treatments-xml/data/03/EA/99/03EA9979CA4CFFEF62E3B40EFD83FECE.xml

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<mods:title id="6132BC9384E23EE1B862B65EB71B1D5B">On the identity of Cancer urania Herbst, 1801 (Crustacea: Decapoda: Brachyura: Leucosiidae)</mods:title>
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<heading id="D0B49F03CA4CFFE562E3B40EFDAAF94D" bold="true" box="[151,531,1817,1843]" fontSize="11" level="1" pageId="2" pageNumber="126" reason="1">
<taxonomicName id="4C4353ECCA4CFFE562E3B40EFDAAF94D" ID-CoL="X2CB" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[151,531,1817,1843]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4CFFE562E3B40EFDAAF94D" bold="true" box="[151,531,1817,1843]" pageId="2" pageNumber="126">
<emphasis id="B937F47DCA4CFFE562E3B40EFEE0F94D" bold="true" box="[151,345,1817,1843]" italics="true" pageId="2" pageNumber="126">Coleusia urania</emphasis>
(Herbst, 1801)
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<paragraph id="8BFC286FCA4CFFE562E3B42BFEF3F92B" blockId="2.[151,531,1817,1877]" box="[151,330,1852,1877]" pageId="2" pageNumber="126">
(
<figureCitation id="137834EACA4CFFE562EBB42BFF55F92A" box="[159,236,1852,1877]" captionStart="FIGURE 1" captionStartId="4.[151,250,1950,1972]" captionTargetBox="[162,1424,221,1928]" captionTargetId="figure@4.[162,1426,195,1928]" captionTargetPageId="4" captionText="FIGURE 1. Coleusia urania (Herbst, 1801). A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T. - Y. Chan)." httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="2" pageNumber="126">Figs. 1</figureCitation>
, 
<figureCitation id="137834EACA4CFFE5628EB42BFEB3F92A" box="[250,266,1852,1876]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="2" pageNumber="126">2</figureCitation>
, 
<figureCitation id="137834EACA4CFFE56362B42BFEF8F92B" box="[278,321,1852,1877]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="7.[151,250,1609,1631]" captionStartId-1="8.[151,250,1592,1614]" captionStartId-2="9.[151,250,1233,1255]" captionTargetBox-0="[159,1418,176,1584]" captionTargetBox-1="[204,1383,231,1547]" captionTargetBox-2="[160,1429,202,1204]" captionTargetId-0="figure@7.[159,1419,175,1585]" captionTargetId-1="figure@8.[204,1384,231,1547]" captionTargetId-2="figure@9.[151,1436,195,1211]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." captionText-1="FIGURE 5. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), intact left G 1. A, dorsal view; B, ventral view." captionText-2="FIGURE 6. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), gonopods. A, left G 1; B, C, distal half of left G 1; D, left G 2; E, distal part of left G 2. Setae of all structures denuded. Scales: A, D = 3.0 mm; B, C, E = 1.0 mm." httpUri-0="https://zenodo.org/record/230320/files/figure.png" httpUri-1="https://zenodo.org/record/230321/files/figure.png" httpUri-2="https://zenodo.org/record/230322/files/figure.png" pageId="2" pageNumber="126">4–6</figureCitation>
)
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<taxonomicName id="4C4353ECCA4CFFE562E3B469FE65F9ED" authority="Herbst, 1801: 17" authorityName="Herbst" authorityPageNumber="17" authorityYear="1801" box="[151,476,1917,1939]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4CFFE562E3B469FE92F9ED" box="[151,299,1918,1939]" italics="true" pageId="2" pageNumber="126">Cancer urania</emphasis>
Herbst, 1801: 17
</taxonomicName>
, pl. 53, fig. 3.
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<treatmentCitationGroup id="AB530F41CA4CFFE562E3B48AFC78F991" pageId="2" pageNumber="126">
<taxonomicName id="4C4353ECCA4CFFE562E3B48AFE84F9CC" ID-CoL="84FXW" box="[151,317,1949,1970]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4CFFE562E3B48AFE84F9CC" box="[151,317,1949,1970]" italics="true" pageId="2" pageNumber="126">Leucosia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4CFFE56322B48BFD86F9CC" author="Lichtenstein" box="[342,575,1948,1970]" page="140" pageId="2" pageNumber="126" year="1816">
<bibRefCitation id="EFD2559ECA4CFFE56322B48BFDA9F9CC" author="Lichtenstein" box="[342,528,1948,1970]" pageId="2" pageNumber="126" refString="Lichtenstein, M. H. C. (1816) Die Gattung Leucosia: als Probe einer neuer Bearbeitung der Krabben und Krebse. Magazin der Gesellschaft Naturforschender Freunde zu Berlin, 7, 135 - 144." type="journal article" year="1816">
<collectingCountry id="F35468FFCA4CFFE56322B48BFE6AF9CC" box="[342,467,1948,1970]" name="Liechtenstein" pageId="2" pageNumber="126">Lichtenstein</collectingCountry>
1816
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: 140
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.— 
<treatmentCitation id="0AE20E7ECA4CFFE56029B48BFD45F9CC" author="Leach" box="[605,764,1948,1970]" page="21" pageId="2" pageNumber="126" year="1817">
<bibRefCitation id="EFD2559ECA4CFFE56029B48BFD62F9CC" author="Leach" box="[605,731,1948,1970]" pageId="2" pageNumber="126" refString="Leach, W. E. (1817) The Zoological Miscellany, being descriptions of new or interesting animals. Printed by B. McMillan for E. Nodder &amp; Son, 3, i - vi, 1 - 151." type="journal article" year="1817">Leach 1817</bibRefCitation>
: 21
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.— 
<treatmentCitation id="0AE20E7ECA4CFFE5616EB48BFC4DF9CC" author="Desmarest" box="[794,1012,1948,1970]" page="167" pageId="2" pageNumber="126" year="1825">
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: 167
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.— 
<treatmentCitation id="0AE20E7ECA4CFFE56667B48BFB0DF9CC" author="White" box="[1043,1204,1948,1970]" page="48" pageId="2" pageNumber="126" year="1847">
<bibRefCitation id="EFD2559ECA4CFFE56667B48BFB2BF9CC" author="White" box="[1043,1170,1948,1970]" pageId="2" pageNumber="126" refString="White, A. (1847) List of species in the collections of the British Museum. British Museum, London, viii + 1 - 143 pp." type="book chapter" year="1847">White 1847</bibRefCitation>
: 48
</treatmentCitation>
(part).— 
<treatmentCitation id="0AE20E7ECA4CFFE56765B48BFF55F9AE" author="Gibbes" page="185" pageId="2" pageNumber="126" year="1850">
<bibRefCitation id="EFD2559ECA4CFFE56765B48BFA21F9CC" author="Gibbes" box="[1297,1432,1948,1970]" pageId="2" pageNumber="126" refString="Gibbes, L. R. (1850) On the carcinological collections of the United States, and on enumeration of species contained in them, with notes on the most remarkable, and descriptions of new species. Proceedings of the American Association for the Advancement of Science, 3, 167 - 201." type="journal article" year="1850">Gibbes 1850</bibRefCitation>
: 185
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4CFFE5637FB4ADFE16F9AE" author="Bell" box="[267,431,1978,2000]" page="362" pageId="2" pageNumber="126" year="1855">
<bibRefCitation id="EFD2559ECA4CFFE5637FB4ADFEC6F9AE" author="Bell" box="[267,383,1978,2000]" pageId="2" pageNumber="126" refString="Bell, T. (1855 a) Horae carcinologicae, or notices of Crustacea. I. A monograph of the Leucosiadae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species. Annals and Magazine of Natural History, 16, 361 - 367." type="journal article" year="1855" yearSuffix="a">Bell 1855a</bibRefCitation>
: 362
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4CFFE563B9B4ADFDC8F9AE" author="Bell" box="[461,625,1978,2000]" page="283" pageId="2" pageNumber="126" year="1855">
<bibRefCitation id="EFD2559ECA4CFFE563B9B4ADFDF8F9AE" author="Bell" box="[461,577,1978,2000]" pageId="2" pageNumber="126" refString="Bell, T. (1855 b) Horae carcinologicae, or notices of Crustacea. I. A monograph of the Leucosiadae, with observations on the relations, structure, habits and distribution of the family; a revision of the generic characters; and descriptions of new genera and species. Transactions of the Linnean Society, 21, 277 - 314. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1852. tb 00464. x" type="journal article" year="1855" yearSuffix="b">Bell 1855b</bibRefCitation>
: 283
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4CFFE560FBB4ADFCAEF9AE" author="Bell" box="[655,791,1978,2000]" page="5" pageId="2" pageNumber="126" year="1855">
<bibRefCitation id="EFD2559ECA4CFFE560FBB4ADFCBBF9AE" author="Bell" box="[655,770,1978,2000]" pageId="2" pageNumber="126" refString="Bell, T. (1855 c) Catalogue of Crustacea in the collections of British Museum. Part I. Leucosiadae. Order of the trustees, London, 24 pp." type="book" year="1855" yearSuffix="c">Bell 1855c</bibRefCitation>
: 5
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4CFFE56141B4ADFC66F9AE" author="Serene" box="[821,991,1978,2000]" page="47" pageId="2" pageNumber="126" year="1968">
<bibRefCitation id="EFD2559ECA4CFFE56141B4ADFC03F9AE" author="Serene" box="[821,954,1978,2000]" pageId="2" pageNumber="126" refString="Serene, R. (1968) Prodromus for a check-list of the non-planctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication 1, 1 - 122." type="journal article" year="1968">Serène 1968</bibRefCitation>
: 47
</treatmentCitation>
. 
<taxonomicName id="4C4353ECCA4CFFE562E3B4CDFEFCF991" ID-CoL="6PTZP" box="[151,325,2010,2031]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="anatum">
<emphasis id="B937F47DCA4CFFE562E3B4CDFEFCF991" box="[151,325,2010,2031]" italics="true" pageId="2" pageNumber="126">Leucosia anatum</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AE20E7ECA4CFFE56323B4CDFDA9F991" author="Sakai" box="[343,528,2009,2031]" page="19" pageId="2" pageNumber="126" year="1999">
<bibRefCitation id="EFD2559ECA4CFFE56323B4CDFE55F991" author="Sakai" box="[343,492,2009,2031]" pageId="2" pageNumber="126" refString="Sakai, K. (1999) J. F. W. Herbst collection of decapod Crustacea of the Berlin Zoological Museum with remarks on certain species. Naturalists, Publications of Tokushima Biological Laboratory, Shikoku University, 6, 1 - 45." type="journal article" year="1999">K. Sakai 1999</bibRefCitation>
: 19
</treatmentCitation>
, pl. 7F (not 
<taxonomicName id="4C4353ECCA4CFFE560FFB4CDFC0CF991" authority="Herbst, 1783" authorityName="Herbst" authorityYear="1783" box="[651,949,2009,2031]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="anatum">
<emphasis id="B937F47DCA4CFFE560FFB4CDFC91F991" box="[651,808,2010,2031]" italics="true" pageId="2" pageNumber="126">Cancer anatum</emphasis>
Herbst, 1783
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).
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</paragraph>
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<treatmentCitationGroup id="AB530F41CA4DFFE462E3B38EFBE4FED0" box="[151,1117,152,174]" pageId="3" pageNumber="127">
<taxonomicName id="4C4353ECCA4DFFE462E3B38EFEFCFED0" box="[151,325,152,174]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B937F47DCA4DFFE462E3B38EFEFCFED0" box="[151,325,152,174]" italics="true" pageId="3" pageNumber="127">Leucosia grandis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AE20E7ECA4DFFE4632DB38FFDFCFED0" author="Chen" box="[345,581,152,174]" page="422" pageId="3" pageNumber="127" year="2002">
<bibRefCitation id="EFD2559ECA4DFFE4632DB38FFDABFED0" author="Chen" box="[345,530,152,174]" pageId="3" pageNumber="127" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp." type="book" year="2002">Chen &amp; Sun 2002</bibRefCitation>
: 422
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, fig. 190, pl. 16.5, 6 [name not available, see above].
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</paragraph>
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<treatmentCitationGroup id="AB530F41CA4DFFE462E3B3A1FC34FEB2" box="[151,909,182,205]" pageId="3" pageNumber="127">
<taxonomicName id="4C4353ECCA4DFFE462E3B3A1FE82FEB3" ID-CoL="X2CB" box="[151,315,182,205]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462E3B3A1FE82FEB3" box="[151,315,182,205]" italics="true" pageId="3" pageNumber="127">Coleusia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE46321B3A1FE50FEB2" author="Galil" box="[341,489,182,204]" page="64" pageId="3" pageNumber="127" year="2006">
<bibRefCitation id="EFD2559ECA4DFFE46321B3A1FE7AFEB2" author="Galil" box="[341,451,182,204]" pageId="3" pageNumber="127" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 80, 55 - 69." type="journal article" year="2006">Galil 2006</bibRefCitation>
: 64
</treatmentCitation>
, figs. 2E, F, 3E, 4D.—Ng 
<emphasis id="B937F47DCA4DFFE46080B3AFFC9EFEB2" box="[756,807,182,205]" italics="true" pageId="3" pageNumber="127">et al.</emphasis>
2008: 94.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC286FCA4DFFE462E3B3C1FEB3FF74" blockId="3.[151,1436,152,389]" pageId="3" pageNumber="127">
<treatmentCitationGroup id="AB530F41CA4DFFE462E3B3C1FEB3FF74" pageId="3" pageNumber="127">
Not 
<taxonomicName id="4C4353ECCA4DFFE462BCB3C1FECBFE95" ID-CoL="84FXW" box="[200,370,214,235]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462BCB3C1FECBFE95" box="[200,370,214,235]" italics="true" pageId="3" pageNumber="127">Leucosia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE463F8B3C2FDE2FE95" author="Bianconi" box="[396,603,213,235]" page="341" pageId="3" pageNumber="127" year="1867">
<bibRefCitation id="EFD2559ECA4DFFE463F8B3C2FD91FE95" author="Bianconi" box="[396,552,213,235]" pageId="3" pageNumber="127" refString="Bianconi, G. G. (1867) Specimina Zoologica Mosambicana Fasciculus 19 - 20. Typ. Academiae Scientiarum, Bononiae, 333 - 363." type="book chapter" year="1867">Bianconi 1867</bibRefCitation>
: 341
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4DFFE4600CB3C2FCD8FE95" author="Hilgendorf" box="[632,865,213,235]" page="110" pageId="3" pageNumber="127" year="1869">
<bibRefCitation id="EFD2559ECA4DFFE4600CB3C2FC94FE95" author="Hilgendorf" box="[632,813,213,235]" pageId="3" pageNumber="127" refString="Hilgendorf, F. (1869) Crustaceen. In: Baron Carl Claus von der Decken Reisen in Ost-Afrika in den Jahren 1859 - 1865, 3, 67 - 116, 147." type="journal article" year="1869">Hilgendorf 1869</bibRefCitation>
: 110
</treatmentCitation>
.—De 
<treatmentCitation id="0AE20E7ECA4DFFE461DDB3C1FBF4FE95" author="Man" box="[937,1101,213,235]" page="256" pageId="3" pageNumber="127" year="1881">
<bibRefCitation id="EFD2559ECA4DFFE461DDB3C1FBA0FE95" author="Man" box="[937,1049,213,235]" pageId="3" pageNumber="127" refString="Man, J. G. De (1881) Note XXVIII. Carcinological studies in the Leyden Museum. Notes from the Leyden Museum, 3, 245 - 256." type="journal article" year="1881">Man 1881</bibRefCitation>
: 256
</treatmentCitation>
[= 
<taxonomicName id="4C4353ECCA4DFFE4660FB3C2FA92FE95" ID-CoL="5ZLQH" box="[1147,1323,213,235]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B937F47DCA4DFFE4660FB3C2FA92FE95" box="[1147,1323,213,235]" italics="true" pageId="3" pageNumber="127">Coleusia signata</emphasis>
</taxonomicName>
(Paul’son, 1875)]
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC286FCA4DFFE462E3B204FAAAFF56" blockId="3.[151,1436,152,389]" box="[151,1299,274,296]" pageId="3" pageNumber="127">
<treatmentCitationGroup id="AB530F41CA4DFFE462E3B204FAAAFF56" box="[151,1299,274,296]" pageId="3" pageNumber="127">
Not 
<taxonomicName id="4C4353ECCA4DFFE462B7B205FEC4FF56" box="[195,381,274,296]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462B7B205FEC4FF56" box="[195,381,274,296]" italics="true" pageId="3" pageNumber="127">Leucosides urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE463E3B205FDE2FF56" author="Rathbun" box="[407,603,274,296]" page="309" pageId="3" pageNumber="127" year="1910">
<bibRefCitation id="EFD2559ECA4DFFE463E3B205FD90FF56" author="Rathbun" box="[407,553,274,296]" pageId="3" pageNumber="127" refString="Rathbun, M. J. (1910) Brachyura. The Danish Expedition to Siam 1899 - 1900, V. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 7. Raekke, Naturvidenskabelig og Mathematisk Afdeling, 4, 301 - 367." type="journal article" year="1910">Rathbun 1910</bibRefCitation>
: 309
</treatmentCitation>
, pl. 1, fig. 1.— 
<treatmentCitation id="0AE20E7ECA4DFFE4609BB205FC12FF56" author="Suvatti" box="[751,939,274,296]" page="143" pageId="3" pageNumber="127" year="1950">
<bibRefCitation id="EFD2559ECA4DFFE4609BB205FCC1FF56" author="Suvatti" box="[751,888,274,296]" pageId="3" pageNumber="127" refString="Suvatti, C. (1950) Fauna of Thailand. Department of Fisheries, Bangkok, 1100 pp." type="book" year="1950">Suvatti 1950</bibRefCitation>
: 143
</treatmentCitation>
. [= 
<taxonomicName id="4C4353ECCA4DFFE461A6B205FAB0FF56" ID-CoL="4XYP4" authority="Miers, 1877" authorityName="Miers" authorityYear="1877" box="[978,1289,274,296]" class="Malacostraca" family="Leucosiidae" genus="Soceulia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B937F47DCA4DFFE461A6B205FB3AFF56" box="[978,1155,274,296]" italics="true" pageId="3" pageNumber="127">Soceulia brunnea</emphasis>
Miers, 1877
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC286FCA4DFFE462E3B225FC0BFF39" blockId="3.[151,1436,152,389]" box="[151,946,305,327]" pageId="3" pageNumber="127">
<treatmentCitationGroup id="AB530F41CA4DFFE462E3B225FC0BFF39" box="[151,946,305,327]" pageId="3" pageNumber="127">
Not 
<taxonomicName id="4C4353ECCA4DFFE462B7B225FEDEFF39" ID-CoL="84FXW" box="[195,359,306,327]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462B7B225FEDEFF39" box="[195,359,306,327]" italics="true" pageId="3" pageNumber="127">Leucosia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE463F6B226FDE5FF39" author="Hilgendorf" box="[386,604,305,327]" page="811" pageId="3" pageNumber="127" year="1879">
<bibRefCitation id="EFD2559ECA4DFFE463F6B226FD94FF39" author="Hilgendorf" box="[386,557,305,327]" pageId="3" pageNumber="127" refString="Hilgendorf, F. (1879) Die von Hrn. W. Peters in Mocambique gesammelten Crustaceen. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, 1878, 782 - 851." type="journal article" year="1879">Hilgendorf 1879</bibRefCitation>
: 811
</treatmentCitation>
[= 
<taxonomicName id="4C4353ECCA4DFFE4600BB226FC11FF39" ID-CoL="X2C9" authority="Galil, 2006" authorityName="Galil" authorityYear="2006" box="[639,936,305,327]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="rangita">
<emphasis id="B937F47DCA4DFFE4600BB226FC92FF39" box="[639,811,305,327]" italics="true" pageId="3" pageNumber="127">Coleusia rangita</emphasis>
<bibRefCitation id="EFD2559ECA4DFFE46145B226FC11FF39" author="Galil" box="[817,936,305,327]" pageId="3" pageNumber="127" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 80, 55 - 69." type="journal article" year="2006">Galil, 2006</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC286FCA4DFFE462E3B246FB34FF18" blockId="3.[151,1436,152,389]" box="[151,1165,336,358]" pageId="3" pageNumber="127">
<treatmentCitationGroup id="AB530F41CA4DFFE462E3B246FB34FF18" box="[151,1165,336,358]" pageId="3" pageNumber="127">
Not 
<taxonomicName id="4C4353ECCA4DFFE462B7B246FEDEFF18" ID-CoL="84FXW" box="[195,359,337,358]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462B7B246FEDEFF18" box="[195,359,337,358]" italics="true" pageId="3" pageNumber="127">Leucosia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE463F6B247FDFDFF18" author="Naiyanetr" box="[386,580,336,358]" page="58" pageId="3" pageNumber="127" year="1998">
<bibRefCitation id="EFD2559ECA4DFFE463F6B247FD9BFF18" author="Naiyanetr" box="[386,546,336,358]" pageId="3" pageNumber="127" refString="Naiyanetr, P. (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr 1998</bibRefCitation>
: 58
</treatmentCitation>
.— 
<treatmentCitation id="0AE20E7ECA4DFFE46015B247FC91FF18" author="Naiyanetr" box="[609,808,336,358]" page="69" pageId="3" pageNumber="127" year="2007">
<bibRefCitation id="EFD2559ECA4DFFE46015B247FCBCFF18" author="Naiyanetr" box="[609,773,336,358]" pageId="3" pageNumber="127" refString="Naiyanetr, P. (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, 196 pp." type="book" year="2007">Naiyanetr 2007</bibRefCitation>
: 69
</treatmentCitation>
[= 
<taxonomicName id="4C4353ECCA4DFFE46138B247FB3AFF18" ID-CoL="4XYP4" authority="Miers, 1877" authorityName="Miers" authorityYear="1877" box="[844,1155,336,358]" class="Malacostraca" family="Leucosiidae" genus="Soceulia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B937F47DCA4DFFE46138B247FC44FF18" box="[844,1021,336,358]" italics="true" pageId="3" pageNumber="127">Soceulia brunnea</emphasis>
Miers, 1877
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC286FCA4DFFE462E3B278FB92FFFA" blockId="3.[151,1436,152,389]" box="[151,1067,366,389]" pageId="3" pageNumber="127">
<treatmentCitationGroup id="AB530F41CA4DFFE462E3B278FB92FFFA" box="[151,1067,366,389]" pageId="3" pageNumber="127">
Not 
<taxonomicName id="4C4353ECCA4DFFE462B7B267FED1FFFB" ID-CoL="84FXW" box="[195,360,368,389]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4DFFE462B7B267FED1FFFB" box="[195,360,368,389]" italics="true" pageId="3" pageNumber="127">Leucosia urania</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AE20E7ECA4DFFE463F6B279FD8EFFFA" author="Alcock" box="[386,567,366,388]" page="220" pageId="3" pageNumber="127" year="1896">
<bibRefCitation id="EFD2559ECA4DFFE463F6B279FDBEFFFA" author="Alcock" box="[386,519,366,388]" pageId="3" pageNumber="127" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, 65 (2), 134 - 296." type="journal article" year="1896">Alcock 1896</bibRefCitation>
: 220
</treatmentCitation>
[= 
<taxonomicName id="4C4353ECCA4DFFE4602EB279FD0DFFFA" ID-CoL="7P3KK" box="[602,692,366,388]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B937F47DCA4DFFE4602EB279FD0DFFFA" box="[602,692,366,388]" italics="true" pageId="3" pageNumber="127">Coleusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2044906CA4DFFE460CFB278FC8EFFFA" box="[699,823,366,388]" pageId="3" pageNumber="127" rank="species">new species</taxonomicNameLabel>
, 
<bibRefCitation id="EFD2559ECA4DFFE46135B279FB98FFFA" author="Promdam" box="[833,1057,366,389]" pageId="3" pageNumber="127" refString="Promdam, R., Nabhitabhata, J. &amp; Galil, B. S. (2014) A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia. Zootaxa, 3786 (2), 135 - 140. http: // dx. doi. org / 10.11646 / zootaxa. 3786.2.3" type="journal article" year="2014">
Promdam 
<emphasis id="B937F47DCA4DFFE461DEB267FC64FFFA" box="[938,989,366,389]" italics="true" pageId="3" pageNumber="127">et al.</emphasis>
, 2014
</bibRefCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3597BE4CA4DFFE462E3B2BDFDC1FC04" pageId="3" pageNumber="127" type="materials_examined">
<paragraph id="8BFC286FCA4DFFE462E3B2BDFDC1FC04" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
<emphasis id="B937F47DCA4DFFE462E3B2BDFEC4FFBD" bold="true" box="[151,381,426,451]" pageId="3" pageNumber="127">Material examined</emphasis>
. 
<collectingCountry id="F35468FFCA4DFFE463FAB2BDFE6EFFBD" box="[398,471,426,451]" name="China" pageId="3" pageNumber="127">
<emphasis id="B937F47DCA4DFFE463FAB2BDFE6EFFBD" bold="true" box="[398,471,426,451]" pageId="3" pageNumber="127">China</emphasis>
</collectingCountry>
. Guangdong Province, Haimen, 07– 
<date id="FFFD0EAFCA4DFFE461F4B2BDFBB9FFBD" box="[896,1024,426,451]" pageId="3" pageNumber="127" value="1957-05-15">15.05.1957</date>
, coll. Z. Fan, 
<typeStatus id="54F896CDCA4DFFE466DEB2BCFAB2FFBD" box="[1194,1291,427,451]" pageId="3" pageNumber="127" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C4353ECCA4DFFE46743B2BCFE0AFF99" authority="Chen &amp; Turkay" authorityName="Chen &amp; Turkay" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B937F47DCA4DFFE46743B2BCFF54FF98" italics="true" pageId="3" pageNumber="127">Leucosia grandis</emphasis>
Chen &amp; Türkay
</taxonomicName>
, 
<specimenCount id="9D45E3E6CA4DFFE463B7B2D8FE4BFF98" box="[451,498,463,487]" pageId="3" pageNumber="127" type="generic">1 ♂</specimenCount>
(42.6 × 
<quantity id="4CBB858ACA4DFFE46015B2D8FD72FF99" box="[609,715,463,487]" metricMagnitude="-2" metricUnit="m" metricValue="3.87" pageId="3" pageNumber="127" unit="mm" value="38.7">38.7 mm</quantity>
) (
<collectionCode id="ED52B0AACA4DFFE46092B2D8FC99FF99" LSID="urn:lsid:biocol.org:col:34838" box="[742,800,463,487]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="127">SMF</collectionCode>
25700); Guangdong Province, Shan Wei, 
<geoCoordinate id="EE774EA8CA4DFFE4676CB2D9FA22FF99" box="[1304,1435,462,487]" direction="north" orientation="latitude" pageId="3" pageNumber="127" precision="9" value="22.776667">22°46.60’N</geoCoordinate>
<geoCoordinate id="EE774EA8CA4DFFE462E3B2E5FE9EFC75" box="[151,295,498,523]" direction="east" orientation="longitude" pageId="3" pageNumber="127" precision="9" value="115.34333">115°20.60’E</geoCoordinate>
, donated H.L. Chen, 
<collectionCode id="ED52B0AACA4DFFE46063B2E5FDD7FC74" box="[535,622,498,522]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="3" pageNumber="127">IOCAS</collectionCode>
, 
<specimenCount id="9D45E3E6CA4DFFE4600DB2E5FD1DFC74" box="[633,676,498,522]" pageId="3" pageNumber="127" type="generic">1 ♂</specimenCount>
(38.7 × 
<quantity id="4CBB858ACA4DFFE46170B2E5FCD3FC74" box="[772,874,498,522]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="3" pageNumber="127" unit="mm" value="34.8">34.8 mm</quantity>
) (
<collectionCode id="ED52B0AACA4DFFE461F4B2E5FC01FC74" box="[896,952,498,522]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="127">ZRC</collectionCode>
2013.1295). 
<collectingCountry id="F35468FFCA4DFFE4663AB2E5FB1AFC75" box="[1102,1187,498,523]" name="Taiwan" pageId="3" pageNumber="127">
<emphasis id="B937F47DCA4DFFE4663AB2E5FB1AFC75" bold="true" box="[1102,1187,498,523]" pageId="3" pageNumber="127">Taiwan</emphasis>
</collectingCountry>
. Su-Ao, Nangfangao, 0 
<date id="FFFD0EAFCA4DFFE462D1B101FEAFFC51" box="[165,278,534,559]" pageId="3" pageNumber="127" value="2000-05-11">5.11.2000</date>
, coll. P.K.L. Ng, 
<specimenCount id="9D45E3E6CA4DFFE46393B100FDACFC50" box="[487,533,535,559]" pageId="3" pageNumber="127" type="generic">2 ♂</specimenCount>
(40.9 × 
<quantity id="4CBB858ACA4DFFE4600BB101FD50FC50" box="[639,745,534,559]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="3" pageNumber="127" unit="mm" value="36.8">36.8 mm</quantity>
, 40.3 × 
<quantity id="4CBB858ACA4DFFE4613AB101FC0EFC51" box="[846,951,534,559]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" pageId="3" pageNumber="127" unit="mm" value="36.5">36.5 mm</quantity>
) (
<collectionCode id="ED52B0AACA4DFFE461A6B100FBB3FC51" box="[978,1034,535,559]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="127">ZRC</collectionCode>
2001.0045); Su-Ao, Nangfangao, 
<date id="FFFD0EAFCA4DFFE462E3B12CFEA4FC2B" box="[151,285,571,597]" pageId="3" pageNumber="127" value="1999-03-23">23.03.1999</date>
, coll. M.-S. Hung, 
<specimenCount id="9D45E3E6CA4DFFE46074B12CFD95FC2B" box="[512,556,571,597]" pageId="3" pageNumber="127" type="generic">1 ♂</specimenCount>
(35.7 × 
<quantity id="4CBB858ACA4DFFE460E5B12CFD42FC2B" box="[657,763,571,597]" metricMagnitude="-2" metricUnit="m" metricValue="4.01" pageId="3" pageNumber="127" unit="mm" value="40.1">40.1 mm</quantity>
) (
<collectionCode id="ED52B0AACA4DFFE46167B12CFCF7FC2B" box="[787,846,571,597]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="127">ZRC</collectionCode>
2014.0175); Tahsi, 0 6.04.1986, coll. J.-F. Huang, 
<specimenCount id="9D45E3E6CA4DFFE462E3B177FF7AFC04" box="[151,195,608,634]" pageId="3" pageNumber="127" type="generic">1 ♂</specimenCount>
(35.8 × 
<quantity id="4CBB858ACA4DFFE46352B177FE29FC04" box="[294,400,608,634]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="3" pageNumber="127" unit="mm" value="39.4">39.4 mm</quantity>
) (
<collectionCode id="ED52B0AACA4DFFE463DCB177FE5AFC04" box="[424,483,608,634]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="127">ZRC</collectionCode>
2014.0174).
</paragraph>
</subSubSection>
<subSubSection id="C3597BE4CA4DFFE262B3B193FD65F903" lastPageId="5" lastPageNumber="129" pageId="3" pageNumber="127" type="description">
<paragraph id="8BFC286FCA4DFFE462B3B193FC07FDE6" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
<emphasis id="B937F47DCA4DFFE462B3B193FEE8FCE3" bold="true" box="[199,337,644,669]" pageId="3" pageNumber="127">Description</emphasis>
. Carapace (
<figureCitation id="137834EACA4DFFE463A9B193FD94FCE3" box="[477,557,644,669]" captionStart="FIGURE 1" captionStartId="4.[151,250,1950,1972]" captionTargetBox="[162,1424,221,1928]" captionTargetId="figure@4.[162,1426,195,1928]" captionTargetPageId="4" captionText="FIGURE 1. Coleusia urania (Herbst, 1801). A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T. - Y. Chan)." httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="3" pageNumber="127">Figs. 1</figureCitation>
, 
<figureCitation id="137834EACA4DFFE4604AB192FDF4FCE3" box="[574,589,645,669]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">2</figureCitation>
B, 4A) subpentagonal, globose, 1.1 times as long as wide; dorsal surface glabrous, smooth, sparsely punctate; regions indistinct. Frontal region produced, upcurved, deeply concave medially, laterally constricted; frontal margin rounded. Anterolateral margin sinuous, with obsolescent granules distally (
<figureCitation id="137834EACA4DFFE4628CB1E7FEFFFD76" box="[248,326,752,777]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
B, 4A). Lateral angle of carapace rounded, overhanging thoracic sinus, margin prominently beaded, granules lozenge-shaped. Posterolateral margins rounded, smooth (
<figureCitation id="137834EACA4DFFE461E9B003FC56FD52" box="[925,1007,788,813]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
B, 4A). Epimeral margin invisible in dorsal view, gutter-like anteriorly, narrowing posteriorly, closely beaded marginal granules decreasing in size posteriorly, meeting posterior margin at an angle. Posterior margin minutely granular, sinuous in male (
<figureCitation id="137834EACA4DFFE46747B04CFAC1FD0A" box="[1331,1400,859,884]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
A), slightly rounded in female (
<figureCitation id="137834EACA4DFFE463B9B068FDA8FDE9" box="[461,529,895,920]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Fig. 2</figureCitation>
B); deflexed posterior surface smooth.
</paragraph>
<paragraph id="8BFC286FCA4DFFE462B3B0B4FC4DFA34" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
Antennule obliquely folded into fossa, fossa sealed by basal antennular segment. Antennae short, inserted between antennular fossa, orbit. Orbits small, rounded, outer orbital margin with single suture, anterior margin of efferent branchial channel forms part of lower orbital margin. Eyes retractable. External maxillipeds completely concealing trapezoid buccal opening, smooth; endopod merus triangular, as long as ischium; in female, endopod with setose fringe lengthwise, ischium medially elevated, keel-like (
<figureCitation id="137834EACA4DFFE461E5B726FC6CFA34" box="[913,981,1073,1098]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
F).
</paragraph>
<paragraph id="8BFC286FCA4DFFE462B3B742FAAFFBAD" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
Thoracic sinus deep, tomentose, anterior margin level with base of coxopodite of third maxillipeds, anteriorly defined by overhanging beaded margin of pterygostomian region; ventral margin medially with row of oval granules, decreasing in size posteriorly (
<figureCitation id="137834EACA4DFFE46014B78AFD09FACB" box="[608,688,1181,1206]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
E, 4B). Thoracic sternites smooth, shiny; sternites 1–3 completely fused, trapezoidal, separated from sternite 4 by sinuous suture; sternites 4–8 with medially interrupted sutures between them (
<figureCitation id="137834EACA4DFFE4633DB7F3FE29FA83" box="[329,400,1252,1277]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
F); distinct peg-like abdominal locking mechanism on sternite 5, adjacent to sternite 
<quantity id="4CBB858ACA4DFFE4671AB7F2FA25FA83" box="[1390,1436,1252,1277]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="127" unit="in" value="4.0">4 in</quantity>
male. Abdominal cavity nearly reaching buccal cavity, lateral margins prominently ridged medially, ridge fitting into notch between somites 5, 6 (
<figureCitation id="137834EACA4DFFE4606DB63BFDE6FB3A" box="[537,607,1324,1349]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
F). Male abdomen smooth; somite 1 narrow, transverse; somite 2 small, narrow, spindle-shaped; somites 3–6 fused, proximally inflated, furrowed lengthwise, suture between somites 5, 6 distinct but somites immovable; somite 6 elongated, lateral margins evenly curved, narrowing distally, with prominent triangular denticle subproximally; telson triangular, lateral margins gently convex, as wide as long (
<figureCitation id="137834EACA4DFFE4671AB680FF1EFBAD" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
F). Female abdomen with somites 3–6 fused, greatly enlarged, shield-like, telson triangular (
<figureCitation id="137834EACA4DFFE466C5B6ACFB4DFBAD" box="[1201,1268,1467,1492]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="8BFC286FCA4DFFE462B3B6C8FA20F865" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
G1 elongated, shaft stout, sinuous, tightly coiled 3 times on its axis, ultimate coil swollen, setose; apical process cornute, beak-like, curved interiorly (
<figureCitation id="137834EACA4DFFE460E0B515FD5DF865" box="[660,740,1538,1563]" captionStart="FIGURE 5" captionStartId="8.[151,250,1592,1614]" captionTargetBox="[204,1383,231,1547]" captionTargetId="figure@8.[204,1384,231,1547]" captionTargetPageId="8" captionText="FIGURE 5. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), intact left G 1. A, dorsal view; B, ventral view." httpUri="https://zenodo.org/record/230321/files/figure.png" pageId="3" pageNumber="127">Figs. 5</figureCitation>
, 
<figureCitation id="137834EACA4DFFE46084B515FCB9F865" box="[752,768,1538,1563]" captionStart="FIGURE 6" captionStartId="9.[151,250,1233,1255]" captionTargetBox="[160,1429,202,1204]" captionTargetId="figure@9.[151,1436,195,1211]" captionTargetPageId="9" captionText="FIGURE 6. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), gonopods. A, left G 1; B, C, distal half of left G 1; D, left G 2; E, distal part of left G 2. Setae of all structures denuded. Scales: A, D = 3.0 mm; B, C, E = 1.0 mm." httpUri="https://zenodo.org/record/230322/files/figure.png" pageId="3" pageNumber="127">6</figureCitation>
). G2 relatively short, slender, apex scoop-like (
<figureCitation id="137834EACA4DFFE46767B515FAECF865" box="[1299,1365,1538,1563]" captionStart="FIGURE 6" captionStartId="9.[151,250,1233,1255]" captionTargetBox="[160,1429,202,1204]" captionTargetId="figure@9.[151,1436,195,1211]" captionTargetPageId="9" captionText="FIGURE 6. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), gonopods. A, left G 1; B, C, distal half of left G 1; D, left G 2; E, distal part of left G 2. Setae of all structures denuded. Scales: A, D = 3.0 mm; B, C, E = 1.0 mm." httpUri="https://zenodo.org/record/230322/files/figure.png" pageId="3" pageNumber="127">Fig. 6</figureCitation>
D, E).
</paragraph>
<paragraph id="8BFC286FCA4DFFE462B3B531FCC8F96B" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
Chelipeds subequal, robust, relatively longer in adult male (
<figureCitation id="137834EACA4DFFE46117B531FC1CF841" box="[867,933,1574,1599]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
A) than in females (
<figureCitation id="137834EACA4DFFE466F7B531FB7EF841" box="[1155,1223,1574,1599]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Fig. 2</figureCitation>
B). Merus trigonal; anterior, posterior margins lined along their length with perliform granules, diminishing in size distally; upper surface with coalesced cluster of granules proximally, followed by 4 large perliform granules, distally smooth; lower surface pitted, proximally granular, laterally with rows of small granules (
<figureCitation id="137834EACA4DFFE4664AB586FB37F8D4" box="[1086,1166,1681,1706]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
F, 4C). Carpus inflated, smooth (
<figureCitation id="137834EACA4DFFE4628AB5A2FEE8F8B3" box="[254,337,1717,1742]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
B, 4A). Propodus lenticular, slightly longer than wide, upper margin rounded, smooth; lower margin with row of granules, lower external surface with small, rounded granules (
<figureCitation id="137834EACA4DFFE46620B5CFFB1DF88F" box="[1108,1188,1752,1777]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">Figs. 2</figureCitation>
B, 4A, D). Fingers as long as propodus, gaping, cutting edges, finely teethed (
<figureCitation id="137834EACA4DFFE4617DB5EBFCF4F96B" box="[777,845,1788,1813]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BFC286FCA4DFFE462B3B437FADCF995" blockId="3.[151,1436,426,2028]" pageId="3" pageNumber="127">
Ambulatory legs similar in shape, gradually decreasing in length from first to fourth (
<figureCitation id="137834EACA4DFFE466E5B437FB59F946" box="[1169,1248,1824,1849]" captionStart="FIGURE 1" captionStartId="4.[151,250,1950,1972]" captionTargetBox="[162,1424,221,1928]" captionTargetId="figure@4.[162,1426,195,1928]" captionTargetPageId="4" captionText="FIGURE 1. Coleusia urania (Herbst, 1801). A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T. - Y. Chan)." httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="3" pageNumber="127">Figs. 1</figureCitation>
, 
<figureCitation id="137834EACA4DFFE4669BB437FB47F946" box="[1263,1278,1824,1848]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="3" pageNumber="127">2</figureCitation>
B, 4A). Coxal condyles rounded. Meri subcylindrical; first 3 meri with anterior parts bearing 2 granular rows on respective dorsal margin, posterior parts distally obsolescent, 2 granular rows on ventral margins, anterior parts proximally obsolescent; merus of last leg with single granular row dorsally, prominently granular crest subproximally on ventral margin (
<figureCitation id="137834EACA4DFFE4633BB4B8FE2CF9B6" box="[335,405,1967,1992]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
E). Carpi rounded, minute denticle on upper surface (
<figureCitation id="137834EACA4DFFE46189B4B8FBFDF9B6" box="[1021,1092,1967,1992]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
E). Propodi dorsally, ventrally carinate, expanded, foliaceous (
<figureCitation id="137834EACA4DFFE4638DB4C4FD85F995" box="[505,572,2003,2028]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
A, E). Dactyli longer than propodi, lanceolate, subspatuliform (
<figureCitation id="137834EACA4DFFE46775B4C4FAFDF995" box="[1281,1348,2003,2028]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="3" pageNumber="127">Fig. 4</figureCitation>
E).
</paragraph>
<caption id="DF3C78E7CA4AFFE362E3B489FAC8F9AD" httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="4" pageNumber="128" targetBox="[162,1424,221,1928]" targetPageId="4">
<paragraph id="8BFC286FCA4AFFE362E3B489FAC8F9AD" blockId="4.[151,1435,1950,2003]" pageId="4" pageNumber="128">
<emphasis id="B937F47DCA4AFFE362E3B489FEB7F9CD" bold="true" box="[151,270,1950,1972]" pageId="4" pageNumber="128">FIGURE 1</emphasis>
. 
<taxonomicName id="4C4353ECCA4AFFE36369B489FDDCF9CA" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[285,613,1950,1972]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="128" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4AFFE36369B489FE7AF9CA" box="[285,451,1950,1972]" italics="true" pageId="4" pageNumber="128">Coleusia urania</emphasis>
(Herbst, 1801)
</taxonomicName>
. A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T.-Y. Chan).
</paragraph>
</caption>
<caption id="DF3C78E7CA4BFFE262E3B54DFD23F8B5" httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="5" pageNumber="129" targetBox="[151,1436,193,1603]" targetPageId="5">
<paragraph id="8BFC286FCA4BFFE262E3B54DFD23F8B5" blockId="5.[151,1436,1626,1739]" pageId="5" pageNumber="129">
<emphasis id="B937F47DCA4BFFE262E3B54DFEB5F811" bold="true" box="[151,268,1626,1648]" pageId="5" pageNumber="129">FIGURE 2</emphasis>
. 
<taxonomicName id="4C4353ECCA4BFFE2636EB54DFD82F80E" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[282,571,1626,1648]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4BFFE2636EB54DFE16F80E" box="[282,431,1626,1648]" italics="true" pageId="5" pageNumber="129">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman.
</paragraph>
</caption>
<paragraph id="8BFC286FCA4BFFE262B3B5EFFD65F903" blockId="5.[151,1436,1784,2025]" pageId="5" pageNumber="129">
<emphasis id="B937F47DCA4BFFE262B3B5EFFEA3F96F" bold="true" box="[199,282,1784,1809]" pageId="5" pageNumber="129">Colour</emphasis>
. Dorsal surface of carapace yellowish orange to white, paler laterally and posteriorly; a broad whitish stripe proceeding backwards from the front; two large spots on the posterior part of the carapace, and two smaller ones on each lateral margin, the basal portion of the fingers, and a ring on each segment of the ambulatory legs, are all rust coloured (
<figureCitation id="137834EACA4BFFE26328B473FE24F903" box="[348,413,1892,1917]" captionStart="FIGURE 1" captionStartId="4.[151,250,1950,1972]" captionTargetBox="[162,1424,221,1928]" captionTargetId="figure@4.[162,1426,195,1928]" captionTargetPageId="4" captionText="FIGURE 1. Coleusia urania (Herbst, 1801). A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T. - Y. Chan)." httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="5" pageNumber="129">Fig. 1</figureCitation>
). Ventral surface paler grey.
</paragraph>
</subSubSection>
<subSubSection id="C3597BE4CA4BFFE062B3B49FFDE1F9A0" lastPageId="7" lastPageNumber="131" pageId="5" pageNumber="129" type="discussion">
<paragraph id="8BFC286FCA4BFFE062B3B49FFDE1F9A0" blockId="5.[151,1436,1784,2025]" lastBlockId="7.[151,1436,1737,2014]" lastPageId="7" lastPageNumber="131" pageId="5" pageNumber="129">
<emphasis id="B937F47DCA4BFFE262B3B49FFE8DF9DF" bold="true" box="[199,308,1928,1953]" pageId="5" pageNumber="129">Remarks</emphasis>
. 
<taxonomicName id="4C4353ECCA4BFFE2633CB49FFD00F9DF" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[328,697,1928,1953]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA4BFFE2633CB49FFDBAF9DE" box="[328,515,1928,1952]" italics="true" pageId="5" pageNumber="129">Coleusia urania</emphasis>
(Herbst, 1801)
</taxonomicName>
differs from its congeners— 
<taxonomicName id="4C4353ECCA4BFFE26660B49EFE82F9BB" authority="Tyndale-Biscoe &amp; George, 1962" authorityName="Tyndale-Biscoe &amp; George" authorityYear="1962" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="biannulata">
<emphasis id="B937F47DCA4BFFE26660B49EFB0AF9DE" box="[1044,1203,1928,1952]" italics="true" pageId="5" pageNumber="129">C. biannulata</emphasis>
(
<bibRefCitation id="EFD2559ECA4BFFE266BCB49FFE8AF9BB" author="Tyndale-Biscoe" pageId="5" pageNumber="129" refString="Tyndale-Biscoe, M. &amp; George, R. W. (1962) The Oxystomata and Gymnopleura (Crustacea, Brachyura) of Western Australia with descriptions of two new species from Western Australia and one from India. Journal of the Royal Society of Western Australia, 45, 65 - 96." type="journal article" year="1962">Tyndale-Biscoe &amp; George, 1962</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C4353ECCA4BFFE26333B4BAFC85F9BB" authority="Tyndale-Biscoe &amp; George, 1962" authorityName="Tyndale-Biscoe &amp; George" authorityYear="1962" box="[327,828,1964,1989]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="magna">
<emphasis id="B937F47DCA4BFFE26333B4BAFE0DF9BA" box="[327,436,1965,1988]" italics="true" pageId="5" pageNumber="129">C. magna</emphasis>
(
<bibRefCitation id="EFD2559ECA4BFFE263B0B4BBFC8AF9BB" author="Tyndale-Biscoe" box="[452,819,1964,1989]" pageId="5" pageNumber="129" refString="Tyndale-Biscoe, M. &amp; George, R. W. (1962) The Oxystomata and Gymnopleura (Crustacea, Brachyura) of Western Australia with descriptions of two new species from Western Australia and one from India. Journal of the Royal Society of Western Australia, 45, 65 - 96." type="journal article" year="1962">Tyndale-Biscoe &amp; George, 1962</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C4353ECCA4BFFE2613EB4BAFBFEF9BB" authority="Galil, 2006" authorityName="Galil" authorityYear="2006" box="[842,1095,1964,1989]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="rangita">
<emphasis id="B937F47DCA4BFFE2613EB4BAFC06F9BA" box="[842,959,1965,1988]" italics="true" pageId="5" pageNumber="129">C. rangita</emphasis>
<bibRefCitation id="EFD2559ECA4BFFE261B1B4BBFBFEF9BB" author="Galil" box="[965,1095,1964,1989]" pageId="5" pageNumber="129" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 80, 55 - 69." type="journal article" year="2006">Galil, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C4353ECCA4BFFE26627B4BAFAC7F9BB" authority="Paulson, 1875" authorityName="Paulson" authorityYear="1875" box="[1107,1406,1964,1989]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="129" phylum="Arthropoda" rank="species" species="signata">
<emphasis id="B937F47DCA4BFFE26627B4BAFB7EF9BA" box="[1107,1223,1965,1988]" italics="true" pageId="5" pageNumber="129">C. signata</emphasis>
(Paulson, 1875)
</taxonomicName>
— in having a prominently granular rather than smooth fourth ambulatory merus (
<figureCitation id="137834EACA4BFFE2665DB4C7FBD5F997" box="[1065,1132,2000,2025]" captionStart="FIGURE 4" captionStartId="7.[151,250,1609,1631]" captionTargetBox="[159,1418,176,1584]" captionTargetId="figure@7.[159,1419,175,1585]" captionTargetPageId="7" captionText="FIGURE 4. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax." httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="5" pageNumber="129">Fig. 4</figureCitation>
E); an expanded, distinctly foliaceous ambulatory propodus (
<figureCitation id="137834EACA49FFE06008B5DEFD70F89F" box="[636,713,1737,1762]" captionStart="FIGURE 2" captionStartId="5.[151,250,1626,1648]" captionTargetBox="[151,1436,193,1603]" captionTargetId="figure@5.[151,1436,193,1604]" captionTargetPageId="5" captionText="FIGURE 2. Cancer urania Herbst, 1801, lectotype female (34.0 × 32.0 mm) (ZMB Herbst 2183). A, box in ZMB where type female is kept; B, overall dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace. All photographs by O. Coleman." httpUri="https://zenodo.org/record/230318/files/figure.png" pageId="7" pageNumber="131">Figs. 2</figureCitation>
B, C, 4A, E); the interiorly curved beak-like apical process of the G1 (
<figureCitation id="137834EACA49FFE062BCB5FBFEAFF97B" box="[200,278,1772,1797]" captionStart="FIGURE 5" captionStartId="8.[151,250,1592,1614]" captionTargetBox="[204,1383,231,1547]" captionTargetId="figure@8.[204,1384,231,1547]" captionTargetPageId="8" captionText="FIGURE 5. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), intact left G 1. A, dorsal view; B, ventral view." httpUri="https://zenodo.org/record/230321/files/figure.png" pageId="7" pageNumber="131">Figs. 5</figureCitation>
, 
<figureCitation id="137834EACA49FFE06356B5FBFE89F97B" box="[290,304,1772,1797]" captionStart="FIGURE 6" captionStartId="9.[151,250,1233,1255]" captionTargetBox="[160,1429,202,1204]" captionTargetId="figure@9.[151,1436,195,1211]" captionTargetPageId="9" captionText="FIGURE 6. Coleusia urania (Herbst, 1801), male (40.3 × 36.5 mm) (ZRC 2001.0045), gonopods. A, left G 1; B, C, distal half of left G 1; D, left G 2; E, distal part of left G 2. Setae of all structures denuded. Scales: A, D = 3.0 mm; B, C, E = 1.0 mm." httpUri="https://zenodo.org/record/230322/files/figure.png" pageId="7" pageNumber="131">6</figureCitation>
A–C); and in its distinctive color pattern (
<figureCitation id="137834EACA49FFE0608BB5FBFCF9F97B" box="[767,832,1772,1797]" captionStart="FIGURE 1" captionStartId="4.[151,250,1950,1972]" captionTargetBox="[162,1424,221,1928]" captionTargetId="figure@4.[162,1426,195,1928]" captionTargetPageId="4" captionText="FIGURE 1. Coleusia urania (Herbst, 1801). A, male (46.0 × 37.7 mm) (after Herbst, 1801: pl. 53, fig. 3; B, sex uncertain (specimen not examined), Fangliao fishing port, Pingtung County, Taiwan, coll. 28 May 1999 (photograph by T. - Y. Chan)." httpUri="https://zenodo.org/record/230317/files/figure.png" pageId="7" pageNumber="131">Fig. 1</figureCitation>
). The record of “ 
<taxonomicName id="4C4353ECCA49FFE06675B5F9FB0EF97B" box="[1025,1207,1774,1797]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA49FFE06675B5F9FB0EF97B" box="[1025,1207,1774,1797]" italics="true" pageId="7" pageNumber="131">Leucosia urania</emphasis>
</taxonomicName>
” from the Andaman Sea by 
<bibRefCitation id="EFD2559ECA49FFE0629EB406FE7FF957" author="Alcock" box="[234,454,1809,1834]" pageId="7" pageNumber="131" refString="Alcock, A. (1896) Materials for a carcinological fauna of India. No. 2. The Brachyura Oxystoma. Journal of the Asiatic Society of Bengal, 65 (2), 134 - 296." type="journal article" year="1896">Alcock (1896: 220)</bibRefCitation>
must be referred to a new species of 
<taxonomicName id="4C4353ECCA49FFE0611FB406FB61F957" authority="Promdam et al. 2014" authorityName="Promdam et al." authorityYear="2014" box="[875,1240,1809,1834]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="B937F47DCA49FFE0611FB406FC76F957" box="[875,975,1809,1833]" italics="true" pageId="7" pageNumber="131">Coleusia</emphasis>
(
<bibRefCitation id="EFD2559ECA49FFE061A9B406FB69F957" author="Promdam" box="[989,1232,1809,1834]" pageId="7" pageNumber="131" refString="Promdam, R., Nabhitabhata, J. &amp; Galil, B. S. (2014) A new species of Coleusia Galil, 2006 (Decapoda: Brachyura: Leucosiidae) from southern Asia. Zootaxa, 3786 (2), 135 - 140. http: // dx. doi. org / 10.11646 / zootaxa. 3786.2.3" type="journal article" year="2014">
Promdam 
<emphasis id="B937F47DCA49FFE06627B405FB35F957" box="[1107,1164,1809,1833]" italics="true" pageId="7" pageNumber="131">et al.</emphasis>
2014
</bibRefCitation>
)
</taxonomicName>
. The same is true for some specimens identified as &quot; 
<taxonomicName id="4C4353ECCA49FFE06055B421FD5AF933" box="[545,739,1845,1869]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B937F47DCA49FFE06055B421FD5AF933" box="[545,739,1845,1869]" italics="true" pageId="7" pageNumber="131">Leucosia grandis</emphasis>
</taxonomicName>
&quot; by 
<bibRefCitation id="EFD2559ECA49FFE06168B423FB80F933" author="Chen" box="[796,1081,1844,1869]" pageId="7" pageNumber="131" refString="Chen, H. &amp; Sun, H. (2002) Arthropoda Crustacea. Brachyura. Marine primitive crabs. Fauna Sinica. Invertebrata, 30, Science Press, Beijing, 597 pp." type="book" year="2002">Chen &amp; Sun (2002: 422)</bibRefCitation>
from the Gulf of 
<collectingCountry id="F35468FFCA49FFE06773B423FAD2F933" box="[1287,1387,1844,1869]" name="Thailand" pageId="7" pageNumber="131">Thailand</collectingCountry>
and 
<collectingCountry id="F35468FFCA49FFE062E3B44EFEB5F90C" box="[151,268,1881,1906]" name="Singapore" pageId="7" pageNumber="131">Singapore</collectingCountry>
. This new species is superficially similar to 
<taxonomicName id="4C4353ECCA49FFE06177B44EFCD6F90F" box="[771,879,1881,1905]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA49FFE06177B44EFCD6F90F" box="[771,879,1881,1905]" italics="true" pageId="7" pageNumber="131">C. urania</emphasis>
</taxonomicName>
but differs in the form of the chelipeds and most significantly, in a different G1 structure. The records from the Gulf of 
<collectingCountry id="F35468FFCA49FFE06199B46AFBE8F9E8" box="[1005,1105,1917,1942]" name="Thailand" pageId="7" pageNumber="131">Thailand</collectingCountry>
by 
<bibRefCitation id="EFD2559ECA49FFE066F2B46AFECBF9C7" author="Rathbun" pageId="7" pageNumber="131" refString="Rathbun, M. J. (1910) Brachyura. The Danish Expedition to Siam 1899 - 1900, V. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 7. Raekke, Naturvidenskabelig og Mathematisk Afdeling, 4, 301 - 367." type="journal article" year="1910">
Rathbun (1910: 309, as 
<taxonomicName id="4C4353ECCA49FFE062E3B4B6FED0F9C7" box="[151,361,1953,1977]" class="Malacostraca" family="Leucosiidae" genus="Leucosides" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA49FFE062E3B4B6FED0F9C7" box="[151,361,1953,1977]" italics="true" pageId="7" pageNumber="131">Leucosides urania</emphasis>
</taxonomicName>
)
</bibRefCitation>
and 
<bibRefCitation id="EFD2559ECA49FFE063C5B4B6FDC9F9C7" author="Naiyanetr" box="[433,624,1953,1978]" pageId="7" pageNumber="131" refString="Naiyanetr, P. (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, 161 pp." type="book" year="1998">Naiyanetr (1998</bibRefCitation>
: 58; 2007: 69, as 
<taxonomicName id="4C4353ECCA49FFE0613DB4B5FBBDF9C7" box="[841,1028,1954,1977]" class="Malacostraca" family="Leucosiidae" genus="Leucosia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA49FFE0613DB4B5FBBDF9C7" box="[841,1028,1954,1977]" italics="true" pageId="7" pageNumber="131">Leucosia urania</emphasis>
</taxonomicName>
) are misidentifications of 
<taxonomicName id="4C4353ECCA49FFE06748B4B6FDEDF9A0" authority="(Miers, 1877) (Galil 2006: 64)" authorityName="(Miers, 1877) (Galil" authorityPageNumber="64" authorityYear="2006" class="Malacostraca" family="Leucosiidae" genus="Soceulia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="B937F47DCA49FFE06748B4B6FF4DF9A3" italics="true" pageId="7" pageNumber="131">Soceulia brunnea</emphasis>
(Miers, 1877) (
<bibRefCitation id="EFD2559ECA49FFE063D0B4D2FDF2F9A0" author="Galil" box="[420,587,1989,2014]" pageId="7" pageNumber="131" refString="Galil, B. S. (2006) Contributions to the knowledge of Leucosiidae V. Coleusia gen. nov. (Crustacea: Brachyura). Zoologische Mededelingen, 80, 55 - 69." type="journal article" year="2006">Galil 2006: 64</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF3C78E7CA48FFE162E3B43BFBD6F9A5" httpUri="https://zenodo.org/record/230319/files/figure.png" pageId="6" pageNumber="130" targetBox="[151,1436,193,1814]" targetPageId="6">
<paragraph id="8BFC286FCA48FFE162E3B43BFBD6F9A5" blockId="6.[151,1436,1836,2011]" pageId="6" pageNumber="130">
<emphasis id="B937F47DCA48FFE162E3B43BFEB4F93F" bold="true" box="[151,269,1836,1858]" pageId="6" pageNumber="130">FIGURE 3</emphasis>
. A–G, erroneously labeled male “syntype” of 
<taxonomicName id="4C4353ECCA48FFE16092B43AFBB0F93C" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[742,1033,1836,1858]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA48FFE16092B43AFCC2F93C" box="[742,891,1837,1858]" italics="true" pageId="6" pageNumber="130">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
, (21.0 × 16.5 mm) (ZMB Herbst 2184) (= here identified as 
<taxonomicName id="4C4353ECCA48FFE16305B45CFCABF91F" authority="Shen &amp; Chen, 1978" authorityName="Shen &amp; Chen" authorityYear="1978" box="[369,786,1867,1889]" class="Malacostraca" family="Leucosiidae" genus="Seulocia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B937F47DCA48FFE16305B45CFD99F91F" box="[369,544,1867,1889]" italics="true" pageId="6" pageNumber="130">Seulocia pulchra</emphasis>
(Shen &amp; Chen, 1978))
</taxonomicName>
; H, 
<taxonomicName id="4C4353ECCA48FFE16134B45CFBDEF91F" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[832,1127,1867,1889]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA48FFE16134B45CFC6FF91F" box="[832,982,1867,1889]" italics="true" pageId="6" pageNumber="130">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
, label. A, box in ZMB where supposed type male of 
<taxonomicName id="4C4353ECCA48FFE163F6B47DFD1AF901" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[386,675,1897,1919]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA48FFE163F6B47DFDAEF901" box="[386,535,1898,1919]" italics="true" pageId="6" pageNumber="130">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
, is kept, with detached ambulatory legs that do not belong to specimen; B, dorsal view; C, ventral view of cephalothorax; D, dorsal view of front; E, lateral view of carapace showing thoracic sinus; F, lateral view of carapace showing thoracic sinus and outer view of merus of cheliped; G, posterior view of carapace; H, original label in box of lectotype female of 
<taxonomicName id="4C4353ECCA48FFE1638EB4D1FCA3F9A5" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[506,794,1989,2011]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="130" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA48FFE1638EB4D1FD36F9A5" box="[506,655,1990,2011]" italics="true" pageId="6" pageNumber="130">Cancer urania</emphasis>
Herbst, 1801
</taxonomicName>
. All photographs by O. Coleman.
</paragraph>
</caption>
<caption id="DF3C78E7CA49FFE062E3B55EFD35F8E2" httpUri="https://zenodo.org/record/230320/files/figure.png" pageId="7" pageNumber="131" targetBox="[159,1418,176,1584]" targetPageId="7">
<paragraph id="8BFC286FCA49FFE062E3B55EFD35F8E2" blockId="7.[151,1436,1609,1692]" pageId="7" pageNumber="131">
<emphasis id="B937F47DCA49FFE062E3B55EFEB4F820" bold="true" box="[151,269,1609,1631]" pageId="7" pageNumber="131">FIGURE 4</emphasis>
. 
<taxonomicName id="4C4353ECCA49FFE0636DB55EFDE2F821" authority="Herbst, 1801" authorityName="Herbst" authorityYear="1801" box="[281,603,1609,1631]" class="Malacostraca" family="Leucosiidae" genus="Coleusia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="131" phylum="Arthropoda" rank="species" species="urania">
<emphasis id="B937F47DCA49FFE0636DB55EFE07F821" box="[281,446,1609,1631]" italics="true" pageId="7" pageNumber="131">Coleusia urania</emphasis>
(Herbst, 1801)
</taxonomicName>
, male (40.3 × 36.5 mm) (ZRC 2001.0045). A, overall dorsal view; B, lateral view of carapace showing thoracic sinus; C, outer view of merus of cheliped; D, view of lower margin of palm; E, left fourth ambulatory leg; F, ventral view of cephalothorax.
</paragraph>
</caption>
<subSubSection id="C3597BE4CA46FFEF62B3B380FD83FECE" box="[199,570,151,176]" pageId="8" pageNumber="132" type="distribution">
<paragraph id="8BFC286FCA46FFEF62B3B380FD83FECE" blockId="8.[199,570,151,176]" box="[199,570,151,176]" pageId="8" pageNumber="132">
<emphasis id="B937F47DCA46FFEF62B3B380FEE0FECE" bold="true" box="[199,345,151,176]" pageId="8" pageNumber="132">Distribution</emphasis>
. 
<collectingCountry id="F35468FFCA46FFEF6313B380FE13FECE" box="[359,426,151,176]" name="China" pageId="8" pageNumber="132">China</collectingCountry>
and 
<collectingCountry id="F35468FFCA46FFEF6394B380FD8FFECE" box="[480,566,151,176]" name="Taiwan" pageId="8" pageNumber="132">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>