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<document id="53AF056FCF72A38C934F69C1D6102658" ID-DOI="10.11646/zootaxa.3941.3.1" ID-GBIF-Dataset="67bc3c9d-ec99-4b47-81df-3fca7bd584ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="243860" ID-ZooBank="9907D081-A413-44F1-ABF6-76CC13DC8813" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461026017375" checkinUser="plazi" docAuthor="Wang, Cheng-Bin &amp; Zhou, Hong-Zhang" docDate="2015" docId="03EC87E6FF87881F9DBF1D67FB210F50" docLanguage="en" docName="zt03941p338.pdf" docOrigin="Zootaxa 3941 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptomaphaginus Portevin 1914" docType="treatment" docVersion="9" lastPageNumber="303" masterDocId="FFD5FF9EFF85881A9D281927FFF50E4C" masterDocTitle="Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species" masterLastPageNumber="338" masterPageNumber="301" pageNumber="303" updateTime="1698572131055" updateUser="plazi">
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<mods:title id="4EA244828399FDA163B9819BDEA4811B">Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) from China, with description of eleven new species</mods:title>
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<mods:namePart id="734754E390DDD36F11ABD1CCC991A910">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="73CC5382AFF665C6D6B7062CC9C0CFD8">Zhou, Hong-Zhang</mods:namePart>
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<treatment id="03EC87E6FF87881F9DBF1D67FB210F50" ID-DOI="http://doi.org/10.5281/zenodo.5691653" ID-GBIF-Taxon="119606333" ID-Zenodo-Dep="5691653" LSID="urn:lsid:plazi:treatment:03EC87E6FF87881F9DBF1D67FB210F50" httpUri="http://treatment.plazi.org/id/03EC87E6FF87881F9DBF1D67FB210F50" lastPageId="5" lastPageNumber="303" pageId="2" pageNumber="303">
<subSubSection id="C35F657BFF8788189DBF1D67FBDC0AF7" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="8BFA36F0FF8788189DBF1D67FD9C0A16" blockId="2.[151,617,1088,1114]" box="[151,617,1088,1114]" pageId="2" pageNumber="303">
<heading id="D0B2819CFF8788189DBF1D67FD9C0A16" bold="true" box="[151,617,1088,1114]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
<emphasis id="B931EAE2FF8788189DBF1D67FD9C0A16" bold="true" box="[151,617,1088,1114]" pageId="2" pageNumber="303">
Genus
<taxonomicName id="4C454D73FF8788189DC61D67FD9C0A16" ID-CoL="8X7S4" authority="Portevin, 1914" authorityName="Portevin" authorityYear="1914" box="[238,617,1088,1114]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8788189DC61D67FE590A16" bold="true" box="[238,428,1088,1114]" italics="true" pageId="2" pageNumber="303">Ptomaphaginus</emphasis>
Portevin, 1914
</taxonomicName>
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</paragraph>
<paragraph id="8BFA36F0FF8788189DBF1DA1FBDC0AF7" blockId="2.[151,1436,1158,1303]" pageId="2" pageNumber="303">
<bibRefCitation id="EFD44B01FF8788189DBF1DA1FEC90AD0" author="Portevin" box="[151,316,1158,1180]" pageId="2" pageNumber="303" refString="Portevin, G. (1914 a) Silphides et Liodides nouveaux. Annales de la Societe Entomologique de Belgique Bruxelles, 58, 190 - 198." type="journal article" year="1914" yearSuffix="a">Portevin, 1914a</bibRefCitation>
: 194 (species included:
<taxonomicName id="4C454D73FF8788189F181DA0FD690AD0" box="[560,668,1159,1180]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="longitarsis">
<emphasis id="B931EAE2FF8788189F181DA0FD690AD0" box="[560,668,1159,1180]" italics="true" pageId="2" pageNumber="303">longitarsis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFD44B01FF8788189F9B1DA1FCB70AD0" author="Jeannel" box="[691,834,1158,1180]" pageId="2" pageNumber="303" refString="Jeannel, R. (1936) Monographie des Catopidae (Insectes Coleopteres). Memoires du Museum National d'Histoire Naturelle, (N. S.) 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
: 54 (characters; key to species);
<bibRefCitation id="EFD44B01FF87881899B81DA1FA8D0AD0" author="Szymczakowski" box="[1168,1400,1158,1180]" pageId="2" pageNumber="303" refString="Szymczakowski, W. (1964) Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale. Acta Zoologica Cracoviensia, 9 (2), 55 - 289." type="journal article" year="1964">Szymczakowski, 1964</bibRefCitation>
: 66 (phylogeny; key to species; revision);
<bibRefCitation id="EFD44B01FF8788189F621D81FD2E0AF7" author="Perreau" box="[586,731,1189,1211]" pageId="2" pageNumber="303" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae. Memories de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau, 2000</bibRefCitation>
: 370 (world catalog; 72 species).
</paragraph>
</subSubSection>
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<paragraph id="8BFA36F0FF8788189DBF1DE3FCC70A96" blockId="2.[151,1436,1158,1303]" box="[151,818,1220,1242]" pageId="2" pageNumber="303">
<emphasis id="B931EAE2FF8788189DBF1DE3FED30A96" bold="true" box="[151,294,1220,1242]" pageId="2" pageNumber="303">
<typeStatus id="54FE8852FF8788189DBF1DE3FF390A96" box="[151,204,1220,1242]" pageId="2" pageNumber="303">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C454D73FF8788189C031DE3FD6E0A96" authority="Portevin" authorityName="Portevin" box="[299,667,1220,1242]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="longitarsis">
<emphasis id="B931EAE2FF8788189C031DE3FDCB0A95" box="[299,574,1220,1241]" italics="true" pageId="2" pageNumber="303">Ptomaphaginus longitarsis</emphasis>
Portevin
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8BFA36F0FF8788189DBF1DC4FD870B5B" blockId="2.[151,1436,1158,1303]" pageId="2" pageNumber="303">
<emphasis id="B931EAE2FF8788189DBF1DC4FEF80AB5" bold="true" box="[151,269,1251,1273]" pageId="2" pageNumber="303">Synonymy.</emphasis>
<taxonomicName id="4C454D73FF8788189C3C1DC4FD9A0AB5" authority="Hatch, 1928: 158" authorityName="Hatch" authorityPageNumber="158" authorityYear="1928" box="[276,623,1251,1273]" class="Insecta" family="Leiodidae" genus="Euptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8788189C3C1DC4FE4C0AB4" box="[276,441,1251,1272]" italics="true" pageId="2" pageNumber="303">Euptomaphagus</emphasis>
<bibRefCitation id="EFD44B01FF8788189C971DC4FDCA0AB5" author="Hatch" box="[447,575,1251,1273]" pageId="2" pageNumber="303" refString="Hatch, M. H. (1928) Coleopterum Catalogus, Pars 95: Silphidae II. W. Junk, Berlin, 244 pp." type="book" year="1928">Hatch, 1928</bibRefCitation>
: 158
</taxonomicName>
(
<typeStatus id="54FE8852FF8788189F541DC4FD450AB5" box="[636,688,1251,1273]" pageId="2" pageNumber="303">Type</typeStatus>
species:
<taxonomicName id="4C454D73FF8788189E251DC4FBCA0AB5" authority="Portevin" authorityName="Portevin" box="[781,1087,1251,1273]" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="cilipes">
<emphasis id="B931EAE2FF8788189E251DC4FC140AB4" box="[781,993,1251,1272]" italics="true" pageId="2" pageNumber="303">Ptomaphagus cilipes</emphasis>
Portevin
</taxonomicName>
, by original designation);
<bibRefCitation id="EFD44B01FF87881898621DC4FF080B5B" author="Jeannel" pageId="2" pageNumber="303" refString="Jeannel, R. (1936) Monographie des Catopidae (Insectes Coleopteres). Memoires du Museum National d'Histoire Naturelle, (N. S.) 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
: 54 (synonym with
<taxonomicName id="4C454D73FF8788189CED1C26FD960B5A" box="[453,611,1281,1302]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAE2FF8788189CED1C26FD960B5A" box="[453,611,1281,1302]" italics="true" pageId="2" pageNumber="303">Ptomaphaginus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BFA36F0FF8788189DBF1C63FF1A08A5" blockId="2.[151,1437,1348,2021]" pageId="2" pageNumber="303">
The members of this genus can be distinguished from all other genera within the tribe
<taxonomicName id="4C454D73FF87881899821C63FAB30B11" box="[1194,1350,1348,1373]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="tribe" tribe="Ptomaphagini">Ptomaphagini</taxonomicName>
by the combination of the following characters (according to
<bibRefCitation id="EFD44B01FF8788189FD41C4FFC6F0BCD" author="Jeannel" box="[764,922,1384,1409]" pageId="2" pageNumber="303" refString="Jeannel, R. (1936) Monographie des Catopidae (Insectes Coleopteres). Memoires du Museum National d'Histoire Naturelle, (N. S.) 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
;
<bibRefCitation id="EFD44B01FF8788189E8D1C4FFB500BCD" author="Szymczakowski" box="[933,1189,1384,1409]" pageId="2" pageNumber="303" refString="Szymczakowski, W. (1964) Analyse systematique et zoogeographique des Catopidae (Coleoptera) de la region orientale. Acta Zoologica Cracoviensia, 9 (2), 55 - 289." type="journal article" year="1964">Szymczakowski, 1964</bibRefCitation>
;
<bibRefCitation id="EFD44B01FF87881899991C4FFAC50BCC" author="Peck" box="[1201,1328,1384,1409]" pageId="2" pageNumber="303" refString="Peck, S. B. (1973) A systematic revision and the evolutionary biology of the Ptomaphagus (Adelops) beetles of North America (Coleoptera; Leiodidae; Catopinae), with emphasis on the cave-inhabiting species. Bulletin of the Museum of Comparative Zoology, 145 (2), 29 - 162." type="journal article" year="1973">Peck, 1973</bibRefCitation>
;
<bibRefCitation id="EFD44B01FF87881898141C4EFF270BE8" author="Newton" pageId="2" pageNumber="303" refString="Newton, A. F. (1998) Phylogenetic problems, current classification and generic catalog of world Leiodidae (including Cholevidae). In: Gnaspini, P. &amp; Peck, S. B. (Eds.), Phylogeny and evolution of subterranean and endogean Cholevidae (= Leiodidae Cholevinae), Proceedings of XX International Congress of Entomology, Firenze, 1996. Atti del Museo regionale di Scienze naturali, Torino, pp. 41 - 178." type="journal article" year="1998">Newton, 1998</bibRefCitation>
;
<bibRefCitation id="EFD44B01FF8788189DCC1CABFE6C0BE8" author="Gnaspini" box="[228,409,1420,1445]" pageId="2" pageNumber="303" refString="Gnaspini, P. (1998) Subterranean evolution and phylogenetic relationships within the tribe Ptomaphagini (Coleoptera, Leiodidae, Cholevinae). In: Gnaspini, P. &amp; Peck, S. B. (Eds.), Phylogeny and evolution of subterranean and endogean Cholevidae (= Leiodidae Cholevinae), Proceedings of XX International Congress of Entomology, Firenze, 1996. Atti del Museo regionale di Scienze naturali, Torino, pp. 227 - 234." type="journal article" year="1998">Gnaspini, 1998</bibRefCitation>
;
<bibRefCitation id="EFD44B01FF8788189C841CAAFDA60BE8" author="Perreau" box="[428,595,1420,1444]" pageId="2" pageNumber="303" refString="Perreau, M. (2000) Catalogue des Coleopteres Leiodidae, Cholevinae et Platypsyllinae. Memories de la Societe entomologique de France, 4, 1 - 460." type="journal article" year="2000">Perreau, 2000</bibRefCitation>
): (1) size small,
<quantity id="4CBD9B15FF8788189E0D1CABFC430BE8" box="[805,950,1420,1445]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.5" metricValueMin="1.8" pageId="2" pageNumber="303" unit="mm" value="2.65" valueMax="3.5" valueMin="1.8">1.8–3.5 mm</quantity>
; (2) compound eyes always present; (3) pronotum with transverse striolations, interspaces generally narrower than that on elytra, widest at hind corners of base (
<figureCitation id="137E2A75FF8788189DF31CF3FEDD0BA0" box="[219,296,1492,1517]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
A; 2A); (4) male basal three protarsomeres expanded; (5) protibiae of fossorial
<typeStatus id="54FE8852FF87881899E41CF2FB0A0BA1" box="[1228,1279,1493,1517]" pageId="2" pageNumber="303">type</typeStatus>
, with distinct comb of short spines along outer margin as well as around apex; (6) mesepimera transverse, as wide at inner margin as at outer margin, anterior lateral margin straight, and posterior lateral corner pointing posteriorly; (7) mesosternal medial carina relatively low; (8) metasternal lateral carinae almost parallel to each other and to the axis of the body; (9) winged or wingless; (10) spiculum gastrale with anterior margin aligned with that of genital plates; (11) aedeagus subcylindrical, broad and short, stout in most species, sometimes more or less slender and flattened, apex variable in shape, genital orifice on ventral surface or cutting right margin of median lobe in shift to dorsal surface.
</paragraph>
<paragraph id="8BFA36F0FF87881F9DEF1FD3FB210F50" blockId="2.[151,1437,1348,2021]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="306" pageId="2" pageNumber="303">
Some other morphological characters had also found to be of taxonomic significance:
<emphasis id="B931EAE2FF87881899981FD3FAC70941" box="[1200,1330,1780,1805]" italics="true" pageId="2" pageNumber="303">Mouthparts</emphasis>
. Labrum transverse, anterior margin rounded; epipharyngeal area with torma (
<figureCitation id="137E2A75FF8788189EEA1E3FFBE7097C" box="[962,1042,1816,1841]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
B; 2B). Left and right mandibles distinctly asymmetrical, each with a tooth on inner margin of subapex; subapical orthogonal penicillus made up of slender, fine and dense setae (
<figureCitation id="137E2A75FF8788189CCA1E47FDD90934" box="[482,556,1888,1913]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
C, D; 2C, D). Maxillary palp only penultimate palpomere wholly covered with setae, length more than twice its width, and slightly swollen towards apex; ultimate palpomere quite slender, feebly arched, a little shorter than penultimate palpomere, base slightly thickened, apex narrowly rounded (
<figureCitation id="137E2A75FF87881899D71E8FFABC098C" box="[1279,1353,1960,1985]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
E; 2E). Hypopharynx and labial palps as shown in
<figureCitation id="137E2A75FF8788189F521EEBFD3609A8" box="[634,707,1996,2021]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="2" pageNumber="303">Figs. 1</figureCitation>
F; 2F, premier labial palpomere about as long as succeeding two palpomeres together.
<emphasis id="B931EAE2FF80881F9CBB19B0FDAB0EFC" box="[403,606,151,176]" italics="true" pageId="5" pageNumber="306">Cervical sclerites</emphasis>
. Cervical sclerites with end acuminate, pistol-like (
<figureCitation id="137E2A75FF80881F99E219B0FAE50EFC" box="[1226,1296,151,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="5" pageNumber="306">Fig. 1</figureCitation>
G).
<emphasis id="B931EAE2FF80881F986C19B0FA600EFC" box="[1348,1429,151,176]" italics="true" pageId="5" pageNumber="306">Thorax</emphasis>
. Prosternum with integumental foramina (where prolegs inserted) widely separated (
<figureCitation id="137E2A75FF80881F998A199BFB180E98" box="[1186,1261,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="5" pageNumber="306">
Fig.
<date id="FFFB1030FF80881F99C9199BFB180E98" box="[1249,1261,188,212]" pageId="5" pageNumber="306">1</date>
</figureCitation>
I). Meso- and metasternum as shown in
<figureCitation id="137E2A75FF80881F9CF719F8FDD30EB4" box="[479,550,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="5" pageNumber="306">Fig. 1</figureCitation>
K, mesosternal apophysis strongly developed, edge somewhat thickened. Metatergal apparatus generally broad (
<figureCitation id="137E2A75FF80881F9F6F1823FD720F50" box="[583,647,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="5" pageNumber="306">Fig. 1</figureCitation>
J). Metendosternite with wide furcal arms (
<figureCitation id="137E2A75FF80881F99451823FB590F50" box="[1133,1196,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[167,1420,205,1853]" captionTargetId="figure@3.[151,1436,195,1867]" captionTargetPageId="3" captionText="FIGURE 1. Ptomaphaginus gutianshanicus sp. nov. ♂: A, extended habitus (dorsal view); B, labrum (dorsal view); C, left mandible (dorsal view); D, right mandible (dorsal view); E, right maxilla (dorsal view); F, hypopharynx and labial palps (ventral view); G, cervical sclerite (dorsal view); H, metendosternite (dorsal view); I, prothorax (ventral view); J, metatergal apparatus (dorsal view); K, pterothorax (ventral view). Scales: A = 1 mm; B – K = 0.1 mm." httpUri="https://zenodo.org/record/243861/files/figure.png" pageId="5" pageNumber="306">Fig. 1</figureCitation>
H).
</paragraph>
</subSubSection>
</treatment>
</document>
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