312 lines
49 KiB
XML
312 lines
49 KiB
XML
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<mods:title id="0DAD6CAFBF56E1196CF58C8DF75342F7">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="DF1B6E495A2E3E409B420B34F17B4E4C">Inkhavilay, Khamla</mods:namePart>
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<mods:nameIdentifier id="FBF5033D99AEE2CB47F02FB2F43972B3" type="email">kinkhavilay@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0796DF146ABCF54FCA0378C8D801BD2D">Sutcharit, Chirasak</mods:namePart>
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<mods:nameIdentifier id="E5EB1BAEB3D527EA29282DC0B19630CB" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="22A46365D0A9FE0986F090C202794E47">Panha, Somsak</mods:namePart>
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<mods:title id="BF3D42DA4188CAAE967926F08A716A0C">European Journal of Taxonomy</mods:title>
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<treatment id="03DAD35F6939EF29FE30A9E6FEA02644" ID-DOI="http://doi.org/10.5281/zenodo.3848209" ID-GBIF-Taxon="164161056" ID-Zenodo-Dep="3848209" LSID="urn:lsid:plazi:treatment:03DAD35F6939EF29FE30A9E6FEA02644" httpUri="http://treatment.plazi.org/id/03DAD35F6939EF29FE30A9E6FEA02644" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
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<subSubSection id="C36931C26939EF17FE30A9E6FBB12391" box="[443,1145,1752,1779]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BCC62496939EF17FE30A9E6FBB12391" blockId="31.[443,1145,1752,1816]" box="[443,1145,1752,1779]" pageId="31" pageNumber="32">
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<heading id="D084D5256939EF17FE30A9E6FBB12391" box="[443,1145,1752,1779]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="32" reason="2">
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<taxonomicName id="4C7319CA6939EF17FE30A9E6FBB12391" ID-CoL="D2LH" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[443,1145,1752,1779]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fuscolabris" subGenus="Syndromus">
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<emphasis id="B907BE5B6939EF17FE30A9E6FC512390" bold="true" box="[443,921,1752,1779]" pageId="31" pageNumber="32">
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<emphasis id="B907BE5B6939EF17FE30A9E6FDA02390" bold="true" box="[443,616,1752,1779]" italics="true" pageId="31" pageNumber="32">Amphidromus</emphasis>
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(
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<emphasis id="B907BE5B6939EF17FDF2A9E6FCCA2390" bold="true" box="[633,770,1752,1779]" italics="true" pageId="31" pageNumber="32">Syndromus</emphasis>
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)
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<emphasis id="B907BE5B6939EF17FC98A9E6FC512390" bold="true" box="[787,921,1752,1779]" italics="true" pageId="31" pageNumber="32">fuscolabris</emphasis>
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</emphasis>
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<bibRefCitation id="EFE21FB86939EF17FC2BA9E6FBB12391" author="Mollendorff O. F. von" box="[928,1145,1752,1778]" pageId="31" pageNumber="32" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26939EF17FD85A9C0FBEE227B" box="[526,1062,1790,1816]" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BCC62496939EF17FD85A9C0FBEE227B" blockId="31.[443,1145,1752,1816]" box="[526,1062,1790,1816]" pageId="31" pageNumber="32">
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<figureCitation id="13487ECC6939EF17FD85A9C0FD9F227B" box="[526,599,1790,1816]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="31" pageNumber="32">Figs 1</figureCitation>
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,
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<figureCitation id="13487ECC6939EF17FDEEA9C0FDBA227B" box="[613,626,1790,1816]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="31" pageNumber="32">9</figureCitation>
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E–F, 12G–I, 13J–M, 14C–D;
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<tableCitation id="C6F157F26939EF17FC44A9C0FBEE227B" box="[975,1062,1790,1816]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="31" pageNumber="32" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
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</paragraph>
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<subSubSection id="C36931C26939EF17FF36A87FFC532288" pageId="31" pageNumber="32" type="reference_group">
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<treatmentCitationGroup id="AB6345676939EF17FF36A87FFD5E22E1" pageId="31" pageNumber="32">
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<treatmentCitation id="0AD244586939EF17FF36A87FFCBA223F" author="Mollendorff O. F. von" box="[189,882,1857,1884]" page="75" pageId="31" pageNumber="32" year="1898">
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<taxonomicName id="4C7319CA6939EF17FF36A87FFCBA223F" ID-CoL="D2LH" authority="Mollendorff, 1898: 75" authorityName="Mollendorff" authorityPageNumber="75" authorityYear="1898" box="[189,882,1857,1884]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="zebrinus" subSpecies="fuscolabris">
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<emphasis id="B907BE5B6939EF17FF36A87FFD932238" box="[189,603,1857,1883]" italics="true" pageId="31" pageNumber="32">Amphidromus zebrinus fuscolabris</emphasis>
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<bibRefCitation id="EFE21FB86939EF17FDEDA87FFCBA223F" author="Mollendorff O. F. von" box="[614,882,1857,1884]" pageId="31" pageNumber="32" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898: 75</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="54C8DCEB6939EF17FC08A87CFC75223F" box="[899,957,1858,1884]" pageId="31" pageNumber="32">Type</typeStatus>
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locality: Boloven (Boloven Plateau, Paksong District,
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<collectingRegion id="49B7ACAB6939EF17FE3FA859FD8822E2" box="[436,576,1895,1921]" country="Laos" name="Champasak" pageId="31" pageNumber="32">Champasak</collectingRegion>
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,
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<collectingCountry id="F36422D96939EF17FDC7A856FD4222E1" box="[588,650,1896,1922]" name="Laos" pageId="31" pageNumber="32">Laos</collectingCountry>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCC62496939EF17FF36A895FC532288" blockId="31.[189,1398,1963,2027]" pageId="31" pageNumber="32">
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<treatmentCitationGroup id="AB6345676939EF17FF36A895FC532288" pageId="31" pageNumber="32">
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<taxonomicName id="4C7319CA6939EF17FF36A895FD9322A6" ID-CoL="D2LH" authorityName="Mollendorff" authorityYear="1898" box="[189,603,1963,1989]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="zebrinus" subSpecies="fuscolabris">
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<emphasis id="B907BE5B6939EF17FF36A895FD9322A6" box="[189,603,1963,1989]" italics="true" pageId="31" pageNumber="32">Amphidromus zebrinus fuscolabris</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AD244586939EF17FD0BA895FC5E22A6" author="Pilsbry H. A." box="[640,918,1963,1989]" page="199" pageId="31" pageNumber="32" year="1900">
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<bibRefCitation id="EFE21FB86939EF17FD0BA895FC5E22A6" author="Pilsbry H. A." box="[640,918,1963,1989]" pageId="31" pageNumber="32" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 199–200</bibRefCitation>
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</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AD244586939EF17FC47A895FB5922A6" author="Zilch A." box="[972,1169,1963,1990]" page="134" pageId="31" pageNumber="32" year="1953">
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<bibRefCitation id="EFE21FB86939EF17FC47A895FB5922A6" author="Zilch A." box="[972,1169,1963,1990]" pageId="31" pageNumber="32" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch 1953: 134</bibRefCitation>
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</treatmentCitation>
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||
, pl. 23, fig. 22. —
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||
<treatmentCitation id="0AD244586939EF17FF68A8EFFDFC2288" author="Laidlaw F. F. & Solem A." box="[227,564,2000,2027]" page="564" pageId="31" pageNumber="32" year="1961">
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<bibRefCitation id="EFE21FB86939EF17FF68A8EFFDFC2288" author="Laidlaw F. F. & Solem A." box="[227,564,2000,2027]" pageId="31" pageNumber="32" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. & Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw & Solem 1961: 564</bibRefCitation>
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</treatmentCitation>
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||
, 621. —
|
||
<treatmentCitation id="0AD244586939EF17FD14A8EFFC5F2288" author="Richardson L." box="[671,919,2000,2027]" page="49" pageId="31" pageNumber="32" year="1985">
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<bibRefCitation id="EFE21FB86939EF17FD14A8EFFC5F2288" author="Richardson L." box="[671,919,2000,2027]" pageId="31" pageNumber="32" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 49</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36931C26906EF28FF36AE35FD672738" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BCC62496906EF28FF36AE35FE622446" blockId="32.[189,426,267,293]" box="[189,426,267,293]" pageId="32" pageNumber="33">
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<heading id="D084D5256906EF28FF36AE35FE622446" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
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<emphasis id="B907BE5B6906EF28FF36AE35FE622446" bold="true" box="[189,426,267,293]" pageId="32" pageNumber="33">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCC62496906EF28FF61AE03FE512435" blockId="32.[189,643,316,381]" box="[234,409,316,343]" pageId="32" pageNumber="33">
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<emphasis id="B907BE5B6906EF28FF61AE03FE512435" bold="true" box="[234,409,316,343]" pageId="32" pageNumber="33">
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<typeStatus id="54C8DCEB6906EF28FF61AE03FEEF2434" box="[234,295,317,343]" pageId="32" pageNumber="33">Type</typeStatus>
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material
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</emphasis>
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</paragraph>
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<paragraph id="8BCC62496906EF28FF36AE5DFD4B241E" blockId="32.[189,643,316,381]" box="[189,643,355,381]" pageId="32" pageNumber="33">
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<materialsCitation id="3B1B68146906EF28FF36AE5DFD4B241E" ID-GBIF-Occurrence="2620312302" box="[189,643,355,381]" collectingDate="2013-01" collectionCode="SMF" country="Laos" location="LAOS" pageId="32" pageNumber="33" specimenCode="SMF 7641" specimenCount="1" typeStatus="holotype">
|
||
<collectingCountry id="F36422D96906EF28FF36AE5DFEC6241E" box="[189,270,355,381]" name="Laos" pageId="32" pageNumber="33">LAOS</collectingCountry>
|
||
:
|
||
<typeStatus id="54C8DCEB6906EF28FE92AE5DFE48241E" box="[281,384,355,381]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
|
||
<specimenCode id="DBD5CA326906EF28FE0CAE5DFDCC241E" box="[391,516,355,381]" pageId="32" pageNumber="33">SMF 7641</specimenCode>
|
||
(
|
||
<figureCitation id="13487ECC6906EF28FD9FAE5DFDBD241E" box="[532,629,355,381]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="32" pageNumber="33">Fig. 13I</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF61AE99FE6E24A2" blockId="32.[189,1399,423,603]" box="[234,422,423,449]" pageId="32" pageNumber="33">
|
||
<emphasis id="B907BE5B6906EF28FF61AE99FE6E24A2" bold="true" box="[234,422,423,449]" pageId="32" pageNumber="33">Other material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36AEF0FD672738" blockId="32.[189,1399,423,603]" pageId="32" pageNumber="33">
|
||
<collectingCountry id="F36422D96906EF28FF36AEF0FEC6248B" box="[189,270,462,488]" name="Laos" pageId="32" pageNumber="33">LAOS</collectingCountry>
|
||
: 34 shells, typical form (
|
||
<figureCitation id="13487ECC6906EF28FDB7AEF0FD58248B" box="[572,656,462,488]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="32" pageNumber="33">Fig. 13</figureCitation>
|
||
J–K,
|
||
<collectionCode id="ED62FA8C6906EF28FD58AEF0FCE0248B" box="[723,808,462,488]" country="0" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
|
||
7040);
|
||
<specimenCount id="9D75A9C06906EF28FC03AEF0FBD5248B" box="[904,1053,462,488]" pageId="32" pageNumber="33" type="generic">2 specimens</specimenCount>
|
||
, in ethanol (
|
||
<figureCitation id="13487ECC6906EF28FB3FAEF0FADD248B" box="[1204,1301,462,488]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="32" pageNumber="33">Figs 9E</figureCitation>
|
||
,
|
||
<figureCitation id="13487ECC6906EF28FAAFAEF0FA88248B" box="[1316,1344,462,488]" captionStart="Fig" captionStartId="24.[189,232,1824,1850]" captionTargetBox="[194,1391,713,1777]" captionTargetId="figure@24.[189,1398,706,1783]" captionTargetPageId="24" captionText="Fig. 12. SEM images of the radula. A–C. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). D–F. Amphidromus xiengensis Morlet, 1891 from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037). G–I. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="32" pageNumber="33">12</figureCitation>
|
||
G–I, 14C−D,
|
||
<collectionCode id="ED62FA8C6906EF28FEAEAECAFEB2276D" box="[293,378,500,526]" country="0" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
|
||
7041); 73 +
|
||
<specimenCount id="9D75A9C06906EF28FD90AECAFD67276D" box="[539,687,500,526]" pageId="32" pageNumber="33" type="juvenile">10 juveniles</specimenCount>
|
||
, yellowish form (
|
||
<figureCitation id="13487ECC6906EF28FC0EAECAFC11276D" box="[901,985,500,526]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="32" pageNumber="33">Fig. 13</figureCitation>
|
||
L–M,
|
||
<collectionCode id="ED62FA8C6906EF28FBACAECAFBB4276D" box="[1063,1148,500,526]" country="0" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
|
||
7042);
|
||
<specimenCount id="9D75A9C06906EF28FB57AECAFABA276D" box="[1244,1394,500,526]" pageId="32" pageNumber="33" type="generic">2 specimens</specimenCount>
|
||
, in ethanol (
|
||
<figureCitation id="13487ECC6906EF28FECDAD25FE562756" box="[326,414,539,565]" captionStart="Fig" captionStartId="20.[189,232,1823,1849]" captionTargetBox="[189,1399,656,1782]" captionTargetId="figure@20.[189,1399,656,1782]" captionTargetPageId="20" captionText="Fig. 9. Living snails of Amphidromus (Syndromus) spp. A. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapeu, Laos (CUMZ 7023). B. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). C–D. Amphidromus xiengensis Morlet, 1891 from Thad Kacham, Luang Phrabang, Laos (CUMZ 7035), the typical form, and var. “multifasciata” Fulton, 1896, respectively. E–F. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041–7042). E. Typical color form. F. Monochrome yellowish color form." figureDoi="http://doi.org/10.5281/zenodo.3829762" httpUri="https://zenodo.org/record/3829762/files/figure.png" pageId="32" pageNumber="33">Fig. 9F</figureCitation>
|
||
,
|
||
<collectionCode id="ED62FA8C6906EF28FE27AD25FDC92756" box="[428,513,539,565]" country="0" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
|
||
7043),
|
||
<collectingRegion id="49B7ACAB6906EF28FDD4AD25FD2C2756" box="[607,740,539,565]" country="Laos" name="Xekong" pageId="32" pageNumber="33">Ban Phone</collectingRegion>
|
||
, La-Marm District,
|
||
<collectingRegion id="49B7ACAB6906EF28FC5FAD25FBF82756" box="[980,1072,539,565]" country="Laos" name="Xekong" pageId="32" pageNumber="33">Sekong</collectingRegion>
|
||
; 2 shells, Ban Xai Na Pho, Phatumphone,
|
||
<collectingRegion id="49B7ACAB6906EF28FEE6AD7FFE3E2738" box="[365,502,577,603]" country="Laos" name="Champasak" pageId="32" pageNumber="33">Champasak</collectingRegion>
|
||
(
|
||
<collectionCode id="ED62FA8C6906EF28FD8EAD7FFD942738" box="[517,604,577,603]" country="0" name="Cameroon University, Museum of Zoology" pageId="32" pageNumber="33">CUMZ</collectionCode>
|
||
7044).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26906EF28FF36ADBBFB9823B8" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BCC62496906EF28FF36ADBBFE8627FC" blockId="32.[189,334,645,671]" box="[189,334,645,671]" pageId="32" pageNumber="33">
|
||
<heading id="D084D5256906EF28FF36ADBBFE8627FC" bold="true" box="[189,334,645,671]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
|
||
<emphasis id="B907BE5B6906EF28FF36ADBBFE8627FC" bold="true" box="[189,334,645,671]" pageId="32" pageNumber="33">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36AD89FB5926D4" blockId="32.[189,1399,695,951]" pageId="32" pageNumber="33">Shell sinistral, elongate conical, rather solid and glossy. Spire elongate conical; apex acute, with pinkish tint to brownish without black spot on tip. Whorls 6 to 7 nearly smooth; suture wide and shallow; last whorl rounded. Periostracum transparent to thin corneous, with greenish streaks on lower half of last whorl. Last whorl shows yellow to orange Band 1; Bands 2 to 5 merged and become six or seven black or brown slanted blotches; Band 6 yellow; varix absent. Parietal callus thickened, bright purplishpink. Aperture elongated auriform; peristome thickened, expanded and not reflected; lip purplish-pink. Columella rather thick, straight, dilated and purplish-pink. Umbilicus imperforated.</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF61ACDCFE8D269F" blockId="32.[189,1399,994,1136]" box="[234,325,994,1020]" pageId="32" pageNumber="33">
|
||
<emphasis id="B907BE5B6906EF28FF61ACDCFE8D269F" bold="true" box="[234,325,994,1020]" pageId="32" pageNumber="33">Radula</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36AB36FD01210C" blockId="32.[189,1399,994,1136]" pageId="32" pageNumber="33">
|
||
Each row contains about 161 (80-(35-31)-1-(31-35)-80) teeth. Central tooth unicuspid, with curved cusp. Lateral and marginal teeth similar to that described in
|
||
<taxonomicName id="4C7319CA6906EF28FC29AB11FBFC212A" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[930,1076,1071,1097]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="areolatus">
|
||
<emphasis id="B907BE5B6906EF28FC29AB11FBFC212A" box="[930,1076,1071,1097]" italics="true" pageId="32" pageNumber="33">A. areolatus</emphasis>
|
||
</taxonomicName>
|
||
. Marginal teeth start from lateral tooth 17 to 21 outwards (
|
||
<figureCitation id="13487ECC6906EF28FDBFAB6BFD40210C" box="[564,648,1109,1135]" captionStart="Fig" captionStartId="24.[189,232,1824,1850]" captionTargetBox="[194,1391,713,1777]" captionTargetId="figure@24.[189,1398,706,1783]" captionTargetPageId="24" captionText="Fig. 12. SEM images of the radula. A–C. Amphidromus flavus (Pfeiffer, 1861) from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7027). D–F. Amphidromus xiengensis Morlet, 1891 from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7037). G–I. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos (CUMZ 7041). A, D, G = central tooth with the first to fifth or sixth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829768" httpUri="https://zenodo.org/record/3829768/files/figure.png" pageId="32" pageNumber="33">Fig. 12</figureCitation>
|
||
G−I).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF61ABA4FE6C21D7" blockId="32.[189,1398,1178,1357]" box="[234,420,1178,1204]" pageId="32" pageNumber="33">
|
||
<emphasis id="B907BE5B6906EF28FF61ABA4FE6C21D7" bold="true" box="[234,420,1178,1204]" pageId="32" pageNumber="33">Genital organs</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36ABFEFC8E202E" blockId="32.[189,1398,1178,1357]" pageId="32" pageNumber="33">
|
||
Atrium (at) rather long (n = 10). Penis (p) long, cylindrical and slightly swollen at distal end. Epiphallus (e) short, half of penis length; flagellum (fl) of about same length as epiphallus and terminated with pointed tip. Appendix absent. Penial retractor muscle (pr) thin and long. Vas deferens (vd) very narrow tube connecting epiphallus and free oviduct (
|
||
<figureCitation id="13487ECC6906EF28FD46AA0DFCF1202E" box="[717,825,1331,1357]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="32" pageNumber="33">Fig. 14C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36AA46FB9E20DB" blockId="32.[189,1399,1400,1465]" pageId="32" pageNumber="33">
|
||
Internal penial wall corrugated into strong penial pilasters, which form fringe around penial verge. Penial verge conical, with smooth surface and orifice open near tip (
|
||
<figureCitation id="13487ECC6906EF28FC50AAA0FB8120DB" box="[987,1097,1438,1464]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="32" pageNumber="33">Fig. 14D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36AADDFDC12313" blockId="32.[189,1399,1507,1648]" pageId="32" pageNumber="33">
|
||
Vagina cylindrical, long, slender, about two times as long as penis. Gametolytic duct long, proximal to genital orifice of a relatively larger diameter than vagina and distally tapering; distal to genital orifice small tube of about same length as proximal part and connected to gametolytic sac. Oviduct and albumin gland very small (
|
||
<figureCitation id="13487ECC6906EF28FE1AA968FE342313" box="[401,508,1622,1648]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="32" pageNumber="33">Fig. 14C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36A9A4FB9823B8" blockId="32.[189,1399,1690,1755]" pageId="32" pageNumber="33">
|
||
Internally, vaginal wall shows longitudinal vaginal pilasters (vp); proximal to genital orifice with thin wall and smooth surface, distally pilasters show continuous ridges (
|
||
<figureCitation id="13487ECC6906EF28FC5EA9FFFB8B23B8" box="[981,1091,1729,1755]" captionStart="Fig" captionStartId="29.[189,232,1858,1884]" captionTargetBox="[189,1399,310,1811]" captionTargetId="figure@29.[189,1399,310,1811]" captionTargetPageId="29" captionText="Fig. 14. Genitalia of Amphidromus (Syndromus) spp. A–B. Amphidromus xiengensis Morlet, 1891 from Luang Phrabang showing the reproductive system and interior structures of the penis and vaginal chamber (CUMZ 7035). C–D. Amphidromus fuscolabris Möllendorff, 1898 from Ban Phone, Sekong, Laos showing the general characteristics of the genital system and the interior structures of the penis, atrium and vaginal chamber (CUMZ 7041)." figureDoi="http://doi.org/10.5281/zenodo.3829772" httpUri="https://zenodo.org/record/3829772/files/figure.png" pageId="32" pageNumber="33">Fig. 14D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26906EF28FF36A83BFABF2288" pageId="32" pageNumber="33" type="distribution">
|
||
<paragraph id="8BCC62496906EF28FF36A83BFE9E227C" blockId="32.[189,342,1797,1823]" box="[189,342,1797,1823]" pageId="32" pageNumber="33">
|
||
<heading id="D084D5256906EF28FF36A83BFE9E227C" bold="true" box="[189,342,1797,1823]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
|
||
<emphasis id="B907BE5B6906EF28FF36A83BFE9E227C" bold="true" box="[189,342,1797,1823]" pageId="32" pageNumber="33">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496906EF28FF36A809FABF2288" blockId="32.[189,1399,1847,2027]" pageId="32" pageNumber="33">
|
||
The original description of this species was given by
|
||
<bibRefCitation id="EFE21FB86906EF28FCDBA809FBF12232" author="Mollendorff O. F. von" box="[848,1081,1847,1873]" pageId="32" pageNumber="33" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff (1898)</bibRefCitation>
|
||
and the
|
||
<typeStatus id="54C8DCEB6906EF28FB23A806FB132231" box="[1192,1243,1848,1874]" pageId="32" pageNumber="33">type</typeStatus>
|
||
locality was Boloven,
|
||
<collectingCountry id="F36422D96906EF28FEB9A860FEA7221B" box="[306,367,1886,1912]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
|
||
. Unfortunately, we couldn’t obtain any specimen from the
|
||
<typeStatus id="54C8DCEB6906EF28FBB6A860FBB8221B" box="[1085,1136,1886,1912]" pageId="32" pageNumber="33">type</typeStatus>
|
||
locality in our recent survey. The specimens were collected from a mountain at
|
||
<collectingRegion id="49B7ACAB6906EF28FCEAA8BAFC2822FD" box="[865,992,1924,1950]" country="Laos" name="Xekong" pageId="32" pageNumber="33">Ban Phone</collectingRegion>
|
||
(Phone village),
|
||
<collectingRegion id="49B7ACAB6906EF28FB23A8BAFABA22FD" box="[1192,1394,1924,1950]" country="Laos" name="Xekong" pageId="32" pageNumber="33">Sekong Province</collectingRegion>
|
||
, and Ban Xai Na Pho (Xai Na Pho village) Phatumphone,
|
||
<collectingRegion id="49B7ACAB6906EF28FCD5A894FC2222A7" box="[862,1002,1962,1988]" country="Laos" name="Champasak" pageId="32" pageNumber="33">Champasak</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36422D96906EF28FC7EA895FBF922A6" box="[1013,1073,1963,1989]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
|
||
, which are both located at a lower altitude than the Boloven Plateau. It seems that this species is only found in low altitude habitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36931C26907EF29FF36AE34FEA02644" pageId="33" pageNumber="34" type="discussion">
|
||
<paragraph id="8BCC62496907EF29FF36AE34FEE72447" blockId="33.[189,303,266,292]" box="[189,303,266,292]" pageId="33" pageNumber="34">
|
||
<heading id="D084D5256907EF29FF36AE34FEE72447" bold="true" box="[189,303,266,292]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
|
||
<emphasis id="B907BE5B6907EF29FF36AE34FEE72447" bold="true" box="[189,303,266,292]" pageId="33" pageNumber="34">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCC62496907EF29FF36AE04FEF5274D" blockId="33.[189,1399,314,558]" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C7319CA6907EF29FF36AE04FE242437" authorityName="Mollendorff" authorityYear="1898" box="[189,492,314,340]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="fuscolabris">
|
||
<emphasis id="B907BE5B6907EF29FF36AE04FE242437" box="[189,492,314,340]" italics="true" pageId="33" pageNumber="34">Amphidromus fuscolabris</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7319CA6907EF29FD06AE04FCDB2437" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[653,787,314,340]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="zebrinus">
|
||
<emphasis id="B907BE5B6907EF29FD06AE04FCDB2437" box="[653,787,314,340]" italics="true" pageId="33" pageNumber="34">A. zebrinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7319CA6907EF29FCDBAE04FC732437" authorityName="Ancey" authorityYear="1897" box="[848,955,314,340]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="eudeli">
|
||
<emphasis id="B907BE5B6907EF29FCDBAE04FC732437" box="[848,955,314,340]" italics="true" pageId="33" pageNumber="34">A. eudeli</emphasis>
|
||
</taxonomicName>
|
||
in having a large and elongate shell, apex tinted pink, purplish-pink parietal callus, Band 1 yellowish and Bands 2 to 5 merged and becoming 6 to 7 slanted blotches (on last whorl). In comparison,
|
||
<taxonomicName id="4C7319CA6907EF29FCC2AEBDFC0624FF" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[841,974,386,413]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="zebrinus">
|
||
<emphasis id="B907BE5B6907EF29FCC2AEBDFC0624FF" box="[841,974,386,413]" italics="true" pageId="33" pageNumber="34">A. zebrinus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFE21FB86907EF29FB84AEBCFACD24FE" author="Sutcharit C. & Ablett J. & Tongkerd P. & Naggs F. & Panha S." box="[1039,1285,386,413]" pageId="33" pageNumber="34" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. & Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
|
||
Sutcharit
|
||
<emphasis id="B907BE5B6907EF29FB09AEBDFB7E24FF" box="[1154,1206,386,412]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
|
||
. 2015
|
||
</bibRefCitation>
|
||
: fig. 15k) has a smaller shell, Band 1 absent, Bands 2 and 5 merged and becoming fifteen narrow slanted stripes on last whorl, and reddish band in between Bands 5 and 6. On the other hand,
|
||
<taxonomicName id="4C7319CA6907EF29FBA9AEF2FB422486" authorityName="Ancey" authorityYear="1897" box="[1058,1162,459,486]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="eudeli">
|
||
<emphasis id="B907BE5B6907EF29FBA9AEF2FB422486" box="[1058,1162,459,486]" italics="true" pageId="33" pageNumber="34">A. eudeli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13487ECC6907EF29FB12AEF5FACC2485" box="[1177,1284,459,486]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13N</figureCitation>
|
||
,
|
||
<typeStatus id="54C8DCEB6907EF29FA84AEF5FABE2486" box="[1295,1398,459,485]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
RBINS 617427) has a smaller shell, thin parietal callus, Bands 2 and 3 merged and Bands 4 and 5 well developed.
|
||
</paragraph>
|
||
<paragraph id="8BCC62496907EF29FF36AD66FEA02644" blockId="33.[189,1398,599,808]" pageId="33" pageNumber="34">
|
||
Two color forms were observed from
|
||
<collectingRegion id="49B7ACAB6907EF29FD0DAD69FD292712" box="[646,737,599,625]" country="Laos" name="Xekong" pageId="33" pageNumber="34">Sekong</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36422D96907EF29FD64AD66FCE42711" box="[751,812,600,626]" name="Laos" pageId="33" pageNumber="34">Laos</collectingCountry>
|
||
. These are the typical form (
|
||
<figureCitation id="13487ECC6907EF29FB0CAD69FB132712" box="[1159,1243,599,625]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13</figureCitation>
|
||
I–K) and the yellowish form that is monochrome yellowish, with Bands 2 to 5 absent and stained with pale yellowish color, and Band 6 sometimes present as a brownish color (
|
||
<figureCitation id="13487ECC6907EF29FCD5AD9EFC6627D9" box="[862,942,672,698]" captionStart="Fig" captionStartId="28.[189,232,1613,1639]" captionTargetBox="[263,1324,265,1582]" captionTargetId="figure@28.[263,1324,265,1582]" captionTargetPageId="28" captionText="Fig. 13. Shells of Amphidromus (Syndromus) spp. A−C. Amphidromus xiengensis Morlet, 1891. A. Lectotype (MNHN-IM-2000-5249). B. Specimen from Thailand (CUMZ 7050). C. Specimen from Tam Chiang Dao, Chiangmai, Thailand (CUMZ 7034). D−F. Amphidromus xiengensis var. “clausus” Pilsbry, 1900. D. Lectotype (ANSP 31496). E. Specimen from Ban Na Deauy, Luang Phrabang, Laos (CUMZ 7036). F. Specimen from Pha Tang, Prayao, Thailand (CUMZ 7052). G–H. Amphidromus xiengensis var. “multifasciata” Fulton, 1896 from Phu Nang National Park, Phayao, Thailand (CUMZ 7038). I−M. Amphidromus fuscolabris Möllendorff, 1898. I. Holotype (SMF 7641). J−M. Specimens from Ban Phon, Sekong, Laos (CUMZ 7041–7042). N. Amphidromus eudeli Ancey, 1897, syntype from Binh Dinh, Annam (RBINS 617427). O–P. Amphidromus haematostoma Möllendorff, 1898. O. Lectotype (SMF 7559). P. Paralectotype (SMF 7560). Q–R. Amphidromus haematostoma var. “varians” Möllendorff, 1898. Q. Lectotype (SMF 7561). R. Paralectotype (SMF 7562). S–T. Amphidromus xiengkhaungensis sp. nov. S. Holotype (CUMZ 7045). T. Paratype (CUMZ 7046)." figureDoi="http://doi.org/10.5281/zenodo.3829770" httpUri="https://zenodo.org/record/3829770/files/figure.png" pageId="33" pageNumber="34">Fig. 13</figureCitation>
|
||
L–M). However, these two color forms show a pinkish to purplish-pink parietal callus and columella, and have identical genital structures and sculptures inside the penis and vagina. In addition, they occur sympatrically and so we recognize them as conspecific.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |