481 lines
69 KiB
XML
481 lines
69 KiB
XML
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<treatment id="03EC879FFFADFFB2AB77ADC2FDDD1402" ID-DOI="http://doi.org/10.5281/zenodo.7909513" ID-GBIF-Taxon="209589373" ID-Zenodo-Dep="7909513" LSID="urn:lsid:plazi:treatment:03EC879FFFADFFB2AB77ADC2FDDD1402" httpUri="http://treatment.plazi.org/id/03EC879FFFADFFB2AB77ADC2FDDD1402" lastPageId="66" lastPageNumber="67" pageId="34" pageNumber="35">
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<taxonomicName id="4C454D0AFFADFFD2AB77ADC2FB231698" authority="Nakahara & Willmott, 2023" authorityName="Nakahara & Willmott" authorityYear="2023" box="[888,1160,1721,1741]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus" status="genus novum">
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<emphasis id="B931EA9BFFADFFD2AB77ADC2FC151698" bold="true" box="[888,958,1721,1741]" italics="true" pageId="34" pageNumber="35">Occulta</emphasis>
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Nakahara & Willmott
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,
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<emphasis id="B931EA9BFFADFFD2AC9FADC2FABF1698" bold="true" box="[1168,1300,1721,1741]" pageId="34" pageNumber="35">genus novum.</emphasis>
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species:
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<emphasis id="B931EA9BFFADFFD2ABF4AD84FB271746" box="[1019,1164,1791,1811]" italics="true" pageId="34" pageNumber="35">Euptychia ocnus</emphasis>
|
||
<bibRefCitation id="EFD44B78FFADFFD2AC9DAD84FAAC1746" author="Butler" box="[1170,1287,1791,1811]" firstAuthor="Butler" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="34" pageNumber="35" pagination="104 - 110" part="1867" refId="ref35367" refString="Butler, A. G. (1867) Descriptions of some new species of Satyridae belonging to the genus Euptychia. Proceedings of the Zoological Society of London, 1867, 104 - 110." title="Descriptions of some new species of Satyridae belonging to the genus Euptychia" type="journal article" year="1867">Butler, 1867</bibRefCitation>
|
||
</taxonomicName>
|
||
, by present designation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFADFFD2AB77AC1DFAE617C8" pageId="34" pageNumber="35" type="description">
|
||
<paragraph id="8BFA3689FFADFFD2AB77AC1DFAE617C8" blockId="34.[888,1396,1894,1949]" pageId="34" pageNumber="35">
|
||
<emphasis id="B931EA9BFFADFFD2AB77AC1DFB99172F" bold="true" box="[888,1074,1894,1914]" pageId="34" pageNumber="35">Zoobank registration</emphasis>
|
||
: https://zoobank.org/Nomenclatural Acts/7B2BBC71-C2FC-40B8-A24A-14A986151D85
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFACFFD3A8ADAF32FB231526" pageId="35" pageNumber="36" type="diagnosis">
|
||
<paragraph id="8BFA3689FFACFFD3A8ADAF32FB231526" blockId="35.[122,771,1097,1951]" lastBlockId="35.[818,1467,1097,1949]" pageId="35" pageNumber="36">
|
||
<emphasis id="B931EA9BFFACFFD3A8ADAF32FE501408" bold="true" box="[162,507,1097,1117]" pageId="35" pageNumber="36">Systematic placement and diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C454D0AFFACFFD3AA07AF32FDE61408" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[520,589,1097,1117]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AA07AF32FDE61408" box="[520,589,1097,1117]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AA42AF32FD2D1408" bold="true" box="[589,646,1097,1117]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
is a member of the ‘
|
||
<taxonomicName id="4C454D0AFFACFFD3A8CFAF17FEFA14D5" authorityName="Forster" authorityYear="1964" box="[192,337,1132,1152]" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3A8CFAF17FEFA14D5" box="[192,337,1132,1152]" italics="true" pageId="35" pageNumber="36">Splendeuptychia</emphasis>
|
||
</taxonomicName>
|
||
clade’, and it is rather distantly related to the remainder of the clade. Currently,
|
||
<taxonomicName id="4C454D0AFFACFFD3A9DAAFF4FDB114F6" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[469,538,1167,1187]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A9DAAFF4FDB114F6" box="[469,538,1167,1187]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AA15AFF4FDF814F6" bold="true" box="[538,595,1167,1187]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
is modestly supported (SH-aLRT 100,
|
||
<collectionCode id="ED54AE4CFFACFFD3A958AFCAFE2B1490" box="[343,384,1201,1221]" pageId="35" pageNumber="36">UFB</collectionCode>
|
||
75) as sister to a large clade including species-rich genera such as
|
||
<taxonomicName id="4C454D0AFFACFFD3A996AFAFFD8314BD" authorityName="Forster" authorityYear="1964" box="[409,552,1236,1256]" class="Insecta" family="Nymphalidae" genus="Caeruleuptychia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3A996AFAFFD8314BD" box="[409,552,1236,1256]" italics="true" pageId="35" pageNumber="36">Caeruleuptychia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D0AFFACFFD3AA35AFAFFD6614BD" authorityName="Forster" authorityYear="1964" box="[570,717,1236,1256]" class="Insecta" family="Nymphalidae" genus="Paryphthimoides" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3AA35AFAFFD6614BD" box="[570,717,1236,1256]" italics="true" pageId="35" pageNumber="36">Paryphthimoides</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C454D0AFFACFFD3A875AF8CFEA0155E" authorityName="Forster" authorityYear="1964" box="[122,267,1271,1291]" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3A875AF8CFEA0155E" box="[122,267,1271,1291]" italics="true" pageId="35" pageNumber="36">Splendeuptychia</emphasis>
|
||
</taxonomicName>
|
||
, among others (
|
||
<figureCitation id="137E2A0CFFACFFD3A9A6AF8CFDAE155E" box="[425,517,1271,1291]" captionStart="FIGURE 10" captionStartId="16.[818,915,1826,1845]" captionTargetBox="[809,1455,113,1785]" captionTargetId="graphics-563@16.[917,1163,140,1782]" captionTargetPageId="16" captionText="FIGURE 10 Phylogeny of the ’Splendeuptychia clade’ based on the FULL dataset. Interpretation of support values can be found in the legend. The scale bars under the butterfly images equal 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909417" httpUri="https://zenodo.org/record/7909417/files/figure.png" pageId="35" pageNumber="36">Figure 10</figureCitation>
|
||
). Based on hybrid enrichment data,
|
||
<bibRefCitation id="EFD44B78FFACFFD3A8EAAE62FE171578" author="Espeland" box="[229,444,1305,1325]" etAl="et al." firstAuthor="Espeland" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="35" pageNumber="36" pagination="116 - 124" part="131" refId="ref36456" refString="Espeland, M., Breinholt, J., Barbosa, E. P., Casagrande, M., Huertas, B., Lamas, G. et al. (2019 a) Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data. Molecular Phylogenetics and Evolution, 131, 116 - 124." title="Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data" type="journal article" year="2019" yearSuffix="a">Espeland et al. (2019a)</bibRefCitation>
|
||
recovered
|
||
<taxonomicName id="4C454D0AFFACFFD3AA28AE62FDC71578" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[551,620,1305,1325]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AA28AE62FDC71578" box="[551,620,1305,1325]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AA63AE62FD0E1578" bold="true" box="[620,677,1305,1325]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
(as ‘
|
||
<taxonomicName id="4C454D0AFFACFFD3AADCAE62FEBD1505" authorityName="Butler" authorityYear="1866" class="Insecta" family="Nymphalidae" genus="Magneuptychia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ocnus">
|
||
<emphasis id="B931EA9BFFACFFD3AADCAE62FEBD1505" italics="true" pageId="35" pageNumber="36">Magneuptychia ocnus</emphasis>
|
||
</taxonomicName>
|
||
’) as sister to ‘
|
||
<taxonomicName id="4C454D0AFFACFFD3A9AFAE47FDAC1505" authorityName="Forster" authorityYear="1964" box="[416,519,1340,1360]" class="Insecta" family="Nymphalidae" genus="Erichthodes" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3A9AFAE47FDAC1505" box="[416,519,1340,1360]" italics="true" pageId="35" pageNumber="36">Erichthodes</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C454D0AFFACFFD3AA1AAE47FDF81505" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Schaus" baseAuthorityYear="1902" box="[533,595,1340,1360]" class="Insecta" family="Nymphalidae" genus="Capronnieria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="narapa">
|
||
<emphasis id="B931EA9BFFACFFD3AA1AAE47FDF81505" box="[533,595,1340,1360]" italics="true" pageId="35" pageNumber="36">narapa</emphasis>
|
||
</taxonomicName>
|
||
(now
|
||
<taxonomicName id="4C454D0AFFACFFD3AA9FAE47FF131526" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Schaus" baseAuthorityYear="1902" class="Insecta" family="Nymphalidae" genus="Capronnieria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="narapa" status="comb.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AA9FAE47FF131526" italics="true" pageId="35" pageNumber="36">Capronnieria narapa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A8B7AE25FEA91527" bold="true" box="[184,258,1374,1394]" pageId="35" pageNumber="36">comb.n.</emphasis>
|
||
) and
|
||
<taxonomicName id="4C454D0AFFACFFD3A93AAE24FE401526" baseAuthorityName="Godart" baseAuthorityYear="1824" box="[309,491,1375,1395]" class="Insecta" family="Nymphalidae" genus="Capronnieria" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="galesus">
|
||
<emphasis id="B931EA9BFFACFFD3A93AAE24FE401526" box="[309,491,1375,1395]" italics="true" pageId="35" pageNumber="36">Capronnieria galesus</emphasis>
|
||
</taxonomicName>
|
||
, although with only moderate support (
|
||
<collectionCode id="ED54AE4CFFACFFD3A8C1AEFAFF4315C0" box="[206,232,1409,1429]" pageId="35" pageNumber="36">BS</collectionCode>
|
||
and
|
||
<collectionCode id="ED54AE4CFFACFFD3A914AEFAFE9F15C0" box="[283,308,1409,1429]" pageId="35" pageNumber="36">PP</collectionCode>
|
||
>0.75 <0.95). This relationship, with similar low support, was also found in the backbone tree in this study (
|
||
<figureCitation id="137E2A0CFFACFFD3AA91AEDFFD5C15ED" box="[670,759,1444,1464]" captionStart="FIGURE 1" captionStartId="5.[122,218,1354,1373]" captionTargetBox="[155,1432,126,1307]" captionTargetId="graphics-326@5.[251,1339,165,1253]" captionText="FIGURE 1 Summary tree based on the FULL dataset showing the major clades supported in this study. Colours are the same as in Espeland et al. (2019a). Support for major clades is shown. Interpretation of support values can be found in the legend." figureDoi="http://doi.org/10.5281/zenodo.7968436" httpUri="https://zenodo.org/record/7968436/files/figure.png" pageId="35" pageNumber="36">
|
||
Figure
|
||
<collectionCode id="ED54AE4CFFACFFD3AAD0AEDFFD5B15ED" box="[735,752,1444,1464]" pageId="35" pageNumber="36">S</collectionCode>
|
||
1
|
||
</figureCitation>
|
||
). The single species of
|
||
<taxonomicName id="4C454D0AFFACFFD3A947AEBCFE26158E" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[328,397,1479,1499]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A947AEBCFE26158E" box="[328,397,1479,1499]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A982AEBDFE6D158F" bold="true" box="[397,454,1478,1498]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
(
|
||
<figureCitation id="137E2A0CFFACFFD3A9DDAEBCFD86158E" box="[466,557,1479,1499]" captionStart="FIGURE 30" captionStartId="35.[122,218,975,994]" captionTargetBox="[163,1432,129,924]" captionTargetId="figure-591@35.[155,1433,123,934]" captionTargetPageId="35" captionText="FIGURE 30 Occulta ocnus comb.n. (a) male wing venation; (b and c) male (LEP-10408) dorsal (b) and ventral (c); (d and e) female (ST ocnus) dorsal (d) and ventral (e), with specimen labels. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909477" httpUri="https://zenodo.org/record/7909477/files/figure.png" pageId="35" pageNumber="36">Figure 30</figureCitation>
|
||
) resembles some species of
|
||
<taxonomicName id="4C454D0AFFACFFD3A8B4AE92FEAB15A8" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[187,256,1513,1533]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A8B4AE92FEAB15A8" box="[187,256,1513,1533]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A90FAE92FE9215A8" bold="true" box="[256,313,1513,1533]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
(see
|
||
<figureCitation id="137E2A0CFFACFFD3A968AE92FE6315A8" box="[359,456,1513,1533]" captionStart="FIGURE 33" captionStartId="38.[122,218,917,936]" captionTargetBox="[158,1432,132,876]" captionTargetId="figure-612@38.[154,1434,123,877]" captionTargetPageId="38" captionText="FIGURE 33 Deltaya ocypete comb.n. (a) male wing venation; (b and c) male (FLMNH-MGCL-209681) dorsal (b) and ventral (c); (d and e) female dorsal (d) and ventral (e). Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909485" httpUri="https://zenodo.org/record/7909485/files/figure.png" pageId="35" pageNumber="36">Figures 33</figureCitation>
|
||
and
|
||
<figureCitation id="137E2A0CFFACFFD3A9FAAE92FDA615A8" box="[501,525,1513,1533]" captionStart="FIGURE 34" captionStartId="39.[122,218,1083,1102]" captionTargetBox="[155,1424,136,1042]" captionTargetId="figure-515@39.[153,1436,123,1043]" captionTargetPageId="39" captionText="FIGURE 34 Deltaya gen.n. representative species. (a) D. opima comb.n; (b) D. louisammour comb.n (FLMNH-MGCL-111522); (c) D. andrei comb.n; (d) D. probata comb.n. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909489" httpUri="https://zenodo.org/record/7909489/files/figure.png" pageId="35" pageNumber="36">34</figureCitation>
|
||
below), but may be distinguished by possessing elongate, ‘smudge-like’ orangish spots with a silver marking in the middle in VHW cells M
|
||
<subScript id="17C134CCFFACFFD3AA29AD4CFD841611" attach="left" box="[550,559,1591,1604]" fontSize="5" pageId="35" pageNumber="36">2</subScript>
|
||
and M
|
||
<subScript id="17C134CCFFACFFD3AA79AD4CFDD41611" attach="left" box="[630,639,1591,1604]" fontSize="5" pageId="35" pageNumber="36">3</subScript>
|
||
, whereas the ocelli in these cells in
|
||
<taxonomicName id="4C454D0AFFACFFD3A947AD2AFE261630" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[328,397,1617,1637]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A947AD2AFE261630" box="[328,397,1617,1637]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A982AD2AFE6E1630" bold="true" box="[397,453,1617,1637]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
are more rounded and have double silver spots encircled by a better defined yellowish ring. Furthermore, the dark ventral discal and postdiscal bands are typically narrower in
|
||
<taxonomicName id="4C454D0AFFACFFD3A8E5ADC2FE841698" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[234,303,1721,1741]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A8E5ADC2FE841698" box="[234,303,1721,1741]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A920ADC2FEC31698" bold="true" box="[303,360,1721,1741]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
compared to
|
||
<taxonomicName id="4C454D0AFFACFFD3A9E7ADC2FD861698" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[488,557,1721,1741]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A9E7ADC2FD861698" box="[488,557,1721,1741]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AA23ADC2FDCE1698" bold="true" box="[556,613,1721,1741]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
species, perhaps with the exception of
|
||
<taxonomicName id="4C454D0AFFACFFD3A944ADA7FE0016A5" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" baseAuthorityYear="2023" box="[331,427,1756,1776]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ocypete">
|
||
<emphasis id="B931EA9BFFACFFD3A944ADA7FEF116A5" box="[331,346,1756,1776]" italics="true" pageId="35" pageNumber="36">D</emphasis>
|
||
.
|
||
<emphasis id="B931EA9BFFACFFD3A969ADA7FE0016A5" box="[358,427,1756,1776]" italics="true" pageId="35" pageNumber="36">ocypete</emphasis>
|
||
</taxonomicName>
|
||
, whose ventral bands are often narrower in comparison with other closely related species. In particular, in
|
||
<taxonomicName id="4C454D0AFFACFFD3A89EAC5AFF7D1760" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[145,214,1825,1845]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A89EAC5AFF7D1760" box="[145,214,1825,1845]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A8DAAC5AFEA51760" bold="true" box="[213,270,1825,1845]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
the VHW marginal band is of even width throughout, whereas this band forms a ‘wedge-shaped’ swelling at the tornus in
|
||
<taxonomicName id="4C454D0AFFACFFD3A875AC1CFF14172E" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[122,191,1895,1915]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A875AC1CFF14172E" box="[122,191,1895,1915]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A8B0AC1DFF5C172F" bold="true" box="[191,247,1894,1914]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
In addition,
|
||
<taxonomicName id="4C454D0AFFACFFD3A98BAC1CFE62172E" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[388,457,1895,1915]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3A98BAC1CFE62172E" box="[388,457,1895,1915]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3A9C6AC1DFDA9172F" bold="true" box="[457,514,1894,1914]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
possesses a tiny, rather incomplete ocellus in VHW cell Cu
|
||
<subScript id="17C134CCFFACFFD3A9CFACE9FE6217CA" attach="left" box="[448,457,1938,1951]" fontSize="5" pageId="35" pageNumber="36">2</subScript>
|
||
, which is absent in
|
||
<taxonomicName id="4C454D0AFFACFFD3AA8BACF2FD6217C8" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[644,713,1929,1949]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AA8BACF2FD6217C8" box="[644,713,1929,1949]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AAC6ACF2FCA917C8" bold="true" box="[713,770,1929,1949]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
The male genitalia also differ between
|
||
<taxonomicName id="4C454D0AFFACFFD3ACAAAF32FB411408" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[1189,1258,1097,1117]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3ACAAAF32FB411408" box="[1189,1258,1097,1117]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3ACE5AF32FA881408" bold="true" box="[1258,1315,1097,1117]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
(
|
||
<figureCitation id="137E2A0CFFACFFD3AD20AF32FA211408" box="[1327,1418,1097,1117]" captionStart="FIGURE 31" captionStartId="36.[122,218,1273,1292]" captionTargetBox="[155,1433,134,1231]" captionTargetId="figure-301@36.[154,1434,123,1233]" captionTargetPageId="36" captionText="FIGURE 31 Occulta ocnus comb.n. (a and b) male genitalia (dissection SN-19-157), lateral (a) with posterior view juxta, aedeagus lateral (b); (c–e) female genitalia (dissection NHMUK010402850), lateral view exterior tip abdomen (c), ventral view exterior tip abdomen (d), dorsal view interior abdomen (e), corpus bursae perpendicular to signa (f). Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7909481" httpUri="https://zenodo.org/record/7909481/files/figure.png" pageId="35" pageNumber="36">Figure 31</figureCitation>
|
||
) and the
|
||
<typeStatus id="54FE882BFFACFFD3AB56AF17FC2914D5" box="[857,898,1132,1152]" pageId="35" pageNumber="36">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C454D0AFFACFFD3ABFCAF17FB9314D5" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[1011,1080,1132,1152]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3ABFCAF17FB9314D5" box="[1011,1080,1132,1152]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AC37AF17FBDA14D5" bold="true" box="[1080,1137,1132,1152]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
and close relatives (referred to as ‘core’
|
||
<taxonomicName id="4C454D0AFFACFFD3AB7FAFF4FC1E14F6" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[880,949,1167,1187]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AB7FAFF4FC1E14F6" box="[880,949,1167,1187]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3ABBAAFF4FC4514F6" bold="true" box="[949,1006,1167,1187]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
) (
|
||
<figureCitation id="137E2A0CFFACFFD3ABF0AFF4FBF114F6" box="[1023,1114,1167,1187]" captionStart="FIGURE 34" captionStartId="39.[122,218,1083,1102]" captionTargetBox="[155,1424,136,1042]" captionTargetId="figure-515@39.[153,1436,123,1043]" captionTargetPageId="39" captionText="FIGURE 34 Deltaya gen.n. representative species. (a) D. opima comb.n; (b) D. louisammour comb.n (FLMNH-MGCL-111522); (c) D. andrei comb.n; (d) D. probata comb.n. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909489" httpUri="https://zenodo.org/record/7909489/files/figure.png" pageId="35" pageNumber="36">Figure 34</figureCitation>
|
||
), at least, by the lack of a developed ‘hump’ on the dorsal margin of the valva, and instead only have a slightly serrated region at the dorsal margin distal of the costa. In lateral view, the costa appears as a narrow plate in
|
||
<taxonomicName id="4C454D0AFFACFFD3AD37AF8CFAD6155E" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[1336,1405,1271,1291]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AD37AF8CFAD6155E" box="[1336,1405,1271,1291]" italics="true" pageId="35" pageNumber="36">Occulta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AD72AF8DFA1D155F" bold="true" box="[1405,1462,1270,1290]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
, whereas the costa appears as a somewhat trapezoidal plate in lateral view in ‘core’
|
||
<taxonomicName id="4C454D0AFFACFFD3ABCFAE47FBAE1505" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[960,1029,1340,1360]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3ABCFAE47FBAE1505" box="[960,1029,1340,1360]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AC0BAE47FB961505" bold="true" box="[1028,1085,1340,1360]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
See description of
|
||
<taxonomicName id="4C454D0AFFACFFD3ACF3AE47FAEA1505" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[1276,1345,1340,1360]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3ACF3AE47FAEA1505" box="[1276,1345,1340,1360]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AD4FAE47FAD21505" bold="true" box="[1344,1401,1340,1360]" pageId="35" pageNumber="36">gen.n.</emphasis>
|
||
below for details about ‘core’
|
||
<taxonomicName id="4C454D0AFFACFFD3AC04AE24FBFB1526" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[1035,1104,1375,1395]" class="Insecta" family="Nymphalidae" genus="Deltaya" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFACFFD3AC04AE24FBFB1526" box="[1035,1104,1375,1395]" italics="true" pageId="35" pageNumber="36">Deltaya</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EA9BFFACFFD3AC5FAE25FB281527" bold="true" box="[1104,1155,1374,1394]" pageId="35" pageNumber="36">gen.n</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFACFFD3AB55AEFAFBEF1675" pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph id="8BFA3689FFACFFD3AB55AEFAFBEF1675" blockId="35.[818,1467,1097,1949]" pageId="35" pageNumber="36">
|
||
<emphasis id="B931EA9BFFACFFD3AB55AEFAFC1515C0" bold="true" box="[858,958,1409,1429]" pageId="35" pageNumber="36">Etymology</emphasis>
|
||
. The generic name is a Latin feminine adjective treated as a noun in the nominative singular, ‘occulta’, meaning a ‘hidden’ or ‘secret’ thing, in reference to the former concealment of this taxon within
|
||
<taxonomicName id="4C454D0AFFACFFD3AB7BAE92FC5715A8" baseAuthorityName="Lamas" baseAuthorityYear="2004" box="[884,1020,1513,1533]" class="Insecta" family="Nymphalidae" genus="Magneuptychia" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFACFFD3AB7BAE92FC5715A8" box="[884,1020,1513,1533]" italics="true" pageId="35" pageNumber="36">Magneuptychia</emphasis>
|
||
</taxonomicName>
|
||
. The generic name is also coined in alliteration with the species-group name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFACFFD4AB55AD55FD1B1698" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="8BFA3689FFACFFD4AB55AD55FD1B1698" blockId="35.[818,1467,1097,1949]" lastBlockId="36.[122,771,1444,1741]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
<emphasis id="B931EA9BFFACFFD3AB55AD55FC6C1617" bold="true" box="[858,967,1582,1602]" pageId="35" pageNumber="36">Description</emphasis>
|
||
(
|
||
<figureCitation id="137E2A0CFFACFFD3ABDCAD54FB9E1616" box="[979,1077,1583,1603]" captionStart="FIGURE 30" captionStartId="35.[122,218,975,994]" captionTargetBox="[163,1432,129,924]" captionTargetId="figure-591@35.[155,1433,123,934]" captionTargetPageId="35" captionText="FIGURE 30 Occulta ocnus comb.n. (a) male wing venation; (b and c) male (LEP-10408) dorsal (b) and ventral (c); (d and e) female (ST ocnus) dorsal (d) and ventral (e), with specimen labels. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909477" httpUri="https://zenodo.org/record/7909477/files/figure.png" pageId="35" pageNumber="36">Figures 30</figureCitation>
|
||
and
|
||
<figureCitation id="137E2A0CFFACFFD3AC6BAD54FBD71616" box="[1124,1148,1583,1603]" captionStart="FIGURE 31" captionStartId="36.[122,218,1273,1292]" captionTargetBox="[155,1433,134,1231]" captionTargetId="figure-301@36.[154,1434,123,1233]" captionTargetPageId="36" captionText="FIGURE 31 Occulta ocnus comb.n. (a and b) male genitalia (dissection SN-19-157), lateral (a) with posterior view juxta, aedeagus lateral (b); (c–e) female genitalia (dissection NHMUK010402850), lateral view exterior tip abdomen (c), ventral view exterior tip abdomen (d), dorsal view interior abdomen (e), corpus bursae perpendicular to signa (f). Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7909481" httpUri="https://zenodo.org/record/7909481/files/figure.png" pageId="35" pageNumber="36">31</figureCitation>
|
||
). Wing pattern and shape as illustrated (
|
||
<figureCitation id="137E2A0CFFACFFD3AB7AAD2AFC651630" box="[885,974,1617,1637]" captionStart="FIGURE 30" captionStartId="35.[122,218,975,994]" captionTargetBox="[163,1432,129,924]" captionTargetId="figure-591@35.[155,1433,123,934]" captionTargetPageId="35" captionText="FIGURE 30 Occulta ocnus comb.n. (a) male wing venation; (b and c) male (LEP-10408) dorsal (b) and ventral (c); (d and e) female (ST ocnus) dorsal (d) and ventral (e), with specimen labels. Scale bars 1 cm." figureDoi="http://doi.org/10.5281/zenodo.7909477" httpUri="https://zenodo.org/record/7909477/files/figure.png" pageId="35" pageNumber="36">Figure 30</figureCitation>
|
||
). Other notable characters include eyes setose; pterothoracic legs dorsally darker, tibia with two principal longitudinal rows of spines ventrally, in addition to some spines laterally, pair of spurs of similar length at distal end of tibia, first tarsomere with three principal longitudinal rows of spines ventrally, remaining distal tarsomeres with four principal longitudinal rows of spines ventrally. Medium-sized
|
||
<taxonomicName id="4C454D0AFFACFFD3ABB4AC5AFB8C1760" box="[955,1063,1825,1845]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="53" phylum="Arthropoda" rank="subTribe" subTribe="Euptychiina">Euptychiina</taxonomicName>
|
||
(
|
||
<collectionCode id="ED54AE4CFFACFFD3AC3DAC5AFBFF1760" box="[1074,1108,1825,1845]" pageId="35" pageNumber="36">FW</collectionCode>
|
||
length typically
|
||
<quantity id="4CBD9B6CFFACFFD3ACE4AC5AFAF81760" box="[1259,1363,1825,1845]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="35" pageNumber="36" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">
|
||
23
|
||
<emphasis id="B931EA9BFFACFFD3AD0CAC59FAA41760" box="[1283,1295,1826,1845]" italics="true" pageId="35" pageNumber="36">–</emphasis>
|
||
26 mm
|
||
</quantity>
|
||
), DFW and DHW of males without obvious androconial scales, but long setiform scales visible in discal cell and adjacent areas, especially extending along inner margin. Male with 8th tergite a narrow stripe at basal side of the eighth abdominal segment, broad weakly sclerotized patch present at posterior side of eighth abdominal segment; genitalia as illustrated (
|
||
<figureCitation id="137E2A0CFFABFFD4A8B0AE92FEB215A8" box="[191,281,1513,1533]" captionStart="FIGURE 31" captionStartId="36.[122,218,1273,1292]" captionTargetBox="[155,1433,134,1231]" captionTargetId="figure-301@36.[154,1434,123,1233]" captionTargetPageId="36" captionText="FIGURE 31 Occulta ocnus comb.n. (a and b) male genitalia (dissection SN-19-157), lateral (a) with posterior view juxta, aedeagus lateral (b); (c–e) female genitalia (dissection NHMUK010402850), lateral view exterior tip abdomen (c), ventral view exterior tip abdomen (d), dorsal view interior abdomen (e), corpus bursae perpendicular to signa (f). Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7909481" httpUri="https://zenodo.org/record/7909481/files/figure.png" pageId="36" pageNumber="37">Figure 31</figureCitation>
|
||
); costa appearing as a narrow plate in lateral view; cornuti present, otherwise genitalia as illustrated (
|
||
<figureCitation id="137E2A0CFFABFFD4AA5EAD77FD071675" box="[593,684,1548,1568]" captionStart="FIGURE 31" captionStartId="36.[122,218,1273,1292]" captionTargetBox="[155,1433,134,1231]" captionTargetId="figure-301@36.[154,1434,123,1233]" captionTargetPageId="36" captionText="FIGURE 31 Occulta ocnus comb.n. (a and b) male genitalia (dissection SN-19-157), lateral (a) with posterior view juxta, aedeagus lateral (b); (c–e) female genitalia (dissection NHMUK010402850), lateral view exterior tip abdomen (c), ventral view exterior tip abdomen (d), dorsal view interior abdomen (e), corpus bursae perpendicular to signa (f). Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7909481" httpUri="https://zenodo.org/record/7909481/files/figure.png" pageId="36" pageNumber="37">Figure 31</figureCitation>
|
||
). Female genitalia (
|
||
<figureCitation id="137E2A0CFFABFFD4A8DCAD54FE851616" box="[211,302,1583,1603]" captionStart="FIGURE 31" captionStartId="36.[122,218,1273,1292]" captionTargetBox="[155,1433,134,1231]" captionTargetId="figure-301@36.[154,1434,123,1233]" captionTargetPageId="36" captionText="FIGURE 31 Occulta ocnus comb.n. (a and b) male genitalia (dissection SN-19-157), lateral (a) with posterior view juxta, aedeagus lateral (b); (c–e) female genitalia (dissection NHMUK010402850), lateral view exterior tip abdomen (c), ventral view exterior tip abdomen (d), dorsal view interior abdomen (e), corpus bursae perpendicular to signa (f). Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7909481" httpUri="https://zenodo.org/record/7909481/files/figure.png" pageId="36" pageNumber="37">Figure 31</figureCitation>
|
||
) with intersegmental membrane of between seventh and eighth abdominal segment somewhat pleated and expandable with very weak sclerotization ventrally; lamella antevaginalis membranous; eighth segment with irregular lateral sclerotized plate narrowing slightly dorsally; pair of well-defined signa present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFCDFFB2A875AF63FE30147D" box="[122,411,1048,1064]" pageId="66" pageNumber="67" type="taxon_list">
|
||
<paragraph id="8BFA3689FFCDFFB2A875AF63FE30147D" blockId="66.[122,1218,205,1687]" box="[122,411,1048,1064]" pageId="66" pageNumber="67">
|
||
<taxonomicName id="4C454D0AFFCDFFB2A875AF63FEF4147D" authority="Nakahara & Willmott" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[122,351,1048,1064]" class="Insecta" family="Nymphalidae" genus="Occulta" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen.n.">
|
||
<emphasis id="B931EA9BFFCDFFB2A875AF63FF1D147D" box="[122,182,1048,1064]" italics="true" pageId="66" pageNumber="67">Occulta</emphasis>
|
||
Nakahara & Willmott
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B931EA9BFFCDFFB2A969AF63FE30147D" bold="true" box="[358,411,1048,1064]" pageId="66" pageNumber="67">gen.n.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6502FFCDFFB2A8ADAF4BFDDD1402" pageId="66" pageNumber="67" type="taxon_list">
|
||
<paragraph id="8BFA3689FFCDFFB2A8ADAF4BFD9F1415" blockId="66.[122,1218,205,1687]" box="[162,564,1072,1088]" pageId="66" pageNumber="67">
|
||
<taxonomicName id="4C454D0AFFCDFFB2A8ADAF4BFEE81415" authority="(Butler, 1867)" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Butler" baseAuthorityYear="1867" box="[162,323,1072,1088]" class="Insecta" family="Nymphalidae" genus="Euptychia" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="ocnus" status="comb.n.">
|
||
<emphasis id="B931EA9BFFCDFFB2A8ADAF4BFF7A1415" box="[162,209,1072,1088]" italics="true" pageId="66" pageNumber="67">ocnus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FFCDFFB2A8D5AF4BFE951415" author="Butler, A. G." box="[218,318,1072,1088]" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="66" pageNumber="67" pagination="104 - 110" part="1867" refId="ref35367" refString="Butler, A. G. (1867) Descriptions of some new species of Satyridae belonging to the genus Euptychia. Proceedings of the Zoological Society of London, 1867, 104 - 110." title="Descriptions of some new species of Satyridae belonging to the genus Euptychia" type="journal article" year="1867">Butler, 1867</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B931EA9BFFCDFFB2A945AF4BFE241415" bold="true" box="[330,399,1072,1088]" pageId="66" pageNumber="67">comb.n.</emphasis>
|
||
, was
|
||
<taxonomicName id="4C454D0AFFCDFFB2A9B3AF4BFD9F1415" authorityName="Forster" authorityYear="1964" box="[444,564,1072,1088]" class="Insecta" family="Nymphalidae" genus="Magneuptychia" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B931EA9BFFCDFFB2A9B3AF4BFD9F1415" box="[444,564,1072,1088]" italics="true" pageId="66" pageNumber="67">Magneuptychia</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BFA3689FFCDFFB2A8B0AF3CFDDD1402" blockId="66.[122,1218,205,1687]" box="[191,630,1095,1111]" pageId="66" pageNumber="67">
|
||
=
|
||
<taxonomicName id="4C454D0AFFCDFFB2A8C6AF3CFE2F1402" authority="(Weymer, 1911)" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" baseAuthorityName="Weymer" baseAuthorityYear="1911" box="[201,388,1095,1111]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="66" pageNumber="67" phylum="Tracheophyta" rank="species" species="gracilis" status="comb.n.">
|
||
<emphasis id="B931EA9BFFCDFFB2A8C6AF3CFEAB1402" box="[201,256,1095,1111]" italics="true" pageId="66" pageNumber="67">gracilis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B78FFCDFFB2A905AF3CFE2B1402" author="Weymer, G." box="[266,384,1095,1111]" editor="Seitz, A." journalOrPublisher="Stuttgart: A. Kernen" pageId="66" pageNumber="67" pagination="173 - 280" refId="ref43751" refString="Weymer, G. (1911) 4. Familie: Satyridae. In: Seitz, A. (Ed.) Die GrossSchmetterlinge der Erde. Stuttgart: A. Kernen, pp. 173 - 280." title="Familie: Satyridae" type="book chapter" volumeTitle="Die Gross- Schmetterlinge der Erde" year="1911">Weymer, 1911</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B931EA9BFFCDFFB2A984AF3CFE7B1402" bold="true" box="[395,464,1095,1111]" pageId="66" pageNumber="67">comb.n.</emphasis>
|
||
, was
|
||
<taxonomicName id="4C454D0AFFCDFFB2A9F1AF3CFDDD1402" authority="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott, 2023" authorityName="Espeland1 & Nakahara & Zacca & Barbosa & Huertas & Marín & Lamas & Benmesbah & Brévignon & Casagrande & Fåhraeus & Grishin & Kawahara1 & Mielke & Miller & Nakamura & Navas & Patrusky & Pyrcz & Richards & Tan & Tyler & Viloria & Warren1 & Xiao & Freitas & Willmott" authorityYear="2023" box="[510,630,1095,1111]" class="Insecta" family="Nymphalidae" genus="Magneuptychia" kingdom="Animalia" order="Lepidoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis id="B931EA9BFFCDFFB2A9F1AF3CFDDD1402" box="[510,630,1095,1111]" italics="true" pageId="66" pageNumber="67">Magneuptychia</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |