230 lines
32 KiB
XML
230 lines
32 KiB
XML
<document id="F6802B020EE82B72B21D0FA4AF8AA92D" ID-DOI="10.11646/zootaxa.3946.3.2" ID-GBIF-Dataset="d57d5862-5fc5-480c-84c9-3b598b597197" ID-ISSN="1175-5326" ID-Zenodo-Dep="244741" ID-ZooBank="24E2F41D-89BF-473F-ACE0-ED951BCB2699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034391373" checkinUser="plazi" docAuthor="Grinang, Jongkar & Ng, Peter K. L." docDate="2015" docId="03DF87F3FFCB6E79FF4FFB28D20B2C0D" docLanguage="en" docName="zt03946p346.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terrathelphusa aglaia Grinang & Ng, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="335" masterDocId="FFE6FF8BFFC86E7DFFD8FF92D3052E7E" masterDocTitle="The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia" masterLastPageNumber="346" masterPageNumber="331" pageNumber="334" updateTime="1698574262036" updateUser="plazi">
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<mods:titleInfo id="A089FA25E9BB6489EA71902E2C1AEE13">
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<mods:title id="6DC277AF12222F31660EA78AB2B18274">The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="7D48BEDE6D3813D9B53AC7E870C752AF" type="personal">
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<mods:roleTerm id="55480148E73AD9F15054E9E987B032EF">Author</mods:roleTerm>
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<mods:namePart id="011D76C1DA8AF6DD214AAE254972F64E">Grinang, Jongkar</mods:namePart>
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</mods:name>
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<mods:name id="A6925A10428CEE22C370322E6F20E996" type="personal">
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<mods:roleTerm id="1AF66D96681F0195AFA8E112CA0BA515">Author</mods:roleTerm>
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<mods:namePart id="857ADF41D283F92A12D159A98C62289B">Ng, Peter K. L.</mods:namePart>
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<mods:typeOfResource id="634C1AE4904C52E65F3E44EBEDE07130">text</mods:typeOfResource>
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<mods:title id="44114545AC5615FBAD87DA34DAE18C64">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A45FB16B7717BDFC87520F606AB27018">
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<mods:date id="FE68379D02F484F92E07330CE1779D7E">2015</mods:date>
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<mods:detail id="0064CF55740463D0D6CD890005FB7A7B" type="volume">
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<mods:number id="AB6951E8F565754FD3CC7E8AB550FD34">3946</mods:number>
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</mods:detail>
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<mods:detail id="4011499A733EEA6C4891E3D5E7BBD94A" type="issue">
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<mods:number id="18C6A8E9DFC468EF3805EF63C6E7DA18">3</mods:number>
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<mods:classification id="443EB2A8D8947AE2F287F7C3497956A1">journal article</mods:classification>
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<mods:identifier id="224C806D2C5EF245DC2C0C0CC66CEC58" type="DOI">10.11646/zootaxa.3946.3.2</mods:identifier>
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<mods:identifier id="26D1A605B5387E0A987EBFA8511561C1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B262FF0AAAFAFC4DB92BB81EC9907A14" type="Zenodo-Dep">244741</mods:identifier>
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</mods:mods>
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<treatment id="03DF87F3FFCB6E79FF4FFB28D20B2C0D" ID-DOI="http://doi.org/10.5281/zenodo.5695387" ID-GBIF-Taxon="119608781" ID-Zenodo-Dep="5695387" LSID="urn:lsid:plazi:treatment:03DF87F3FFCB6E79FF4FFB28D20B2C0D" httpUri="http://treatment.plazi.org/id/03DF87F3FFCB6E79FF4FFB28D20B2C0D" lastPageId="4" lastPageNumber="335" pageId="3" pageNumber="334">
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<subSubSection id="C36C656EFFCB6E7EFF4FFB28D2DC2A8B" pageId="3" pageNumber="334" type="nomenclature">
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<paragraph id="8BC936E5FFCB6E7EFF4FFB28D2EE2AAA" blockId="3.[151,491,1210,1269]" box="[151,491,1210,1236]" pageId="3" pageNumber="334">
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<heading id="D0818189FFCB6E7EFF4FFB28D2EE2AAA" bold="true" box="[151,491,1210,1236]" fontSize="11" level="1" pageId="3" pageNumber="334" reason="1">
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<emphasis id="B902EAF7FFCB6E7EFF4FFB28D2EE2AAA" bold="true" box="[151,491,1210,1236]" pageId="3" pageNumber="334">
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<taxonomicName id="4C764D66FFCB6E7EFF4FFB28D2A72AAA" ID-CoL="7C3MD" box="[151,418,1210,1236]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
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<emphasis id="B902EAF7FFCB6E7EFF4FFB28D2A72AAA" bold="true" box="[151,418,1210,1236]" italics="true" pageId="3" pageNumber="334">Terrathelphusa aglaia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A231578CFFCB6E7EFE71FB28D2EE2AAA" box="[425,491,1210,1236]" pageId="3" pageNumber="334" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936E5FFCB6E7EFF4FFB4ED2DC2A8B" blockId="3.[151,491,1210,1269]" box="[151,473,1244,1269]" pageId="3" pageNumber="334">
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(
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<figureCitation id="134D2A60FFCB6E7EFF47FB4ED3EF2A8B" box="[159,234,1244,1269]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Figs. 3</figureCitation>
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A–H, 4A–E, 11C, D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C656EFFCB6E7EFF4FFAB0D25A286F" pageId="3" pageNumber="334" type="materials_examined">
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<paragraph id="8BC936E5FFCB6E7EFF4FFAB0D25A286F" blockId="3.[151,1437,1314,2038]" pageId="3" pageNumber="334">
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<emphasis id="B902EAF7FFCB6E7EFF4FFAB0D2852B45" bold="true" box="[151,384,1314,1339]" pageId="3" pageNumber="334">Material examined</emphasis>
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.
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<typeStatus id="54CD8847FFCB6E7EFE4CFAB0D1052B45" box="[404,512,1314,1339]" pageId="3" pageNumber="334" type="holotype">Holotype</typeStatus>
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: male (35.9 ×
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<quantity id="4C8E9B00FFCB6E7EFD64FAB0D02D2B45" box="[700,808,1314,1339]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="3" pageNumber="334" unit="mm" value="26.6">26.6 mm</quantity>
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) (
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<collectionCode id="ED67AE20FFCB6E7EFC9DFAB1D0782B45" box="[837,893,1315,1339]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="334">ZRC</collectionCode>
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2015.0002), open green golf course, Borneo Highlands Resort, Gunung Penrissen, Kuching,
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<collectingCountry id="F3617675FFCB6E7EFD6FFAD5D01F2B1E" box="[695,794,1351,1376]" name="Malaysia" pageId="3" pageNumber="334">Sarawak</collectingCountry>
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||
, 1O07’43.8”N 110O13’07.3”E,
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<quantity id="4C8E9B00FFCB6E7EFBB7FAD5D7B22B21" box="[1135,1207,1351,1376]" metricMagnitude="2" metricUnit="m" metricValue="8.52" pageId="3" pageNumber="334" unit="m" value="852.0">852 m</quantity>
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asl, coll. J. Grinang
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<emphasis id="B902EAF7FFCB6E7EFF4FFAFED3CD2BFD" box="[151,200,1386,1411]" italics="true" pageId="3" pageNumber="334">et al</emphasis>
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.,
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<date id="FFC81025FFCB6E7EFF04FAF8D2BC2BFD" box="[220,441,1386,1411]" pageId="3" pageNumber="334" value="2013-09-16">16 September 2013</date>
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||
.
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||
<typeStatus id="54CD8847FFCB6E7EFE1BFAF9D12C2BFD" box="[451,553,1387,1411]" pageId="3" pageNumber="334" type="paratype">Paratype</typeStatus>
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:
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||
<specimenCount id="9D70FD6CFFCB6E7EFDEBFAF9D1782BFD" box="[563,637,1386,1411]" pageId="3" pageNumber="334" type="male">1 male</specimenCount>
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||
(32.3 ×
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||
<quantity id="4C8E9B00FFCB6E7EFD04FAF9D0462BFD" box="[732,835,1387,1411]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="3" pageNumber="334" unit="mm" value="23.8">23.8 mm</quantity>
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),
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<specimenCount id="9D70FD6CFFCB6E7EFC8EFAF9D0B32BFD" box="[854,950,1386,1411]" pageId="3" pageNumber="334" type="female">1 female</specimenCount>
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||
(37.3 ×
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||
<quantity id="4C8E9B00FFCB6E7EFBCDFAF8D77F2BFD" box="[1045,1146,1386,1411]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="334" unit="mm" value="27.5">27.5 mm</quantity>
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||
) (SBC.C.00361–62), data same as
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<typeStatus id="54CD8847FFCB6E7EFF2DFA1FD25E2BD8" box="[245,347,1421,1446]" pageId="3" pageNumber="334" type="holotype">holotype</typeStatus>
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||
;
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||
<specimenCount id="9D70FD6CFFCB6E7EFEBDFA1CD2D12BD8" box="[357,468,1421,1446]" pageId="3" pageNumber="334" type="juvenile">1 juvenile</specimenCount>
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||
(17.9 ×
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||
<quantity id="4C8E9B00FFCB6E7EFDEAFA1CD1932BD8" box="[562,662,1422,1446]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="334" unit="mm" value="14.2">14.2 mm</quantity>
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||
) (SBC.C.00363), steep soil bank of Sungai Biduan, based of Gunung Penrissen,
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||
<collectingCountry id="F3617675FFCB6E7EFEC8FA20D2762BB5" box="[272,371,1458,1483]" name="Malaysia" pageId="3" pageNumber="334">Sarawak</collectingCountry>
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||
, 1O08’48.6”N 110O14’23.9”E,
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||
<quantity id="4C8E9B00FFCB6E7EFD1AFA21D00C2BB4" box="[706,777,1459,1483]" metricMagnitude="2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="334" unit="m" value="172.0">172 m</quantity>
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||
asl, coll. J. Grinang
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<emphasis id="B902EAF7FFCB6E7EFC36FA26D71B2BB5" box="[1006,1054,1458,1483]" italics="true" pageId="3" pageNumber="334">et al</emphasis>
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||
.,
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||
<date id="FFC81025FFCB6E7EFBEBFA21D7BD2BB5" box="[1075,1208,1458,1483]" pageId="3" pageNumber="334" value="2013-07-08">8 July 2013</date>
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||
;
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||
<specimenCount id="9D70FD6CFFCB6E7EFB1BFA21D6082BB5" box="[1219,1293,1458,1483]" pageId="3" pageNumber="334" type="male">1 male</specimenCount>
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||
(27.5 ×
|
||
<quantity id="4C8E9B00FFCB6E7EFAB3FA21D3C22B90" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="3" pageNumber="334" unit="mm" value="21.1">21.1 mm</quantity>
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||
),
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||
<specimenCount id="9D70FD6CFFCB6E7EFF04FA44D24F2B90" box="[220,330,1493,1518]" pageId="3" pageNumber="334" type="female">3 females</specimenCount>
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||
(largest 35.5 ×
|
||
<quantity id="4C8E9B00FFCB6E7EFDDBFA47D16E2B90" box="[515,619,1493,1518]" metricMagnitude="-2" metricUnit="m" metricValue="2.62" pageId="3" pageNumber="334" unit="mm" value="26.2">26.2 mm</quantity>
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||
) (
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||
<collectionCode id="ED67AE20FFCB6E7EFD5BFA44D1BE2B90" box="[643,699,1494,1518]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="334">ZRC</collectionCode>
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2014.0844), coll. J. Grinang
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<emphasis id="B902EAF7FFCB6E7EFBD7FA44D7472B90" box="[1039,1090,1493,1518]" italics="true" pageId="3" pageNumber="334">et al</emphasis>
|
||
.,
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||
<date id="FFC81025FFCB6E7EFB82FA47D63E2B90" box="[1114,1339,1493,1518]" pageId="3" pageNumber="334" value="2013-09-15">15 September 2013</date>
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, locality same as
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<typeStatus id="54CD8847FFCB6E7EFF2EFA6AD259286F" box="[246,348,1528,1553]" pageId="3" pageNumber="334" type="holotype">holotype</typeStatus>
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||
.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36C656EFFCB6E7EFF1FF988D29329AD" pageId="3" pageNumber="334" type="diagnosis">
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||
<paragraph id="8BC936E5FFCB6E7EFF1FF988D07D29F0" blockId="3.[151,1437,1314,2038]" pageId="3" pageNumber="334">
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<emphasis id="B902EAF7FFCB6E7EFF1FF988D244284D" bold="true" box="[199,321,1562,1587]" pageId="3" pageNumber="334">Diagnosis.</emphasis>
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Carapace broader than long, conspicuously swollen, surface convex, smooth; cervical grooves broad, deep; epigastric cristae high, separated by deep epigastric groove; H-groove deep, long almost confluent with cervical grooves; epibranchial tooth indistinct but can be separated from external orbital angle by notch; anterolateral, frontal regions appearing compressed; supraorbital margin almost parallel with frontal margin; epistome median lobe blunt to pointed (
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<figureCitation id="134D2A60FFCB6E7EFD8BF937D19028C0" box="[595,661,1701,1726]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
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A–C). Male abdomen with elongated somite 6, lateral margins gently concave (
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<figureCitation id="134D2A60FFCB6E7EFEDAF95AD241289E" box="[258,324,1736,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
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D). Ischium of third maxilliped rectangular, about 1.9 times maximum width (
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<figureCitation id="134D2A60FFCB6E7EFB73F95AD7E8289E" box="[1195,1261,1736,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
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E). Ambulatory legs smooth, relatively slender, second pair longest; length of fourth merus about 3.9 times width (
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<figureCitation id="134D2A60FFCB6E7EFAEDF978D679297D" box="[1333,1404,1770,1795]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
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F). Cheliped carpus rugose, without granules (
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<figureCitation id="134D2A60FFCB6E7EFD52F89FD1CB2958" box="[650,718,1805,1830]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
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G, H), inner angle with acutely triangular tooth; gap between fingers of major chela wide (
|
||
<figureCitation id="134D2A60FFCB6E7EFE2EF8A2D13A2936" box="[502,575,1840,1865]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="3" pageNumber="334">Fig. 3</figureCitation>
|
||
A). G1 gently curving outwards; terminal segment cone-shaped, tapered, slightly curving upwards, about 0.3 times length of subterminal segment (
|
||
<figureCitation id="134D2A60FFCB6E7EFC00F8C0D7192915" box="[984,1052,1874,1899]" captionStart="FIGURE 4" captionStartId="5.[151,250,919,941]" captionTargetBox="[158,1430,198,893]" captionTargetId="figure@5.[151,1436,193,898]" captionTargetPageId="5" captionText="FIGURE 4. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244745/files/figure.png" pageId="3" pageNumber="334">Fig. 4</figureCitation>
|
||
A, B, D, E). Distal segment of G2 relatively long, about 0.8 times length of basal segment (
|
||
<figureCitation id="134D2A60FFCB6E7EFCCBF8E7D05229F0" box="[787,855,1909,1934]" captionStart="FIGURE 4" captionStartId="5.[151,250,919,941]" captionTargetBox="[158,1430,198,893]" captionTargetId="figure@5.[151,1436,193,898]" captionTargetPageId="5" captionText="FIGURE 4. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244745/files/figure.png" pageId="3" pageNumber="334">Fig. 4</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFCB6E7EFF1FF80AD29329AD" blockId="3.[151,1437,1314,2038]" pageId="3" pageNumber="334">
|
||
<emphasis id="B902EAF7FFCB6E7EFF1FF80AD24A29CF" bold="true" box="[199,335,1944,1969]" pageId="3" pageNumber="334">Life colour.</emphasis>
|
||
The carapace is dark crimson, chelipeds and legs reddish to purplish (
|
||
<figureCitation id="134D2A60FFCB6E7EFB84F80AD7AE29CE" box="[1116,1195,1944,1969]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="3" pageNumber="334">Fig. 11</figureCitation>
|
||
C, D). The juvenile is purple-reddish overall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCB6E79FF1FF84FD24F2F45" lastPageId="4" lastPageNumber="335" pageId="3" pageNumber="334" type="description">
|
||
<paragraph id="8BC936E5FFCB6E79FF1FF84FD23A2E88" blockId="3.[151,1437,1314,2038]" lastBlockId="4.[151,1437,151,627]" lastPageId="4" lastPageNumber="335" pageId="3" pageNumber="334">
|
||
<emphasis id="B902EAF7FFCB6E7EFF1FF84FD23E2988" bold="true" box="[199,315,2013,2038]" pageId="3" pageNumber="334">Variation.</emphasis>
|
||
The additional specimens agree very well with the
|
||
<typeStatus id="54CD8847FFCB6E7EFC47F84FD7042988" box="[927,1025,2013,2038]" pageId="3" pageNumber="334" type="holotype">holotype</typeStatus>
|
||
male, differing only in minor nonsexual characters: the epistome median lobe varies from blunt to pointed, which is not associated with size or sex; and in larger specimens, the cervical grooves are proportionately broader, with the ambulatory legs relatively slightly longer.
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFCC6E79FF1FFF6DD24F2F45" blockId="4.[151,1437,151,627]" pageId="4" pageNumber="335">
|
||
<emphasis id="B902EAF7FFCC6E79FF1FFF6DD22F2F66" bold="true" box="[199,298,255,280]" pageId="4" pageNumber="335">Habitat.</emphasis>
|
||
Bases of sandstone outcrops from low to high elevations. Their burrows in moist compact soil are about
|
||
<quantity id="4C8E9B00FFCC6E79FF04FEB1D2022F44" box="[220,263,291,315]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="335" unit="m" value="1.0">1 m</quantity>
|
||
deep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCC6E79FF1FFED7D7322FFE" pageId="4" pageNumber="335" type="etymology">
|
||
<paragraph id="8BC936E5FFCC6E79FF1FFED7D7322FFE" blockId="4.[151,1437,151,627]" pageId="4" pageNumber="335">
|
||
<emphasis id="B902EAF7FFCC6E79FF1FFED7D24B2F20" bold="true" box="[199,334,325,350]" pageId="4" pageNumber="335">Etymology.</emphasis>
|
||
The name is derived from the Latin
|
||
<taxonomicName id="4C764D66FFCC6E79FD32FED7D0372F20" box="[746,818,325,350]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="aglaia">
|
||
<emphasis id="B902EAF7FFCC6E79FD32FED7D0372F20" box="[746,818,325,350]" italics="true" pageId="4" pageNumber="335">aglaia</emphasis>
|
||
</taxonomicName>
|
||
for beauty or splendor alluding to the striking purplish carapace and reddish legs of the species. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFCC6E79FF1FFE18D20B2C0D" pageId="4" pageNumber="335" type="discussion">
|
||
<paragraph id="8BC936E5FFCC6E79FF1FFE18D20B2C0D" blockId="4.[151,1437,151,627]" pageId="4" pageNumber="335">
|
||
<emphasis id="B902EAF7FFCC6E79FF1FFE18D23E2FDD" bold="true" box="[199,315,394,419]" pageId="4" pageNumber="335">Remarks.</emphasis>
|
||
<taxonomicName id="4C764D66FFCC6E79FE9CFE18D1452FDD" box="[324,576,394,419]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCC6E79FE9CFE18D1452FDD" box="[324,576,394,419]" italics="true" pageId="4" pageNumber="335">Terrathelphusa aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCC6E79FD91FE18D18F2FDD" box="[585,650,394,419]" pageId="4" pageNumber="335" rank="species">
|
||
<emphasis id="B902EAF7FFCC6E79FD91FE18D18F2FDD" bold="true" box="[585,650,394,419]" pageId="4" pageNumber="335">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is easily distinguishable from its
|
||
<collectingCountry id="F3617675FFCC6E79FBC0FE18D77D2FDD" box="[1048,1144,394,419]" name="Malaysia" pageId="4" pageNumber="335">Sarawak</collectingCountry>
|
||
congeners by its striking colour: dark crimson carapace, reddish to purplish chelipeds and legs (
|
||
<figureCitation id="134D2A60FFCC6E79FC1CFE3FD7112FB8" box="[964,1044,429,454]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="4" pageNumber="335">Fig. 11</figureCitation>
|
||
C–D). Other than in coloration,
|
||
<taxonomicName id="4C764D66FFCC6E79FA5FFE3CD3DA2F96" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCC6E79FA5FFE3CD3DA2F96" italics="true" pageId="4" pageNumber="335">T. aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCC6E79FF3FFE5DD22D2F96" box="[231,296,463,488]" pageId="4" pageNumber="335" rank="species">
|
||
<emphasis id="B902EAF7FFCC6E79FF3FFE5DD22D2F96" bold="true" box="[231,296,463,488]" pageId="4" pageNumber="335">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is relatively close to
|
||
<taxonomicName id="4C764D66FFCC6E79FDF8FE43D1CB2F96" box="[544,718,463,488]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCC6E79FDF8FE43D1CB2F96" box="[544,718,463,488]" italics="true" pageId="4" pageNumber="335">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
but has a distinctly less swollen and convex carapace than the latter (
|
||
<figureCitation id="134D2A60FFCC6E79FF07FE60D21B2C75" box="[223,286,498,523]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="4" pageNumber="335">Fig. 1</figureCitation>
|
||
A vs.
|
||
<figureCitation id="134D2A60FFCC6E79FEBBFE60D2A12C75" box="[355,420,498,523]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="4" pageNumber="335">Fig. 3</figureCitation>
|
||
A). The G1 subterminal segment of
|
||
<taxonomicName id="4C764D66FFCC6E79FCEFFE66D09F2C75" box="[823,922,498,523]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCC6E79FCEFFE66D09F2C75" box="[823,922,498,523]" italics="true" pageId="4" pageNumber="335">T. aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCC6E79FC79FE60D0DA2C75" box="[929,991,498,523]" pageId="4" pageNumber="335" rank="species">
|
||
<emphasis id="B902EAF7FFCC6E79FC79FE60D0DA2C75" bold="true" box="[929,991,498,523]" pageId="4" pageNumber="335">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is also relatively more slender, with the terminal segment straight (
|
||
<figureCitation id="134D2A60FFCC6E79FE13FD87D1152C50" box="[459,528,533,558]" captionStart="FIGURE 4" captionStartId="5.[151,250,919,941]" captionTargetBox="[158,1430,198,893]" captionTargetId="figure@5.[151,1436,193,898]" captionTargetPageId="5" captionText="FIGURE 4. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244745/files/figure.png" pageId="4" pageNumber="335">Fig. 4</figureCitation>
|
||
A, B, D, E) (subterminal segment relatively broader with the terminal segment curving in
|
||
<taxonomicName id="4C764D66FFCC6E79FEC9FDABD2B82C2E" box="[273,445,567,592]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCC6E79FEC9FDABD2B82C2E" box="[273,445,567,592]" italics="true" pageId="4" pageNumber="335">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="134D2A60FFCC6E79FE14FDA5D1082C2E" box="[460,525,567,592]" captionStart="FIGURE 2" captionStartId="3.[151,250,1052,1074]" captionTargetBox="[159,1427,198,1024]" captionTargetId="figure@3.[153,1434,193,1030]" captionTargetPageId="3" captionText="FIGURE 2. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244743/files/figure.png" pageId="4" pageNumber="335">Fig. 2</figureCitation>
|
||
A, B, D, E). Differences between
|
||
<taxonomicName id="4C764D66FFCC6E79FC52FDABD0EB2C2E" box="[906,1006,567,592]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="335" phylum="Arthropoda" rank="species" species="aglaia" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCC6E79FC52FDABD0EB2C2E" box="[906,1006,567,592]" italics="true" pageId="4" pageNumber="335">T. aglaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCC6E79FC2DFDA5D7312C2E" box="[1013,1076,567,592]" pageId="4" pageNumber="335" rank="species">
|
||
<emphasis id="B902EAF7FFCC6E79FC2DFDA5D7312C2E" bold="true" box="[1013,1076,567,592]" pageId="4" pageNumber="335">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and congeners are summarised in
|
||
<tableCitation id="C6F4035EFFCC6E79FF6CFDC8D2022C0D" box="[180,263,602,627]" captionStart="TABLE 1" captionStartId="13.[125,210,534,556]" captionTargetBox="[125,1879,736,1664]" captionTargetPageId="13" captionText="TABLE 1. Differences between eight Terrathelphusa species from Borneo [* from Ng (1997)]" httpUri="http://table.plazi.org/id/DF09666DFFC56E70FFA5FD84D7322C52" pageId="4" pageNumber="335" tableUuid="DF09666DFFC56E70FFA5FD84D7322C52">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |