242 lines
31 KiB
XML
242 lines
31 KiB
XML
<document id="7E9EDE79A855B23D0682316829165F62" ID-DOI="10.11646/zootaxa.3946.3.2" ID-GBIF-Dataset="d57d5862-5fc5-480c-84c9-3b598b597197" ID-ISSN="1175-5326" ID-Zenodo-Dep="244741" ID-ZooBank="24E2F41D-89BF-473F-ACE0-ED951BCB2699" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461034391373" checkinUser="plazi" docAuthor="Grinang, Jongkar & Ng, Peter K. L." docDate="2015" docId="03DF87F3FFC96E7FFF4FFC12D1792C66" docLanguage="en" docName="zt03946p346.pdf" docOrigin="Zootaxa 3946 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Terrathelphusa kuchingensis Nobili 1901, s. str." docType="treatment" docVersion="9" lastPageNumber="333" masterDocId="FFE6FF8BFFC86E7DFFD8FF92D3052E7E" masterDocTitle="The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia" masterLastPageNumber="346" masterPageNumber="331" pageNumber="332" updateTime="1698574262036" updateUser="plazi">
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<mods:title id="173E9677CFA4A7BDBD29BC20EAB463B2">The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia</mods:title>
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<mods:namePart id="B280C53C91A7C501CAAB75881B028416">Grinang, Jongkar</mods:namePart>
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<mods:namePart id="FB2BD86DFEE41EF07B0EB7FC74590B51">Ng, Peter K. L.</mods:namePart>
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<mods:title id="7753B5BC3DD821360225D371E82CBD2A">Zootaxa</mods:title>
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<mods:part id="36A5AC9F13512A8BF2F6B1D2BB362356">
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<mods:date id="3A804F79568F98F6D84D0C9CD5B3F948">2015</mods:date>
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<mods:number id="BD18CB9E18DC7958DA05263E1DA59BAE">3946</mods:number>
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<treatment id="03DF87F3FFC96E7FFF4FFC12D1792C66" ID-DOI="http://doi.org/10.5281/zenodo.5695385" ID-GBIF-Taxon="119608782" ID-Zenodo-Dep="5695385" LSID="urn:lsid:plazi:treatment:03DF87F3FFC96E7FFF4FFC12D1792C66" httpUri="http://treatment.plazi.org/id/03DF87F3FFC96E7FFF4FFC12D1792C66" lastPageId="2" lastPageNumber="333" pageId="1" pageNumber="332">
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<subSubSection id="C36C656EFFC96E7CFF4FFC12D2DF2DC5" pageId="1" pageNumber="332" type="nomenclature">
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<paragraph id="8BC936E5FFC96E7CFF4FFC12D1EA2DE4" blockId="1.[151,751,896,955]" box="[151,751,896,922]" pageId="1" pageNumber="332">
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<heading id="D0818189FFC96E7CFF4FFC12D1EA2DE4" bold="true" box="[151,751,896,922]" fontSize="11" level="1" pageId="1" pageNumber="332" reason="1">
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<emphasis id="B902EAF7FFC96E7CFF4FFC12D1EA2DE4" bold="true" box="[151,751,896,922]" pageId="1" pageNumber="332">
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<taxonomicName id="4C764D66FFC96E7CFF4FFC12D1A32DE4" ID-CoL="55HXH" authority="Nobili, 1901" authorityName="Nobili" authorityYear="1901" box="[151,678,896,922]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" sensu="stricto" species="kuchingensis">
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<emphasis id="B902EAF7FFC96E7CFF4FFC12D2F32DE4" bold="true" box="[151,502,896,922]" italics="true" pageId="1" pageNumber="332">Terrathelphusa kuchingensis</emphasis>
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(Nobili, 1901)
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</taxonomicName>
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<taxonomicNameLabel id="A231578CFFC96E7CFD76FC12D1EA2DE4" box="[686,751,896,922]" pageId="1" pageNumber="332" sensu="stricto">s. str.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936E5FFC96E7CFF4FFC30D2DF2DC5" blockId="1.[151,751,896,955]" box="[151,474,930,955]" pageId="1" pageNumber="332">
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(
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<figureCitation id="134D2A60FFC96E7CFF47FC30D3E22DC5" box="[159,231,930,955]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Figs. 1</figureCitation>
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A–H, 2A–E, 11A, B)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36C656EFFC96E7CFF4FFC79D6412A05" pageId="1" pageNumber="332" type="reference_group">
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||
<paragraph id="8BC936E5FFC96E7CFF4FFC79D0932A7E" blockId="1.[151,1436,1002,1147]" box="[151,918,1002,1024]" pageId="1" pageNumber="332">
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<taxonomicName id="4C764D66FFC96E7CFF4FFC79D1C72A7E" authority="Nobili, 1901: 5" authorityName="Nobili" authorityPageNumber="5" authorityYear="1901" box="[151,706,1002,1024]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="kuchingensis" subGenus="Geothelphusa">
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<emphasis id="B902EAF7FFC96E7CFF4FFC79D3F62A7E" box="[151,243,1003,1024]" italics="true" pageId="1" pageNumber="332">Potamon</emphasis>
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(
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<emphasis id="B902EAF7FFC96E7CFED9FC78D2882A7E" box="[257,397,1002,1024]" italics="true" pageId="1" pageNumber="332">Geothelphusa</emphasis>
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)
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<emphasis id="B902EAF7FFC96E7CFE44FC78D11A2A7E" box="[412,543,1002,1024]" italics="true" pageId="1" pageNumber="332">kuchingensis</emphasis>
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Nobili, 1901: 5
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</taxonomicName>
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; Rathbun 1905: 217.
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</paragraph>
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<paragraph id="8BC936E5FFC96E7CFF4FFB9BD0152A61" blockId="1.[151,1436,1002,1147]" box="[151,784,1033,1055]" pageId="1" pageNumber="332">
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<taxonomicName id="4C764D66FFC96E7CFF4FFB9BD2C22A61" ID-CoL="7PSC5" authority="(Liothelphusa)" baseAuthorityName="Liothelphusa" box="[151,455,1033,1055]" class="Malacostraca" family="Gecarcinucidae" genus="Parathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">
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<emphasis id="B902EAF7FFC96E7CFF4FFB9BD2292A61" box="[151,300,1033,1055]" italics="true" pageId="1" pageNumber="332">Parathelphusa</emphasis>
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(
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<emphasis id="B902EAF7FFC96E7CFEE3FB9BD2BA2A61" box="[315,447,1033,1055]" italics="true" pageId="1" pageNumber="332">Liothelphusa</emphasis>
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)
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</taxonomicName>
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<emphasis id="B902EAF7FFC96E7CFE15FB9BD1632A61" box="[461,614,1033,1055]" italics="true" pageId="1" pageNumber="332">kuchingensis—</emphasis>
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Colosi 1920: 27.
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</paragraph>
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<paragraph id="8BC936E5FFC96E7CFF4FFBBAD7EB2A40" blockId="1.[151,1436,1002,1147]" box="[151,1262,1064,1086]" pageId="1" pageNumber="332">
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<emphasis id="B902EAF7FFC96E7CFF4FFBBAD2F22A40" box="[151,503,1064,1086]" italics="true" pageId="1" pageNumber="332">
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<taxonomicName id="4C764D66FFC96E7CFF4FFBBAD2812A40" box="[151,388,1064,1086]" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="melanippe">Thelphusula melanippe</taxonomicName>
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buergeri—
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</emphasis>
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Bott 1970: 61 (part) (not
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<taxonomicName id="4C764D66FFC96E7CFD2EFBBBD7E72A40" authority="De Man, 1899" authorityName="De Man" authorityYear="1899" box="[758,1250,1064,1086]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="burgeri" subGenus="Geothelphusa">
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<emphasis id="B902EAF7FFC96E7CFD2EFBBBD0572A40" box="[758,850,1065,1086]" italics="true" pageId="1" pageNumber="332">Potamon</emphasis>
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(
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<emphasis id="B902EAF7FFC96E7CFCB8FBBAD0E82A40" box="[864,1005,1064,1086]" italics="true" pageId="1" pageNumber="332">Geothelphusa</emphasis>
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)
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<emphasis id="B902EAF7FFC96E7CFC24FBBAD7422A40" box="[1020,1095,1064,1086]" italics="true" pageId="1" pageNumber="332">bürgeri</emphasis>
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De Man, 1899
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</taxonomicName>
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).
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</paragraph>
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<paragraph id="8BC936E5FFC96E7CFF4FFBD4D6412A05" blockId="1.[151,1436,1002,1147]" pageId="1" pageNumber="332">
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<emphasis id="B902EAF7FFC96E7CFF4FFBD4D2D42A22" box="[151,465,1094,1116]" italics="true" pageId="1" pageNumber="332">
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<taxonomicName id="4C764D66FFC96E7CFF4FFBD4D2342A22" ID-CoL="7Q3ZV" box="[151,305,1094,1116]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="genus">Terrathelphusa</taxonomicName>
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kuchingensis—
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</emphasis>
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Ng 1989: 127, pl. 4, fig. 3, colour pl. 2B; Ng 1995: 195; Ng & Grinang 2004: 314, fig. 7; Ng 2004: 327; Ng & Yeo 2007: 111; Ng
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<emphasis id="B902EAF7FFC96E7CFDE6FBF4D16E2A05" box="[574,619,1125,1147]" italics="true" pageId="1" pageNumber="332">et al</emphasis>
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. 2008: 73; Cumberlidge
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<emphasis id="B902EAF7FFC96E7CFCB0FBF4D0902A05" box="[872,917,1125,1147]" italics="true" pageId="1" pageNumber="332">et al</emphasis>
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. 2009: appendix 1; Grinang 2013: 6, pl. 3.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C656EFFC96E7CFF4FFB35D0A82B2D" pageId="1" pageNumber="332" type="materials_examined">
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<paragraph id="8BC936E5FFC96E7CFF4FFB35D0A82B2D" blockId="1.[151,1437,1191,2011]" pageId="1" pageNumber="332">
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<emphasis id="B902EAF7FFC96E7CFF4FFB35D2792ABE" bold="true" box="[151,380,1191,1216]" pageId="1" pageNumber="332">Material examined</emphasis>
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.
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<typeStatus id="54CD8847FFC96E7CFE56FB35D2FE2ABE" box="[398,507,1191,1216]" pageId="1" pageNumber="332" type="holotype">Holotype</typeStatus>
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: female (25.0 ×
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<quantity id="4C8E9B00FFC96E7CFD66FB35D0222ABE" box="[702,807,1191,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="332" unit="mm" value="19.5">19.5 mm</quantity>
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) (
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<collectionCode id="ED67AE20FFC96E7CFC99FB3AD0802ABE" box="[833,901,1192,1216]" httpUri="http://grbio.org/cool/0ffb-hhk1" name="Mycotheca Universitatis Taurinensis" pageId="1" pageNumber="332">MUT</collectionCode>
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), Kuching, coll. R. Shelford,
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<date id="FFC81025FFC96E7CFB07FB35D69D2ABE" box="[1247,1432,1191,1216]" pageId="1" pageNumber="332" value="1891-12">December 1891</date>
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.
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<typeStatus id="54CD8847FFC96E7CFF4FFB5FD3FB2A9B" box="[151,254,1229,1253]" pageId="1" pageNumber="332" type="paratype">Paratype</typeStatus>
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||
:
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<specimenCount id="9D70FD6CFFC96E7CFED0FB5ED2562A9B" box="[264,339,1228,1253]" pageId="1" pageNumber="332" type="male">1 male</specimenCount>
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||
(22.5 ×
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<quantity id="4C8E9B00FFC96E7CFE6CFB5ED1182A9A" box="[436,541,1228,1252]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="1" pageNumber="332" unit="mm" value="18.4">18.4 mm</quantity>
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),
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<specimenCount id="9D70FD6CFFC96E7CFDE8FB5ED1982A9B" box="[560,669,1228,1253]" pageId="1" pageNumber="332" type="female">3 females</specimenCount>
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||
(largest 34.6 × 26.0 mm) (SM Cru 1986.62-65), Kuching,
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<collectingCountry id="F3617675FFC96E7CFAEDFB5ED69D2A9B" box="[1333,1432,1228,1253]" name="Malaysia" pageId="1" pageNumber="332">Sarawak</collectingCountry>
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||
, as
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<taxonomicName id="4C764D66FFC96E7CFF6FFB7DD2E22B76" box="[183,487,1263,1288]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="kuchingense">
|
||
<emphasis id="B902EAF7FFC96E7CFF6FFB7DD2E22B76" box="[183,487,1263,1288]" italics="true" pageId="1" pageNumber="332">Geothelphusa kuchingense</emphasis>
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||
</taxonomicName>
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by G. Nobili, same data as
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||
<typeStatus id="54CD8847FFC96E7CFCF3FB7DD0942B76" box="[811,913,1263,1288]" pageId="1" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="9D70FD6CFFC96E7CFC46FB62D0E92B76" box="[926,1004,1263,1288]" pageId="1" pageNumber="332" type="male">1 male</specimenCount>
|
||
(36.0 ×
|
||
<quantity id="4C8E9B00FFC96E7CFB8AFB7DD7BF2B76" box="[1106,1210,1263,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="1" pageNumber="332" unit="mm" value="26.4">26.4 mm</quantity>
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||
) (
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||
<collectionCode id="ED67AE20FFC96E7CFB0BFB62D60E2B76" box="[1235,1291,1264,1288]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="332">ZRC</collectionCode>
|
||
2015.0001), farming plots, foothill of Gunung Temurang, Kuching,
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||
<collectingCountry id="F3617675FFC96E7CFCDFFA84D06F2B51" box="[775,874,1302,1327]" name="Malaysia" pageId="1" pageNumber="332">Sarawak</collectingCountry>
|
||
, 1O12’34.4”N 110O16’10.1”E,
|
||
<quantity id="4C8E9B00FFC96E7CFB66FA84D7FD2B51" box="[1214,1272,1302,1327]" metricMagnitude="1" metricUnit="m" metricValue="6.3" pageId="1" pageNumber="332" unit="m" value="63.0">63 m</quantity>
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||
asl, Padawan- Penrissen limestone formation, coll. J. Grinang
|
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<emphasis id="B902EAF7FFC96E7CFD75FAAED1D82B2D" box="[685,733,1338,1363]" italics="true" pageId="1" pageNumber="332">et al</emphasis>
|
||
.,
|
||
<date id="FFC81025FFC96E7CFD2BFAA8D0AF2B2D" box="[755,938,1338,1363]" pageId="1" pageNumber="332" value="2009-08-16">16 August 2009</date>
|
||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFC96E7CFF1FFACCD2CF2959" pageId="1" pageNumber="332" type="diagnosis">
|
||
<paragraph id="8BC936E5FFC96E7CFF1FFACCD1C528C5" blockId="1.[151,1437,1191,2011]" pageId="1" pageNumber="332">
|
||
<emphasis id="B902EAF7FFC96E7CFF1FFACCD23F2B09" bold="true" box="[199,314,1374,1399]" pageId="1" pageNumber="332">Diagnosis</emphasis>
|
||
. Carapace evenly oval transversely, longitudinally, surface convex, conspicuously smooth; epigastric cristae separated by deep epigastric groove; H-groove deep, almost confluent with cervical grooves; epibranchial tooth indistinct but can be separated from external orbital angle by notch, anterolateral, frontal regions appearing compressed; supraorbital margin almost parallel with frontal margin; epistome median lobe pointed to bifurcated (
|
||
<figureCitation id="134D2A60FFC96E7CFEC4FA7CD25A2879" box="[284,351,1518,1543]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
A–C; Ng, 1989: pls. 4A–C). Male abdomen with elongated somite 6, lateral margins gently concave (
|
||
<figureCitation id="134D2A60FFC96E7CFEDAF980D2442855" box="[258,321,1554,1579]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
D). Ischium of third maxilliped rectangular, about 1.8 times maximum width (
|
||
<figureCitation id="134D2A60FFC96E7CFB73F980D7EF2855" box="[1195,1258,1554,1579]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
E). Ambulatory legs smooth, long, about 3.9 times width (
|
||
<figureCitation id="134D2A60FFC96E7CFDA6F9A4D1C52831" box="[638,704,1590,1615]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
F; Ng, 1989: pl. 4A). Cheliped carpus rugose, without granules (
|
||
<figureCitation id="134D2A60FFC96E7CFF47F9C8D3D8280D" box="[159,221,1626,1651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
G, H), inner angle with acutely triangular tooth (
|
||
<figureCitation id="134D2A60FFC96E7CFD21F9C8D03D280D" box="[761,824,1626,1651]" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
A); gap between major chela conspicuously wide (
|
||
<figureCitation id="134D2A60FFC96E7CFAB7F9C8D3A728E9" captionStart="FIGURE 1" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[302,1284,591,1881]" captionTargetId="figure@2.[302,1284,591,1881]" captionTargetPageId="2" captionText="FIGURE 1. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244742/files/figure.png" pageId="1" pageNumber="332">Fig. 1</figureCitation>
|
||
H). G1 terminal segment cone-shaped, tapered, gently curving outwards, subequal to half-length of subterminal segment (
|
||
<figureCitation id="134D2A60FFC96E7CFEDCF930D24328C5" box="[260,326,1698,1723]" captionStart="FIGURE 2" captionStartId="3.[151,250,1052,1074]" captionTargetBox="[159,1427,198,1024]" captionTargetId="figure@3.[153,1434,193,1030]" captionTargetPageId="3" captionText="FIGURE 2. Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), Gunung Temurang, Kuching, Sarawak, Padawan-Penrissen limestone formation. A, ventral view of left G 1; B, ventral view of distal part of left G 1; C, dorsal view of left G 2; D, dorsal view of G 1; E, dorsal view of distal part of left G 1. Scales: A, C, D = 1 mm; B, E = 0.5 mm." httpUri="https://zenodo.org/record/244743/files/figure.png" pageId="1" pageNumber="332">Fig. 2</figureCitation>
|
||
A, B, D, E; Ng 1989:
|
||
<figureCitation id="134D2A60FFC96E7CFDE3F930D17928C5" box="[571,636,1698,1723]" captionStart="FIGURE 3" captionStartId="4.[151,250,1902,1924]" captionTargetBox="[356,1228,673,1884]" captionTargetId="figure@4.[356,1228,673,1884]" captionTargetPageId="4" captionText="FIGURE 3. Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), sandstone outcrop, Gunung Penrissen, Kuching, Sarawak. A, overall dorsal view; B, dorsal view of carapace; C, frontal view of carapace; D, ventral view showing anterior thoracic sternum and male abdomen; E, left third maxilliped; F, outer view of right fourth ambulatory leg; G, outer view of right chela; H, outer view of left chela." httpUri="https://zenodo.org/record/244744/files/figure.png" pageId="1" pageNumber="332">Fig. 3</figureCitation>
|
||
B, C).
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFC96E7CFF1FF954D7C028A1" blockId="1.[151,1437,1191,2011]" box="[199,1221,1734,1760]" pageId="1" pageNumber="332">
|
||
<emphasis id="B902EAF7FFC96E7CFF1FF954D24A28A1" bold="true" box="[199,335,1734,1759]" pageId="1" pageNumber="332">Life colour.</emphasis>
|
||
Carapace is dark purple with orange to white legs and chelipeds (
|
||
<figureCitation id="134D2A60FFC96E7CFBE8F954D77A28A1" box="[1072,1151,1734,1759]" captionStart="FIGURE 11" captionStartId="12.[151,250,1856,1878]" captionTargetBox="[157,1429,195,1832]" captionTargetId="figure@12.[157,1430,195,1834]" captionTargetPageId="12" captionText="FIGURE 11. Life colour of five species of Terrathelphusa from Kuching Division, Sarawak. A, B, Terrathelphusa kuchingensis (Nobili, 1901), male (36.0 × 26.4 mm) (ZRC 2015.0001), shortly after collection, Gunung Temurang; C, D, Terrathelphusa aglaia n. sp., holotype male (35.9 × 26.6 mm) (ZRC 2015.0002), Gunung Penrissen; E, F, Terrathelphusa cerina n. sp., female (24.2 × 20.1 mm), captured-marked-released, Gunung Singai; G, H, Terrathelphusa kundong n. sp., female (30.8 × 23.0 mm) (SBC. C. 00364 - 65), Gunung Serambu; I, J, Terrathelphusa mas n. sp., holotype male (35.5 × 26.4 mm) (ZRC 2015.0005), Gunung Penrissen." httpUri="https://zenodo.org/record/244752/files/figure.png" pageId="1" pageNumber="332">Fig. 11</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFC96E7CFF1FF978D2CF2959" blockId="1.[151,1437,1191,2011]" pageId="1" pageNumber="332">
|
||
<emphasis id="B902EAF7FFC96E7CFF1FF978D22F297D" bold="true" box="[199,298,1770,1795]" pageId="1" pageNumber="332">Habitat.</emphasis>
|
||
Foothills of limestone outcrops and adjacent farming plots; in shaded and moist soil. Individuals live in burrows about
|
||
<quantity id="4C8E9B00FFC96E7CFE84F89DD2822959" box="[348,391,1807,1831]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="332" unit="m" value="1.0">1 m</quantity>
|
||
deep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C656EFFC96E7FFF1FF8A0D1792C66" lastPageId="2" lastPageNumber="333" pageId="1" pageNumber="332" type="discussion">
|
||
<paragraph id="8BC936E5FFC96E7FFF1FF8A0D1652F3E" blockId="1.[151,1437,1191,2011]" lastBlockId="2.[151,1436,151,536]" lastPageId="2" lastPageNumber="333" pageId="1" pageNumber="332">
|
||
<emphasis id="B902EAF7FFC96E7CFF1FF8A0D23E2935" bold="true" box="[199,315,1842,1867]" pageId="1" pageNumber="332">Remarks.</emphasis>
|
||
The taxonomic problems associated with
|
||
<taxonomicName id="4C764D66FFC96E7CFCCAF8A6D0BB2935" box="[786,958,1842,1867]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="332" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFC96E7CFCCAF8A6D0BB2935" box="[786,958,1842,1867]" italics="true" pageId="1" pageNumber="332">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
, mostly on the absence of fresh specimens and their precise provenance, have been discussed at length by Ng (1989) and Ng & Grinang (2004). Nobili (1901) only recorded that the
|
||
<typeStatus id="54CD8847FFC96E7CFE4AF8E8D2F129ED" box="[402,500,1914,1939]" pageId="1" pageNumber="332" type="holotype">holotype</typeStatus>
|
||
specimen was passed to him by Robert Shelford, curator of zoology in the
|
||
<collectingCountry id="F3617675FFC96E7CFAE4F8E8D69929ED" box="[1340,1436,1914,1939]" name="Malaysia" pageId="1" pageNumber="332">Sarawak</collectingCountry>
|
||
Museum, who obtained it from Kuching in
|
||
<collectingCountry id="F3617675FFC96E7CFD43F80CD1FA29C9" box="[667,767,1950,1975]" name="Malaysia" pageId="1" pageNumber="332">Sarawak</collectingCountry>
|
||
. Recent collections in the Kuching region have provided further knowledge on the taxonomy and precise distribution of the species. The recent male specimen agrees very well with description of the species by Ng (1989) and the SM specimens collected at around the same time as the
|
||
<typeStatus id="54CD8847FFCA6E7FFF4FFF2FD3CE2EAB" box="[151,203,189,213]" pageId="2" pageNumber="333">type</typeStatus>
|
||
. It was collected from Gunung Temurang in Padawan-Penrissen limestone formation area. The limestone hill is located about
|
||
<quantity id="4C8E9B00FFCA6E7FFE83FF4DD29C2E86" box="[347,409,223,248]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="333" unit="km" value="5.0">5 km</quantity>
|
||
from Pangkalan Ampat, a site where Shelford stayed during his expedition to Gunung Penrissen (Shelford 1900), during which he and/or his field assistants probably collected the
|
||
<typeStatus id="54CD8847FFCA6E7FFB3AFE96D6412F63" box="[1250,1348,260,285]" pageId="2" pageNumber="333" type="holotype">holotype</typeStatus>
|
||
female specimen (see also Bott 1970; Ng 1989).
|
||
</paragraph>
|
||
<paragraph id="8BC936E5FFCA6E7FFF1FFEDFD1792C66" blockId="2.[151,1436,151,536]" pageId="2" pageNumber="333">
|
||
Ng (1989) had identified a red crab photographed by Gerald Cubitt in Bako National Park as possibly being
|
||
<taxonomicName id="4C764D66FFCA6E7FFA5FFEDFD22D2FF6" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCA6E7FFA5FFEDFD22D2FF6" italics="true" pageId="2" pageNumber="333">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
, but recent collections have showed all known
|
||
<taxonomicName id="4C764D66FFCA6E7FFC95FEFDD0F22FF6" box="[845,1015,367,392]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EAF7FFCA6E7FFC95FEFDD0F22FF6" box="[845,1015,367,392]" italics="true" pageId="2" pageNumber="333">Terrathelphusa</emphasis>
|
||
</taxonomicName>
|
||
species from the Kuching region do not have this appearance or colour (see below). The Bako crab is actually very similar to
|
||
<taxonomicName id="4C764D66FFCA6E7FFB54FE06D28A2FAE" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" class="Malacostraca" family="Gecarcinucidae" genus="Thelphusula" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="cristicervix">
|
||
<emphasis id="B902EAF7FFCA6E7FFB54FE06D6992FD2" box="[1164,1436,404,429]" italics="true" pageId="2" pageNumber="333">Thelphusula cristicervix</emphasis>
|
||
(Ng & Grinang, 2004)
|
||
</taxonomicName>
|
||
from Sama Jaya Nature Reserve in Kuching (see Grinang & Ng 2014). In light of the present revision, most of the specimens that had been identified as
|
||
<taxonomicName id="4C764D66FFCA6E7FFC8BFE4FD7012F8B" box="[851,1028,476,501]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="kuchingensis">
|
||
<emphasis id="B902EAF7FFCA6E7FFC8BFE4FD7012F8B" box="[851,1028,476,501]" italics="true" pageId="2" pageNumber="333">T. kuchingensis</emphasis>
|
||
</taxonomicName>
|
||
by Ng & Grinang (2004) are here referred to another taxon,
|
||
<taxonomicName id="4C764D66FFCA6E7FFE62FD93D1322C66" box="[442,567,511,536]" class="Malacostraca" family="Gecarcinucidae" genus="Terrathelphusa" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="333" phylum="Arthropoda" rank="species" species="kundong" status="sp. nov.">
|
||
<emphasis id="B902EAF7FFCA6E7FFE62FD93D1322C66" box="[442,567,511,536]" italics="true" pageId="2" pageNumber="333">T. kundong</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A231578CFFCA6E7FFDE5FE6DD1792C66" box="[573,636,511,536]" pageId="2" pageNumber="333" rank="species">
|
||
<emphasis id="B902EAF7FFCA6E7FFDE5FE6DD1792C66" bold="true" box="[573,636,511,536]" pageId="2" pageNumber="333">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |