treatments-xml/data/03/DF/87/03DF8791FFFBFFB9B4DA5C7CFA9566B6.xml

<document id="A3F30682FD1E83EE2D7890432F1BBD95" ID-DOI="10.5281/zenodo.12767696" ID-ISSN="1989-6581" ID-Zenodo-Dep="12767696" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721250947469" checkinUser="felipe" docAuthor="Petrulevičius, Julián F." docDate="2017" docId="03DF8791FFFBFFB9B4DA5C7CFA9566B6" docLanguage="en" docName="ArqEntomol.18.367-374.pdf" docOrigin="Arquivos Entomolóxicos 18" docStyle="DocumentStyle:DF23C849EA72F80F6DF49FA5382909D5.2:ArqEntomol.2008-.journal_article.1column" docStyleId="DF23C849EA72F80F6DF49FA5382909D5" docStyleName="ArqEntomol.2008-.journal_article.1column" docStyleVersion="2" docTitle="Treintamilun vuelvenlucha Petrulevičius, 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="370" masterDocId="FFE6FFE9FFF9FFBAB5205D0DFF97602E" masterDocTitle="First Frenguelliidae (Insecta: Odonata) from the middle Eocene of Río Pichileuf ̇, Patagonia, Argentina." masterLastPageNumber="374" masterPageNumber="367" pageNumber="369" updateTime="1721260506247" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="17F074666B3E223D97AB75110C28597F">First Frenguelliidae (Insecta: Odonata) from the middle Eocene of Río Pichileuf ̇, Patagonia, Argentina.</mods:title>
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<mods:namePart id="B530CA3F777315BB02E233DE222F4CAD">Petrulevičius, Julián F.</mods:namePart>
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<mods:date id="4123447721DE154F32E6C44E2A2DAEB3">2017</mods:date>
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<subSubSection id="C36C650CFFFBFFB8B4DA5C7CFBAD61A4" box="[506,1082,369,394]" pageId="2" pageNumber="369" type="nomenclature">
<paragraph id="8BC93687FFFBFFB8B4DA5C7CFBAD61A4" blockId="2.[506,1082,369,394]" box="[506,1082,369,394]" pageId="2" pageNumber="369">
<heading id="D08181EBFFFBFFB8B4DA5C7CFBAD61A4" bold="true" box="[506,1082,369,394]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="369" reason="6">
<emphasis id="B902EA95FFFBFFB8B4DA5C7CFBAD61A4" bold="true" box="[506,1082,369,394]" pageId="2" pageNumber="369">
<taxonomicName id="4C764D04FFFBFFB8B4DA5C7CFCD761A4" authorityName="Petrulevičius" authorityYear="2017" box="[506,832,369,394]" class="Insecta" family="Frenguelliidae" genus="Treintamilun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="vuelvenlucha" status="sp. nov.">
<emphasis id="B902EA95FFFBFFB8B4DA5C7CFCD761A4" bold="true" box="[506,832,369,394]" italics="true" pageId="2" pageNumber="369">Treintamilun vuelvenlucha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157EEFFFBFFB8B66B5C7CFC2561A4" box="[843,946,369,394]" pageId="2" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="134D2A02FFFBFFB8B6E55C7CFBA761A4" box="[965,1072,369,394]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[157,193,1999,2019]" captionStartId-1="7.[157,193,652,672]" captionStartId-2="7.[159,195,1884,1904]" captionTargetBox-0="[155,1449,973,1966]" captionTargetBox-1="[196,1400,276,562]" captionTargetBox-2="[163,1450,850,1827]" captionTargetId-0="figure-147@6.[145,1451,963,1972]" captionTargetId-1="figure-24@7.[159,1437,229,605]" captionTargetId-2="figure-84@7.[159,1458,846,1831]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionText-0="Fig. 1.- Photographs of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Part. b.- Counterpart. Scale bar = 3 mm." captionText-1="Fig. 2.- Composite camera lucida drawing of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. Scale bar = 2 mm." captionText-2="Fig. 3.- Photographs of details of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Discoidal cell, scale bar = 0.5 mm. b.- Basal part, scale bar = 1 mm. c.- Nodus, scale bar = 0.25 mm. d.- Pterostigma, scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12767698" figureDoi-1="http://doi.org/10.5281/zenodo.12767700" figureDoi-2="http://doi.org/10.5281/zenodo.12767702" httpUri-0="https://zenodo.org/record/12767698/files/figure.png" httpUri-1="https://zenodo.org/record/12767700/files/figure.png" httpUri-2="https://zenodo.org/record/12767702/files/figure.png" pageId="2" pageNumber="369">Figs. 1-3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C650CFFFBFFB8B5B05CB9FD1761E3" box="[144,640,436,461]" pageId="2" pageNumber="369" type="diagnosis">
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<emphasis id="B902EA95FFFBFFB8B5B05CB9FD1761E3" bold="true" box="[144,640,436,461]" pageId="2" pageNumber="369">Diagnosis. As for the genus (see above).</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36C650CFFFBFFB8B5B05CE2FE1E662B" pageId="2" pageNumber="369" type="description">
<paragraph id="8BC93687FFFBFFB8B5B05CE2FE1E662B" blockId="2.[144,1444,495,1541]" pageId="2" pageNumber="369">
<emphasis id="B902EA95FFFBFFB8B5B05CE2FE1E662B" bold="true" pageId="2" pageNumber="369">
Description. A complete hindwing (?) with two dark zones crossing the wing (
<figureCitation id="134D2A02FFFBFFB8B1705CE2FB286226" box="[1104,1215,495,520]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[157,193,1999,2019]" captionStartId-1="7.[157,193,652,672]" captionTargetBox-0="[155,1449,973,1966]" captionTargetBox-1="[196,1400,276,562]" captionTargetId-0="figure-147@6.[145,1451,963,1972]" captionTargetId-1="figure-24@7.[159,1437,229,605]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 1.- Photographs of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Part. b.- Counterpart. Scale bar = 3 mm." captionText-1="Fig. 2.- Composite camera lucida drawing of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. Scale bar = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12767698" figureDoi-1="http://doi.org/10.5281/zenodo.12767700" httpUri-0="https://zenodo.org/record/12767698/files/figure.png" httpUri-1="https://zenodo.org/record/12767700/files/figure.png" pageId="2" pageNumber="369">Figs. 1-2</figureCitation>
), one beneath the pterostigma and the other covering the area distal to the nodus to the height of RP2; other areas hyaline; petiole short, about 
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long; wing 
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long, 
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wide; petiole short and broad, 
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long and 
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wide; wing 
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long, 
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wide; distance between base and arculus, 
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, between arculus and nodus, 
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, between nodus and pterostigma, 
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, between pterostigma and apex, 
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; nodus basally recessed; pterostigma long (
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) and broad (
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), covering four and a half cells (
<figureCitation id="134D2A02FFFBFFB8B4355FDFFEE662C2" box="[277,369,722,748]" captionStart="Fig" captionStartId="7.[159,195,1884,1904]" captionTargetBox="[163,1450,850,1827]" captionTargetId="figure-84@7.[159,1458,846,1831]" captionTargetPageId="7" captionText="Fig. 3.- Photographs of details of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Discoidal cell, scale bar = 0.5 mm. b.- Basal part, scale bar = 1 mm. c.- Nodus, scale bar = 0.25 mm. d.- Pterostigma, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12767702" httpUri="https://zenodo.org/record/12767702/files/figure.png" pageId="2" pageNumber="369">Fig. 3C</figureCitation>
); pterostigmal brace reduced, anterior side of pterostigma sligthly oblique; Ax2 just distal to arculus; Ax1 
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basally; discoidal cell basally closed, broad, distinctly widened distally, anterior side, 
<quantity id="4C8E9B62FFFBFFB8B4625E13FE0C6319" box="[322,411,798,823]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="369" unit="mm" value="0.7">0.7 mm</quantity>
long, posterior side, 
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long, basal side, 
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long, distal side, 
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long; no antesubnodal cross-veins; discoidal cell basally closed, broad, distinctly widened distally, anterior side (MA), 
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long, posterior side (cross-vein = ddcv), 
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long, basal side (cross-vein = bdcv), 
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long, distal side, 
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long; arculus short; RP get free nearer anterior side of arculus; ddcv about 80º to MA; MP + CuA with a strong angle just distal of the base of CuP; CuP curved (
<figureCitation id="134D2A02FFFBFFB8B1D55EB8FAC463E0" box="[1269,1363,949,974]" captionStart="Fig" captionStartId="7.[159,195,1884,1904]" captionTargetBox="[163,1450,850,1827]" captionTargetId="figure-84@7.[159,1458,846,1831]" captionTargetPageId="7" captionText="Fig. 3.- Photographs of details of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Discoidal cell, scale bar = 0.5 mm. b.- Basal part, scale bar = 1 mm. c.- Nodus, scale bar = 0.25 mm. d.- Pterostigma, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12767702" httpUri="https://zenodo.org/record/12767702/files/figure.png" pageId="2" pageNumber="369">Fig. 3A</figureCitation>
); base of RP3+4 between arculus and nodus, about 
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basal to subnodus, 
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distal to arculus; base of IR2 below subnodus; base of RP2 six cells, 
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distal of subnodus; base of IR1 2-3 veins distal to RP2; nodal cross-vein (Cr) sub-vertical (
<figureCitation id="134D2A02FFFBFFB8B765592BFD356411" box="[581,674,1062,1087]" captionStart="Fig" captionStartId="7.[159,195,1884,1904]" captionTargetBox="[163,1450,850,1827]" captionTargetId="figure-84@7.[159,1458,846,1831]" captionTargetPageId="7" captionText="Fig. 3.- Photographs of details of Treintamilun vuelvenlucha gen. et sp. nov., holotype MAPBAR 4139 from Río Pichileuḟ (Río Negro, Argentina); Lutetian, middle Eocene. a.- Discoidal cell, scale bar = 0.5 mm. b.- Basal part, scale bar = 1 mm. c.- Nodus, scale bar = 0.25 mm. d.- Pterostigma, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12767702" httpUri="https://zenodo.org/record/12767702/files/figure.png" pageId="2" pageNumber="369">Fig. 3B</figureCitation>
), 
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distal of point of fusion of ScP with costal margin; subnodus vertical; posterior bent of CP not aligned with Cr but in a slightly distal position, at the point of fusion between ScP and costal margin; 17 postnodal cross-veins between C and RA, only aligned the four basal ones with the corresponding cross-veins between RA and RP1; cubito-anal area broad, with three rows of cells between CuA and posterior wing margin; CuA zigzagged reaching posterior wing margin well distal (about 
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) of nodus level; postdiscoidal area with only one row of cells and distally narrowed; area between MA and RP3+4 distally widened; area between RP3+4 and IR2 narrow, with one row of cells; areas between IR2 and RP2 and between RP2 and IR1 distally widened with two long secondary longitudinal veins; area between IR1 and RP1 with only one row of cells, broader than long; MA and CuA distally zigzagged; MP, RP3+4, IR2 and RP2 more or less straight or slightly curved; IR1 with a distinct but smooth curve opposite pterostigma, corresponding to a narrowing of the area between it and RP1 and a broadening of the area between it and RP2; no significant increase of spine-density at the apical costal margin.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36C650CFFFBFFB8B5B05B25FE8466A5" pageId="2" pageNumber="369" type="etymology">
<paragraph id="8BC93687FFFBFFB8B5B05B25FE8466A5" blockId="2.[144,1444,1575,1675]" pageId="2" pageNumber="369">
<emphasis id="B902EA95FFFBFFB8B5B05B25FE8466A5" bold="true" pageId="2" pageNumber="369">
Etymology. From the Castilian “ 
<emphasis id="B902EA95FFFBFFB8B7025B25FD76666F" bold="true" box="[546,737,1576,1601]" italics="true" pageId="2" pageNumber="369">vuelve en lucha</emphasis>
”, meAning returning in fight. In homage to the 30,001, alive in the dreams and commitment of the People. The specific epithet is to be considered as a noun in apposition.
</emphasis>
</paragraph>
</subSubSection>
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<emphasis id="B902EA95FFFBFFB8B5B05BA3FD8966C2" bold="true" pageId="2" pageNumber="369">
<materialsCitation id="3B1E3CDAFFFBFFB8B5B05BA3FD8C66C2" country="Argentina" location="Museo" pageId="2" pageNumber="369" specimenCount="1" stateProvince="Asociacion Paleontologica de Bariloche" typeStatus="holotype">
<typeStatus id="54CD8825FFFBFFB8B5B05BA3FF5966E9" box="[144,206,1710,1735]" pageId="2" pageNumber="369">Type</typeStatus>
material. MAPBAR 4139, 
<location id="8EA9605CFFFBFFB8B7125BA3FD1666E9" LSID="urn:lsid:plazi:treatment:03DF8791FFFBFFB9B4DA5C7CFA9566B6:8EA9605CFFFBFFB8B7125BA3FD1666E9" box="[562,641,1710,1735]" country="Argentina" name="Museo" pageId="2" pageNumber="369" stateProvince="Asociacion Paleontologica de Bariloche">Museo</location>
de la 
<collectingRegion id="49B2F865FFFBFFB8B7C25BA0FB4066E9" box="[738,1239,1709,1735]" country="Argentina" name="Asociacion Paleontologica de Bariloche" pageId="2" pageNumber="369">Asociación Paleontológica de Bariloche</collectingRegion>
, 
<collectingRegion id="49B2F865FFFBFFB8B1CA5BA0FE9066C2" country="Argentina" name="San Carlos de Bariloche" pageId="2" pageNumber="369">San Carlos de Bariloche</collectingRegion>
, 
<collectingRegion id="49B2F865FFFBFFB8B4335BDFFE0666C2" box="[275,401,1746,1772]" country="Argentina" name="Rio Negro" pageId="2" pageNumber="369">Río Negro</collectingRegion>
, 
<collectingCountry id="F3617617FFFBFFB8B4BE5BDEFD8C66C2" box="[414,539,1747,1772]" name="Argentina" pageId="2" pageNumber="369">Argentina</collectingCountry>
</materialsCitation>
.
</emphasis>
</paragraph>
<paragraph id="8BC93687FFFBFFB8B5B05A03FAA5675C" blockId="2.[144,1443,1806,1906]" pageId="2" pageNumber="369">
<emphasis id="B902EA95FFFBFFB8B5B05A03FAA5675C" bold="true" pageId="2" pageNumber="369">
<materialsCitation id="3B1E3CDAFFFBFFB8B5B05A03FABA675C" collectionCode="RP" country="Argentina" county="Patagonia" latitude="-46.0" location="Volcanic" municipality="And" pageId="2" pageNumber="369" specimenCode="RP4, RP3" specimenCount="1" stateProvince="Rio Pichileufu" typeStatus="holotype">
<typeStatus id="54CD8825FFFBFFB8B5B05A03FF5A6709" box="[144,205,1806,1831]" pageId="2" pageNumber="369">Type</typeStatus>
locality. 
<location id="8EA9605CFFFBFFB8B46B5A03FE276709" LSID="urn:lsid:plazi:treatment:03DF8791FFFBFFB9B4DA5C7CFA9566B6:8EA9605CFFFBFFB8B46B5A03FE276709" box="[331,432,1806,1831]" country="Argentina" county="Patagonia" latitude="-46.0" municipality="And" name="Volcanic" pageId="2" pageNumber="369" stateProvince="Rio Pichileufu">Volcanic</location>
caldera-lAKe beds, 
<collectingRegion id="49B2F865FFFBFFB8B78D5A03FCCC6706" box="[685,859,1806,1832]" country="Argentina" name="Rio Pichileufu" pageId="2" pageNumber="369">Río Pichileufú</collectingRegion>
, quArry 
<specimenCode id="DBD09EFCFFFBFFB8B6E75A03FC616709" box="[967,1014,1806,1831]" collectionCode="RP" pageId="2" pageNumber="369">RP4</specimenCode>
(= to field numbers “PichiPrem”), new locality discovered by AriAnA “Premgi” PAulinA CArAbAJAl in 2016 
<collectingMunicipality id="6BADACFDFFFBFFB8B6D25A39FB8A6763" box="[1010,1053,1844,1869]" pageId="2" pageNumber="369">And</collectingMunicipality>
lAterAl equivAlent to 
<specimenCode id="DBD09EFCFFFBFFB8B00C5A39FACC6763" box="[1324,1371,1844,1869]" collectionCode="RP" pageId="2" pageNumber="369">RP3</specimenCode>
from 
<bibRefCitation id="EFE74B76FFFBFFB8B5B05A54FEF9675C" author="Wilf, P. &amp; Johnson, K. R. &amp; Smith, M. E. &amp; Singer, B. S. &amp; Gandolfo, M. A." box="[144,366,1881,1906]" pageId="2" pageNumber="369" pagination="634 - 650" refId="ref4587" refString="Wilf, P.; Johnson, K. R.; C ˙ neo, N. R.; Smith, M. E.; Singer, B. S. &amp; Gandolfo, M. A. 2005. Eocene Plant Diversity at Laguna del Hunco and Rio Pichileuf ˙, Patagonia, Argentina. The American Naturalist 165: 634 - 650." type="journal article" year="2005">Wilf et al. (2005)</bibRefCitation>
, 
<collectingRegion id="49B2F865FFFBFFB8B45A5A54FE62675C" box="[378,501,1881,1906]" country="Argentina" name="Pilcaniyeu" pageId="2" pageNumber="369">Pilcaniyeu</collectingRegion>
, province of 
<collectingRegion id="49B2F865FFFBFFB8B7B55A55FC84675C" box="[661,787,1880,1906]" country="Argentina" name="Rio Negro" pageId="2" pageNumber="369">Río Negro</collectingRegion>
, 
<collectingCounty id="62A84E0BFFFBFFB8B63F5A54FC03675C" box="[799,916,1881,1906]" pageId="2" pageNumber="369">Patagonia</collectingCounty>
<collectingCountry id="F3617617FFFBFFB8B6BC5A54FB8D675C" box="[924,1050,1881,1906]" name="Argentina" pageId="2" pageNumber="369">Argentina</collectingCountry>
, palaeolatitude ~ 
<geoCoordinate id="EE425040FFFBFFB8B1CC5A55FABA675C" box="[1260,1325,1880,1906]" degrees="46" direction="south" orientation="latitude" pageId="2" pageNumber="369" precision="55555" value="-46.0">46°S</geoCoordinate>
</materialsCitation>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36C650CFFFBFFB9B5B05A99FA9566B6" lastPageId="3" lastPageNumber="370" pageId="2" pageNumber="369" type="discussion">
<paragraph id="8BC93687FFFBFFB9B5B05A99FD07620B" blockId="2.[144,1443,1940,2042]" lastBlockId="3.[144,1444,221,1688]" lastPageId="3" lastPageNumber="370" pageId="2" pageNumber="369">
<emphasis id="B902EA95FFFBFFB9B5B05A99FD07620B" bold="true" lastPageId="3" lastPageNumber="370" pageId="2" pageNumber="369">
Discussion. The new specimen could be included into 
<taxonomicName id="4C764D04FFFBFFB8B6025A99FC476783" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[802,976,1940,1965]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="369" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
because they share some characters as the terminal kink of the CP very weak, not aligned with nodal Cr; nodal furrow reduced; ScP reaching costal very obliquely at nodus; nodal Cr sub-vertical; subnodus vertical; midfork symmetrical and recessed basally to a position between 12 and 26% of wing length; pterostigma elongate and broad; discoidal cell basally closed in hindwing, quite broad, distinctly widened distally; all secondary antenodal cross-veins between ScP and RA suppressed; antesubnodal space without cross-veins; cubito-anal area broad, with three rows of cells between CuA and posterior wing margin; nodus in the basal third of the wing, postnodal area very elongate; postnodal and postsubnodal cross-veins very numerous; and the petiole short and broad. The new genus could be distinguished from 
<taxonomicName id="4C764D04FFFAFFB9B1325C97FF4661F7" authority="Petrulevicius &amp; Nel, 2003" authorityName="Petrulevicius &amp; Nel" authorityYear="2003" class="Insecta" family="Frenguelliidae" genus="Frenguellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA95FFFAFFB9B1325C97FB0D619D" bold="true" box="[1042,1178,410,435]" italics="true" pageId="3" pageNumber="370">Frenguellia</emphasis>
<bibRefCitation id="EFE74B76FFFAFFB9B1865C94FF4661F7" author="Petrulevicius, J. F. &amp; Nel, A." pageId="3" pageNumber="370" pagination="2909 - 2918" refId="ref3978" refString="Petrulevicius, J. F. &amp; Nel, A. 2003. FrenguelliidAe, A new fAmily of drAgonflies from the eArliest Eocene of Argentina (Insecta: Odonata). Phylogenetic relationships within Odonata. Journal of Natural History 37: 2909 - 2918." type="journal article" year="2003">Petrulevičius &amp; Nel, 2003</bibRefCitation>
</taxonomicName>
, the other genus of 
<taxonomicName id="4C764D04FFFAFFB9B4C45CCDFD0261F7" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[484,661,448,473]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
, because it has the terminal kink of the CP not aligned with nodal Cr but in a slightly distal position (contra in a very distal position in 
<taxonomicName id="4C764D04FFFAFFB9B11A5CEBFB5561D1" authorityName="Petrulevicius &amp; Nel" authorityYear="2003" box="[1082,1218,486,511]" class="Insecta" family="Frenguelliidae" genus="Frenguellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA95FFFAFFB9B11A5CEBFB5561D1" bold="true" box="[1082,1218,486,511]" italics="true" pageId="3" pageNumber="370">Frenguellia</emphasis>
</taxonomicName>
); and the base of RP2 8 cells from nodus (contra 2-4 cells).
</emphasis>
</paragraph>
<paragraph id="8BC93687FFFAFFB9B5CE5F3FFC8062CD" blockId="3.[144,1444,221,1688]" pageId="3" pageNumber="370">
<emphasis id="B902EA95FFFAFFB9B5CE5F3FFC8062CD" bold="true" pageId="3" pageNumber="370">
The presence of a discoidal cell distinctly widened distally and a much less oblique and more transverse distal side (ddcv) like in the forewing of 
<taxonomicName id="4C764D04FFFAFFB9B6635F55FC5C625F" authorityName="Petrulevicius &amp; Nel" authorityYear="2003" box="[835,971,600,625]" class="Insecta" family="Frenguelliidae" genus="Frenguellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA95FFFAFFB9B6635F55FC5C625F" bold="true" box="[835,971,600,625]" italics="true" pageId="3" pageNumber="370">Frenguellia</emphasis>
</taxonomicName>
correspond to an apomorphy of the clade Epiproctophora 
<bibRefCitation id="EFE74B76FFFAFFB9B4BF5F73FDA862B9" author="Bechly, G." box="[415,575,638,663]" pageId="3" pageNumber="370" refId="ref3049" refString="Bechly, G. 1996. Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2: 402 pp. (Revised edition of the 1995 publication, with an English appendix including a new phylogenetic system of fossil and Recent Odonata)." type="book" year="1996">Bechly, 1996</bibRefCitation>
. Also, the presence of a CuP strongly curved and apparently beginning on AA (
<bibRefCitation id="EFE74B76FFFAFFB9B5CE5FAEFDAE6293" author="Petrulevicius, J. F. &amp; Nel, A." box="[238,569,675,701]" pageId="3" pageNumber="370" pagination="2909 - 2918" refId="ref3978" refString="Petrulevicius, J. F. &amp; Nel, A. 2003. FrenguelliidAe, A new fAmily of drAgonflies from the eArliest Eocene of Argentina (Insecta: Odonata). Phylogenetic relationships within Odonata. Journal of Natural History 37: 2909 - 2918." type="journal article" year="2003">Petrulevičius &amp; Nel, 2003</bibRefCitation>
), potentiAl synApomorphy of the group And Also discussed in 
<bibRefCitation id="EFE74B76FFFAFFB9B01A5FA9FE9F62CD" author="Fleck, G. &amp; Bechly, G. &amp; Martinez-Delclos, X. &amp; Jarzembowski, E. A. &amp; Nel, A." pageId="3" pageNumber="370" pagination="33 - 60" refId="ref3282" refString="Fleck, G.; Bechly, G.; Martinez-Delclos, X.; Jarzembowski, E. A. &amp; Nel, A. 2004. A revision of the Mesozoic dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Odonatoptera: Panodonata). Geodiversitas 26: 33 - 60." type="journal article" year="2004">
FlecK 
<emphasis id="B902EA95FFFAFFB9B0A95FA9FF3B62CD" bold="true" italics="true" pageId="3" pageNumber="370">et al.</emphasis>
(2004)
</bibRefCitation>
support its attribution to Epiproctophora.
</emphasis>
</paragraph>
<paragraph id="8BC93687FFFAFFB9B5CE5FFDFE7F6417" blockId="3.[144,1444,221,1688]" pageId="3" pageNumber="370">
<emphasis id="B902EA95FFFAFFB9B5CE5FFDFE7F6417" bold="true" pageId="3" pageNumber="370">
The base of the vein IR2 of 
<taxonomicName id="4C764D04FFFAFFB9B7585FFDFCB16327" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[632,806,752,777]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
is below the subnodus while it is nearly midway between the nodus and the arculus in all the Epiproctophora. This is probably an autapomorphy of 
<taxonomicName id="4C764D04FFFAFFB9B5B05E31FEAA637B" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[144,317,828,853]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">FrenguelliidAe</taxonomicName>
within this clAde (
<bibRefCitation id="EFE74B76FFFAFFB9B70B5E36FCE1637B" author="Petrulevicius, J. F. &amp; Nel, A." box="[555,886,827,853]" pageId="3" pageNumber="370" pagination="341 - 347" refId="ref4071" refString="Petrulevicius, J. F. &amp; Nel, A. 2007. Enigmatic and little known Odonata (Insecta) from the Paleogene of Patagonia and northwest Argentina. Annales de la Societe Entomologique de France (n. s.) 43: 341 - 347." type="journal article" year="2007">Petrulevičius &amp; Nel, 2007</bibRefCitation>
). This structure could be correlAted to the basal recession of the nodus. Also, the short petiolation together with the broadened wing and the sub-vertical nodal cross-vein and subnodus is a character present in some Zygoptera (
<taxonomicName id="4C764D04FFFAFFB9B1935E85FA08638F" baseAuthorityName="Tillyard &amp; Fraser" baseAuthorityYear="1938" box="[1203,1439,904,929]" class="Insecta" family="Thaumatoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Thaumatoneuridae</taxonomicName>
: Dysagrioninae) and in the Oligocene-Miocene family 
<taxonomicName id="4C764D04FFFAFFB9B63F5EA0FC2363E9" authorityName="Handlirsch" authorityYear="1907" box="[799,948,941,967]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
of enigmatic phylogenetic position, but considered Epiproctophora in several publications (
<bibRefCitation id="EFE74B76FFFAFFB9B6345EDEFC6A63C2" author="Fleck, G. &amp; Bechly, G. &amp; Martinez-Delclos, X. &amp; Jarzembowski, E. A. &amp; Nel, A." box="[788,1021,979,1004]" pageId="3" pageNumber="370" pagination="33 - 60" refId="ref3282" refString="Fleck, G.; Bechly, G.; Martinez-Delclos, X.; Jarzembowski, E. A. &amp; Nel, A. 2004. A revision of the Mesozoic dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Odonatoptera: Panodonata). Geodiversitas 26: 33 - 60." type="journal article" year="2004">
Fleck 
<emphasis id="B902EA95FFFAFFB9B6455EDEFC3063C2" bold="true" box="[869,935,979,1004]" italics="true" pageId="3" pageNumber="370">et al.</emphasis>
, 2004
</bibRefCitation>
; 
<bibRefCitation id="EFE74B76FFFAFFB9B12D5EDEFB6263C2" author="Nel, A. &amp; Fleck, G." box="[1037,1269,979,1004]" pageId="3" pageNumber="370" pagination="64 - 68" refId="ref3592" refString="Nel, A. &amp; Fleck, G. 2012. A new genus and species of Sieblosiidae from the Middle Miocene of Germany (Odonata: Epiproctophora). Zootaxa 3582: 64 - 68." type="journal article" year="2012">Nel &amp; Fleck, 2012</bibRefCitation>
). Differences with the 
<taxonomicName id="4C764D04FFFAFFB9B4225EF4FE00643D" authorityName="Handlirsch" authorityYear="1907" box="[258,407,1017,1043]" class="Insecta" family="Sieblosiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Sieblosiidae</taxonomicName>
are in the shape of the discoidal cells and position of the bases of RP3+4 and IR2 (
<bibRefCitation id="EFE74B76FFFAFFB9B5B75912FE4C6417" author="Petrulevicius, J. F. &amp; Nel, A." box="[151,475,1055,1081]" pageId="3" pageNumber="370" pagination="341 - 347" refId="ref4071" refString="Petrulevicius, J. F. &amp; Nel, A. 2007. Enigmatic and little known Odonata (Insecta) from the Paleogene of Patagonia and northwest Argentina. Annales de la Societe Entomologique de France (n. s.) 43: 341 - 347." type="journal article" year="2007">Petrulevičius &amp; Nel, 2007</bibRefCitation>
).
</emphasis>
</paragraph>
<paragraph id="8BC93687FFFAFFB9B5CE5948FEAB659A" blockId="3.[144,1444,221,1688]" pageId="3" pageNumber="370">
<emphasis id="B902EA95FFFAFFB9B5CE5948FEAB659A" bold="true" pageId="3" pageNumber="370">
The 
<taxonomicName id="4C764D04FFFAFFB9B40A5948FC356470" authority="Petrulevicius &amp; Nel, 2005 Also" authorityName="Petrulevicius &amp; Nel" authorityYear="2005" box="[298,930,1093,1119]" class="Insecta" family="Austroperilestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">
AustroperilestidAe 
<bibRefCitation id="EFE74B76FFFAFFB9B73F5948FCF16470" author="Petrulevicius, J. F. &amp; Nel, A." box="[543,870,1093,1119]" pageId="3" pageNumber="370" pagination="658 - 662" refId="ref4025" refString="Petrulevicius, J. F. &amp; Nel, A. 2005. AustroperilestidAe, A new fAmily of dAmselflies from the eArliest Eocene of Argentina (Insecta: Odonata). Phylogenetic relationships within odonata. Journal of Paleontology 79: 658 - 662." type="journal article" year="2005">Petrulevičius &amp; Nel, 2005</bibRefCitation>
Also
</taxonomicName>
present in the Eocene of PAtAgoniA Are based in a single wrinkled specimen that shows some shared characters with 
<taxonomicName id="4C764D04FFFAFFB9B17B5966FA9F64AA" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[1115,1288,1131,1156]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
which could indicate its synonymy with 
<taxonomicName id="4C764D04FFFAFFB9B4D4599CFD336484" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[500,676,1169,1194]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
. Only the discovery of new and more complete material is awaited to a fine comparison. Both families differ strongly in the characters of the discoidal cell placing 
<taxonomicName id="4C764D04FFFAFFB9B5B059D0FEEE64D8" authorityName="Petrulevicius &amp; Nel" authorityYear="2005" box="[144,377,1245,1270]" class="Insecta" family="Austroperilestidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">AustroperilestidAe</taxonomicName>
into the ZygopterA (
<bibRefCitation id="EFE74B76FFFAFFB9B7BF59D1FC6A64D8" author="Petrulevicius, J. F. &amp; Nel, A." box="[671,1021,1244,1270]" pageId="3" pageNumber="370" pagination="658 - 662" refId="ref4025" refString="Petrulevicius, J. F. &amp; Nel, A. 2005. AustroperilestidAe, A new fAmily of dAmselflies from the eArliest Eocene of Argentina (Insecta: Odonata). Phylogenetic relationships within odonata. Journal of Paleontology 79: 658 - 662." type="journal article" year="2005">Petrulevičius &amp; Nel, 2005</bibRefCitation>
). In this way, 
<taxonomicName id="4C764D04FFFAFFB9B1F259D0FE4B6532" authority="Petrulevicius &amp; Nel, 2005" authorityName="Petrulevicius &amp; Nel" authorityYear="2005" class="Insecta" family="Austroperilestidae" genus="Austroperilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B902EA95FFFAFFB9B1F259D0FA3364D8" bold="true" box="[1234,1444,1245,1270]" italics="true" pageId="3" pageNumber="370">Austroperilestes</emphasis>
<bibRefCitation id="EFE74B76FFFAFFB9B5B0580FFE4B6532" author="Petrulevicius, J. F. &amp; Nel, A." box="[144,476,1282,1308]" pageId="3" pageNumber="370" pagination="658 - 662" refId="ref4025" refString="Petrulevicius, J. F. &amp; Nel, A. 2005. AustroperilestidAe, A new fAmily of dAmselflies from the eArliest Eocene of Argentina (Insecta: Odonata). Phylogenetic relationships within odonata. Journal of Paleontology 79: 658 - 662." type="journal article" year="2005">Petrulevičius &amp; Nel, 2005</bibRefCitation>
</taxonomicName>
could be differentiAted with 
<taxonomicName id="4C764D04FFFAFFB9B64A580EFB906532" authorityName="Petrulevičius" authorityYear="2017" box="[874,1031,1283,1308]" class="Insecta" family="Frenguelliidae" genus="Treintamilun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B902EA95FFFAFFB9B64A580EFB906532" bold="true" box="[874,1031,1283,1308]" italics="true" pageId="3" pageNumber="370">Treintamilun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157EEFFFAFFB9B133580EFB156532" box="[1043,1154,1283,1308]" pageId="3" pageNumber="370" rank="genus">gen. nov.</taxonomicNameLabel>
by having the arcular cross-vein reaching RP+MA basal to its separation (contra after RP separation in 
<taxonomicName id="4C764D04FFFAFFB9B1A45824FAB7656C" authorityName="Petrulevičius" authorityYear="2017" box="[1156,1312,1321,1346]" class="Insecta" family="Frenguelliidae" genus="Treintamilun" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B902EA95FFFAFFB9B1A45824FAB7656C" bold="true" box="[1156,1312,1321,1346]" italics="true" pageId="3" pageNumber="370">Treintamilun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157EEFFFAFFB9B0095824FA00656C" box="[1321,1431,1321,1346]" pageId="3" pageNumber="370" rank="genus">gen. nov.</taxonomicNameLabel>
); the RP and MA strongly approximated at their base in discoidal quadrangle (contra very distant); the pterostigma braced (contra not braced); and a curved vein between and near MA and ddcv forming a little triangle.
</emphasis>
</paragraph>
<paragraph id="8BC93687FFFAFFB9B5CE58CCFA9566B6" blockId="3.[144,1444,221,1688]" pageId="3" pageNumber="370">
<emphasis id="B902EA95FFFAFFB9B5CE58CCFA9566B6" bold="true" pageId="3" pageNumber="370">
If 
<taxonomicName id="4C764D04FFFAFFB9B43C58CCFE5E65F4" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[284,457,1473,1498]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
are Epiproctophorans, they are not Euepiproctophorans (
<taxonomicName id="4C764D04FFFAFFB9B1F058CCFA1265F4" baseAuthorityName="Tillyard" baseAuthorityYear="1917" box="[1232,1413,1473,1498]" class="Insecta" family="Epiophlebiidae" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Epiophlebiidae</taxonomicName>
+ Anisopteromorpha 
<bibRefCitation id="EFE74B76FFFAFFB9B45B58EAFD8B662E" author="Bechly, G." box="[379,540,1511,1536]" pageId="3" pageNumber="370" refId="ref3049" refString="Bechly, G. 1996. Morphologische Untersuchungen am Flugelgeader der rezenten Libellen und deren Stammgruppenvertreter (Insecta; Pterygota; Odonata), unter besonderer Berucksichtigung der Phylogenetischen Systematik und des Grundplanes der Odonata. Petalura, Special Volume 2: 402 pp. (Revised edition of the 1995 publication, with an English appendix including a new phylogenetic system of fossil and Recent Odonata)." type="book" year="1996">Bechly, 1996</bibRefCitation>
) because they have a different, derived pattern of alternating width of wing spaces between the longitudinal veins, and a tendency to a triangular hind wing discoidal cell traversed by one cross-vein. The 
<taxonomicName id="4C764D04FFFAFFB9B76D5B3FFD6C6665" authorityName="Petrulevicius and Nel" authorityYear="2003" box="[589,763,1586,1611]" class="Insecta" family="Frenguelliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="370" phylum="Arthropoda" rank="family">Frenguelliidae</taxonomicName>
are probably in a very inclusive position within the Epiproctophora, either sister group of all other Epiprotophora or sister group of the Isophlebioptera, As it shAres some potentiAl synApomorphies with this lAst group (
<bibRefCitation id="EFE74B76FFFAFFB9B6905B73FB6266B6" author="Petrulevicius, J. F. &amp; Nel, A." box="[944,1269,1662,1688]" pageId="3" pageNumber="370" pagination="341 - 347" refId="ref4071" refString="Petrulevicius, J. F. &amp; Nel, A. 2007. Enigmatic and little known Odonata (Insecta) from the Paleogene of Patagonia and northwest Argentina. Annales de la Societe Entomologique de France (n. s.) 43: 341 - 347." type="journal article" year="2007">Petrulevičius &amp; Nel, 2007</bibRefCitation>
).
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>