203 lines
30 KiB
XML
203 lines
30 KiB
XML
<document id="0D65A28B708463322D3CFCE9265AC0F2" ID-DOI="10.11646/zootaxa.3946.3.2" ID-GBIF-Dataset="d57d5862-5fc5-480c-84c9-3b598b597197" ID-ISSN="1175-5326" ID-Zenodo-Dep="244741" ID-ZooBank="24E2F41D-89BF-473F-ACE0-ED951BCB2699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034391373" checkinUser="plazi" docAuthor="Grinang, Jongkar & Ng, Peter K. L." docDate="2015" docId="03DF87F3FFCD6E7BFF4FFBA9D7B72F3E" docLanguage="en" docName="zt03946p346.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terrathelphusa cerina Grinang & Ng, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="336" masterDocId="FFE6FF8BFFC86E7DFFD8FF92D3052E7E" masterDocTitle="The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia" masterLastPageNumber="346" masterPageNumber="331" pageNumber="336" updateTime="1698574262036" updateUser="plazi">
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<mods:title id="BCF5AEF1E452C32D741D2D61DE251F05">The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia</mods:title>
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<mods:roleTerm id="717405570ACEF3D342463814BA357699">Author</mods:roleTerm>
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<mods:namePart id="8DF3593D8E69395CAE852ABF3D9EFEE0">Grinang, Jongkar</mods:namePart>
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<mods:roleTerm id="66F7475AA569C5228B82CB9944BB9085">Author</mods:roleTerm>
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<mods:namePart id="24597A8871B98B3080746A7056B9B8DB">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="B2C8C79FD1A0E39035DB6C57732E9919">text</mods:typeOfResource>
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<mods:title id="3487DD4EB454D4C5C3DB04BF5EC9B73E">Zootaxa</mods:title>
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<mods:part id="91AD2DC0AB9D05ABA47DD9D08EA51879">
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<mods:date id="2F6B7AFB394C82C309EA85B2D13D54B0">2015</mods:date>
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<mods:number id="AD6CED4D8EA88094EB6840DD1761F717">3946</mods:number>
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<mods:detail id="A152BE768F975C2D2E296FB274BCF34A" type="issue">
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<mods:identifier id="FB0054B496B63F5094E46FA064F6558E" type="DOI">10.11646/zootaxa.3946.3.2</mods:identifier>
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<treatment id="03DF87F3FFCD6E7BFF4FFBA9D7B72F3E" ID-DOI="http://doi.org/10.5281/zenodo.5695389" ID-GBIF-Taxon="119608780" ID-Zenodo-Dep="5695389" LSID="urn:lsid:plazi:treatment:03DF87F3FFCD6E7BFF4FFBA9D7B72F3E" httpUri="http://treatment.plazi.org/id/03DF87F3FFCD6E7BFF4FFBA9D7B72F3E" lastPageId="6" lastPageNumber="336" pageId="5" pageNumber="336">
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<subSubSection id="C36C656EFFCD6E78FF4FFBA9D2D62A08" pageId="5" pageNumber="336" type="nomenclature">
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<paragraph id="8BC936E5FFCD6E78FF4FFBA9D2E82A2B" blockId="5.[151,493,1083,1142]" box="[151,493,1083,1109]" pageId="5" pageNumber="336">
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<heading id="D0818189FFCD6E78FF4FFBA9D2E82A2B" bold="true" box="[151,493,1083,1109]" fontSize="11" level="1" pageId="5" pageNumber="336" reason="1">
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<emphasis id="B902EAF7FFCD6E78FF4FFBA9D2E82A2B" bold="true" box="[151,493,1083,1109]" pageId="5" pageNumber="336">
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<taxonomicName id="4C764D66FFCD6E78FF4FFBA9D2A12A2B" ID-CoL="55HXF" box="[151,420,1083,1109]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
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<emphasis id="B902EAF7FFCD6E78FF4FFBA9D2A12A2B" bold="true" box="[151,420,1083,1109]" italics="true" pageId="5" pageNumber="336">Terrathelphusa cerina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A231578CFFCD6E78FE73FBA9D2E82A2B" box="[427,493,1083,1109]" pageId="5" pageNumber="336" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936E5FFCD6E78FF4FFBCFD2D62A08" blockId="5.[151,493,1083,1142]" box="[151,467,1117,1142]" pageId="5" pageNumber="336">
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(
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<figureCitation id="134D2A60FFCD6E78FF47FBCFD3EE2A08" box="[159,235,1117,1142]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Figs. 5</figureCitation>
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A–H, 6A–E, 11E, F)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36C656EFFCD6E78FF4FFB37D6822B77" pageId="5" pageNumber="336" type="materials_examined">
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<paragraph id="8BC936E5FFCD6E78FF4FFB37D6822B77" blockId="5.[151,1437,1189,2009]" pageId="5" pageNumber="336">
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<emphasis id="B902EAF7FFCD6E78FF4FFB37D2812AC0" bold="true" box="[151,388,1189,1214]" pageId="5" pageNumber="336">Material examined.</emphasis>
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<typeStatus id="54CD8847FFCD6E78FE55FB37D2FF2AC0" box="[397,506,1189,1214]" pageId="5" pageNumber="336" type="holotype">Holotype</typeStatus>
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: male (24.7 ×
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<quantity id="4C8E9B00FFCD6E78FD7FFB34D0142AC0" box="[679,785,1190,1214]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="5" pageNumber="336" unit="mm" value="19.3">19.3 mm</quantity>
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) (
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<collectionCode id="ED67AE20FFCD6E78FCF2FB34D0672AC0" box="[810,866,1190,1214]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="336">ZRC</collectionCode>
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2015.0003), damp riverbank of Sungai Sebayak, Gunung Singai, Bau,
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<collectingCountry id="F3617675FFCD6E78FE4EFB5ED2FC2A9B" box="[406,505,1228,1253]" name="Malaysia" pageId="5" pageNumber="336">Sarawak</collectingCountry>
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, 1O30’17.1”N 110O10’40.1”E,
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<quantity id="4C8E9B00FFCD6E78FC8FFB5ED0902A9A" box="[855,917,1228,1253]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="5" pageNumber="336" unit="m" value="66.0">66 m</quantity>
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asl, coll. J. Grinang
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<emphasis id="B902EAF7FFCD6E78FB4AFB5CD7CD2A9B" box="[1170,1224,1228,1253]" italics="true" pageId="5" pageNumber="336">et al</emphasis>
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.,
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<date id="FFC81025FFCD6E78FB39FB5FD69C2A9B" box="[1249,1433,1228,1253]" pageId="5" pageNumber="336" value="2012-03-31">31 March 2012</date>
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.
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<typeStatus id="54CD8847FFCD6E78FF4FFB63D3FB2B77" box="[151,254,1265,1289]" pageId="5" pageNumber="336" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D70FD6CFFCD6E78FED0FB63D2562B77" box="[264,339,1264,1289]" pageId="5" pageNumber="336" type="male">1 male</specimenCount>
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(30.0 ×
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<quantity id="4C8E9B00FFCD6E78FE6AFB63D11F2B77" box="[434,538,1265,1289]" metricMagnitude="-2" metricUnit="m" metricValue="2.28" pageId="5" pageNumber="336" unit="mm" value="22.8">22.8 mm</quantity>
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),
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<specimenCount id="9D70FD6CFFCD6E78FDF5FB63D19C2B77" box="[557,665,1264,1289]" pageId="5" pageNumber="336" type="female">2 females</specimenCount>
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(larger 31.0 ×
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||
<quantity id="4C8E9B00FFCD6E78FC98FB62D0A02B77" box="[832,933,1264,1289]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="5" pageNumber="336" unit="mm" value="26.2">26.2 mm</quantity>
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) (
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<collectionCode id="ED67AE20FFCD6E78FC64FB63D0F12B77" box="[956,1012,1265,1289]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="336">ZRC</collectionCode>
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2014.0842), data same as
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<typeStatus id="54CD8847FFCD6E78FAC6FB62D6812B77" box="[1310,1412,1264,1289]" pageId="5" pageNumber="336" type="holotype">holotype</typeStatus>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36C656EFFCD6E78FF1FFA86D0A828C7" pageId="5" pageNumber="336" type="diagnosis">
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<paragraph id="8BC936E5FFCD6E78FF1FFA86D3CD28EB" blockId="5.[151,1437,1189,2009]" pageId="5" pageNumber="336">
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||
<emphasis id="B902EAF7FFCD6E78FF1FFA86D2442B53" bold="true" box="[199,321,1300,1325]" pageId="5" pageNumber="336">Diagnosis.</emphasis>
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Carapace broader than long, swollen, surface convex, very smooth; cervical grooves broad, deep; epigastric cristae high separated by deep epigastric groove; H-groove deep, long almost confluent with cervical grooves; epibranchial tooth indiscernible, anterolateral, frontal regions appearing compressed, supraorbital margin almost parallel with frontal margin; epistome median lobe blunt to pointed (
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<figureCitation id="134D2A60FFCD6E78FBC3FA12D7642BE7" box="[1051,1121,1408,1433]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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A–C). Male abdomen with elongated somite 6, lateral margins gently concave (
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<figureCitation id="134D2A60FFCD6E78FD36FA36D0362BC3" box="[750,819,1444,1469]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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D). Ischium of third maxilliped rectangular, about 2.0 times maximum width (
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<figureCitation id="134D2A60FFCD6E78FE7CFA5AD2E22B9F" box="[420,487,1480,1505]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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E). Ambulatory legs smooth, relatively slender, second pair longest, length of fourth merus about 3.9 times width (
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<figureCitation id="134D2A60FFCD6E78FE29FA7ED133287B" box="[497,566,1516,1541]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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F). Cheliped carpus rugose, without granules (
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<figureCitation id="134D2A60FFCD6E78FB94FA7ED795287B" box="[1100,1168,1516,1541]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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||
G, H), inner angle with acutely triangular tooth; gap between fingers of major chela relatively narrow (
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<figureCitation id="134D2A60FFCD6E78FB93F982D7962857" box="[1099,1171,1552,1577]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
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A). G1 gently curving outwards; terminal segment cone-shaped, tapered, gently curving upwards, about 0.3 times length of subterminal segment (
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<figureCitation id="134D2A60FFCD6E78FEDEF9CAD24F280F" box="[262,330,1624,1649]" captionStart="FIGURE 6" captionStartId="7.[151,250,1156,1178]" captionTargetBox="[156,1428,203,1126]" captionTargetId="figure@7.[150,1436,193,1134]" captionTargetPageId="7" captionText="FIGURE 6. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244747/files/figure.png" pageId="5" pageNumber="336">Fig. 6</figureCitation>
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||
A, B, D, E). Distal segment of G2 relatively long, about 0.7 times length of basal segment (
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||
<figureCitation id="134D2A60FFCD6E78FAB7F9CAD3A328EB" captionStart="FIGURE 6" captionStartId="7.[151,250,1156,1178]" captionTargetBox="[156,1428,203,1126]" captionTargetId="figure@7.[150,1436,193,1134]" captionTargetPageId="7" captionText="FIGURE 6. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244747/files/figure.png" pageId="5" pageNumber="336">Fig. 6</figureCitation>
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C).
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</paragraph>
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||
<paragraph id="8BC936E5FFCD6E78FF1FF932D0A828C7" blockId="5.[151,1437,1189,2009]" box="[199,941,1696,1721]" pageId="5" pageNumber="336">
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||
<emphasis id="B902EAF7FFCD6E78FF1FF932D24A28C7" bold="true" box="[199,335,1696,1721]" pageId="5" pageNumber="336">Life colour.</emphasis>
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Individuals are bright yellowish overall (
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||
<figureCitation id="134D2A60FFCD6E78FCC7F932D06B28C7" box="[799,878,1696,1721]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="5" pageNumber="336">Fig. 11</figureCitation>
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E, F).
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCD6E78FF1FF956D1532937" pageId="5" pageNumber="336" type="description">
|
||
<paragraph id="8BC936E5FFCD6E78FF1FF956D646297F" blockId="5.[151,1437,1189,2009]" pageId="5" pageNumber="336">
|
||
<emphasis id="B902EAF7FFCD6E78FF1FF956D23E28A3" bold="true" box="[199,315,1732,1757]" pageId="5" pageNumber="336">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="54CD8847FFCD6E78FEA5F957D2EC28A3" box="[381,489,1733,1757]" pageId="5" pageNumber="336" type="paratype">paratypes</typeStatus>
|
||
agree very well with the
|
||
<typeStatus id="54CD8847FFCD6E78FCCFF956D07C28A3" box="[791,889,1732,1757]" pageId="5" pageNumber="336" type="holotype">holotype</typeStatus>
|
||
male except in few non-sexual characters. The cervical grooves are proportionately broader and the ambulatory legs relatively longer in larger specimens.
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFCD6E78FF1FF89ED1532937" blockId="5.[151,1437,1189,2009]" pageId="5" pageNumber="336">
|
||
<emphasis id="B902EAF7FFCD6E78FF1FF89ED22F295B" bold="true" box="[199,298,1804,1829]" pageId="5" pageNumber="336">Habitat.</emphasis>
|
||
Swampy floodplains in slightly disturbed lowland primary forest. Lives in shallow burrows less than
|
||
<quantity id="4C8E9B00FFCD6E78FF4FF8A3D3C72936" box="[151,194,1841,1865]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="336" unit="m" value="1.0">1 m</quantity>
|
||
depth, with dense debris in the soil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCD6E78FF1FF8C6D0DA29EF" pageId="5" pageNumber="336" type="etymology">
|
||
<paragraph id="8BC936E5FFCD6E78FF1FF8C6D0DA29EF" blockId="5.[151,1437,1189,2009]" pageId="5" pageNumber="336">
|
||
<emphasis id="B902EAF7FFCD6E78FF1FF8C6D24B2913" bold="true" box="[199,334,1876,1901]" pageId="5" pageNumber="336">Etymology.</emphasis>
|
||
The name is derived from the Greek
|
||
<emphasis id="B902EAF7FFCD6E78FCF5F8C6D0852913" box="[813,896,1876,1901]" italics="true" pageId="5" pageNumber="336">kerinos</emphasis>
|
||
for “yellowish wax colour”, alluding to the appearance of the species in life. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCD6E7BFF1FF80ED7B72F3E" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336" type="discussion">
|
||
<paragraph id="8BC936E5FFCD6E7BFF1FF80ED7B72F3E" blockId="5.[151,1437,1189,2009]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="337" pageId="5" pageNumber="336">
|
||
<emphasis id="B902EAF7FFCD6E78FF1FF80ED23E29CB" bold="true" box="[199,315,1948,1973]" pageId="5" pageNumber="336">Remarks.</emphasis>
|
||
<taxonomicName id="4C764D66FFCD6E78FE9AF80ED13929CB" box="[322,572,1948,1973]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCD6E78FE9AF80ED13929CB" box="[322,572,1948,1973]" italics="true" pageId="5" pageNumber="336">Terrathelphusa cerina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCD6E78FD9BF80ED18729CB" box="[579,642,1948,1973]" pageId="5" pageNumber="336" rank="species">
|
||
<emphasis id="B902EAF7FFCD6E78FD9BF80ED18729CB" bold="true" box="[579,642,1948,1973]" pageId="5" pageNumber="336">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is distinguished from
|
||
<taxonomicName id="4C764D66FFCD6E78FC5EF80CD73629CB" box="[902,1075,1948,1973]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="336" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCD6E78FC5EF80CD73629CB" box="[902,1075,1948,1973]" italics="true" pageId="5" pageNumber="336">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
and the other three new species by a combination of characters. The carapace is relatively less swollen and convex (
|
||
<figureCitation id="134D2A60FFCD6E78FB9FF852D78F29A7" box="[1095,1162,1984,2009]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="5" pageNumber="336">Fig. 5</figureCitation>
|
||
B) than in its congeners, and the epibranchial tooth is indiscernible (
|
||
<figureCitation id="134D2A60FFCE6E7BFDA7FF05D1C52ECE" box="[639,704,151,176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1723,1745]" captionTargetBox="[341,1248,375,1692]" captionTargetId="figure@6.[341,1248,375,1692]" captionTargetPageId="6" captionText="FIGURE 5. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244746/files/figure.png" pageId="6" pageNumber="337">Fig. 5</figureCitation>
|
||
A–C). In comparison to its closest congener,
|
||
<taxonomicName id="4C764D66FFCE6E7BFB66FF0BD6692ECE" box="[1214,1388,151,176]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCE6E7BFB66FF0BD6692ECE" box="[1214,1388,151,176]" italics="true" pageId="6" pageNumber="337">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
, the G1 subterminal segment of
|
||
<taxonomicName id="4C764D66FFCE6E7BFE04FF2FD1452EAA" box="[476,576,189,212]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCE6E7BFE04FF2FD1452EAA" box="[476,576,189,212]" italics="true" pageId="6" pageNumber="337">T. cerina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCE6E7BFD94FF2ED18B2EAB" box="[588,654,188,213]" pageId="6" pageNumber="337" rank="species">
|
||
<emphasis id="B902EAF7FFCE6E7BFD94FF2ED18B2EAB" bold="true" box="[588,654,188,213]" pageId="6" pageNumber="337">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinctly curved outwards, with the tip of the terminal segment relatively blunter (
|
||
<figureCitation id="134D2A60FFCE6E7BFEB0FF4DD2A92E86" box="[360,428,223,248]" captionStart="FIGURE 6" captionStartId="7.[151,250,1156,1178]" captionTargetBox="[156,1428,203,1126]" captionTargetId="figure@7.[150,1436,193,1134]" captionTargetPageId="7" captionText="FIGURE 6. Terrathelphusa cerina n. sp., holotype male (24.7 × 19.3 mm) (ZRC 2015.0003), sandstone outcrop, Gunung Singai, Bau, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244747/files/figure.png" pageId="6" pageNumber="337">Fig. 6</figureCitation>
|
||
A, B, D, E) (subterminal segment is less prominently curved outwards, with the terminal segment more pointed in
|
||
<taxonomicName id="4C764D66FFCE6E7BFE6DFE97D1642F63" box="[437,609,260,285]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCE6E7BFE6DFE97D1642F63" box="[437,609,260,285]" italics="true" pageId="6" pageNumber="337">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="134D2A60FFCE6E7BFDB7FE96D1B42F62" box="[623,689,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1052,1074]" captionTargetBox="[159,1427,198,1024]" captionTargetId="figure@3.[153,1434,193,1030]" captionTargetPageId="3" captionText="FIGURE 2. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244743/files/figure.png" pageId="6" pageNumber="337">Fig. 2</figureCitation>
|
||
A, B, D, E). Live specimens of
|
||
<taxonomicName id="4C764D66FFCE6E7BFBCDFE97D77C2F62" box="[1045,1145,261,284]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="cerina" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCE6E7BFBCDFE97D77C2F62" box="[1045,1145,261,284]" italics="true" pageId="6" pageNumber="337">T. cerina</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCE6E7BFB58FE96D7BA2F63" box="[1152,1215,260,285]" pageId="6" pageNumber="337" rank="species">
|
||
<emphasis id="B902EAF7FFCE6E7BFB58FE96D7BA2F63" bold="true" box="[1152,1215,260,285]" pageId="6" pageNumber="337">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are more similar to
|
||
<taxonomicName id="4C764D66FFCE6E7BFF4FFEBBD3DA2F3E" box="[151,223,297,320]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="337" phylum="Arthropoda" rank="species" species="mas" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCE6E7BFF4FFEBBD3DA2F3E" box="[151,223,297,320]" italics="true" pageId="6" pageNumber="337">T. mas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCE6E7BFF3EFEB5D2202F3E" box="[230,293,295,320]" pageId="6" pageNumber="337" rank="species">
|
||
<emphasis id="B902EAF7FFCE6E7BFF3EFEB5D2202F3E" bold="true" box="[230,293,295,320]" pageId="6" pageNumber="337">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
but the latter species is of an even more striking yellow colour (
|
||
<figureCitation id="134D2A60FFCE6E7BFC2DFEB5D7412F3E" box="[1013,1092,295,320]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="6" pageNumber="337">Fig. 11</figureCitation>
|
||
E, F, I, J).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |