266 lines
26 KiB
XML
266 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.62245" ID-GBIF-Dataset="71bbf3b9-a613-49cc-8971-e04a31a448b8" ID-PMC="PMC8222211" ID-Pensoft-Pub="1313-2970-1044-41" ID-Pensoft-UUID="535A79F3B89F5699A71E2B83B65C4C21" ID-PubMed="34183875" ID-ZooBank="C6AE8C5BC5D64A09A26E00FAADBF86E1" ModsDocID="1313-2970-1044-41" checkinTime="1623875478834" checkinUser="pensoft" docAuthor="Kavanaugh, David H., Maddison, David R., Simison, W. Brian, Schoville, Sean D., Schmidt, Joachim, Faille, Arnaud, Moore, Wendy, Pflug, James M., Archambeault, Sophie L., Hoang, Tinya & Chen, Jei-Ying" docDate="2021" docId="6226343FCCF3514DB574B8B0341021C8" docLanguage="en" docName="ZooKeys 1044: 41-152" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.62245" docTitle="Nebria (Erwinebria) Kavanaugh 2021, subgen. nov." docType="treatment" docUuid="12F2832A-C3A8-44B3-9909-32482CEBFE6B" docUuidSource="ZooBank" docVersion="5" id="535A79F3B89F5699A71E2B83B65C4C21" lastPageNumber="41" masterDocId="535A79F3B89F5699A71E2B83B65C4C21" masterDocTitle="Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data" masterLastPageNumber="152" masterPageNumber="41" pageNumber="41" updateTime="1668150468173" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Phylogeny of the supertribe Nebriitae (Coleoptera, Carabidae) based on analyses of DNA sequence data</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kavanaugh, David H.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="email">dkavanaugh@calacademy.org</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Maddison, David R.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7152-3824</mods:nameIdentifier>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Simison, W. Brian</mods:namePart>
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<mods:affiliation>Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schoville, Sean D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7364-434X</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Wisconsin, Madison, WI 53706, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schmidt, Joachim</mods:namePart>
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<mods:affiliation>Institute of Biosciences, University of Rostock, Universitaetsplatz 2, D- 18055 Rostock, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Faille, Arnaud</mods:namePart>
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<mods:affiliation>Department of Entomology, Coleoptera, Stuttgart State Museum of Natural History, Rosenstein 1, 70191 Stuttgart, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moore, Wendy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4441-6203</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, University of Arizona, Tucson, AZ 85721 - 0036, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pflug, James M.</mods:namePart>
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<mods:affiliation>Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Archambeault, Sophie L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Berkeley, 142 Weill Hall # 3200, Berkeley, CA 94720, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hoang, Tinya</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Jei-Ying</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA & University of California, Santa Cruz, Long Marine Lab, 117 McAllister Way, Santa Cruz, CA 95060, USA</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-16</mods:number>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>152</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.62245</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.62245</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-41</mods:identifier>
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<mods:identifier type="ZooBank">C6AE8C5BC5D64A09A26E00FAADBF86E1</mods:identifier>
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<treatment ID-GBIF-Taxon="182754136" LSID="urn:lsid:zoobank.org:act:12F2832A-C3A8-44B3-9909-32482CEBFE6B" httpUri="http://treatment.plazi.org/id/6226343FCCF3514DB574B8B0341021C8" lastPageNumber="41" pageId="0" pageNumber="41">
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<subSubSection pageId="0" pageNumber="41" type="nomenclature">
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<paragraph pageId="0" pageNumber="41">
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Subgenus
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<taxonomicName LSID="http://zoobank.org/12F2832A-C3A8-44B3-9909-32482CEBFE6B" authority="Kavanaugh," authorityName="Kavanaugh" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria (Erwinebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Erwinebria">Nebria Erwinebria Kavanaugh,</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="41">subgen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="reference_group">
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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= "
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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group" sensu
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Proceedings of a symposium (28 August, 1996, Florence, Italy). XX International Congress of Entomology. Museo Regionale di Scienze Naturali, Torino" pageId="0" pageNumber="41" refId="B127" refString="Lindroth, CH, 1961. The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No.20: 1-200." title="The ground-beetles (Carabidae, excl. Cicindelinae) of Canada and Alaska. Part 2. Opuscula Entomologica Supplementum No. 20: 1 - 200." year="1961">Lindroth 1961</bibRefCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus" status="subgen. nov.">Reductonebria</taxonomicName>
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Shilenkov, 1975 (in part); = "
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria sahlbergi" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="sahlbergi" status="subgen. nov.">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria sahlbergi</emphasis>
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</taxonomicName>
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" + "
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<taxonomicName authority="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen, 2021" authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Nebria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nebria gregaria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="gregaria" status="subgen. nov.">Nebria gregaria</taxonomicName>
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" groups sensu
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<bibRefCitation author="Ledoux, G" journalOrPublisher="Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" pageId="0" pageNumber="41" refId="B121" refString="Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" title="Nebria (Coleoptera, Nebriidae). Faune mondiale." year="2005">Ledoux and Roux 2005</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="type species">
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<paragraph pageId="0" pageNumber="41">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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<taxonomicName authorityName="Fischer von Waldheim" authorityYear="1828" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria sahlbergii" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="sahlbergii">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria sahlbergii</emphasis>
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</taxonomicName>
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Fischer von Waldheim, 1828:254, by present designation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="diagnosis">
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<paragraph pageId="0" pageNumber="41">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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Body size small to medium, SBL = 7.1 to 11.7 mm. Head width medium to slightly broadened, not constricted behind eyes; vertex with a pair of paramedial pale spots and a single pair of supraorbital setae. Eyes slightly reduced to moderate in size, moderately to markedly convex. Antennal scape with one (two in very few specimens) subapicodorsal seta; antennomeres 3 and 4 not laterally compressed, without extra setae. Labrum with three pairs of apical setae. Maxillary stipes typical for genus, with setae inserted flush on smooth surface. Penultimate labial palpomere with three setae. Pronotum with midlateral setae absent, a single pair of basolateral setae present. Elytral intervals smooth, without macrosculpture, interval 3 with three to eight or more setiferous pores, intervals 5 and 7 without setiferous pores, interval 3 not or only faintly catenate, intervals 5 and 7 not catenate. Hindwings full-sized or slightly to markedly reduced in size. Metepisterna smooth or faintly punctulate. Protarsomeres 1-3 expanded in males; mesotarsomeres 2-4 longer than their apical width; tarsi dorsally glabrous or with a few fine setae. Abdominal sternites IV-VI with two to four pairs of posterior paramedial setae, without paralateral setae. Median lobe of male aedeagus sclerotized dorsally at least to midlength on shaft, moderately to markedly deflected right in dorsal aspect; basal bulb expanded, quadrate, broadly open basally and closed dorsally, without a sagittal aileron present at base or with only a lightly sclerotized collar; mid-shaft parallel-sided or slightly to moderately narrowed basally in lateral aspect, slightly compressed in cross-section, with right lateral face unmodified; apical orifice markedly to extremely deflected right. Right paramere slender and moderately long. Female valvifers with vestiture (a few specimens of some species without vestiture); gonopods VIII fused to dorsomedial bases of gonocoxae; gonocoxae with ventral diagonal row of setiform setae and mediodorsal row of setae present. Bursa copulatrix without dorsal sclerites in vestibular chamber and with longitudinal axis of the bursa moderately to markedly sigmoid dorsally in lateral aspect; spermathecal chamber broadly cordate in dorsal aspect, without (in
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<taxonomicName authorityName="Van Dyke" authorityYear="1925" class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria lyelli" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="lyelli">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria lyelli</emphasis>
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</taxonomicName>
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Van Dyke, 1925 and
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<taxonomicName class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria quileute" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="species" species="quileute">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria quileute</emphasis>
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</taxonomicName>
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Kavanaugh, 1979) or with a small to large midline sclerotized dorsal plate, insemination duct not sclerotized; spermathecal duct medium-length to slightly long, uniform in diameter throughout or nearly so, inserted basodorsally on spermathecal chamber; spermathecal reservoir of medium length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="etymology">
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<paragraph pageId="0" pageNumber="41">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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The subgeneric epithet is a noun of feminine gender and a combination of the surname of Terry
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<taxonomicName genus="L." lsidName="L." pageId="0" pageNumber="41" rank="genus">L.</taxonomicName>
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Erwin, in whose honor we name this subgenus, and the genus name,
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<taxonomicName class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="41" type="remarks">
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<paragraph pageId="0" pageNumber="41">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="41">
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Members of this group are easily identified as members of the
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
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</taxonomicName>
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Complex using the
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<bibRefCitation author="Ledoux, G" journalOrPublisher="Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" pageId="0" pageNumber="41" refId="B121" refString="Ledoux, G, Roux, P, 2005. Nebria (Coleoptera, Nebriidae). Faune mondiale. Museum - Centre de Conservation et d'Etude des Collections et Societe Linneenne de Lyon, Lyon" title="Nebria (Coleoptera, Nebriidae). Faune mondiale." year="2005">
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Ledoux and
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<normalizedToken originalValue="Roux’s">Roux's</normalizedToken>
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(2005)
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</bibRefCitation>
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key to subgenera. Features that distinguish them from members of subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria (Insulanebria)" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="subGenus" subGenus="Insulanebria">Reductonebria Insulanebria</taxonomicName>
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have been discussed above for that taxon. We have not yet identified any external morphological features that consistently distinguish all members of all species of
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
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</taxonomicName>
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from all
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<taxonomicName class="Insecta" family="Carabidae" genus="Erwinebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erwinebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
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</taxonomicName>
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members, but there are several distinguishing internal features evident in both sexes. The mid-shaft is parallel-sided or slightly to moderately narrowed basally and the apical orifice is markedly to extremely deflected right in
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<taxonomicName class="Insecta" family="Carabidae" genus="Erwinebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erwinebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
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</taxonomicName>
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males. In
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<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
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</taxonomicName>
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males, the mid-shaft is markedly narrowed apically and the apical orifice is only moderately deflected right. As in most other
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<taxonomicName class="Insecta" family="Carabidae" genus="Nebria" kingdom="Animalia" lsidName="Nebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="41">Nebria</emphasis>
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</taxonomicName>
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examined, the gonopods VIII in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Erwinebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erwinebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females are fused dorsomedially with the bases of gonocoxae. In
|
||
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females, these gonopods are distinctly separate from, although in the same position in relation to, the gonocoxae. The spermathecal chamber is broadly cordate in dorsal aspect in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Erwinebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erwinebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females, but somewhat broadly ovoid in
|
||
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females. Although most members of both of the subgenera have sclerotized plates in the dorsal or dorsoapical wall of the spermathecal chamber of bursa copulatrix, they are of a different form in the two groups. In
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Erwinebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Erwinebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Erwinebria</emphasis>
|
||
</taxonomicName>
|
||
females (except in
|
||
<taxonomicName lsidName="N. lyelli" pageId="0" pageNumber="41" rank="species" species="lyelli">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">N. lyelli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="N. quileute" pageId="0" pageNumber="41" rank="species" species="quileute">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">N. quileute</emphasis>
|
||
</taxonomicName>
|
||
, which have no dorsal sclerites), the dorsal sclerite is variously formed as a small to large domed, horseshoe- or saddle-shaped plate associated with the point insertion of the spermathecal duct on the chamber. In
|
||
<taxonomicName authorityName="Kavanaugh & Maddison & Simison & Schoville & Schmidt & Faille & Moore & Pflug & Archambeault & Hoang & Chen" authorityYear="2021" class="Insecta" family="Carabidae" genus="Reductonebria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Reductonebria" order="Coleoptera" pageId="0" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">Reductonebria</emphasis>
|
||
</taxonomicName>
|
||
females, the sclerotized area is predominately an apicodorsal cap on the spermathecal chamber. This cap sclerite may be relatively small, round, and smooth, as in
|
||
<taxonomicName lsidName="N. ochotica" pageId="0" pageNumber="41" rank="species" species="ochotica">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">N. ochotica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="N. mannerheimii" pageId="0" pageNumber="41" rank="species" species="mannerheimii">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">N. mannerheimii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="N. darlingtoni" pageId="0" pageNumber="41" rank="species" species="darlingtoni">
|
||
<emphasis italics="true" pageId="0" pageNumber="41">N. darlingtoni</emphasis>
|
||
</taxonomicName>
|
||
females, or larger, crenulated rather than smooth, and/or expanded laterally or basally to or beyond the insertion point of the spermathecal duct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="41" type="distribution">
|
||
<paragraph pageId="0" pageNumber="41">Known distribution and diversity.</paragraph>
|
||
<paragraph pageId="0" pageNumber="41">The geographical range of this clade covers much of western North America, from the outer Aleutian Islands in the northwest, south to eastcentral California, northern Arizona, and New Mexico and east to the eastern edge of the Rocky Mountain system from Yukon Territory in Canada south to northcentral New Mexico.</paragraph>
|
||
<paragraph pageId="0" pageNumber="41">Twenty-one species in this clade have been described.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |