059,1080]" colspan="3" colspanRight="2" gridcol="0" gridrow="5" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BC2803CFF4DED5F" bold="true" box="[148,241,1059,1080]" pageId="7" pageNumber="260">Peduncle</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC28053FB0DED05" box="[148,1201,1100,1122]" gridrow="6" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28053FE6CED05" box="[148,464,1100,1122]" gridcol="0" gridrow="6" pageId="7" pageNumber="260">color</th>
<td id="F1775DD80B69006A59108053FCE8ED05" box="[582,852,1100,1122]" gridcol="1" gridrow="6" pageId="7" pageNumber="260">green, reddish pink at nodes</td>
<td id="F1775DD80B69006A5F588053FB0DED05" box="[1038,1201,1100,1122]" gridcol="2" gridrow="6" pageId="7" pageNumber="260">greenish brown</td>
</tr>
<tr id="B2A634A40B69006A5BC2806BFB0DEDEE" box="[148,1201,1140,1161]" gridrow="7" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC2806BFB0DEDEE" box="[148,1201,1140,1161]" colspan="3" colspanRight="2" gridcol="0" gridrow="7" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BC2806BFF71EDEE" bold="true" box="[148,205,1140,1161]" pageId="7" pageNumber="260">Bract</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC28082FB0DEDD4" box="[148,1201,1181,1203]" gridrow="8" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28082FE6CEDD4" box="[148,464,1181,1203]" gridcol="0" gridrow="8" pageId="7" pageNumber="260">shape</th>
<td id="F1775DD80B69006A59108082FCE8EDD4" box="[582,852,1181,1203]" gridcol="1" gridrow="8" pageId="7" pageNumber="260">obliquely ovate</td>
<td id="F1775DD80B69006A5F588082FB0DEDD4" box="[1038,1201,1181,1203]" gridcol="2" gridrow="8" pageId="7" pageNumber="260">orbicular</td>
</tr>
<tr id="B2A634A40B69006A5BC280D9FB0DEDBB" box="[148,1201,1222,1244]" gridrow="9" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC280D9FE6CEDBB" box="[148,464,1222,1244]" gridcol="0" gridrow="9" pageId="7" pageNumber="260">dimension (mm)</th>
<td id="F1775DD80B69006A591080D9FCE8EDBB" box="[582,852,1222,1244]" gridcol="1" gridrow="9" pageId="7" pageNumber="260">(12–)18–22 × (12–)15–18</td>
<td id="F1775DD80B69006A5F5880D9FB0DEDBB" box="[1038,1201,1222,1244]" gridcol="2" gridrow="9" pageId="7" pageNumber="260">
10–15 × 
<emphasis id="3EE2EAF40B69FF925F3580D8FB3DEDBB" box="[1123,1153,1223,1244]" italics="true" pageId="7" pageNumber="260">ca.</emphasis>
10
</td>
</tr>
<tr id="B2A634A40B69006A5BC280F1FB0DEC63" box="[148,1201,1262,1284]" gridrow="10" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC280F1FB0DEC63" box="[148,1201,1262,1284]" colspan="3" colspanRight="2" gridcol="0" gridrow="10" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BC280F1FE3BEC63" bold="true" box="[148,391,1262,1284]" pageId="7" pageNumber="260">Staminate inflorescence</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC28107FB0DEC49" box="[148,1201,1304,1326]" gridrow="11" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28107FE6CEC49" box="[148,464,1304,1326]" gridcol="0" gridrow="11" pageId="7" pageNumber="260">tepal size (mm)</th>
<td id="F1775DD80B69006A59108107FCE8EC49" box="[582,852,1304,1326]" gridcol="1" gridrow="11" pageId="7" pageNumber="260">12–17 × 10–13</td>
<td id="F1775DD80B69006A5F588107FB0DEC49" box="[1038,1201,1304,1326]" gridcol="2" gridrow="11" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925F588106FB99EC49" box="[1038,1061,1305,1326]" italics="true" pageId="7" pageNumber="260">ca</emphasis>
. 8 × 7
</td>
</tr>
<tr id="B2A634A40B69006A5BC2815EFB0DEC30" box="[148,1201,1345,1367]" gridrow="12" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC2815EFE6CEC30" box="[148,464,1345,1367]" gridcol="0" gridrow="12" pageId="7" pageNumber="260">stamen number</th>
<td id="F1775DD80B69006A5910815EFCE8EC30" box="[582,852,1345,1367]" gridcol="1" gridrow="12" pageId="7" pageNumber="260">18–20</td>
<td id="F1775DD80B69006A5F58815EFB0DEC30" box="[1038,1201,1345,1367]" gridcol="2" gridrow="12" pageId="7" pageNumber="260">20–30</td>
</tr>
<tr id="B2A634A40B69006A5BC28176FB0DECE7" box="[148,1201,1385,1408]" gridrow="13" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28176FE6CECE7" box="[148,464,1385,1408]" gridcol="0" gridrow="13" pageId="7" pageNumber="260">stamen length (mm)</th>
<td id="F1775DD80B69006A59108176FCE8ECE7" box="[582,852,1385,1408]" gridcol="1" gridrow="13" pageId="7" pageNumber="260">1.5–2</td>
<td id="F1775DD80B69006A5F588176FB0DECE7" box="[1038,1201,1385,1408]" gridcol="2" gridrow="13" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925F588175FB99EC18" box="[1038,1061,1386,1407]" italics="true" pageId="7" pageNumber="260">ca</emphasis>
. 1
</td>
</tr>
<tr id="B2A634A40B69006A5BC2818DFB0DECCF" box="[148,1201,1426,1448]" gridrow="14" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC2818DFB0DECCF" box="[148,1201,1426,1448]" colspan="3" colspanRight="2" gridcol="0" gridrow="14" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BC2818DFEC4ECCF" bold="true" box="[148,376,1426,1448]" pageId="7" pageNumber="260">Pistillate inflorescence</emphasis>
</th>
</tr>
<tr id="B2A634A40B69006A5BC281A4FB0DECB6" box="[148,1201,1467,1489]" gridrow="15" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC281A4FE6CECB6" box="[148,464,1467,1489]" gridcol="0" gridrow="15" pageId="7" pageNumber="260">habit</th>
<td id="F1775DD80B69006A591081A4FCE8ECB6" box="[582,852,1467,1489]" gridcol="1" gridrow="15" pageId="7" pageNumber="260">solitary</td>
<td id="F1775DD80B69006A5F5881A4FB0DECB6" box="[1038,1201,1467,1489]" gridcol="2" gridrow="15" pageId="7" pageNumber="260">in pairs</td>
</tr>
<tr id="B2A634A40B69006A5BC281FBFB0DEC9D" box="[148,1201,1508,1530]" gridrow="16" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC281FBFE6CEC9D" box="[148,464,1508,1530]" gridcol="0" gridrow="16" pageId="7" pageNumber="260">pedicel length (mm)</th>
<td id="F1775DD80B69006A591081FBFCE8EC9D" box="[582,852,1508,1530]" gridcol="1" gridrow="16" pageId="7" pageNumber="260">16–17</td>
<td id="F1775DD80B69006A5F5881FBFB0DEC9D" box="[1038,1201,1508,1530]" gridcol="2" gridrow="16" pageId="7" pageNumber="260">5–10</td>
</tr>
<tr id="B2A634A40B69006A5BC28212FB0DEF44" box="[148,1201,1549,1571]" gridrow="17" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28212FE6CEF44" box="[148,464,1549,1571]" gridcol="0" gridrow="17" pageId="7" pageNumber="260">tepal size (mm)</th>
<td id="F1775DD80B69006A59108212FCE8EF44" box="[582,852,1549,1571]" gridcol="1" gridrow="17" pageId="7" pageNumber="260">14–15 × 8–13</td>
<td id="F1775DD80B69006A5F588212FB0DEF44" box="[1038,1201,1549,1571]" gridcol="2" gridrow="17" pageId="7" pageNumber="260">9–10 × 6</td>
</tr>
<tr id="B2A634A40B69006A5BC28229FB0DEF2B" box="[148,1201,1590,1612]" gridrow="18" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28229FE6CEF2B" box="[148,464,1590,1612]" gridcol="0" gridrow="18" pageId="7" pageNumber="260">ovary size (mm)</th>
<td id="F1775DD80B69006A59108229FCE8EF2B" box="[582,852,1590,1612]" gridcol="1" gridrow="18" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF9259108229FDE1EF2C" box="[582,605,1590,1611]" italics="true" pageId="7" pageNumber="260">ca</emphasis>
. 18.5 × 8
</td>
<td id="F1775DD80B69006A5F588229FB0DEF2B" box="[1038,1201,1590,1612]" gridcol="2" gridrow="18" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925F588229FB99EF2C" box="[1038,1061,1590,1611]" italics="true" pageId="7" pageNumber="260">ca</emphasis>
. 10 × 4
</td>
</tr>
<tr id="B2A634A40B69006A5BC28240FB0DEF12" box="[148,1201,1631,1653]" gridrow="19" pageId="7" pageNumber="260">
<th id="F1775DD80B69006A5BC28240FE6CEF12" box="[148,464,1631,1653]" gridcol="0" gridrow="19" pageId="7" pageNumber="260">capsule wing dimension (mm)</th>
<td id="F1775DD80B69006A59108240FCE8EF12" box="[582,852,1631,1653]" gridcol="1" gridrow="19" pageId="7" pageNumber="260">
17–20 × 
<emphasis id="3EE2EAF40B69FF9259CD8240FD0EEF13" box="[667,690,1631,1652]" italics="true" pageId="7" pageNumber="260">ca</emphasis>
. 9
</td>
<td id="F1775DD80B69006A5F588240FB0DEF12" box="[1038,1201,1631,1653]" gridcol="2" gridrow="19" pageId="7" pageNumber="260">10–13 × 4–6</td>
</tr>
</table>
</paragraph>
<subSubSection id="448C656D0B69FF925BEB82BAFF6DEE4C" pageId="7" pageNumber="260" type="etymology">
<paragraph id="0C2936E60B69FF925BEB82BAFF6DEE4C" blockId="7.[136,1452,1701,2087]" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BEB82BAFED8EFD8" bold="true" box="[189,356,1701,1727]" pageId="7" pageNumber="260">Etymology:—</emphasis>
The epithet “ 
<emphasis id="3EE2EAF40B69FF925AAA82BAFDFAEFD8" box="[508,582,1701,1727]" italics="true" pageId="7" pageNumber="260">
<taxonomicName id="CB964D650B69FF925AAA82BAFD81EFD8" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[508,573,1701,1727]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="akaw">akaw</taxonomicName>
”
</emphasis>
is a Balereño word used to convey awe, disbelief, or surprise. The species had been documented and observed on multiple occasions, being misidentified as 
<taxonomicName id="CB964D650B69FF925F7382D5FB27EF84" box="[1061,1179,1738,1763]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="ramosii">
<emphasis id="3EE2EAF40B69FF925F7382D5FB27EF84" box="[1061,1179,1738,1763]" italics="true" pageId="7" pageNumber="260">B. ramosii</emphasis>
</taxonomicName>
. This can be considered as a case of an overlooked undescribed species, which is true for other threatened flora in the country (
<bibRefCitation id="68074B170B69FF925E4182F2FF79EE4C" author="Daipan, B. P. O. &amp; Buot, I. E. Jr. &amp; Pampolina, N. M." pageId="7" pageNumber="260" pagination="307 - 319" refId="ref5956" refString="Daipan, B. P. O., Buot, I. E. Jr. &amp; Pampolina, N. M. (2022) Evaluating the species distribution patterns of the genus Saurauia Willd. in the Philippines using geospatial analysis. Webbia 77 (2): 307 - 319. https: // doi. org / 10.36253 / jopt- 13193" type="journal article" year="2022">
Daipan 
<emphasis id="3EE2EAF40B69FF925E2482F1FA19EE60" box="[1394,1445,1773,1799]" italics="true" pageId="7" pageNumber="260">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="448C656D0B69FF925BEB832AFCE1E164" pageId="7" pageNumber="260" type="distribution">
<paragraph id="0C2936E60B69FF925BEB832AFCE1E164" blockId="7.[136,1452,1701,2087]" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BEB832AFDB7EE28" bold="true" box="[189,523,1845,1871]" pageId="7" pageNumber="260">Distribution and habitat:—</emphasis>
Aside from the locality from which the 
<typeStatus id="D32D88440B69FF92588D8329FBB7EE37" box="[987,1035,1846,1872]" pageId="7" pageNumber="260">type</typeStatus>
was collected in AMNP (
<figureCitation id="94AD2A630B69FF925E6E832AFA21EE28" box="[1336,1437,1845,1871]" captionStart="FIGURE 6" captionStartId="9.[136,229,1072,1094]" captionTargetBox="[187,1400,194,1045]" captionTargetId="figure-19@9.[183,1403,190,1048]" captionTargetPageId="9" captionText="FIGURE 6. Map of Aurora Province showing the locality of B. masilig and B. akaw (star)." figureDoi="http://doi.org/10.5281/zenodo.8297095" httpUri="https://zenodo.org/record/8297095/files/figure.png" pageId="7" pageNumber="260">Figure 6</figureCitation>
), local populations were also found in the Caunayan Falls and Ditumabo Falls areas, San Luis; the vicinity of Bulawan Falls, Dimayuga; and Amro River Protected Landscape, Dilasag. It occurs locally on rock walls with dripping water, streams, waterfall areas, deeply shaded humus-rich forest soil, and along forest trails. This taxon was often observed growing sympatrically with 
<emphasis id="3EE2EAF40B69FF925A9383D9FDD1EEB8" box="[453,621,1989,2015]" italics="true" pageId="7" pageNumber="260">
<taxonomicName id="CB964D650B69FF925A9383D9FDD5EEB8" box="[453,617,1989,2015]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="droseroides">B. droseroides</taxonomicName>
,
</emphasis>
<taxonomicName id="CB964D650B69FF92592283D9FBBAEEB8" authority="Merr. (1914: 378)" authorityName="Merr." authorityPageNumber="378" authorityYear="1914" box="[628,1030,1989,2016]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" isUncertain="true" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="megacarpa">
<emphasis id="3EE2EAF40B69FF92592283D9FD30EEB8" box="[628,652,1990,2015]" italics="true" pageId="7" pageNumber="260">B.</emphasis>
cf. 
<emphasis id="3EE2EAF40B69FF9259E183D9FC89EEB8" box="[695,821,1990,2015]" italics="true" pageId="7" pageNumber="260">megacarpa</emphasis>
Merr. (1914: 378)
</taxonomicName>
, and 
<taxonomicName id="CB964D650B69FF925F1783D9FF02E164" authority="Elmer (1910: 736)" authorityName="Elmer" authorityPageNumber="736" authorityYear="1910" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" isUncertain="true" kingdom="Plantae" order="Cucurbitales" pageId="7" pageNumber="260" phylum="Tracheophyta" rank="species" species="negrosensis">
<emphasis id="3EE2EAF40B69FF925F1783D9FBE5EEB8" box="[1089,1113,1990,2015]" italics="true" pageId="7" pageNumber="260">B.</emphasis>
aff. 
<emphasis id="3EE2EAF40B69FF925FDB83D9FAACEEB8" box="[1165,1296,1990,2015]" italics="true" pageId="7" pageNumber="260">negrosensis</emphasis>
Elmer (1910: 736)
</taxonomicName>
, among other shade-tolerant herbs and rock wall associates.
</paragraph>
</subSubSection>
<subSubSection id="448C656D0B69FF925BEB8C12FC82E140" box="[189,830,2061,2087]" pageId="7" pageNumber="260" type="biology_ecology">
<paragraph id="0C2936E60B69FF925BEB8C12FC82E140" blockId="7.[136,1452,1701,2087]" box="[189,830,2061,2087]" pageId="7" pageNumber="260">
<emphasis id="3EE2EAF40B69FF925BEB8C12FEE3E140" bold="true" box="[189,351,2061,2087]" pageId="7" pageNumber="260">Phenology:—</emphasis>
This species flowers and fruits year-round.
</paragraph>
</subSubSection>
<caption id="58E9666E0B66FF9D5BDE835CFDDDEEA3" ID-DOI="http://doi.org/10.5281/zenodo.8297093" ID-Zenodo-Dep="8297093" httpUri="https://zenodo.org/record/8297093/files/figure.png" pageId="8" pageNumber="261" startId="8.[136,229,1859,1881]" targetBox="[151,1435,190,1834]" targetPageId="8" targetType="figure">
<paragraph id="0C2936E60B66FF9D5BDE835CFDDDEEA3" blockId="8.[136,1452,1858,1989]" pageId="8" pageNumber="261">
<emphasis id="3EE2EAF40B66FF9D5BDE835CFE2CEE3F" bold="true" box="[136,400,1858,1881]" pageId="8" pageNumber="261">
FIGURE 5. 
<taxonomicName id="CB964D650B66FF9D5A55835CFE2CEE3F" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[259,400,1858,1880]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="8" pageNumber="261" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B66FF9D5A55835CFE2CEE3F" bold="true" box="[259,400,1858,1880]" italics="true" pageId="8" pageNumber="261">Begonia akaw</emphasis>
</taxonomicName>
</emphasis>
. 
<emphasis id="3EE2EAF40B66FF9D5ACA835CFE11EE3F" bold="true" box="[412,429,1859,1880]" pageId="8" pageNumber="261">A</emphasis>
. Habitat and habit; 
<emphasis id="3EE2EAF40B66FF9D593A835CFDC0EE3F" bold="true" box="[620,636,1859,1880]" pageId="8" pageNumber="261">B</emphasis>
. Stem; 
<emphasis id="3EE2EAF40B66FF9D5990835CFD6BEE3E" bold="true" box="[710,727,1859,1881]" pageId="8" pageNumber="261">C</emphasis>
. Stipule; 
<emphasis id="3EE2EAF40B66FF9D5862835CFCF9EE3F" bold="true" box="[820,837,1859,1880]" pageId="8" pageNumber="261">D</emphasis>
. Leaf adaxial (left) and abaxial (right) surface; 
<emphasis id="3EE2EAF40B66FF9D5E58835CFAA2EE3F" bold="true" box="[1294,1310,1859,1880]" pageId="8" pageNumber="261">E</emphasis>
. Inflorescence showing bracts; 
<emphasis id="3EE2EAF40B66FF9D5A718378FE8AEE1B" bold="true" box="[295,310,1895,1916]" pageId="8" pageNumber="261">F</emphasis>
. Inflorescence showing peduncle; 
<emphasis id="3EE2EAF40B66FF9D59D08378FD25EE1A" bold="true" box="[646,665,1895,1917]" pageId="8" pageNumber="261">G</emphasis>
. Staminate flower ventral view; 
<emphasis id="3EE2EAF40B66FF9D58818378FC56EE1B" bold="true" box="[983,1002,1895,1916]" pageId="8" pageNumber="261">H</emphasis>
. Staminate flower ventral view, a variant with rounder tepals; 
<emphasis id="3EE2EAF40B66FF9D5A4A8394FE99EEC7" bold="true" box="[284,293,1931,1952]" pageId="8" pageNumber="261">I</emphasis>
. Staminate flower lateral view; 
<emphasis id="3EE2EAF40B66FF9D59038394FDDDEEC7" bold="true" box="[597,609,1931,1952]" pageId="8" pageNumber="261">J</emphasis>
. Pistillate flower ventral view; 
<emphasis id="3EE2EAF40B66FF9D58DD8394FC22EEC7" bold="true" box="[907,926,1931,1952]" pageId="8" pageNumber="261">K</emphasis>
. Pistillate flower lateral view showing ovary; 
<emphasis id="3EE2EAF40B66FF9D5E028394FAD8EEC7" bold="true" box="[1364,1380,1931,1952]" pageId="8" pageNumber="261">L</emphasis>
. Ovary cross section; 
<emphasis id="3EE2EAF40B66FF9D5A4683B0FE9BEEA3" bold="true" box="[272,295,1967,1988]" pageId="8" pageNumber="261">M</emphasis>
. capsule. All photos by Y.P. Ang.
</paragraph>
</caption>
<caption id="58E9666E0B67FF9C5BDE802FFC4FED22" ID-DOI="http://doi.org/10.5281/zenodo.8297095" ID-Zenodo-Dep="8297095" box="[136,1011,1072,1094]" httpUri="https://zenodo.org/record/8297095/files/figure.png" pageId="9" pageNumber="262" startId="9.[136,229,1072,1094]" targetBox="[187,1400,194,1045]" targetPageId="9" targetType="figure">
<paragraph id="0C2936E60B67FF9C5BDE802FFC4FED22" blockId="9.[136,1011,1072,1094]" box="[136,1011,1072,1094]" pageId="9" pageNumber="262">
<emphasis id="3EE2EAF40B67FF9C5BDE802FFF41ED22" bold="true" box="[136,253,1072,1094]" pageId="9" pageNumber="262">FIGURE 6.</emphasis>
Map of Aurora Province showing the locality of 
<taxonomicName id="CB964D650B67FF9C5980802EFC84ED22" authorityName=", J. Collantes, C. J. P. Dela Cruz &amp; Y. P. Ang" authorityYear="2023" box="[726,824,1072,1094]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="262" phylum="Tracheophyta" rank="species" species="masilig">
<emphasis id="3EE2EAF40B67FF9C5980802EFC84ED22" box="[726,824,1072,1094]" italics="true" pageId="9" pageNumber="262">B. masilig</emphasis>
</taxonomicName>
and 
<taxonomicName id="CB964D650B67FF9C5830802EFC08ED22" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[870,948,1072,1094]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="262" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B67FF9C5830802EFC08ED22" box="[870,948,1072,1094]" italics="true" pageId="9" pageNumber="262">B. akaw</emphasis>
</taxonomicName>
(star).
</paragraph>
</caption>
<subSubSection id="448C656D0B67FF9C5BEB806BFDA5EC25" pageId="9" pageNumber="262" type="conservation">
<paragraph id="0C2936E60B67FF9C5BEB806BFDA5EC25" blockId="9.[136,1451,1140,1527]" pageId="9" pageNumber="262">
<emphasis id="3EE2EAF40B67FF9C5BEB806BFDC4EDE9" bold="true" box="[189,632,1140,1166]" pageId="9" pageNumber="262">Proposed conservation assessment:—</emphasis>
Because the localities of 
<taxonomicName id="CB964D650B67FF9C58DF806AFB9BEDE9" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[905,1063,1140,1166]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="262" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B67FF9C58DF806AFB9BEDE9" box="[905,1063,1140,1166]" italics="true" pageId="9" pageNumber="262">Begonia akaw</emphasis>
</taxonomicName>
have developed into tourism spots, the local rock wall populations of Caunayan Falls and Ditumabo Falls, Barangay San Luis have been observed to be largely removed. Furthermore, the creation of logging trails, forest conversion, and clearing also devastate localities where 
<taxonomicName id="CB964D650B67FF9C5B8580FEFE90ED9D" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[211,300,1248,1274]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="262" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B67FF9C5B8580FEFE90ED9D" box="[211,300,1248,1274]" italics="true" pageId="9" pageNumber="262">B. akaw</emphasis>
</taxonomicName>
occurs. From our surveys, we estimate the EOO is to be &lt;
<quantity id="CB6E9B030B67FF9C58E180FFFBB7ED9D" box="[951,1035,1248,1274]" metricMagnitude="5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="262" unit="km" value="800.0">800 km</quantity>
<superScript id="FBE39BAE0B67FF9C5F5D80FFFBAFED89" attach="left" box="[1035,1043,1248,1262]" fontSize="6" pageId="9" pageNumber="262">2</superScript>
with an AOO of about 
<quantity id="CB6E9B030B67FF9C5E4380FFFAE7ED9D" box="[1301,1371,1248,1274]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="262" unit="km" value="20.0">20 km</quantity>
<superScript id="FBE39BAE0B67FF9C5E0D80FFFADFED89" attach="left" box="[1371,1379,1248,1262]" fontSize="6" pageId="9" pageNumber="262">2</superScript>
These threats for 
<taxonomicName id="CB964D650B67FF9C5A56811AFEE5EC79" authorityName="Mansibang, Aumentado &amp; Y. P. Ang" authorityYear="2023" box="[256,345,1284,1310]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="9" pageNumber="262" phylum="Tracheophyta" rank="species" species="akaw">
<emphasis id="3EE2EAF40B67FF9C5A56811AFEE5EC79" box="[256,345,1284,1310]" italics="true" pageId="9" pageNumber="262">B. akaw</emphasis>
</taxonomicName>
satisfy the IUCN 15.1 Red List EN (Endangered) Criteria 
<emphasis id="3EE2EAF40B67FF9C58B2811BFB65EC79" box="[996,1241,1284,1310]" italics="true" pageId="9" pageNumber="262">B1ab (ii,iii)+2ab(ii,iii</emphasis>
) (
<bibRefCitation id="68074B170B67FF9C5FA6811BFDB1EC25" author="IUCN Standards &amp; Petitions Subcommittee" pageId="9" pageNumber="262" refId="ref6407" refString="IUCN Standards and Petitions Subcommittee (2022) Guidelines for using the IUCN Red List Categories and Criteria. Version 15.1. Available from: https: // nc. iucnredlist. org / redlist / content / attachment _ files / RedListGuidelines. pdf (accessed 1 October 2022)" type="book" year="2022">IUCN Standards and Petitions Subcommittee, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="448C656D0B67FF9C5BEB8153FD13ECED" pageId="9" pageNumber="262" type="discussion">
<paragraph id="0C2936E60B67FF9C5BEB8153FD13ECED" blockId="9.[136,1451,1140,1527]" pageId="9" pageNumber="262">
<emphasis id="3EE2EAF40B67FF9C5BEB8153FE9AEC01" bold="true" box="[189,294,1356,1382]" pageId="9" pageNumber="262">Notes:—</emphasis>
The stem and stipule may appear glabrous in fresh specimens. However, there are minute trichomes that can only be seen with a dissecting scope at 160×.
</paragraph>
</subSubSection>
<subSubSection id="448C656D0B67FF9C5BEB818BFDFBEC91" pageId="9" pageNumber="262" type="materials_examined">
<paragraph id="0C2936E60B67FF9C5BEB818BFDFBEC91" blockId="9.[136,1451,1140,1527]" pageId="9" pageNumber="262">
<emphasis id="3EE2EAF40B67FF9C5BEB818BFDA5ECC9" bold="true" box="[189,537,1428,1454]" pageId="9" pageNumber="262">Other specimen examined:—</emphasis>
<materialsCitation id="BCFE3CBB0B67FF9C594F818BFDFFEC91" ID-GBIF-Occurrence="4402053306" collectingDate="2022-02-26" collectionCode="PNH, CAHUP, PPC" country="Philippines" county="Luzon" elevation="535" latitude="15.681015" location="Barangay Villa" longLatPrecision="1" longitude="121.39388" municipality="Municipality of Maria" pageId="9" pageNumber="262" specimenCount="1" stateProvince="Aurora">
<collectingCountry id="748176760B67FF9C594F818BFD7EECC9" box="[537,706,1428,1454]" name="Philippines" pageId="9" pageNumber="262">PHILIPPINES</collectingCountry>
. 
<collectingCounty id="E5484E6A0B67FF9C599D818AFCA9ECC8" box="[715,789,1429,1455]" pageId="9" pageNumber="262">Luzon</collectingCounty>
, 
<collectingRegion id="CE52F8040B67FF9C5849818AFC67ECC9" box="[799,987,1428,1455]" country="Philippines" name="Aurora" pageId="9" pageNumber="262">Aurora Province</collectingRegion>
, 
<collectingMunicipality id="EC4DAC9C0B67FF9C58B0818BFB62ECC9" box="[998,1246,1428,1454]" pageId="9" pageNumber="262">Municipality of Maria</collectingMunicipality>
Aurora, 
<location id="0949603D0B67FF9C5E16818AFF02ECB5" LSID="urn:lsid:plazi:treatment:843F87F00B6BFF9C5BDE827BFDFBEC91:0949603D0B67FF9C5E16818AFF02ECB5" country="Philippines" county="Luzon" latitude="15.681015" longLatPrecision="1" longitude="121.39388" municipality="Municipality of Maria" name="Barangay Villa" pageId="9" pageNumber="262" stateProvince="Aurora">Barangay Villa</location>
Aurora, 
<location id="0949603D0B67FF9C5A7481A7FE09ECB5" LSID="urn:lsid:plazi:treatment:843F87F00B6BFF9C5BDE827BFDFBEC91:0949603D0B67FF9C5A7481A7FE09ECB5" box="[290,437,1464,1490]" country="Philippines" county="Luzon" latitude="15.681015" longLatPrecision="1" longitude="121.39388" municipality="Municipality of Maria" name="Sitio Dimani" pageId="9" pageNumber="262" stateProvince="Aurora">Sitio Dimani</location>
, primary forest, 
<elevation id="87BBD1D50B67FF9C593981A7FC9AECB4" box="[623,806,1464,1491]" metricMagnitude="2" metricUnit="m" metricValue="5.35" pageId="9" pageNumber="262" unit="m" value="535.0">
elevation 
<quantity id="CB6E9B030B67FF9C598B81A7FC9AECB4" box="[733,806,1464,1491]" metricMagnitude="2" metricUnit="m" metricValue="5.35" pageId="9" pageNumber="262" unit="m" value="535.0">535 m</quantity>
</elevation>
, 
<geoCoordinate id="69A250210B67FF9C586481A7FC53ECB4" box="[818,1007,1464,1491]" degrees="15" direction="north" minutes="40" orientation="latitude" pageId="9" pageNumber="262" precision="1" seconds="51.654" value="15.681015">15°40’51.654”N</geoCoordinate>
, 
<geoCoordinate id="69A250210B67FF9C58AD81A7FB72ECB4" box="[1019,1230,1464,1491]" degrees="121" direction="east" minutes="23" orientation="longitude" pageId="9" pageNumber="262" precision="1" seconds="37.9932" value="121.39388">121°23’37.9932”E</geoCoordinate>
, 
<date id="782810260B67FF9C5F8A81A7FA1BECB5" box="[1244,1447,1464,1490]" pageId="9" pageNumber="262" value="2022-02-26">
<collectingDate id="686CE9CE0B67FF9C5F8A81A7FA1BECB5" box="[1244,1447,1464,1490]" pageId="9" pageNumber="262" value="2022-02-26">26 February 2022</collectingDate>
</date>
, 
<emphasis id="3EE2EAF40B67FF9C5BDE81C3FEF8EC91" box="[136,324,1500,1526]" italics="true" pageId="9" pageNumber="262">PTI-AMNP-1-46</emphasis>
(
<collectionCode id="6A87AE230B67FF9C5A0581C3FE31EC91" box="[339,397,1500,1526]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="9" pageNumber="262" type="Herbarium">PNH</collectionCode>
, 
<collectionCode id="6A87AE230B67FF9C5ACD81C3FE46EC91" box="[411,506,1500,1526]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15301" lsid="urn:lsid:biocol.org:col:15301" name="University of the Philippines Los Baños" pageId="9" pageNumber="262" type="Herbarium">CAHUP</collectionCode>
, 
<collectionCode id="6A87AE230B67FF9C595281C3FD87EC91" box="[516,571,1500,1526]" country="Philippines" httpUri="http://grbio.org/cool/07j4-sdqm" name="Palawan State University" pageId="9" pageNumber="262" type="Herbarium">PPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>