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<document id="62C6DDA9CC22855DE90D43CA09962C14" ID-DOI="10.5281/zenodo.280870" ID-GBIF-Dataset="59423779-337f-41b0-8993-702ba050a79a" ID-ISSN="1175-5326" ID-Zenodo-Dep="280870" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460476674350" checkinUser="plazi" docAuthor="Matallanas, Jesús &amp; Corbella, Cecília" docDate="2012" docId="03DF87D68D1DD644FF48E2F9FF4BFC43" docLanguage="en" docName="zt03296p018.pdf" docOrigin="Zootaxa 3296" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Iluocoetes Jenyns 1842" docType="treatment" docVersion="7" lastPageNumber="6" masterDocId="FFE6FFAE8D19D641FFDFE453FF94FFAF" masterDocTitle="Redescription of Iluocoetes Jenyns, 1842; proposal of a new genus, Argentinolycus, for Iluocoetes elongatus (Smitt, 1898), and description of Patagolycus melastomus gen. et sp. nov. (Teleostei, Zoarcidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="5" updateTime="1698306862796" updateUser="plazi">
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<mods:title id="40AD99AC405E39B50B017E984261E97A">Redescription of Iluocoetes Jenyns, 1842; proposal of a new genus, Argentinolycus, for Iluocoetes elongatus (Smitt, 1898), and description of Patagolycus melastomus gen. et sp. nov. (Teleostei, Zoarcidae)</mods:title>
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<mods:namePart id="87BE43532823F161F9DFCDFDC88A7E7D">Matallanas, Jesús</mods:namePart>
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<mods:namePart id="2273BA19B31F3E7D56FE82021E51D1CD">Corbella, Cecília</mods:namePart>
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<mods:date id="8D4B2F28FD50C9632C30780A20DCB49F">2012</mods:date>
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<paragraph id="8BC936C08D1DD645FF48E2F9FD1BF96B" blockId="4.[151,655,1706,1765]" box="[151,655,1706,1732]" pageId="4" pageNumber="5">
<heading id="D08181AC8D1DD645FF48E2F9FD1BF96B" bold="true" box="[151,655,1706,1732]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B902EAD28D1DD645FF48E2F9FD1BF96B" bold="true" box="[151,655,1706,1732]" pageId="4" pageNumber="5">
Redescription of
<taxonomicName id="4C764D438D1DD645FEB3E2F9FD1BF96B" ID-CoL="53WT" ID-ENA="458578" authority="Jenyns, 1842" authorityName="Jenyns" authorityYear="1842" box="[364,655,1706,1732]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
<emphasis id="B902EAD28D1DD645FEB3E2F9FE7CF96B" bold="true" box="[364,488,1706,1732]" italics="true" pageId="4" pageNumber="5">Iluocoetes</emphasis>
<bibRefCitation id="EFE74B318D1DD645FE30E2F9FD1BF96B" author="Jenyns" box="[495,655,1706,1732]" pageId="4" pageNumber="5" refString="Jenyns, L. (1842) The zoology of the voyage of H. M. S. Beagle, under the command of Captain Fitzroy, R. N., during the years 1832 to 1836. IV. Fish. Smith, Elder &amp; Co., London, v. 4, pp. 97 - 172." type="journal article" year="1842">Jenyns, 1842</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC936C08D1DD645FF48E29EFE4FF94A" blockId="4.[151,655,1706,1765]" box="[151,475,1741,1765]" pageId="4" pageNumber="5">
(
<figureCitation id="134D2A458D1DD645FF40E29EFF79F94A" box="[159,237,1741,1765]" captionStart="FIGURE 1" captionStartId="4.[151,250,1522,1545]" captionTargetBox="[167,1422,207,1490]" captionTargetId="figure@4.[152,1437,195,1493]" captionTargetPageId="4" captionText="FIGURE 1. Neurocranium of Iluocoetes fimbriatus Jenyns, 1842. (UAB. P 47, 224 mm TL). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; ETM CART, ethmoid cartilage; EXO, exoccipital; FR, frontal; INT, intercalary; LAT ETM, lateral ethmoid; MES mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V vomer." httpUri="https://zenodo.org/record/280871/files/figure.png" pageId="4" pageNumber="5">Figs. 1</figureCitation>
,
<figureCitation id="134D2A458D1DD645FF25E29EFE9CF94A" box="[250,264,1741,1765]" captionStart="FIGURE 2" captionStartId="5.[151,250,1463,1486]" captionTargetBox="[166,1435,1058,1426]" captionTargetId="figure@5.[151,1436,1023,1441]" captionTargetPageId="5" captionText="FIGURE 2. Iluocoetes fimbriatus BMNH 1936.2.26.988 991, 203 mm TL. Left lateral view showing a dark band between the antero-inferior edge of the eye and the upper jaw." httpUri="https://zenodo.org/record/280872/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
&amp;
<figureCitation id="134D2A458D1DD645FEF3E29EFEDFF94A" box="[300,331,1741,1765]" captionStart="FIGURE 10" captionStartId="14.[151,250,1539,1562]" captionTargetBox="[427,1167,506,1511]" captionTargetId="figure@14.[414,1185,485,1528]" captionTargetPageId="14" captionText="FIGURE 10. Interrelationships among the Magellanic endemic lycodine genera. Consensus tree resulting from the parsimony analysis of 78 morphological characters (Length = 105; CI = 0.53; RI = 0.50). Bootstrap support values (rep = 1000) are indicated on the nodes. White boxes on branches: homoplastic apomorphies; black boxes: synapomorphies." httpUri="https://zenodo.org/record/280881/files/figure.png" pageId="4" pageNumber="5">10</figureCitation>
; Tables 24)
</paragraph>
<paragraph id="8BC936C08D1DD644FF48E346FF51FECB" blockId="4.[151,1437,1813,2017]" lastBlockId="5.[151,1436,151,1004]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B902EAD28D1DD645FF48E346FCEDF881" bold="true" box="[151,889,1813,1838]" pageId="4" pageNumber="5">
Diagnostic characters found in the
<typeStatus id="54CD88628D1DD645FD95E345FD25F881" box="[586,689,1814,1838]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(+radiographs):
</emphasis>
Submental crests present, not fused anteriorly; neurocranium elongate, narrowed; frontal bones fused totally with no trace of a suture; frontal corner squared; frontal ramus long; sphenotic and parietal separated by pterotic; parietal bones meeting in the mid-line; parasphenoid wing height high; ceratohyal epihyal juncture smooth; five branchiostegal rays; posterior hyomandibular ramus elongate; palatal arch well developed; posttemporal ventral ramus well developed; scapular foramen enclosed by bone; scapular strut present; four notched radials (=actinosts); postcleithrum present; dorsal fin-rays 85 (last 7 counted from dorsal-fin pterygiophores); anal-fin rays, 71 (last 10 counted from anal-fin pterygiophores); pectoralfin rays 18; pelvic bone present; vertebrae asymmetrical (21+68=89); ribs on 421 abdominal vertebrae; one epural; dorsal-fin origin associated with vertebrae 4; oral valve well developed; gill slit extending ventrally to lower end of pectoral-fin base; gill rakers blunt; pyloric caeca nub-like; scales, palatine and vomerine teeth present; oral cavity pale; peritoneum, black; a blurred brown band between the anteroventral edge of the eye and the upper jaw.
</paragraph>
<paragraph id="8BC936C08D1CD644FF16E53CFDA8FD63" blockId="5.[151,1436,151,1004]" pageId="5" pageNumber="6">
All these characters are present also in the other specimens of
<taxonomicName id="4C764D438D1CD644FC58E522FB85FE27" box="[903,1041,368,392]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B902EAD28D1CD644FC58E522FB85FE27" box="[903,1041,368,392]" italics="true" pageId="5" pageNumber="6">I. fimbriatus</emphasis>
</taxonomicName>
examined, and additional diagnostic characters of
<taxonomicName id="4C764D438D1CD644FE89E5C7FE5DFE03" box="[342,457,404,428]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B902EAD28D1CD644FE89E5C7FE5DFE03" box="[342,457,404,428]" italics="true" pageId="5" pageNumber="6">Iluocoetes</emphasis>
</taxonomicName>
based on cleared and stained specimens of
<taxonomicName id="4C764D438D1CD644FC1EE5C6FBD9FE03" box="[961,1101,404,428]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B902EAD28D1CD644FC1EE5C6FBD9FE03" box="[961,1101,404,428]" italics="true" pageId="5" pageNumber="6">I. fimbriatus</emphasis>
</taxonomicName>
for anatomical study, are the following (
<figureCitation id="134D2A458D1CD644FECBE5E4FEC7FE60" box="[276,339,439,463]" captionStart="FIGURE 1" captionStartId="4.[151,250,1522,1545]" captionTargetBox="[167,1422,207,1490]" captionTargetId="figure@4.[152,1437,195,1493]" captionTargetPageId="4" captionText="FIGURE 1. Neurocranium of Iluocoetes fimbriatus Jenyns, 1842. (UAB. P 47, 224 mm TL). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; ETM CART, ethmoid cartilage; EXO, exoccipital; FR, frontal; INT, intercalary; LAT ETM, lateral ethmoid; MES mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V vomer." httpUri="https://zenodo.org/record/280871/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
a, b): frontal and parasphenoid not separated by pterosphenoid; supraoccipital exoccipìtal articulation excluded by epioccipitals; supratemporal commissure and occipital pores absent (
<figureCitation id="134D2A458D1CD644FBB3E58FFB3FFE5B" box="[1132,1195,476,500]" captionStart="FIGURE 1" captionStartId="4.[151,250,1522,1545]" captionTargetBox="[167,1422,207,1490]" captionTargetId="figure@4.[152,1437,195,1493]" captionTargetPageId="4" captionText="FIGURE 1. Neurocranium of Iluocoetes fimbriatus Jenyns, 1842. (UAB. P 47, 224 mm TL). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; ETM CART, ethmoid cartilage; EXO, exoccipital; FR, frontal; INT, intercalary; LAT ETM, lateral ethmoid; MES mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V vomer." httpUri="https://zenodo.org/record/280871/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
b); ascending rami of the parasphenoid reaches the upper margin of the trigeminofacialis foramen; intercalar well developed, not reaching prootic (
<figureCitation id="134D2A458D1CD644FEFDE677FEF6FD93" box="[290,354,548,572]" captionStart="FIGURE 1" captionStartId="4.[151,250,1522,1545]" captionTargetBox="[167,1422,207,1490]" captionTargetId="figure@4.[152,1437,195,1493]" captionTargetPageId="4" captionText="FIGURE 1. Neurocranium of Iluocoetes fimbriatus Jenyns, 1842. (UAB. P 47, 224 mm TL). Lateral view (a) and dorsal view (b). BAS, basioccipital; EPO, epioccipital; ETM CART, ethmoid cartilage; EXO, exoccipital; FR, frontal; INT, intercalary; LAT ETM, lateral ethmoid; MES mesethmoid; PAR, parietal; PAS, parasphenoid; PRO, prootic; PTO, pterotic; PTS, pterosphenoid; SO, supraoccipital; SPH, sphenotic; TGF, trigeminofacialis foramen; V vomer." httpUri="https://zenodo.org/record/280871/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
a); suborbital bones 89, canal with 7 pores; postorbital pores 1 and 4; posterior nasal pore single; cartilaginous basal plate of pectoral girdle with three foramina; vertebrae asymmetrical (1921+6477=83 97); dorsal-fin origin associated with vertebrae 24; oral valve well developed; gill rakers blunt; pelvic-fin rays ensheathed; squamation extensive, but head and pectoral-fin base and axil scaleless; lateral line mediolateral, palatine teeth 923; vomerine teeth 717.
</paragraph>
<paragraph id="8BC936C08D1CD644FF16E684FB51FCD0" blockId="5.[151,1436,151,1004]" pageId="5" pageNumber="6">
Colouration in juvenile preserved specimens of
<taxonomicName id="4C764D438D1CD644FCD1E68BFB96FD5F" box="[782,1026,728,752]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B902EAD28D1CD644FCD1E68BFB96FD5F" box="[782,1026,728,752]" italics="true" pageId="5" pageNumber="6">Iluocoetes fimbriatus</emphasis>
</taxonomicName>
(UAB.P
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TL; BMNH 1936.8.26.962972:
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TL; BMNH 1912.7.1.84,
<quantity id="4C8E9B258D1CD644FCF1E6AFFC1FFCBB" box="[814,907,764,788]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="mm" value="140.0">140 mm</quantity>
TL; BMNH 1936.8.
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TL). Head and body mid-brown coloured with dull white spots on both, head (nape, interorbital area, cheeks) and body, mainly in its dorsolateral part. A darker band on head between the anteroventral edge of the eye and the upper jaw. Ventrolateral part of both head and body light brown. Edge of dorsal fin with 56 darker bands.
</paragraph>
<paragraph id="8BC936C08D1CD644FF16E7DFFF4BFC43" blockId="5.[151,1436,151,1004]" pageId="5" pageNumber="6">
The character state of other diagnostic features of this genus are given in Tables 24. Other descriptive characters of
<taxonomicName id="4C764D438D1CD644FF38E7E2FEE7FC67" box="[231,371,944,968]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B902EAD28D1CD644FF38E7E2FEE7FC67" box="[231,371,944,968]" italics="true" pageId="5" pageNumber="6">I. fimbriatus</emphasis>
</taxonomicName>
can be found in
<bibRefCitation id="EFE74B318D1CD644FDEAE7FCFCE5FC68" author="Gosztonyi" box="[565,881,943,967]" pageId="5" pageNumber="6" refString="Gosztonyi, A. E. (1977) Results of the research cruises of FRV &quot; Walther Herwig &quot; to South America. XLVIII. Revision of the South American Zoarcidae (Osteichthyes, Blennioidei) with the description of three new genera and five new species. Archiv fur Fischereiwissenschaft, 27, 191 - 249." type="journal article" year="1977">Gosztonyi (1977: 211215)</bibRefCitation>
. The general shape of
<taxonomicName id="4C764D438D1CD644FBABE7E3FAF6FC67" box="[1140,1378,944,968]" class="Actinopterygii" family="Zoarcidae" genus="Iluocoetes" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="fimbriatus">
<emphasis id="B902EAD28D1CD644FBABE7E3FAF6FC67" box="[1140,1378,944,968]" italics="true" pageId="5" pageNumber="6">Iluocoetes fimbriatus</emphasis>
</taxonomicName>
is in
<figureCitation id="134D2A458D1CD644FF48E787FF4FFC43" box="[151,219,980,1004]" captionStart="FIGURE 2" captionStartId="5.[151,250,1463,1486]" captionTargetBox="[166,1435,1058,1426]" captionTargetId="figure@5.[151,1436,1023,1441]" captionTargetPageId="5" captionText="FIGURE 2. Iluocoetes fimbriatus BMNH 1936.2.26.988 991, 203 mm TL. Left lateral view showing a dark band between the antero-inferior edge of the eye and the upper jaw." httpUri="https://zenodo.org/record/280872/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
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