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<document id="991B050D798653FEC7601744F50C292D" ID-DOI="10.11646/zootaxa.3946.3.2" ID-GBIF-Dataset="d57d5862-5fc5-480c-84c9-3b598b597197" ID-ISSN="1175-5326" ID-Zenodo-Dep="244741" ID-ZooBank="24E2F41D-89BF-473F-ACE0-ED951BCB2699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034391373" checkinUser="plazi" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2015" docId="03DF87F3FFCF6E77FF4FFABAD15E2F20" docLanguage="en" docName="zt03946p346.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terrathelphusa kundong Grinang &amp; Ng, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="340" masterDocId="FFE6FF8BFFC86E7DFFD8FF92D3052E7E" masterDocTitle="The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia" masterLastPageNumber="346" masterPageNumber="331" pageNumber="338" updateTime="1698574262036" updateUser="plazi">
<mods:mods id="310602F8EA2FE8A5EA98281A12DB2863" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1DD1495146A4C498391065DE7B7798B6">
<mods:title id="464DFFD765B60A662A1AEAC3ADC49A77">The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="A320AF12B764C54EDC2FEC31325F232C" type="personal">
<mods:role id="EA015794D97F0540C159D0D13617D73D">
<mods:roleTerm id="E513F4761842B1FC0609F6AFEFF0AB2E">Author</mods:roleTerm>
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<mods:namePart id="0AD7D649DE712C59E35E06A953D9A090">Grinang, Jongkar</mods:namePart>
</mods:name>
<mods:name id="4E4605C44D9E00CD653A58AEDC6DA2E5" type="personal">
<mods:role id="AA1AA7E8D4A162B82B9B1A966749011E">
<mods:roleTerm id="9D502F6A9F72E41E5611A18D38ED406F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA6E018F28E8E1C617528C593E240D9D">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EE1916E00C0E67E5663622F88A68E20E">text</mods:typeOfResource>
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<mods:title id="222EA405E49157C1908B89B2E8FD0D34">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="32EA000608B0599B59C787665C3B22B7">
<mods:date id="72A7F66313EB92505BFBB55DC0F87810">2015</mods:date>
<mods:detail id="A0DB7E7EB4F7DD33688E95D4BE099D50" type="volume">
<mods:number id="65F2DB4EC88612F6A27952AD0FB0DF98">3946</mods:number>
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<mods:detail id="A2BCA6615B98D7FC0F04882F3C25F9D8" type="issue">
<mods:number id="EB798F687D33985321CD9EEB07663C7B">3</mods:number>
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<mods:start id="AAFB46B4E73AE1D734B2CBD531D97D0B">331</mods:start>
<mods:end id="CF2AE7445FEECE9514084626F54E4AE7">346</mods:end>
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<mods:classification id="817B86A464004B5D311A0FA6425CD146">journal article</mods:classification>
<mods:identifier id="723F09C242C26BA120EDF6A65885A1DA" type="DOI">10.11646/zootaxa.3946.3.2</mods:identifier>
<mods:identifier id="4F9C668E5739CFD13B3D1DD2CDB74A23" type="GBIF-Dataset">d57d5862-5fc5-480c-84c9-3b598b597197</mods:identifier>
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<mods:identifier id="5E771789C6BD3BD9224C796004E22520" type="Zenodo-Dep">244741</mods:identifier>
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<treatment id="03DF87F3FFCF6E77FF4FFABAD15E2F20" ID-DOI="http://doi.org/10.5281/zenodo.5695391" ID-GBIF-Taxon="119608776" ID-Zenodo-Dep="5695391" LSID="urn:lsid:plazi:treatment:03DF87F3FFCF6E77FF4FFABAD15E2F20" httpUri="http://treatment.plazi.org/id/03DF87F3FFCF6E77FF4FFABAD15E2F20" lastPageId="10" lastPageNumber="340" pageId="7" pageNumber="338">
<subSubSection id="C36C656EFFCF6E7AFF4FFABAD6992BDB" pageId="7" pageNumber="338" type="nomenclature">
<paragraph id="8BC936E5FFCF6E7AFF4FFABAD1092B3C" blockId="7.[151,524,1320,1379]" box="[151,524,1320,1346]" pageId="7" pageNumber="338">
<heading id="D0818189FFCF6E7AFF4FFABAD1092B3C" bold="true" box="[151,524,1320,1346]" fontSize="11" level="1" pageId="7" pageNumber="338" reason="1">
<emphasis id="B902EAF7FFCF6E7AFF4FFABAD1092B3C" bold="true" box="[151,524,1320,1346]" pageId="7" pageNumber="338">
<taxonomicName id="4C764D66FFCF6E7AFF4FFABAD2C62B3C" ID-CoL="55HXK" box="[151,451,1320,1346]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
<emphasis id="B902EAF7FFCF6E7AFF4FFABAD2C62B3C" bold="true" box="[151,451,1320,1346]" italics="true" pageId="7" pageNumber="338">Terrathelphusa kundong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231578CFFCF6E7AFE11FABAD1092B3C" box="[457,524,1320,1346]" pageId="7" pageNumber="338" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936E5FFCF6E7AFF4FFAD8D2D22B1D" blockId="7.[151,524,1320,1379]" box="[151,471,1354,1379]" pageId="7" pageNumber="338">
(
<figureCitation id="134D2A60FFCF6E7AFF47FAD8D3EE2B1D" box="[159,235,1354,1379]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="7" pageNumber="338">Figs. 7</figureCitation>
A–H, 8A–E, 11G, H)
</paragraph>
<paragraph id="8BC936E5FFCF6E7AFF4FFA1DD6992BDB" blockId="7.[151,1436,1423,1446]" box="[151,1436,1423,1446]" pageId="7" pageNumber="338">
<emphasis id="B902EAF7FFCF6E7AFF4FFA1DD2CA2BDB" box="[151,463,1423,1445]" italics="true" pageId="7" pageNumber="338">
<taxonomicName id="4C764D66FFCF6E7AFF4FFA1DD2342BDB" box="[151,305,1423,1445]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="338" phylum="Arthropoda" rank="genus">Terrathelphusa</taxonomicName>
kuchingensis—
</emphasis>
Ng &amp; Grinang 2004: 314 (part), fig. 7 (not
<taxonomicName id="4C764D66FFCF6E7AFC58FA03D6942BDB" authority="Nobili, 1901" authorityName="Nobili" authorityYear="1901" box="[896,1425,1423,1446]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="338" phylum="Arthropoda" rank="species" species="kuchingensis" subGenus="Geothelphusa">
<emphasis id="B902EAF7FFCF6E7AFC58FA03D0D92BD8" box="[896,988,1425,1446]" italics="true" pageId="7" pageNumber="338">Potamon</emphasis>
(
<emphasis id="B902EAF7FFCF6E7AFC32FA1DD7722BDB" box="[1002,1143,1423,1445]" italics="true" pageId="7" pageNumber="338">Geothelphusa</emphasis>
)
<emphasis id="B902EAF7FFCF6E7AFB5DFA1DD60D2BDB" box="[1157,1288,1423,1445]" italics="true" pageId="7" pageNumber="338">kuchingensis</emphasis>
Nobili, 1901
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C656EFFCF6E74FF4FFA46D1022A93" lastPageId="9" lastPageNumber="340" pageId="7" pageNumber="338" type="materials_examined">
<paragraph id="8BC936E5FFCF6E7AFF4FFA46D690299E" blockId="7.[151,1436,1492,2017]" pageId="7" pageNumber="338">
<emphasis id="B902EAF7FFCF6E7AFF4FFA46D2872B93" bold="true" box="[151,386,1492,1517]" pageId="7" pageNumber="338">Material examined.</emphasis>
<typeStatus id="54CD8847FFCF6E7AFE52FA46D2F22B93" box="[394,503,1492,1517]" pageId="7" pageNumber="338" type="holotype">Holotype</typeStatus>
: male (29.4 ×
<quantity id="4C8E9B00FFCF6E7AFD44FA47D0072B93" box="[668,770,1493,1517]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" pageId="7" pageNumber="338" unit="mm" value="21.9">21.9 mm</quantity>
) (
<collectionCode id="ED67AE20FFCF6E7AFCC2FA47D0572B93" box="[794,850,1493,1517]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="338">ZRC</collectionCode>
2015.0004), middle elevation of Gunung Serambu, Bau,
<collectingCountry id="F3617675FFCF6E7AFF0CFA68D232286D" box="[212,311,1530,1555]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, 1O25’50.4”N 110O13’04.3”E,
<quantity id="4C8E9B00FFCF6E7AFD54FA69D1D0286C" box="[652,725,1531,1555]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="338" unit="m" value="340.0">340 m</quantity>
asl, coll. J. Grinang
<emphasis id="B902EAF7FFCF6E7AFC1CFA6ED0F3286D" box="[964,1014,1530,1555]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFBD4FA69D7B5286D" box="[1036,1200,1530,1555]" pageId="7" pageNumber="338" value="2013-04-24">24 April 2013</date>
.
<typeStatus id="54CD8847FFCF6E7AFB64FA69D627286D" box="[1212,1314,1531,1555]" pageId="7" pageNumber="338" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D70FD6CFFCF6E7AFAF6FA69D699286D" box="[1326,1436,1530,1555]" pageId="7" pageNumber="338" type="female">2 females</specimenCount>
(larger 30.8 × 23.0 mm) (SBC.C.00364–65), data same as
<typeStatus id="54CD8847FFCF6E7AFCEDF98FD09E2848" box="[821,923,1565,1590]" pageId="7" pageNumber="338" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D70FD6CFFCF6E7AFC70F98CD70E2848" box="[936,1035,1565,1590]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(34.9 ×
<quantity id="4C8E9B00FFCF6E7AFBA8F98FD7D32848" box="[1136,1238,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="7" pageNumber="338" unit="mm" value="25.3">25.3 mm</quantity>
) (SBC.C.00366), flat bank of Sungai Bukan Kagap, base of Gunung Serambu at Podam area, Bau,
<collectingCountry id="F3617675FFCF6E7AFB41F9D0D7F92825" box="[1177,1276,1602,1627]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, 1O25’38.4”N 110O12’29.4”E,
<quantity id="4C8E9B00FFCF6E7AFE91F9FAD28428FE" box="[329,385,1640,1664]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="338" unit="m" value="34.0">34 m</quantity>
asl, coll. J. Grinang
<emphasis id="B902EAF7FFCF6E7AFDBEF9FBD19328FF" box="[614,662,1640,1665]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFD73F9FAD05D28FE" box="[683,856,1640,1665]" pageId="7" pageNumber="338" value="2013-03-13">13 March 2013</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFCBBF9FAD0BD28FF" box="[867,952,1640,1665]" pageId="7" pageNumber="338" type="male">3 males</specimenCount>
(largest 36.2 ×
<quantity id="4C8E9B00FFCF6E7AFBBEF9FAD7CB28FE" box="[1126,1230,1640,1665]" metricMagnitude="-2" metricUnit="m" metricValue="2.53" pageId="7" pageNumber="338" unit="mm" value="25.3">25.3 mm</quantity>
),
<specimenCount id="9D70FD6CFFCF6E7AFB38F9FAD64E28FF" box="[1248,1355,1640,1665]" pageId="7" pageNumber="338" type="female">2 females</specimenCount>
(larger 30.0 × 22.0 mm) (SBC.C.00367–71), foothill of Gunung Sewa, Giam area, Padawan-Penrissen limestone formation,
<collectingCountry id="F3617675FFCF6E7AFEC8F922D27428B7" box="[272,369,1712,1737]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, 1O19’01.2”N 110O16’22.2”E,
<quantity id="4C8E9B00FFCF6E7AFD64F922D1F628B6" box="[700,755,1712,1736]" metricMagnitude="1" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="338" unit="m" value="49.0">49 m</quantity>
asl, coll. J. Grinang
<emphasis id="B902EAF7FFCF6E7AFC0DF923D70128B7" box="[981,1028,1712,1737]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFBC1F922D7F728B6" box="[1049,1266,1712,1737]" pageId="7" pageNumber="338" value="2013-09-21">21 September 2013</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFB25F922D65828B7" box="[1277,1373,1712,1737]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(36.3 ×
<quantity id="4C8E9B00FFCF6E7AFF68F944D2132890" box="[176,278,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="7" pageNumber="338" unit="mm" value="24.8">24.8 mm</quantity>
) (SBC.C.0072), secondary forest, sandstone, Bung Bratak, Bau,
<collectingCountry id="F3617675FFCF6E7AFC2AF947D7562890" box="[1010,1107,1749,1774]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, 1O28’17.4”N 110O05’13.5”E,
<quantity id="4C8E9B00FFCF6E7AFF4FF96BD3D8296E" box="[151,221,1785,1809]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="338" unit="m" value="100.0">100 m</quantity>
asl, coll. S.T. Pang
<emphasis id="B902EAF7FFCF6E7AFE66F96BD2F5296F" box="[446,496,1784,1809]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFDDDF96BD1CA296F" box="[517,719,1784,1809]" pageId="7" pageNumber="338" value="2013-12-01">1 December 2013</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFD04F96BD039296F" box="[732,828,1784,1809]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(24.6 ×
<quantity id="4C8E9B00FFCF6E7AFC46F96BD707296F" box="[926,1026,1785,1809]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="7" pageNumber="338" unit="mm" value="17.7">17.7 mm</quantity>
) (SBC.C.00373), foothill of Gunung Sewa, Giam area, Padawan-Penrissen limestone formation,
<collectingCountry id="F3617675FFCF6E7AFCF5F889D08B294A" box="[813,910,1819,1844]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, coll. Anang
<emphasis id="B902EAF7FFCF6E7AFBFAF88ED756294A" box="[1058,1107,1819,1844]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFBB1F889D61B294D" box="[1129,1310,1819,1843]" pageId="7" pageNumber="338" value="2008-08-22">22 August 2008</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFAF2F889D687294A" box="[1322,1410,1819,1844]" pageId="7" pageNumber="338" type="male">2 males</specimenCount>
,
<specimenCount id="9D70FD6CFFCF6E7AFA56F889D3E72928" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(
<collectionCode id="ED67AE20FFCF6E7AFF2BF8ACD22F2928" box="[243,298,1854,1878]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="338">ZRC</collectionCode>
2014.0843), forest next to rubber and cocoa plantations, base of Gunung Sewa, Giam area, Padawan-Penrissen limestone formation,
<collectingCountry id="F3617675FFCF6E7AFD59F8F2D1E72907" box="[641,738,1888,1913]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, coll. J. Grinang
<emphasis id="B902EAF7FFCF6E7AFC15F8F3D70C2907" box="[973,1033,1888,1913]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFBF1F8F3D61C2907" box="[1065,1305,1888,1913]" pageId="7" pageNumber="338" value="2013-09-14">14 September 2013</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFAE9F8F3D6992907" box="[1329,1436,1888,1913]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(SBC.C.00014), Bau limestone formation, Gunung Tai Ton, Bau,
<collectingCountry id="F3617675FFCF6E7AFCBCF811D0C029E2" box="[868,965,1923,1948]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, coll. J. Grinang
<emphasis id="B902EAF7FFCF6E7AFBA6F816D7A829E2" box="[1150,1197,1923,1948]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFB19F811D69D29E5" box="[1217,1432,1923,1948]" pageId="7" pageNumber="338" value="2001-11-13">13 November 2001</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFF4FF834D3FA29C0" box="[151,255,1957,1982]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(SBC.C.00015), Bau limestone formation, Gunung Tai Ton, Bau,
<collectingCountry id="F3617675FFCF6E7AFBF0F837D78C29C0" box="[1064,1161,1957,1982]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, coll. A. Denis
<emphasis id="B902EAF7FFCF6E7AFA8EF835D68B29C0" box="[1366,1422,1957,1982]" italics="true" pageId="7" pageNumber="338">et al</emphasis>
.,
<date id="FFC81025FFCF6E7AFF4FF85BD271299F" box="[151,372,1992,2017]" pageId="7" pageNumber="338" value="2001-11-13">13 November 2001</date>
;
<specimenCount id="9D70FD6CFFCF6E7AFE58F85BD2E6299F" box="[384,483,1992,2017]" pageId="7" pageNumber="338" type="female">1 female</specimenCount>
(SBC.C.00049), Bau limestone formation, Gunung Tai Ton, Bau,
<collectingCountry id="F3617675FFCF6E7AFB01F85AD63B299F" box="[1241,1342,1992,2017]" name="Malaysia" pageId="7" pageNumber="338">Sarawak</collectingCountry>
, coll. J.
</paragraph>
<caption id="DF09666DFFC06E75FF4FF92FD1F3294E" httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="8" pageNumber="339" targetBox="[206,1384,197,1688]" targetPageId="8">
<paragraph id="8BC936E5FFC06E75FF4FF92FD1F3294E" blockId="8.[151,1436,1725,1840]" pageId="8" pageNumber="339">
<emphasis id="B902EAF7FFC06E75FF4FF92FD20B28AD" bold="true" box="[151,270,1725,1747]" pageId="8" pageNumber="339">FIGURE 7</emphasis>
.
<taxonomicName id="4C764D66FFC06E75FEC4F92FD11028AD" box="[284,533,1725,1747]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="339" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
<emphasis id="B902EAF7FFC06E75FEC4F92FD11028AD" box="[284,533,1725,1747]" italics="true" pageId="8" pageNumber="339">Terrathelphusa kundong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231578CFFC06E75FDC5F92FD15D28AD" box="[541,600,1725,1747]" pageId="8" pageNumber="339" rank="species">
<emphasis id="B902EAF7FFC06E75FDC5F92FD15D28AD" bold="true" box="[541,600,1725,1747]" pageId="8" pageNumber="339">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela.
</paragraph>
</caption>
<caption id="DF09666DFFC16E74FF4FFC41D7A72A58" httpUri="https://zenodo.org/record/244749/files/figure.png" pageId="9" pageNumber="340" targetBox="[208,1370,198,959]" targetPageId="9">
<paragraph id="8BC936E5FFC16E74FF4FFC41D7A72A58" blockId="9.[151,1436,979,1062]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF4FFC41D20B2D97" bold="true" box="[151,270,979,1001]" pageId="9" pageNumber="340">FIGURE 8</emphasis>
.
<taxonomicName id="4C764D66FFC16E74FEC4FC41D1102D97" box="[284,533,979,1001]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
<emphasis id="B902EAF7FFC16E74FEC4FC41D1102D97" box="[284,533,979,1001]" italics="true" pageId="9" pageNumber="340">Terrathelphusa kundong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231578CFFC16E74FDC5FC41D15D2D97" box="[541,600,979,1001]" pageId="9" pageNumber="340" rank="species">
<emphasis id="B902EAF7FFC16E74FDC5FC41D15D2D97" bold="true" box="[541,600,979,1001]" pageId="9" pageNumber="340">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, ventral view of left G1; B, ventral view of distal part of left G1; C, dorsal view of left G2; D, dorsal view of G1; E, dorsal view of distal part of left G1. Scales: A, C, D = 1 mm; B, E = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BC936E5FFC16E74FF4FFBFED1022A93" blockId="9.[151,1437,1132,2024]" pageId="9" pageNumber="340">
Grinang
<emphasis id="B902EAF7FFC16E74FF24FBFFD22A2AFB" box="[252,303,1132,1157]" italics="true" pageId="9" pageNumber="340">et al</emphasis>
.,
<date id="FFC81025FFC16E74FE9FFBFFD2E92AFB" box="[327,492,1132,1157]" pageId="9" pageNumber="340" value="2002-04-14">14 April 2002</date>
;
<specimenCount id="9D70FD6CFFC16E74FE22FBFFD15B2AFB" box="[506,606,1132,1157]" pageId="9" pageNumber="340" type="female">1 female</specimenCount>
(21.6 ×
<quantity id="4C8E9B00FFC16E74FD1DFBFED0292AFB" box="[709,812,1132,1157]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="9" pageNumber="340" unit="mm" value="16.2">16.2 mm</quantity>
) (
<collectionCode id="ED67AE20FFC16E74FC9EFBFFD0872AFB" box="[838,898,1133,1157]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="340">ZRC</collectionCode>
), Bau limestone formation, Gunung Aup, Bau,
<collectingCountry id="F3617675FFC16E74FF4FFB1DD3FF2AD6" box="[151,250,1167,1192]" name="Malaysia" pageId="9" pageNumber="340">Sarawak</collectingCountry>
, coll. J. Grinang
<emphasis id="B902EAF7FFC16E74FE1BFB02D2F32AD6" box="[451,502,1167,1192]" italics="true" pageId="9" pageNumber="340">et al</emphasis>
.,
<date id="FFC81025FFC16E74FDD7FB1DD1AC2AD9" box="[527,681,1167,1192]" pageId="9" pageNumber="340" value="2002-05-14">14 May 2002</date>
(det. as
<taxonomicName id="4C764D66FFC16E74FCC8FB1DD7502AD6" box="[784,1109,1167,1192]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kuchingensis">
<emphasis id="B902EAF7FFC16E74FCC8FB1DD7502AD6" box="[784,1109,1167,1192]" italics="true" pageId="9" pageNumber="340">Terrathelphusa kuchingensis</emphasis>
</taxonomicName>
by Ng &amp; Grinang, 2004);
<specimenCount id="9D70FD6CFFC16E74FA56FB1DD3EB2AB4" pageId="9" pageNumber="340" type="female">2 females</specimenCount>
(larger 30.8 ×
<quantity id="4C8E9B00FFC16E74FE4FFB20D2F82AB4" box="[407,509,1202,1226]" metricMagnitude="-2" metricUnit="m" metricValue="2.21" pageId="9" pageNumber="340" unit="mm" value="22.1">22.1 mm</quantity>
) (SM Cru.1986.144–145), Bidi, Bau,
<collectingCountry id="F3617675FFC16E74FC73FB23D70B2AB4" box="[939,1038,1201,1226]" name="Malaysia" pageId="9" pageNumber="340">Sarawak</collectingCountry>
, coll. C. J. Brooke,
<date id="FFC81025FFC16E74FB37FB20D6672AB4" box="[1263,1378,1202,1226]" pageId="9" pageNumber="340" value="1903-06">June 1903</date>
(det. as
<taxonomicName id="4C764D66FFC16E74FF6DFB46D2F22A93" box="[181,503,1236,1261]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kuchingensis">
<emphasis id="B902EAF7FFC16E74FF6DFB46D2F22A93" box="[181,503,1236,1261]" italics="true" pageId="9" pageNumber="340">Terrathelphusa kuchingensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36C656EFFC16E74FF1FFB65D7C028F3" pageId="9" pageNumber="340" type="diagnosis">
<paragraph id="8BC936E5FFC16E74FF1FFB65D00D2814" blockId="9.[151,1437,1132,2024]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FFB65D2442B6E" bold="true" box="[199,321,1271,1296]" pageId="9" pageNumber="340">Diagnosis.</emphasis>
Carapace broader than long, conspicuously swollen, surface convex, smooth; cervical grooves broad, deep; epigastric cristae high, separated by deep epigastric groove; H-groove deep, long almost confluent with cervical grooves; epibranchial tooth indistinct but can be separated from external orbital angle by notch, anterolateral, frontal regions appearing compressed, supraorbital margin almost parallel with frontal margin; epistome median lobe blunt to bifurcated (
<figureCitation id="134D2A60FFC16E74FD5CFA13D1CC2BE4" box="[644,713,1409,1434]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
A–C). Male abdomen with elongated somite 6, lateral margins gently concave (
<figureCitation id="134D2A60FFC16E74FE83FA36D2A12BC3" box="[347,420,1444,1469]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
D). Third maxilliped ischium rectangular, about 1.9 times maximum width (
<figureCitation id="134D2A60FFC16E74FAEBFA36D6792BC3" box="[1331,1404,1444,1469]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
E). Ambulatory legs smooth, relatively slender, second pair longest; length of fourth merus about 3.8 times width (
<figureCitation id="134D2A60FFC16E74FAB7FA55D3A3287C" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
F). Cheliped carpus rugose, without granules (
<figureCitation id="134D2A60FFC16E74FD74FA7BD1E8287C" box="[684,749,1513,1538]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
G, H), inner angle with broadly triangular tooth; gap between fingers of major chela wide (
<figureCitation id="134D2A60FFC16E74FE03F99ED11B285B" box="[475,542,1548,1573]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="9" pageNumber="340">Fig. 7</figureCitation>
A). G1 gently curving outwards; terminal segment cone-shaped, tapered, gently curving upwards, about 0.3 times length of subterminal segment (
<figureCitation id="134D2A60FFC16E74FCA9F9BDD0B72839" box="[881,946,1583,1608]" captionStart="FIGURE 8" captionStartId="9.[151,250,979,1001]" captionTargetBox="[208,1370,198,959]" captionTargetId="figure@9.[204,1378,193,964]" captionTargetPageId="9" captionText="FIGURE 8. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244749/files/figure.png" pageId="9" pageNumber="340">Fig. 8</figureCitation>
A, B, D, E). Distal segment of G2 relatively long, about 0.6 times length of basal segment (
<figureCitation id="134D2A60FFC16E74FD7AF9C3D1E02814" box="[674,741,1617,1642]" captionStart="FIGURE 8" captionStartId="9.[151,250,979,1001]" captionTargetBox="[208,1370,198,959]" captionTargetId="figure@9.[204,1378,193,964]" captionTargetPageId="9" captionText="FIGURE 8. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244749/files/figure.png" pageId="9" pageNumber="340">Fig. 8</figureCitation>
C).
</paragraph>
<paragraph id="8BC936E5FFC16E74FF1FF9E6D7C028F3" blockId="9.[151,1437,1132,2024]" box="[199,1221,1652,1677]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FF9E6D24A28F3" bold="true" box="[199,335,1652,1677]" pageId="9" pageNumber="340">Life colour.</emphasis>
Carapace is grape-purple with whitish cream legs and chelipeds (
<figureCitation id="134D2A60FFC16E74FBF7F9E6D77928F3" box="[1071,1148,1652,1677]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="9" pageNumber="340">Fig. 11</figureCitation>
G–H).
</paragraph>
</subSubSection>
<subSubSection id="C36C656EFFC16E74FF1FF905D7492923" pageId="9" pageNumber="340" type="description">
<paragraph id="8BC936E5FFC16E74FF1FF905D7E6288B" blockId="9.[151,1437,1132,2024]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FF905D23E28CE" bold="true" box="[199,315,1687,1712]" pageId="9" pageNumber="340">Variation.</emphasis>
The
<typeStatus id="54CD8847FFC16E74FEA3F90AD2E328CE" box="[379,486,1688,1712]" pageId="9" pageNumber="340" type="paratype">paratypes</typeStatus>
agree well with the
<typeStatus id="54CD8847FFC16E74FD0AF905D03128CE" box="[722,820,1687,1712]" pageId="9" pageNumber="340" type="holotype">holotype</typeStatus>
male except in some non-sexual characters: the tip of the median lobe of the epistome varies from blunt to bifurcated that was not determined by size or sex. The cervical grooves are proportionately broader and the ambulatory legs relatively longer in larger specimens.
</paragraph>
<paragraph id="8BC936E5FFC16E74FF1FF96DD7492923" blockId="9.[151,1437,1132,2024]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FF96DD22F2966" bold="true" box="[199,298,1791,1816]" pageId="9" pageNumber="340">Habitat.</emphasis>
Widely distributed in limestone and sandstone outcrops, from lowlands to mid- elevations of mountains. The burrows are shallower in sandstone possibly because of the compact soil, whereas burrows are deeper (approximately more than
<quantity id="4C8E9B00FFC16E74FDCAF8D7D13B2923" box="[530,574,1861,1885]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="340" unit="m" value="1.0">1 m</quantity>
) in the loose soil at the base of limestone hills.
</paragraph>
</subSubSection>
<subSubSection id="C36C656EFFC16E74FF1FF8F5D21629BB" pageId="9" pageNumber="340" type="etymology">
<paragraph id="8BC936E5FFC16E74FF1FF8F5D21629BB" blockId="9.[151,1437,1132,2024]" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FF8F5D24B29FE" bold="true" box="[199,334,1895,1920]" pageId="9" pageNumber="340">Etymology.</emphasis>
The name is derived from the Iban word
<taxonomicName id="4C764D66FFC16E74FCEBF8F5D09129FE" box="[819,916,1895,1920]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kundong">
<emphasis id="B902EAF7FFC16E74FCEBF8F5D09129FE" box="[819,916,1895,1920]" italics="true" pageId="9" pageNumber="340">kundong</emphasis>
</taxonomicName>
alluding to the dark crimson carapace that is very similar to the ripe fruit of a plant of the same name (
<taxonomicName id="4C764D66FFC16E74FC98F818D71929DC" box="[832,1052,1929,1954]" class="Magnoliopsida" family="Clusiaceae" genus="Garcinia" kingdom="Plantae" order="Malpighiales" pageId="9" pageNumber="340" phylum="Tracheophyta" rank="species" species="parvifolia">
<emphasis id="B902EAF7FFC16E74FC98F818D71929DC" box="[832,1052,1929,1954]" italics="true" pageId="9" pageNumber="340">Garcinia parvifolia</emphasis>
</taxonomicName>
). The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C36C656EFFC16E77FF1FF85DD15E2F20" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340" type="discussion">
<paragraph id="8BC936E5FFC16E77FF1FF85DD15E2F20" blockId="9.[151,1437,1132,2024]" lastBlockId="10.[151,1436,151,350]" lastPageId="10" lastPageNumber="341" pageId="9" pageNumber="340">
<emphasis id="B902EAF7FFC16E74FF1FF85DD23E2996" bold="true" box="[199,315,1999,2024]" pageId="9" pageNumber="340">Remarks.</emphasis>
<taxonomicName id="4C764D66FFC16E74FE99F85DD1572996" box="[321,594,1999,2024]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
<emphasis id="B902EAF7FFC16E74FE99F85DD1572996" box="[321,594,1999,2024]" italics="true" pageId="9" pageNumber="340">Terrathelphusa kundong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231578CFFC16E74FD81F85DD1922996" box="[601,663,1999,2024]" pageId="9" pageNumber="340" rank="species">
<emphasis id="B902EAF7FFC16E74FD81F85DD1922996" bold="true" box="[601,663,1999,2024]" pageId="9" pageNumber="340">n. sp.</emphasis>
</taxonomicNameLabel>
, can be distinguished from
<taxonomicName id="4C764D66FFC16E74FC14F842D77D2996" box="[972,1144,1999,2024]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="340" phylum="Arthropoda" rank="species" species="kuchingensis">
<emphasis id="B902EAF7FFC16E74FC14F842D77D2996" box="[972,1144,1999,2024]" italics="true" pageId="9" pageNumber="340">T. kuchingensis</emphasis>
</taxonomicName>
and the other new species being described herein by a combination of characters: inner angle of carpus with broadly triangular tooth (
<figureCitation id="134D2A60FFC26E77FAE5FF05D67B2ECE" box="[1341,1406,151,176]" captionStart="FIGURE 7" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[206,1384,197,1688]" captionTargetId="figure@8.[206,1384,197,1689]" captionTargetPageId="8" captionText="FIGURE 7. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244748/files/figure.png" pageId="10" pageNumber="341">Fig. 7</figureCitation>
A) (acutely triangular tooth in other species;
<figureCitation id="134D2A60FFC26E77FDAEFF28D1C52EAD" box="[630,704,186,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="10" pageNumber="341">Figs. 1</figureCitation>
A, 3A, 5A, 9A) and the G1 terminal segment distinctly curving upwards (
<figureCitation id="134D2A60FFC26E77FEDDFF4FD2422E88" box="[261,327,221,246]" captionStart="FIGURE 8" captionStartId="9.[151,250,979,1001]" captionTargetBox="[208,1370,198,959]" captionTargetId="figure@9.[204,1378,193,964]" captionTargetPageId="9" captionText="FIGURE 8. Terrathelphusa kundong n. sp., holotype male (29.4 × 21.9 mm) (ZRC 2015.0004), sandstone outcrop, Gunung Serambu, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244749/files/figure.png" pageId="10" pageNumber="341">Fig. 8</figureCitation>
A, B, C, D) (gently curving upwards in other species;
<figureCitation id="134D2A60FFC26E77FC71FF4FD0F02E88" box="[937,1013,221,246]" captionStart="FIGURE 2" captionStartId="3.[151,250,1052,1074]" captionTargetBox="[159,1427,198,1024]" captionTargetId="figure@3.[153,1434,193,1030]" captionTargetPageId="3" captionText="FIGURE 2. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244743/files/figure.png" pageId="10" pageNumber="341">Figs. 2</figureCitation>
A, B, D, E; 4A, B, D, E; 6A, B, D, E; 8A, B, D, E; 10A, B, D, E). Examination of the specimens previously identified as
<taxonomicName id="4C764D66FFC26E77FB99FE93D7E82F66" box="[1089,1261,255,280]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="kuchingensis">
<emphasis id="B902EAF7FFC26E77FB99FE93D7E82F66" box="[1089,1261,255,280]" italics="true" pageId="10" pageNumber="341">T. kuchingensis</emphasis>
</taxonomicName>
in the
<collectingCountry id="F3617675FFC26E77FAE4FF6DD6992F66" box="[1340,1436,255,280]" name="Malaysia" pageId="10" pageNumber="341">Sarawak</collectingCountry>
Museum and
<collectingCountry id="F3617675FFC26E77FEE1FEB0D29F2F45" box="[313,410,290,315]" name="Malaysia" pageId="10" pageNumber="341">Sarawak</collectingCountry>
Biodiversity Centre from areas adjacent to the
<typeStatus id="54CD8847FFC26E77FC0DFEB1D7002F45" box="[981,1029,291,315]" pageId="10" pageNumber="341">type</typeStatus>
locality show that they should be transferred to
<taxonomicName id="4C764D66FFC26E77FEECFED4D2AA2F20" box="[308,431,325,350]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="341" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
<emphasis id="B902EAF7FFC26E77FEECFED4D2AA2F20" box="[308,431,325,350]" italics="true" pageId="10" pageNumber="341">T. kundong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A231578CFFC26E77FE6FFED7D2F02F20" box="[439,501,325,350]" pageId="10" pageNumber="341" rank="species">
<emphasis id="B902EAF7FFC26E77FE6FFED7D2F02F20" bold="true" box="[439,501,325,350]" pageId="10" pageNumber="341">n. sp.</emphasis>
</taxonomicNameLabel>
, instead.
</paragraph>
</subSubSection>
</treatment>
</document>
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