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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1023.61478" ID-GBIF-Dataset="94123c77-d2c7-4aa7-9bdb-8367f7851b82" ID-PMC="PMC7973068" ID-Pensoft-Pub="1313-2970-1023-81" ID-Pensoft-UUID="8E4EE48C84345AC48B877220002D94B6" ID-PubMed="33776516" ID-ZooBank="F14F12AC47824643929A49941E24BB92" ModsDocID="1313-2970-1023-81" checkinTime="1615560751875" checkinUser="pensoft" docAuthor="Hajek, Jiri, Shaverdo, Helena, Hendrich, Lars &amp; Balke, Michael" docDate="2021" docId="62862C9E0F9E5A779FF692F15E109E59" docLanguage="en" docName="ZooKeys 1023: 81-118" docOrigin="ZooKeys 1023" docPubDate="2021-03-10" docSource="http://dx.doi.org/10.3897/zookeys.1023.61478" docTitle="Copelatus baranensis Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke 2021, sp. nov." docType="treatment" docUuid="4A94D800-F6FB-47A6-A534-33E556BE1775" docUuidSource="ZooBank" docVersion="5" id="8E4EE48C84345AC48B877220002D94B6" lastPageNumber="81" masterDocId="8E4EE48C84345AC48B877220002D94B6" masterDocTitle="A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="118" masterPageNumber="81" pageNumber="81" updateTime="1668149894938" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hajek, Jiri</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5779-1542</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, National Museum, Cirkusova 1740, CZ- 193 00, Prague 9 - Horni Pocernice, Czech Republic</mods:affiliation>
<mods:nameIdentifier type="email">jiri.hajek@nm.cz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010, Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1023</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>118</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1023.61478</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1023.61478</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1023-81</mods:identifier>
<mods:identifier type="ZooBank">F14F12AC47824643929A49941E24BB92</mods:identifier>
<mods:identifier type="Pensoft-UUID">8E4EE48C84345AC48B877220002D94B6</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="180331290" LSID="urn:lsid:zoobank.org:act:4A94D800-F6FB-47A6-A534-33E556BE1775" httpUri="http://treatment.plazi.org/id/62862C9E0F9E5A779FF692F15E109E59" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="http://zoobank.org/4A94D800-F6FB-47A6-A534-33E556BE1775" authority="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke, 2021" authorityName="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baranensis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="baranensis" status="sp. nov.">Copelatus baranensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Habitus of Copelatus 1 C. baranensis sp. nov. (holotype; TL: 5.9 mm) 2 C. bougainvillensis sp. nov. (holotype; TL: 5.2 mm)." figureDoi="10.3897/zookeys.1023.61478.figures1-2" httpUri="https://binary.pensoft.net/fig/519048" pageId="0" pageNumber="81">Figures 1</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type locality">
<paragraph pageId="0" pageNumber="81">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<materialsCitation ID-GBIF-Occurrence="3063630342" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Solomon Islands" latitude="-9.466666" location="Barana Village area" longLatPrecision="129" longitude="159.97333" municipality="Mount Austine" specimenCount="1" typeStatus="holotype">
<collectingCountry name="Solomon Islands">Solomon Islands</collectingCountry>
,
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:3605233E249422882FFD02692792A967" country="Solomon Islands" latitude="-9.466666" longLatPrecision="129" longitude="159.97333" municipality="Mount Austine" name="Barana Village area">Barana Village area</location>
,
<collectingMunicipality>Mount Austine</collectingMunicipality>
,
<geoCoordinate degrees="09" direction="south" minutes="28.0" orientation="latitude" precision="92" value="-9.466666">09°28.0'S</geoCoordinate>
,
<geoCoordinate degrees="159" direction="east" minutes="58.4" orientation="longitude" precision="92" value="159.97333">159°58.4'E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3063630323" collectingDate="2013-11-23" collectingDateMax="2013-12-08" collectingDateMin="2013-11-23" collectionCode="NMPC" collectorName="Jiri Hajek" country="Solomon Islands" determinerName="Balke" elevation="280" latitude="-9.466666" location="Mt. Austine - Barana" longLatPrecision="129" longitude="159.97333" specimenCount="2" typeStatus="holotype">
<specimenCount>♂</specimenCount>
, labelled: &quot;
<collectingCountry name="Solomon Islands">Solomon Islands</collectingCountry>
,
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
/
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:3B0D08EC3132DCA613E4290DEB127D7B" country="Solomon Islands" latitude="-9.466666" longLatPrecision="129" longitude="159.97333" name="Mt. Austine - Barana">Mt. Austine - Barana</location>
vill. env. / (secondary forest, gardens, stream) /
<geoCoordinate degrees="09" direction="south" minutes="28.0" orientation="latitude" precision="92" value="-9.466666">09°28.0'S</geoCoordinate>
,
<geoCoordinate degrees="159" direction="east" minutes="58.4" orientation="longitude" precision="92" value="159.97333">159°58.4'E</geoCoordinate>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</elevation>
</quantity>
/
<collectorName>
<normalizedToken originalValue="Jiří">Jiri</normalizedToken>
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
</collectorName>
leg.,
<date value="2013-11-23" valueMax="2013-12-08" valueMin="2013-11-23">
<collectingDate value="2013-11-23" valueMax="2013-12-08" valueMin="2013-11-23">23.xi.-8.xii.2013</collectingDate>
</date>
[p] //
<typeStatus>HOLOTYPE</typeStatus>
<specimenCount>♂</specimenCount>
/
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="COPELATUS" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baranensis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="baranensis">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus baranensis</emphasis>
</taxonomicName>
sp. nov. /
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
, Shaverdo, Hendrich &amp;
<determinerName>Balke</determinerName>
det. 2020 [red label, p]&quot; (
<collectionCode country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<materialsCitation ID-GBIF-Occurrence="3063630334" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NMPC" collectorName="Jiri Hajek" country="Solomon Islands" determinerName="Balke" elevation="280" latitude="-9.466666" location="Mt. Austine - Barana" longLatPrecision="129" longitude="159.97333" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">3 ♀♀</specimenCount>
, same data as holotype (
<collectionCode country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063630352" collectingDate="2013-11-24" collectingDateMax="2013-12-14" collectingDateMin="2013-11-24" collectionCode="NHMUK, NHMW, NMPC, ZSMG" collectorName="Jiri Hajek" country="Solomon Islands" elevation="190" latitude="-9.496667" location="Barana" longLatPrecision="129" longitude="159.99167" specimenCount="31" specimenCount-female="17" specimenCount-male="14" typeStatus="paratype">
<specimenCount type="male">14 ♂♂</specimenCount>
,
<specimenCount type="female">17 ♀♀</specimenCount>
, labelled: &quot;
<collectingCountry name="Solomon Islands">Solomon Islands</collectingCountry>
,
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
/ ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="km" value="3.5">3.5 km</quantity>
SE of
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:7A4D17A5A1726ABF0AE2E9AA89B0EFFA" country="Solomon Islands" latitude="-9.496667" longLatPrecision="129" longitude="159.99167" name="Barana">Barana</location>
vill. / (drying up stream in shaded gorge) /
<geoCoordinate degrees="09" direction="south" minutes="29.8" orientation="latitude" precision="92" value="-9.496667">09°29.8'S</geoCoordinate>
,
<geoCoordinate degrees="159" direction="east" minutes="59.5" orientation="longitude" precision="92" value="159.99167">159°59.5'E</geoCoordinate>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.9" unit="m" value="190.0">190 m</elevation>
</quantity>
/
<collectorName>
<normalizedToken originalValue="Jiří">Jiri</normalizedToken>
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
</collectorName>
leg.,
<date value="2013-11-24" valueMax="2013-12-14" valueMin="2013-11-24">
<collectingDate value="2013-11-24" valueMax="2013-12-14" valueMin="2013-11-24">24.xi.-14.xii.2013</collectingDate>
</date>
[p]&quot; (
<collectionCode country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
,
<collectionCode country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW</collectionCode>
,
<collectionCode country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
,
<collectionCode>ZSMG</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063630343" collectingDate="2013-12-05" collectingDateMax="2013-12-06" collectingDateMin="2013-12-05" collectionCode="NMPC" collectorName="Jiri Hajek" country="Solomon Islands" elevation="275" latitude="-9.505" location="Moka river" longLatPrecision="129" longitude="159.98167" municipality="Japanese" specimenCount="12" specimenCount-female="6" specimenCount-male="6" typeStatus="paratype">
<specimenCount type="male">6 ♂♂</specimenCount>
,
<specimenCount type="female">6 ♀♀</specimenCount>
, labelled: &quot;
<collectingCountry name="Solomon Islands">Solomon Islands</collectingCountry>
,
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
/ ca
<locationDeviation location="Barana">
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.5" unit="km" value="4.5">4.5 km</quantity>
S of Barana
</locationDeviation>
vill., forest / nr. &quot;
<collectingMunicipality>Japanese</collectingMunicipality>
camp&quot; &amp;
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:22CA62ED17B9504A3535C8FE50995B48" country="Solomon Islands" latitude="-9.505" longLatPrecision="129" longitude="159.98167" municipality="Japanese" name="Moka river">Moka river</location>
/
<geoCoordinate degrees="09" direction="south" minutes="30.3" orientation="latitude" precision="92" value="-9.505">09°30.3'S</geoCoordinate>
,
<geoCoordinate degrees="159" direction="east" minutes="58.9" orientation="longitude" precision="92" value="159.98167">159°58.9'E</geoCoordinate>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.75" unit="m" value="275.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.75" unit="m" value="275.0">275 m</elevation>
</quantity>
/
<collectorName>
<normalizedToken originalValue="Jiří">Jiri</normalizedToken>
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
</collectorName>
leg.,
<date value="2013-12-05" valueMax="2013-12-06" valueMin="2013-12-05">
<collectingDate value="2013-12-05" valueMax="2013-12-06" valueMin="2013-12-05">5.-6.xii.2013</collectingDate>
</date>
[p]&quot; (
<collectionCode country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3063630350" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="LHCM" collectorName="St." country="Solomon Islands" elevation="200" location="Env." municipality="Barana" specimenCount="1" specimenCount-female="1" stateProvince="Honiara" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
, labelled: &quot;
<collectingCountry name="Solomon Islands">Solomon Islands</collectingCountry>
,
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
I. /
<collectingRegion country="Solomon Islands" name="Honiara">Honiara</collectingRegion>
reg.
<collectingMunicipality>Barana</collectingMunicipality>
vill.
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:23D24154A41AB7FEE9D57FE349B1EC7C" country="Solomon Islands" municipality="Barana" name="Env." stateProvince="Honiara">Env.</location>
/
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100-300 m</elevation>
</quantity>
XI-XII.2018 /
<collectorName>St.</collectorName>
Jakl leg.&quot; (
<collectionCode>LHCM</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3063630336" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Solomon Islands" location="All" specimenCount="1" stateProvince="Honiara" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:62862C9E0F9E5A779FF692F15E109E59:23657EDD5E6224E27B9C75E10BA81DD5" country="Solomon Islands" name="All" stateProvince="Honiara">All</location>
<typeStatus>paratypes</typeStatus>
with the respective printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">
Description of male
<typeStatus>holotype</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Habitus</emphasis>
:
</emphasis>
Elongate, oblong-oval, almost parallel-sided, broadest in mid-length of elytra; body moderately convex in lateral view. Body outline continuous, with only indistinct discontinuity between pronotum and elytra. Dorsal surface shiny (Fig.
<figureCitation captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Habitus of Copelatus 1 C. baranensis sp. nov. (holotype; TL: 5.9 mm) 2 C. bougainvillensis sp. nov. (holotype; TL: 5.2 mm)." figureDoi="10.3897/zookeys.1023.61478.figures1-2" httpUri="https://binary.pensoft.net/fig/519048" pageId="0" pageNumber="81">1</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1023.61478.figures1-2" httpUri="https://binary.pensoft.net/fig/519048" pageId="0" pageNumber="81" start="Figures 1, 2" startId="F1">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figures 1, 2.</emphasis>
Habitus of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="81">1</emphasis>
<taxonomicName lsidName="C. baranensis" pageId="0" pageNumber="81" rank="species" species="baranensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. baranensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>holotype</typeStatus>
; TL:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.9" unit="mm" value="5.9">5.9 mm</quantity>
)
<emphasis bold="true" pageId="0" pageNumber="81">2</emphasis>
<taxonomicName lsidName="C. bougainvillensis" pageId="0" pageNumber="81" rank="species" species="bougainvillensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. bougainvillensis</emphasis>
</taxonomicName>
sp. nov. (
<typeStatus>holotype</typeStatus>
; TL:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.2" unit="mm" value="5.2">5.2 mm</quantity>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Colouration</emphasis>
:
</emphasis>
Body colour brown blackish; head, sides of pronotum and appendages paler, ferruginous; base of elytra with irregular transverse testaceous band reaching neither suture nor lateral margin of elytra, band comb-like shaped due to dark colouration of elytral striae; appendages testaceous; ventral part brown blackish.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Head</emphasis>
:
</emphasis>
Moderately broad, ca. 0.64
<normalizedToken originalValue="×">x</normalizedToken>
width of pronotum, trapezoidal. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of moderately deeply impressed polygonal isodiametric meshes. Punctation double, consisting of coarse setigerous punctures, and very small punctures spread sparsely on surface; row of coarse punctures present around inner margin of eyes, several punctures present at frontal level of eyes, and anterolaterally to eyes in fronto-clypeal depressions.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Pronotum</emphasis>
:
</emphasis>
Transverse (width/length ratio = 2.54), broadest between posterior angles, lateral margins moderately curved. Sides with lateral beading very thin and indistinct. Reticulation similar to that of head. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, laterally in longitudinal depression close to sides, several punctures present also in basolateral depressions along basal margin. Disc of pronotum laterally with long, irregularly distributed longitudinal strioles; several strioles present also in depressions close to posterior angles. Centre of disc with medial longitudinal smooth line.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Elytra</emphasis>
:
</emphasis>
Base of elytra as broad as pronotal base; lateral margins of elytra subparallel in basal two thirds, curved in apical third. Eleven longitudinal discal striae present on each elytron: stria 1 short, beginning at a fourth of elytral length and ending approximately before apical fifth of elytral length; striae 2 and 3 absent at base; striae 4-6 beginning at base; stria 7 fragmented basally; stria 8 complete; stria 9 absent at base and ending at apical fifth of elytra; stria 10 fragmented into several small strioles, hardly perceptible; stria 11 fragmented apically; all odd striae generally shortened apically. Submarginal stria absent. Surface reticulation similar to that of head and pronotum, meshes slightly smaller and less impressed. Punctation double, consisting of row of coarse setigerous punctures along elytral striae 4, 6, 8 and along lateral margin of elytra, and very fine sparsely distributed punctures.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Legs</emphasis>
:
</emphasis>
Protibia simple, slightly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive setae on their ventral side.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Ventral side</emphasis>
:
</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation consisting of shallow, hardly perceptible polygonal meshes. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (&quot;metasternal wings&quot;) tongue-shaped, slender. Metacoxal lines well impressed, nearly complete, absent only close to metaventrite. Metacoxal plates covered with long, deep longitudinal strioles; reticulation consisting of elongated, longitudinal polygonal meshes. Metacoxal processes rounded and indistinctly incised at posterior margin. Abdominal ventrites I and II with longitudinal strioles; ventrites III and IV with oblique strioles laterally. Tuft of setae present antero-medially on ventrites III-V; ventrite VI with setigerous punctures laterally on either side. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I and II, oblique on ventrite III and transverse on ventrites IV-VI. Punctation consisting of fine, sparsely distributed punctures; punctures coarser and denser laterally on apical ventrite.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Genitalia</emphasis>
:
</emphasis>
Median lobe of aedeagus (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">15</figureCitation>
) sickle-shaped, with obtuse apex and distinct rugose surface sculpture visible in lateral view (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">15A, B</figureCitation>
); consisting of dorsal and ventral sclerites; dorsal sclerite divided into two parts of different shape in apical half, left part slightly shorter than right one, with lateral margin distinctly curved and apex more or less rounded (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">15C</figureCitation>
); ventral sclerite with left part strongly sclerotised, its apex in shape of a small, weak hook visible in lateral left view (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">15B</figureCitation>
), right part membranous; apexes of dorsal and ventral sclerites more or less pressed together.
</paragraph>
<paragraph pageId="0" pageNumber="81">
Lateral lobes (parameres) of narrow triangular form, with almost straight setigerous dorsal margin; setae numerous, dense, and strong distally, and distinctly less numerous, weaker, and sparser basally (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Male genitalia of Copelatus baranensis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure15" httpUri="https://binary.pensoft.net/fig/519054" pageId="0" pageNumber="81">15D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Female.</emphasis>
Identical to male in habitus. Pro- and mesotarsomeres not broadened, without adhesive setae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="variability">
<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
<paragraph pageId="0" pageNumber="81">
The specimens of the
<typeStatus>type</typeStatus>
series vary in extent of the basal testaceous band on elytra. There is also a variation in number and position of longitudinal strioles on pronotum. The highest variability is however, in elytral striation: from complete eleven striae to largely fragmented striae 1, 2, 4, 6, 7, 9, 10; in rare case, stria 1 completely absent; in several specimens, submarginal stria (or several short striae) present at the level of apical fifth of elytral length, sometimes only on one side.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="81">
TL:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.2" metricValueMin="5.4" unit="mm" value="5.8" valueMax="6.2" valueMin="5.4">5.4-6.2 mm</quantity>
(mean value:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="5.9" metricValueMin="5.5" unit="mm" value="5.7" valueMax="5.9" valueMin="5.5">
5.7
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2 mm
</quantity>
);
<typeStatus>holotype</typeStatus>
:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.9" unit="mm" value="5.9">5.9 mm</quantity>
. TL-h:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.7" metricValueMin="4.9" unit="mm" value="5.3" valueMax="5.7" valueMin="4.9">4.9-5.7 mm</quantity>
(mean value:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.2" metricValueMax="5.4" metricValueMin="5.0" unit="mm" value="5.2" valueMax="5.4" valueMin="5.0">
5.2
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2 mm
</quantity>
);
<typeStatus>holotype</typeStatus>
:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.4" unit="mm" value="5.4">5.4 mm</quantity>
. MW:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4-2.9 mm</quantity>
(mean value:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">
2.6
<normalizedToken originalValue="±">+/-</normalizedToken>
0.1 mm
</quantity>
);
<typeStatus>holotype</typeStatus>
:
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.6" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
Based on the presence of 11 dorsal elytral striae and absence of submarginal stria, the new species can be classified within
<taxonomicName lsidName="C. nigrolineatus" pageId="0" pageNumber="81" rank="species" species="nigrolineatus">
<emphasis italics="true" pageId="0" pageNumber="81">C. nigrolineatus</emphasis>
</taxonomicName>
species group. This group contain worldwide only five species: two in Neotropics, one in southern
<collectingCountry name="China">China</collectingCountry>
, one in
<collectingCountry name="India">India</collectingCountry>
, and one species in
<collectingCountry name="Australia">Australia</collectingCountry>
(
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B16" refString="Nilsson, AN, Hajek, J, 2020. A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2020. Distributed as a PDF file via Internet. http://www.waterbeetles.eu" title="A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2020. Distributed as a PDF file via Internet." url="http://www.waterbeetles.eu" year="2020">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2020
</bibRefCitation>
). On the other hand, presence of a submarginal stria in some specimens indicates affiliation of the new species to the
<taxonomicName lsidName="C. trilobatus" pageId="0" pageNumber="81" rank="species" species="trilobatus">
<emphasis italics="true" pageId="0" pageNumber="81">C. trilobatus</emphasis>
</taxonomicName>
species group. The
<taxonomicName lsidName="C. trilobatus" pageId="0" pageNumber="81" rank="species" species="trilobatus">
<emphasis italics="true" pageId="0" pageNumber="81">C. trilobatus</emphasis>
</taxonomicName>
species group includes up to now 24 species occurring in tropics of all continents; two species of the group are occurring in the
<collectingCountry name="Papua New Guinea">Papua New Guinea</collectingCountry>
, one species in
<collectingCountry name="Fiji">Fiji</collectingCountry>
, and three species are known from
<collectingCountry name="Australia">Australia</collectingCountry>
(
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B16" refString="Nilsson, AN, Hajek, J, 2020. A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2020. Distributed as a PDF file via Internet. http://www.waterbeetles.eu" title="A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2020. Distributed as a PDF file via Internet." url="http://www.waterbeetles.eu" year="2020">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2020
</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
Based on shape of the male genitalia,
<taxonomicName lsidName="C. baranensis" pageId="0" pageNumber="81" rank="species" species="baranensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. baranensis</emphasis>
</taxonomicName>
sp. nov. is not similar to any of the species currently included in the
<taxonomicName lsidName="C. nigrolineatus" pageId="0" pageNumber="81" rank="species" species="nigrolineatus">
<emphasis italics="true" pageId="0" pageNumber="81">C. nigrolineatus</emphasis>
</taxonomicName>
group. On the other hand, it is most likely very close to
<taxonomicName lsidName="C. bougainvillensis" pageId="0" pageNumber="81" rank="species" species="bougainvillensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. bougainvillensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="C. kietensis" pageId="0" pageNumber="81" rank="species" species="kietensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. kietensis</emphasis>
</taxonomicName>
sp. nov., and to the numerous undescribed species of the
<taxonomicName lsidName="C. trilobatus" pageId="0" pageNumber="81" rank="species" species="trilobatus">
<emphasis italics="true" pageId="0" pageNumber="81">C. trilobatus</emphasis>
</taxonomicName>
species group from New
<collectingCountry name="Guinea-Bissau">Guinea</collectingCountry>
(H. Shaverdo et al., in preparation). For separation of
<taxonomicName lsidName="C. baranensis" pageId="0" pageNumber="81" rank="species" species="baranensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. baranensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="C. bougainvillensis" pageId="0" pageNumber="81" rank="species" species="bougainvillensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. bougainvillensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="C. kietensis" pageId="0" pageNumber="81" rank="species" species="kietensis">
<emphasis italics="true" pageId="0" pageNumber="81">C. kietensis</emphasis>
</taxonomicName>
sp. nov., see under the two latter species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="etymology">
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="81">The new species is named after Barana Village, in the vicinity of which the new species was collected. In this way, the first author would like to thank the people of Barana for their company and help with collecting during the 2013 trip. The specific epithet is an adjective in the nominative singular.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">
The species is known so far only from the small area around Barana Village, south from
<collectingRegion country="Solomon Islands" name="Honiara">Honiara</collectingRegion>
City, northern
<taxonomicName genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="habitat">
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="81">
At the
<typeStatus>type</typeStatus>
locality, the species was collected in small shaded pools of a forest stream (Figs
<figureCitation captionStart="Figures 26–28" captionStartId="F16" captionText="Figures 26 - 28. Habitats of Copelatus in Guadalcanal 26 temporary forest stream near Barana village 27 shaded pool on the same stream 28 small temporary puddles in a gorge along the road from Barana to Lungga river." figureDoi="10.3897/zookeys.1023.61478.figures26-28" httpUri="https://binary.pensoft.net/fig/519063" pageId="0" pageNumber="81">26</figureCitation>
,
<figureCitation captionStart="Figures 26–28" captionStartId="F16" captionText="Figures 26 - 28. Habitats of Copelatus in Guadalcanal 26 temporary forest stream near Barana village 27 shaded pool on the same stream 28 small temporary puddles in a gorge along the road from Barana to Lungga river." figureDoi="10.3897/zookeys.1023.61478.figures26-28" httpUri="https://binary.pensoft.net/fig/519063" pageId="0" pageNumber="81">27</figureCitation>
). At the other places, the specimens were collected in puddles/pools with muddy bottom made by a temporary forest stream (Fig.
<figureCitation captionStart="Figures 26–28" captionStartId="F16" captionText="Figures 26 - 28. Habitats of Copelatus in Guadalcanal 26 temporary forest stream near Barana village 27 shaded pool on the same stream 28 small temporary puddles in a gorge along the road from Barana to Lungga river." figureDoi="10.3897/zookeys.1023.61478.figures26-28" httpUri="https://binary.pensoft.net/fig/519063" pageId="0" pageNumber="81">28</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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