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<mods:title id="1985FE151DE1899E90D9E96C4A27D94E">Taxonomic review of the tree snail genus Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Laos, with the description of two new species</mods:title>
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<mods:namePart id="C9E8A40AB09CD0C3A5B6392D2D8BC496">Inkhavilay, Khamla</mods:namePart>
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<mods:namePart id="E2BDB189DCEE33B6C4A987FA864C2DD4">Sutcharit, Chirasak</mods:namePart>
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<taxonomicName id="4C7319CA6924EF0AFE37A834FBBF2246" ID-CoL="D2NM" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[444,1143,1802,1829]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="roseolabiatus" subGenus="Amphidromus">
<emphasis id="B907BE5B6924EF0AFE37A834FC112246" bold="true" box="[444,985,1802,1829]" pageId="2" pageNumber="3">
<emphasis id="B907BE5B6924EF0AFE37A834FDA12246" bold="true" box="[444,617,1802,1829]" italics="true" pageId="2" pageNumber="3">Amphidromus</emphasis>
(
<emphasis id="B907BE5B6924EF0AFDF1A834FCEF2246" bold="true" box="[634,807,1802,1829]" italics="true" pageId="2" pageNumber="3">Amphidromus</emphasis>
) 
<emphasis id="B907BE5B6924EF0AFCB3A834FC112246" bold="true" box="[824,985,1802,1829]" italics="true" pageId="2" pageNumber="3">roseolabiatus</emphasis>
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<bibRefCitation id="EFE21FB86924EF0AFC6AA835FBBF2246" author="Fulton H. C." box="[993,1143,1803,1829]" pageId="2" pageNumber="3" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
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<figureCitation id="13487ECC6924EF0AFE7BA810FDF1222B" box="[496,569,1838,1864]" captionStart="Fig" captionStartId="4.[189,232,1991,2017]" captionTargetBox="[416,1172,981,1949]" captionTargetId="figure@4.[416,1172,981,1949]" captionTargetPageId="4" captionText="Fig. 1. Distribution map of Amphidromus spp. in Laos recognized in this study." figureDoi="http://doi.org/10.5281/zenodo.3829746" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="2" pageNumber="3">Figs 1</figureCitation>
, 
<figureCitation id="13487ECC6924EF0AFDCCA810FD9D222B" box="[583,597,1838,1864]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[189,1399,1098,1849]" captionTargetId="figure@5.[189,1399,1098,1849]" captionTargetPageId="5" captionText="Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="2" pageNumber="3">2</figureCitation>
A–B, 3A–B, 4A−F, 6A–B, 7A−C; 
<tableCitation id="C6F157F26924EF0AFC66A810FB8C222B" box="[1005,1092,1838,1864]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetId="graphics@3.[189,1398,1154,1193]" captionTargetPageId="3" captionText="Table 1 (continued on next page). Shell measurements of nine species of Amphidromus from Laos. Specimen catalogue numbers are indicated in parentheses." httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="2" pageNumber="3" tableUuid="DF0C32C16925EF0BFF36AE32FCA1242B">Table 1</tableCitation>
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<taxonomicName id="4C7319CA6924EF0AFF36A852FD1B22E5" ID-CoL="D2NM" authority="Fulton, 1896: 89" authorityName="Fulton" authorityPageNumber="89" authorityYear="1896" box="[189,723,1900,1926]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6924EF0AFF36A852FDCF22E5" box="[189,519,1900,1926]" italics="true" pageId="2" pageNumber="3">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB86924EF0AFD85A852FD1B22E5" author="Fulton H. C." box="[526,723,1900,1926]" pageId="2" pageNumber="3" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896: 89</bibRefCitation>
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, pl. 6, fig. 8. 
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locality: 
<collectingCountry id="F36422D96924EF0AFB9DA852FB9A22E5" box="[1046,1106,1900,1926]" name="Thailand" pageId="2" pageNumber="3">Siam</collectingCountry>
[
<collectingCountry id="F36422D96924EF0AFBEBA855FB1A22E5" box="[1120,1234,1899,1926]" name="Thailand" pageId="2" pageNumber="3">Thailand</collectingCountry>
].
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<taxonomicName id="4C7319CA6924EF0AFF36A897FDC122A0" ID-CoL="D2NM" authorityName="Fulton" authorityYear="1896" box="[189,521,1961,1987]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6924EF0AFF36A897FDC122A0" box="[189,521,1961,1987]" italics="true" pageId="2" pageNumber="3">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AD244586924EF0AFDA7A897FCCB22A7" author="Pilsbry H. A." box="[556,771,1961,1988]" page="188" pageId="2" pageNumber="3" year="1900">
<bibRefCitation id="EFE21FB86924EF0AFDA7A897FCCB22A7" author="Pilsbry H. A." box="[556,771,1961,1988]" pageId="2" pageNumber="3" refId="ref19586" refString="Pilsbry H. A. 1900. Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Second Series, Volume 13. Academy of Natural Sciences, Philadelphia. Available from http: // biodiversitylibrary. org / item / 16708 [accessed 1 Jan. 2013]." type="url" year="1900">Pilsbry 1900: 188</bibRefCitation>
</treatmentCitation>
, pl. 60, fig. 36. — 
<treatmentCitation id="0AD244586924EF0AFC64A897FA8422A7" author="Laidlaw F. F. &amp; Solem A." box="[1007,1356,1961,1988]" page="527" pageId="2" pageNumber="3" year="1961">
<bibRefCitation id="EFE21FB86924EF0AFC64A897FA8422A7" author="Laidlaw F. F. &amp; Solem A." box="[1007,1356,1961,1988]" pageId="2" pageNumber="3" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem 1961: 527</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD244586924EF0AFF68A8F3FE132284" author="Richardson L." box="[227,475,1996,2023]" page="42" pageId="2" pageNumber="3" year="1985">
<bibRefCitation id="EFE21FB86924EF0AFF68A8F3FE132284" author="Richardson L." box="[227,475,1996,2023]" pageId="2" pageNumber="3" pagination="1 - 479" refId="ref19668" refString="Richardson L. 1985. Camaenidae: catalog of species. Tryonia 12: 1 - 479." type="journal article" year="1985">Richardson 1985: 42</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD244586924EF0AFD80A8F3FD392284" author="Schileyko A. A." box="[523,753,1996,2023]" page="57" pageId="2" pageNumber="3" year="2011">
<bibRefCitation id="EFE21FB86924EF0AFD80A8F3FD392284" author="Schileyko A. A." box="[523,753,1996,2023]" pageId="2" pageNumber="3" pagination="1 - 68" refId="ref19686" refString="Schileyko A. A. 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1 - 68. Available from http: // www. ruthenica. com / documents / vol 21 _ Schileyko _ 1 - 68. pdf [accessed 9 Jan. 2013]." type="journal article" year="2011">Schileyko 2011: 57</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="0AD244586924EF0AFCAAA8F3FBF62284" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[801,1086,1996,2023]" httpUri="http://treatment.plazi.org/id/9168AFE0259F8B1BFD345DC8FBDEC6C5" page="88" pageId="2" pageNumber="3" year="2015">
<bibRefCitation id="EFE21FB86924EF0AFCAAA8F3FBF62284" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[801,1086,1996,2023]" pageId="2" pageNumber="3" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B6924EF0AFC19A8F3FC0E2284" box="[914,966,1997,2023]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
. 2015: 88
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, fig. 13j–k.
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</paragraph>
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<caption id="DF0C32C16925EF0BFF36AE32FCA1242B" ID-Table-UUID="DF0C32C16925EF0BFF36AE32FCA1242B" httpUri="http://table.plazi.org/id/DF0C32C16925EF0BFF36AE32FCA1242B" pageId="3" pageNumber="4" targetBox="[197,1389,369,2003]" targetIsTable="true" targetPageId="3">
<paragraph id="8BCC62496925EF0BFF36AE32FCA1242B" blockId="3.[189,1399,267,329]" pageId="3" pageNumber="4">
<emphasis id="B907BE5B6925EF0BFF36AE32FED42446" bold="true" box="[189,284,267,294]" pageId="3" pageNumber="4">Table 1</emphasis>
(continued on next page). Shell measurements of nine species of 
<taxonomicName id="4C7319CA6925EF0BFBCFAE35FB202446" authorityName="Albers" authorityYear="1850" box="[1092,1256,267,293]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B6925EF0BFBCFAE35FB202446" box="[1092,1256,267,293]" italics="true" pageId="3" pageNumber="4">Amphidromus</emphasis>
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from Laos. Specimen catalogue numbers are indicated in parentheses.
</paragraph>
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<th id="76920977692510F7FF4EAE4FFD3424AE" box="[197,764,369,461]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="B907BE5B6925EF0BFEBEAEACFD5724C9" bold="true" box="[309,671,402,426]" pageId="3" pageNumber="4">Species, locality and CUMZ nos</emphasis>
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<emphasis id="B907BE5B6925EF0BFCA2AEBCFC4D24D9" bold="true" pageId="3" pageNumber="4">No. of specimens</emphasis>
</th>
<th id="76920977692510F7FC13AE4FFB3024AE" box="[920,1272,369,461]" colspan="3" colspanRight="2" gridcol="2" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="B907BE5B6925EF0BFC2EAE4FFB2C24AE" bold="true" pageId="3" pageNumber="4">Range, mean ± S.D. in mm of Shell Shell H/W height width ratio</emphasis>
</th>
<th id="76920977692510F7FA84AE4FFAA524AE" box="[1295,1389,369,461]" gridcol="5" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="B907BE5B6925EF0BFA84AE4CFAAD24A9" bold="true" pageId="3" pageNumber="4">Number of whorls</emphasis>
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<tr id="3543600B692510F7FF4EAEE6FAA52492" box="[197,1389,472,497]" gridrow="1" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAEE6FAA52492" box="[197,1389,472,497]" colspan="6" colspanRight="5" gridcol="0" gridrow="1" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EAEE6FD422492" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[197,650,472,497]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6925EF0BFF4EAEE6FE332493" bold="true" box="[197,507,472,496]" italics="true" pageId="3" pageNumber="4">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB86925EF0BFD89AEE7FD422492" author="Fulton H. C." box="[514,650,473,497]" pageId="3" pageNumber="4" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
</th>
</tr>
<tr id="3543600B692510F7FF4EAEC2FAA52756" box="[197,1389,508,565]" gridrow="2" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAEC2FD342756" box="[197,764,508,565]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Ban Pha Vong, Khammouan (7011, 7012)</th>
<td id="76920977692510F7FC9AAEC2FC4D2756" box="[785,901,508,565]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">21</td>
<td id="76920977692510F7FC13AEC2FBCD2756" box="[920,1029,508,565]" gridcol="2" gridrow="2" pageId="3" pageNumber="4">28.0−41.6 35.7±3.24</td>
<td id="76920977692510F7FB9EAEC2FB4A2756" box="[1045,1154,508,565]" gridcol="3" gridrow="2" pageId="3" pageNumber="4">14.6−21.2 17.9±1.74</td>
<td id="76920977692510F7FB12AEC2FB302756" box="[1177,1272,508,565]" gridcol="4" gridrow="2" pageId="3" pageNumber="4">1.8−2.2 1.9±0.09</td>
<td id="76920977692510F7FA84AEC2FAA52756" box="[1295,1389,508,565]" gridcol="5" gridrow="2" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAD00FAA52715" box="[197,1389,574,630]" gridrow="3" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAD00FD342715" box="[197,764,574,630]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Km 70 Lao-Vietnam border, Khammouan (7008)</th>
<td id="76920977692510F7FC9AAD00FC4D2715" box="[785,901,574,630]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">4</td>
<td id="76920977692510F7FC13AD00FBCD2715" box="[920,1029,574,630]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">28.0−35.3 33.4±3.57</td>
<td id="76920977692510F7FB9EAD00FB4A2715" box="[1045,1154,574,630]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">15.5−19.0 17.1±1.48</td>
<td id="76920977692510F7FB12AD00FB302715" box="[1177,1272,574,630]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">1.4−2.2 1.9±0.34</td>
<td id="76920977692510F7FA84AD00FAA52715" box="[1295,1389,574,630]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EADBEFAA527DB" box="[197,1389,640,696]" gridrow="4" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EADBEFD3427DB" box="[197,764,640,696]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Tam Narng Lod, Khammouan (7009)</th>
<td id="76920977692510F7FC9AADBEFC4D27DB" box="[785,901,640,696]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">5</td>
<td id="76920977692510F7FC13ADBEFBCD27DB" box="[920,1029,640,696]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">33.6−40.1 36.8±3.17</td>
<td id="76920977692510F7FB9EADBEFB4A27DB" box="[1045,1154,640,696]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">16.7−19.1 17.9±1.03</td>
<td id="76920977692510F7FB12ADBEFB3027DB" box="[1177,1272,640,696]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">1.9−2.2 2.0±0.10</td>
<td id="76920977692510F7FA84ADBEFAA527DB" box="[1295,1389,640,696]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EADFFFAA5279A" box="[197,1389,705,761]" gridrow="5" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EADFFFD34279A" box="[197,764,705,761]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Tam Mung Korn, Bolikhamxay (7004, 7005)</th>
<td id="76920977692510F7FC9AADFFFC4D279A" box="[785,901,705,761]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">22</td>
<td id="76920977692510F7FC13ADFFFBCD279A" box="[920,1029,705,761]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">28.2−40.0 37.2±2.40</td>
<td id="76920977692510F7FB9EADFFFB4A279A" box="[1045,1154,705,761]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">14.1−21.7 20.2±1.52</td>
<td id="76920977692510F7FB12ADFFFB30279A" box="[1177,1272,705,761]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">1.7−2.0 1.8±0.10</td>
<td id="76920977692510F7FA84ADFFFAA5279A" box="[1295,1389,705,761]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAC3DFAA52658" box="[197,1389,771,827]" gridrow="6" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAC3DFD342658" box="[197,764,771,827]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Thad Khaungsy, Luang Phrabang (7013, 7047, 7048)</th>
<td id="76920977692510F7FC9AAC3DFC4D2658" box="[785,901,771,827]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">6</td>
<td id="76920977692510F7FC13AC3DFBCD2658" box="[920,1029,771,827]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">34.3−41.8 37.5±2.71</td>
<td id="76920977692510F7FB9EAC3DFB4A2658" box="[1045,1154,771,827]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">17.8−19.8 18.8±0.87</td>
<td id="76920977692510F7FB12AC3DFB302658" box="[1177,1272,771,827]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">1.9−2.1 1.9±0.07</td>
<td id="76920977692510F7FA84AC3DFAA52658" box="[1295,1389,771,827]" gridcol="5" gridrow="6" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAC7AFAA5261E" box="[197,1389,836,893]" gridrow="7" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAC7AFD34261E" box="[197,764,836,893]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Nam Turn Bridge, Khmakert (7006, 707)</th>
<td id="76920977692510F7FC9AAC7AFC4D261E" box="[785,901,836,893]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">9</td>
<td id="76920977692510F7FC13AC7AFBCD261E" box="[920,1029,836,893]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">26.1−34.2 30.3±2.31</td>
<td id="76920977692510F7FB9EAC7AFB4A261E" box="[1045,1154,836,893]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">15.2−17.5 16.3±0.75</td>
<td id="76920977692510F7FB12AC7AFB30261E" box="[1177,1272,836,893]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">1.6−2.0 1.8±0.14</td>
<td id="76920977692510F7FA84AC7AFAA5261E" box="[1295,1389,836,893]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EACB8FAA526DD" box="[197,1389,902,958]" gridrow="8" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EACB8FD3426DD" box="[197,764,902,958]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">Ban Phone Can, Yommalat, Khammouan (7001)</th>
<td id="76920977692510F7FC9AACB8FC4D26DD" box="[785,901,902,958]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">8</td>
<td id="76920977692510F7FC13ACB8FBCD26DD" box="[920,1029,902,958]" gridcol="2" gridrow="8" pageId="3" pageNumber="4">27.2−33.8 30.4±2.39</td>
<td id="76920977692510F7FB9EACB8FB4A26DD" box="[1045,1154,902,958]" gridcol="3" gridrow="8" pageId="3" pageNumber="4">13.4−15.9 14.1±0.91</td>
<td id="76920977692510F7FB12ACB8FB3026DD" box="[1177,1272,902,958]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">1.9−2.3 2.1±0.11</td>
<td id="76920977692510F7FA84ACB8FAA526DD" box="[1295,1389,902,958]" gridcol="5" gridrow="8" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EACF9FAA52163" box="[197,1389,967,1024]" gridrow="9" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EACF9FD342163" box="[197,764,967,1024]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">Wat Paphar, Khamkert (7002)</th>
<td id="76920977692510F7FC9AACF9FC4D2163" box="[785,901,967,1024]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">6</td>
<td id="76920977692510F7FC13ACF9FBCD2163" box="[920,1029,967,1024]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">25.6−34.4 30.7±3.11</td>
<td id="76920977692510F7FB9EACF9FB4A2163" box="[1045,1154,967,1024]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">12.8−17.3 15.5±1.59</td>
<td id="76920977692510F7FB12ACF9FB302163" box="[1177,1272,967,1024]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">1.8−2.0 1.9±0.07</td>
<td id="76920977692510F7FA84ACF9FAA52163" box="[1295,1389,967,1024]" gridcol="5" gridrow="9" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAB37FAA52122" box="[197,1389,1033,1089]" gridrow="10" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAB37FD342122" box="[197,764,1033,1089]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">Limestone Quarry, Bolikhamxay (4013)</th>
<td id="76920977692510F7FC9AAB37FC4D2122" box="[785,901,1033,1089]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">6</td>
<td id="76920977692510F7FC13AB37FBCD2122" box="[920,1029,1033,1089]" gridcol="2" gridrow="10" pageId="3" pageNumber="4">26.9−40.7 30.2±5.25</td>
<td id="76920977692510F7FB9EAB37FB4A2122" box="[1045,1154,1033,1089]" gridcol="3" gridrow="10" pageId="3" pageNumber="4">13.3−19.3 15.4±2.08</td>
<td id="76920977692510F7FB12AB37FB302122" box="[1177,1272,1033,1089]" gridcol="4" gridrow="10" pageId="3" pageNumber="4">1.8−2.1 1.9±0.12</td>
<td id="76920977692510F7FA84AB37FAA52122" box="[1295,1389,1033,1089]" gridcol="5" gridrow="10" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAB75FAA521E0" box="[197,1389,1099,1155]" gridrow="11" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAB75FD3421E0" box="[197,764,1099,1155]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">Ban Phahom, Vang Vieng (4017, 7049)</th>
<td id="76920977692510F7FC9AAB75FC4D21E0" box="[785,901,1099,1155]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">31</td>
<td id="76920977692510F7FC13AB75FBCD21E0" box="[920,1029,1099,1155]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">23.6−30.4 27.4±1.97</td>
<td id="76920977692510F7FB9EAB75FB4A21E0" box="[1045,1154,1099,1155]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">12.9−16.5 14.6±0.8</td>
<td id="76920977692510F7FB12AB75FB3021E0" box="[1177,1272,1099,1155]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">1.6−2.1 1.8±0.12</td>
<td id="76920977692510F7FA84AB75FAA521E0" box="[1295,1389,1099,1155]" gridcol="5" gridrow="11" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EABB0FAA521C4" box="[197,1389,1166,1191]" gridrow="12" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EABB0FAA521C4" box="[197,1389,1166,1191]" colspan="6" colspanRight="5" gridcol="0" gridrow="12" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EABB0FD9E21C4" authority="Geret, 1912" authorityName="Geret" authorityYear="1912" box="[197,598,1166,1191]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6925EF0BFF4EABB0FE1921C5" bold="true" box="[197,465,1166,1190]" italics="true" pageId="3" pageNumber="4">Amphidromus givenchyi</emphasis>
<bibRefCitation id="EFE21FB86925EF0BFE53ABB1FD9E21C4" author="Geret P." box="[472,598,1167,1191]" pageId="3" pageNumber="4" pagination="55 - 56" refId="ref18483" refString="Geret P. 1912. Description d'un nouvel Amphidromus. Journal de Conchyliologie 60: 55 - 56. Available from http: // biodiversitylibrary. org / page / 16358498 [accessed 15 Jul. 2013]." type="journal article" year="1912">Geret, 1912</bibRefCitation>
</taxonomicName>
</th>
</tr>
<tr id="3543600B692510F7FF4EAB8CFAA52188" box="[197,1389,1202,1259]" gridrow="13" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAB8CFD342188" box="[197,764,1202,1259]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">Thad Lor, Salavan (7015, 7016)</th>
<td id="76920977692510F7FC9AAB8CFC4D2188" box="[785,901,1202,1259]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">110</td>
<td id="76920977692510F7FC13AB8CFBCD2188" box="[920,1029,1202,1259]" gridcol="2" gridrow="13" pageId="3" pageNumber="4">23.3−43.6 36.5±3.7</td>
<td id="76920977692510F7FB9EAB8CFB4A2188" box="[1045,1154,1202,1259]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">11.8−22.9 19.6±2.26</td>
<td id="76920977692510F7FB12AB8CFB302188" box="[1177,1272,1202,1259]" gridcol="4" gridrow="13" pageId="3" pageNumber="4">1.7−2.1 1.8±0.09</td>
<td id="76920977692510F7FA84AB8CFAA52188" box="[1295,1389,1202,1259]" gridcol="5" gridrow="13" pageId="3" pageNumber="4">7</td>
</tr>
<tr id="3543600B692510F7FF4EABCAFAA5204F" box="[197,1389,1268,1324]" gridrow="14" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EABCAFD34204F" box="[197,764,1268,1324]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">Wat Phou, Salavan (7017, 7018)</th>
<td id="76920977692510F7FC9AABCAFC4D204F" box="[785,901,1268,1324]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">44</td>
<td id="76920977692510F7FC13ABCAFBCD204F" box="[920,1029,1268,1324]" gridcol="2" gridrow="14" pageId="3" pageNumber="4">27.1−37.8 33.1±2.44</td>
<td id="76920977692510F7FB9EABCAFB4A204F" box="[1045,1154,1268,1324]" gridcol="3" gridrow="14" pageId="3" pageNumber="4">15.6−21.2 18.1±1.35</td>
<td id="76920977692510F7FB12ABCAFB30204F" box="[1177,1272,1268,1324]" gridcol="4" gridrow="14" pageId="3" pageNumber="4">1.6−2.1 1.8±0.08</td>
<td id="76920977692510F7FA84ABCAFAA5204F" box="[1295,1389,1268,1324]" gridcol="5" gridrow="14" pageId="3" pageNumber="4">6−6½</td>
</tr>
<tr id="3543600B692510F7FF4EAA06FAA52033" box="[197,1389,1336,1360]" gridrow="15" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAA06FAA52033" box="[197,1389,1336,1360]" colspan="6" colspanRight="5" gridcol="0" gridrow="15" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EAA06FDC62033" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[197,526,1336,1360]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B6925EF0BFF4EAA06FDC62033" bold="true" box="[197,526,1336,1360]" italics="true" pageId="3" pageNumber="4">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206925EF0BFD9EAA06FDA0202C" box="[533,616,1336,1359]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</th>
</tr>
<tr id="3543600B692510F7FF4EAA62FAA520F7" box="[197,1389,1372,1428]" gridrow="16" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAA62FD3420F7" box="[197,764,1372,1428]" gridcol="0" gridrow="16" pageId="3" pageNumber="4">Tam Narng Ann, Khammouan (7019, 7020, 7021)</th>
<td id="76920977692510F7FC9AAA62FC4D20F7" box="[785,901,1372,1428]" gridcol="1" gridrow="16" pageId="3" pageNumber="4">20</td>
<td id="76920977692510F7FC13AA62FBCD20F7" box="[920,1029,1372,1428]" gridcol="2" gridrow="16" pageId="3" pageNumber="4">21.8−29.1 25.9±1.69</td>
<td id="76920977692510F7FB9EAA62FB4A20F7" box="[1045,1154,1372,1428]" gridcol="3" gridrow="16" pageId="3" pageNumber="4">11.2−14.2 12.7±0.66</td>
<td id="76920977692510F7FB12AA62FB3020F7" box="[1177,1272,1372,1428]" gridcol="4" gridrow="16" pageId="3" pageNumber="4">1.7−2.2 2.0±0.10</td>
<td id="76920977692510F7FA84AA62FAA520F7" box="[1295,1389,1372,1428]" gridcol="5" gridrow="16" pageId="3" pageNumber="4">6</td>
</tr>
<tr id="3543600B692510F7FF4EAAA1FAA520DB" box="[197,1389,1439,1464]" gridrow="17" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAAA1FAA520DB" box="[197,1389,1439,1464]" colspan="6" colspanRight="5" gridcol="0" gridrow="17" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EAAA1FCE120DB" authority="Bavay &amp; Dautzenberg, 1909" authorityName="Bavay &amp; Dautzenberg" authorityYear="1909" box="[197,809,1439,1464]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="pervariabilis">
<emphasis id="B907BE5B6925EF0BFF4EAAA1FE3A20D4" bold="true" box="[197,498,1439,1463]" italics="true" pageId="3" pageNumber="4">Amphidromus pervariabilis</emphasis>
Bavay &amp; Dautzenberg, 1909
</taxonomicName>
</th>
</tr>
<tr id="3543600B692510F7FF4EAAFDFAA52098" box="[197,1389,1475,1531]" gridrow="18" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EAAFDFD342098" box="[197,764,1475,1531]" gridcol="0" gridrow="18" pageId="3" pageNumber="4">Ban Namly, Phongsaly (7014)</th>
<td id="76920977692510F7FC9AAAFDFC4D2098" box="[785,901,1475,1531]" gridcol="1" gridrow="18" pageId="3" pageNumber="4">3</td>
<td id="76920977692510F7FC13AAFDFBCD2098" box="[920,1029,1475,1531]" gridcol="2" gridrow="18" pageId="3" pageNumber="4">31.5−40.9 37.2±5.02</td>
<td id="76920977692510F7FB9EAAFDFB4A2098" box="[1045,1154,1475,1531]" gridcol="3" gridrow="18" pageId="3" pageNumber="4">18.6−19.6 19.0±0.54</td>
<td id="76920977692510F7FB12AAFDFB302098" box="[1177,1272,1475,1531]" gridcol="4" gridrow="18" pageId="3" pageNumber="4">1.6−2.2 1.9±0.31</td>
<td id="76920977692510F7FA84AAFDFAA52098" box="[1295,1389,1475,1531]" gridcol="5" gridrow="18" pageId="3" pageNumber="4">7½</td>
</tr>
<tr id="3543600B692510F7FF4EA939FAA5237C" box="[197,1389,1543,1567]" gridrow="19" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA939FAA5237C" box="[197,1389,1543,1567]" colspan="6" colspanRight="5" gridcol="0" gridrow="19" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EA939FDB2237C" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[197,634,1543,1567]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="areolatus">
<emphasis id="B907BE5B6925EF0BFF4EA939FE06237C" bold="true" box="[197,462,1543,1567]" italics="true" pageId="3" pageNumber="4">Amphidromus areolatus</emphasis>
(Pfeiffer, 1861)
</taxonomicName>
</th>
</tr>
<tr id="3543600B692510F7FF4EA915FAA52300" box="[197,1389,1579,1635]" gridrow="20" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA915FD342300" box="[197,764,1579,1635]" gridcol="0" gridrow="20" pageId="3" pageNumber="4">Thad Fek, Attapeu (7022, 7023)</th>
<td id="76920977692510F7FC9AA915FC4D2300" box="[785,901,1579,1635]" gridcol="1" gridrow="20" pageId="3" pageNumber="4">33</td>
<td id="76920977692510F7FC13A915FBCD2300" box="[920,1029,1579,1635]" gridcol="2" gridrow="20" pageId="3" pageNumber="4">13.3−21.8 16.9±2.19</td>
<td id="76920977692510F7FB9EA915FB4A2300" box="[1045,1154,1579,1635]" gridcol="3" gridrow="20" pageId="3" pageNumber="4">7.1−10.2 8.5±0.74</td>
<td id="76920977692510F7FB12A915FB302300" box="[1177,1272,1579,1635]" gridcol="4" gridrow="20" pageId="3" pageNumber="4">1.5−2.2 1.9±0.13</td>
<td id="76920977692510F7FA84A915FAA52300" box="[1295,1389,1579,1635]" gridcol="5" gridrow="20" pageId="3" pageNumber="4">6½</td>
</tr>
<tr id="3543600B692510F7FF4EA952FAA523C6" box="[197,1389,1644,1701]" gridrow="21" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA952FD3423C6" box="[197,764,1644,1701]" gridcol="0" gridrow="21" pageId="3" pageNumber="4">Thad Phasoam, Champasak (7024)</th>
<td id="76920977692510F7FC9AA952FC4D23C6" box="[785,901,1644,1701]" gridcol="1" gridrow="21" pageId="3" pageNumber="4">9</td>
<td id="76920977692510F7FC13A952FBCD23C6" box="[920,1029,1644,1701]" gridcol="2" gridrow="21" pageId="3" pageNumber="4">13.8−21.8 16.5±2.66</td>
<td id="76920977692510F7FB9EA952FB4A23C6" box="[1045,1154,1644,1701]" gridcol="3" gridrow="21" pageId="3" pageNumber="4">6.9−10.5 8.4±1.08</td>
<td id="76920977692510F7FB12A952FB3023C6" box="[1177,1272,1644,1701]" gridcol="4" gridrow="21" pageId="3" pageNumber="4">1.8−2.0 1.9±0.08</td>
<td id="76920977692510F7FA84A952FAA523C6" box="[1295,1389,1644,1701]" gridcol="5" gridrow="21" pageId="3" pageNumber="4">6½</td>
</tr>
<tr id="3543600B692510F7FF4EA98EFAA523AB" box="[197,1389,1712,1736]" gridrow="22" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA98EFAA523AB" box="[197,1389,1712,1736]" colspan="6" colspanRight="5" gridcol="0" gridrow="22" pageId="3" pageNumber="4">
<taxonomicName id="4C7319CA6925EF0BFF4EA98EFD9D23AB" authority="(Pfeiffer, 1861)" baseAuthorityName="Pfeiffer" baseAuthorityYear="1861" box="[197,597,1712,1736]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="flavus">
<emphasis id="B907BE5B6925EF0BFF4EA98EFE6123AB" bold="true" box="[197,425,1712,1736]" italics="true" pageId="3" pageNumber="4">Amphidromus flavus</emphasis>
(Pfeiffer, 1861)
</taxonomicName>
</th>
</tr>
<tr id="3543600B692510F7FF4EA9EAFAA5226F" box="[197,1389,1748,1804]" gridrow="23" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA9EAFD34226F" box="[197,764,1748,1804]" gridcol="0" gridrow="23" pageId="3" pageNumber="4">Wat Phousy, Luang Phrabang (7025)</th>
<td id="76920977692510F7FC9AA9EAFC4D226F" box="[785,901,1748,1804]" gridcol="1" gridrow="23" pageId="3" pageNumber="4">33</td>
<td id="76920977692510F7FC13A9EAFBCD226F" box="[920,1029,1748,1804]" gridcol="2" gridrow="23" pageId="3" pageNumber="4">23.2−37.4 28.4±3.67</td>
<td id="76920977692510F7FB9EA9EAFB4A226F" box="[1045,1154,1748,1804]" gridcol="3" gridrow="23" pageId="3" pageNumber="4">11.8−17.4 14.4±1.26</td>
<td id="76920977692510F7FB12A9EAFB30226F" box="[1177,1272,1748,1804]" gridcol="4" gridrow="23" pageId="3" pageNumber="4">1.7−2.1 1.9±0.11</td>
<td id="76920977692510F7FA84A9EAFAA5226F" box="[1295,1389,1748,1804]" gridcol="5" gridrow="23" pageId="3" pageNumber="4">6½</td>
</tr>
<tr id="3543600B692510F7FF4EA829FAA5222C" box="[197,1389,1815,1871]" gridrow="24" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA829FD34222C" box="[197,764,1815,1871]" gridcol="0" gridrow="24" pageId="3" pageNumber="4">Ban Na Deauy, Luang Phrabang (7026, 7027)</th>
<td id="76920977692510F7FC9AA829FC4D222C" box="[785,901,1815,1871]" gridcol="1" gridrow="24" pageId="3" pageNumber="4">11</td>
<td id="76920977692510F7FC13A829FBCD222C" box="[920,1029,1815,1871]" gridcol="2" gridrow="24" pageId="3" pageNumber="4">22.2−29.8 25.7±2.27</td>
<td id="76920977692510F7FB9EA829FB4A222C" box="[1045,1154,1815,1871]" gridcol="3" gridrow="24" pageId="3" pageNumber="4">12.1−14.2 12.9±0.68</td>
<td id="76920977692510F7FB12A829FB30222C" box="[1177,1272,1815,1871]" gridcol="4" gridrow="24" pageId="3" pageNumber="4">1.8−2.3 1.9±0.17</td>
<td id="76920977692510F7FA84A829FAA5222C" box="[1295,1389,1815,1871]" gridcol="5" gridrow="24" pageId="3" pageNumber="4">6½</td>
</tr>
<tr id="3543600B692510F7FF4EA867FAA522F2" box="[197,1389,1881,1937]" gridrow="25" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA867FD3422F2" box="[197,764,1881,1937]" gridcol="0" gridrow="25" pageId="3" pageNumber="4">Mouhot’s Tomb, Luang Phrabang (7028)</th>
<td id="76920977692510F7FC9AA867FC4D22F2" box="[785,901,1881,1937]" gridcol="1" gridrow="25" pageId="3" pageNumber="4">5</td>
<td id="76920977692510F7FC13A867FBCD22F2" box="[920,1029,1881,1937]" gridcol="2" gridrow="25" pageId="3" pageNumber="4">24.4−29.0 27.6±1.88</td>
<td id="76920977692510F7FB9EA867FB4A22F2" box="[1045,1154,1881,1937]" gridcol="3" gridrow="25" pageId="3" pageNumber="4">13.1−13.9 13.5±0.36</td>
<td id="76920977692510F7FB12A867FB3022F2" box="[1177,1272,1881,1937]" gridcol="4" gridrow="25" pageId="3" pageNumber="4">1.7−2.2 2.0±0.17</td>
<td id="76920977692510F7FA84A867FAA522F2" box="[1295,1389,1881,1937]" gridcol="5" gridrow="25" pageId="3" pageNumber="4">6½</td>
</tr>
<tr id="3543600B692510F7FF4EA8A4FAA522B0" box="[197,1389,1946,2003]" gridrow="26" pageId="3" pageNumber="4">
<th id="76920977692510F7FF4EA8A4FD3422B0" box="[197,764,1946,2003]" gridcol="0" gridrow="26" pageId="3" pageNumber="4">Tam Pou Kham, Vang Vieng (7029)</th>
<td id="76920977692510F7FC9AA8A4FC4D22B0" box="[785,901,1946,2003]" gridcol="1" gridrow="26" pageId="3" pageNumber="4">12</td>
<td id="76920977692510F7FC13A8A4FBCD22B0" box="[920,1029,1946,2003]" gridcol="2" gridrow="26" pageId="3" pageNumber="4">23.7−35.5 28.3±3.48</td>
<td id="76920977692510F7FB9EA8A4FB4A22B0" box="[1045,1154,1946,2003]" gridcol="3" gridrow="26" pageId="3" pageNumber="4">13.0−17.1 14.3±1.35</td>
<td id="76920977692510F7FB12A8A4FB3022B0" box="[1177,1272,1946,2003]" gridcol="4" gridrow="26" pageId="3" pageNumber="4">1.7−2.2 1.9±0.14</td>
<td id="76920977692510F7FA84A8A4FAA522B0" box="[1295,1389,1946,2003]" gridcol="5" gridrow="26" pageId="3" pageNumber="4">6½</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCC62496922EF0CFF4EAE2BFA8D261B" pageId="4" pageNumber="5">
<table id="F97390E9692210F7FF4EAE2BFA9526EB" box="[197,1373,277,904]" gridcols="6" gridrows="11" pageId="4" pageNumber="5">
<tr id="3543600B692210F7FF4EAE2BFA95244D" box="[197,1373,277,302]" gridrow="0" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAE2BFA95244D" box="[197,1373,277,302]" colspan="6" colspanRight="5" gridcol="0" gridrow="0" pageId="4" pageNumber="5">
<taxonomicName id="4C7319CA6922EF0CFF4EAE2BFDA5244D" authority="Morlet, 1891" authorityName="Morlet" authorityYear="1891" box="[197,621,277,302]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="xiengensis">
<emphasis id="B907BE5B6922EF0CFF4EAE2BFE14244E" bold="true" box="[197,476,277,301]" italics="true" pageId="4" pageNumber="5">Amphidromus xiengensis</emphasis>
Morlet, 1891
</taxonomicName>
</th>
</tr>
<tr id="3543600B692210F7FF4EAE07FA952412" box="[197,1373,313,369]" gridrow="1" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAE07FD392412" box="[197,753,313,369]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">Ban Phon Pai, Champasak (7030)</th>
<td id="76920977692210F7FCBCAE07FC952412" box="[823,861,313,369]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">33</td>
<td id="76920977692210F7FC13AE07FBCD2412" box="[920,1029,313,369]" gridcol="2" gridrow="1" pageId="4" pageNumber="5">21.4−36.2 31.0±3.04</td>
<td id="76920977692210F7FB9EAE07FB4A2412" box="[1045,1154,313,369]" gridcol="3" gridrow="1" pageId="4" pageNumber="5">13.4−16.9 15.1±0.96</td>
<td id="76920977692210F7FB12AE07FB302412" box="[1177,1272,313,369]" gridcol="4" gridrow="1" pageId="4" pageNumber="5">1.4−2.3 2.0±0.14</td>
<td id="76920977692210F7FAABAE07FA952412" box="[1312,1373,313,369]" gridcol="5" gridrow="1" pageId="4" pageNumber="5">6½−7</td>
</tr>
<tr id="3543600B692210F7FF4EAE45FA9524D0" box="[197,1373,379,435]" gridrow="2" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAE45FD3924D0" box="[197,753,379,435]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">Ban Oudom limestone, Oudomxay (7032)</th>
<td id="76920977692210F7FCBCAE45FC9524D0" box="[823,861,379,435]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">6</td>
<td id="76920977692210F7FC13AE45FBCD24D0" box="[920,1029,379,435]" gridcol="2" gridrow="2" pageId="4" pageNumber="5">22.1−29.7 26.5±2.61</td>
<td id="76920977692210F7FB9EAE45FB4A24D0" box="[1045,1154,379,435]" gridcol="3" gridrow="2" pageId="4" pageNumber="5">11.2−13.9 13.0±0.94</td>
<td id="76920977692210F7FB12AE45FB3024D0" box="[1177,1272,379,435]" gridcol="4" gridrow="2" pageId="4" pageNumber="5">1.9−2.1 2.0±0.08</td>
<td id="76920977692210F7FAABAE45FA9524D0" box="[1312,1373,379,435]" gridcol="5" gridrow="2" pageId="4" pageNumber="5">6½−7</td>
</tr>
<tr id="3543600B692210F7FF4EAE82FA952496" box="[197,1373,444,501]" gridrow="3" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAE82FD392496" box="[197,753,444,501]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">Mouhot’s Tomb, Luang Phrabang (7033)</th>
<td id="76920977692210F7FCBCAE82FC952496" box="[823,861,444,501]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">5</td>
<td id="76920977692210F7FC13AE82FBCD2496" box="[920,1029,444,501]" gridcol="2" gridrow="3" pageId="4" pageNumber="5">21.9−35.2 28.1±5.03</td>
<td id="76920977692210F7FB9EAE82FB4A2496" box="[1045,1154,444,501]" gridcol="3" gridrow="3" pageId="4" pageNumber="5">12.0−14.8 13.2±1.43</td>
<td id="76920977692210F7FB12AE82FB302496" box="[1177,1272,444,501]" gridcol="4" gridrow="3" pageId="4" pageNumber="5">1.8−2.3 2.1±0.21</td>
<td id="76920977692210F7FAABAE82FA952496" box="[1312,1373,444,501]" gridcol="5" gridrow="3" pageId="4" pageNumber="5">6½−7</td>
</tr>
<tr id="3543600B692210F7FF4EAEC0FA952755" box="[197,1373,510,566]" gridrow="4" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAEC0FD392755" box="[197,753,510,566]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">Thad Phasoam, Boloven Plateau, Champasak (7031)</th>
<td id="76920977692210F7FCBCAEC0FC952755" box="[823,861,510,566]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">10</td>
<td id="76920977692210F7FC13AEC0FBCD2755" box="[920,1029,510,566]" gridcol="2" gridrow="4" pageId="4" pageNumber="5">17.4−34.4 29.9±5.87</td>
<td id="76920977692210F7FB9EAEC0FB4A2755" box="[1045,1154,510,566]" gridcol="3" gridrow="4" pageId="4" pageNumber="5">8.7−15.9 13.8±2.46</td>
<td id="76920977692210F7FB12AEC0FB302755" box="[1177,1272,510,566]" gridcol="4" gridrow="4" pageId="4" pageNumber="5">2.0−2.2 2.1±0.07</td>
<td id="76920977692210F7FAABAEC0FA952755" box="[1312,1373,510,566]" gridcol="5" gridrow="4" pageId="4" pageNumber="5">6½−7</td>
</tr>
<tr id="3543600B692210F7FF4EAD7EFA95271B" box="[197,1373,576,632]" gridrow="5" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAD7EFD39271B" box="[197,753,576,632]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">Ban Na Deauy, Luang Phrabang (7036, 7037)</th>
<td id="76920977692210F7FCBCAD7EFC95271B" box="[823,861,576,632]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">4</td>
<td id="76920977692210F7FC13AD7EFBCD271B" box="[920,1029,576,632]" gridcol="2" gridrow="5" pageId="4" pageNumber="5">22.4−29.4 26.7±3.32</td>
<td id="76920977692210F7FB9EAD7EFB4A271B" box="[1045,1154,576,632]" gridcol="3" gridrow="5" pageId="4" pageNumber="5">11.4−13.9 13.1±1.16</td>
<td id="76920977692210F7FB12AD7EFB30271B" box="[1177,1272,576,632]" gridcol="4" gridrow="5" pageId="4" pageNumber="5">1.8−2.1 2.0±0.15</td>
<td id="76920977692210F7FAABAD7EFA95271B" box="[1312,1373,576,632]" gridcol="5" gridrow="5" pageId="4" pageNumber="5">6½−7</td>
</tr>
<tr id="3543600B692210F7FF4EADBDFA9527FF" box="[197,1373,643,668]" gridrow="6" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EADBDFA9527FF" box="[197,1373,643,668]" colspan="6" colspanRight="5" gridcol="0" gridrow="6" pageId="4" pageNumber="5">
<taxonomicName id="4C7319CA6922EF0CFF4EADBDFD6527FF" authority="Mollendorff, 1898" authorityName="Mollendorff" authorityYear="1898" box="[197,685,643,668]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="fuscolabris">
<emphasis id="B907BE5B6922EF0CFF4EADBDFE2927F8" bold="true" box="[197,481,643,667]" italics="true" pageId="4" pageNumber="5">Amphidromus fuscolabris</emphasis>
<bibRefCitation id="EFE21FB86922EF0CFE63ADBDFD6527FF" author="Mollendorff O. F. von" box="[488,685,643,668]" pageId="4" pageNumber="5" pagination="65 - 85" refId="ref18807" refString="Mollendorff O. F. von. 1898. Die Binnenmollusken Annams. Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 30: 65 - 85. Available from http: // www. biodiversitylibrary. org / item / 89671 # page / 261 / mode / 1 up [accessed 21 Apr. 2017]." type="journal article" year="1898">Möllendorff, 1898</bibRefCitation>
</taxonomicName>
</th>
</tr>
<tr id="3543600B692210F7FF4EAD99FA9527BC" box="[197,1373,679,735]" gridrow="7" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAD99FD3927BC" box="[197,753,679,735]" gridcol="0" gridrow="7" pageId="4" pageNumber="5">Ban Phone, Sekong (7040, 7041, 7042, 7043)</th>
<td id="76920977692210F7FCBCAD99FC9527BC" box="[823,861,679,735]" gridcol="1" gridrow="7" pageId="4" pageNumber="5">112</td>
<td id="76920977692210F7FC13AD99FBCD27BC" box="[920,1029,679,735]" gridcol="2" gridrow="7" pageId="4" pageNumber="5">26.0−43.5 34.3±3.56</td>
<td id="76920977692210F7FB9EAD99FB4A27BC" box="[1045,1154,679,735]" gridcol="3" gridrow="7" pageId="4" pageNumber="5">13.1−18.1 15.2±1.04</td>
<td id="76920977692210F7FB12AD99FB3027BC" box="[1177,1272,679,735]" gridcol="4" gridrow="7" pageId="4" pageNumber="5">1.8−2.6 2.2±0.13</td>
<td id="76920977692210F7FAABAD99FA9527BC" box="[1312,1373,679,735]" gridcol="5" gridrow="7" pageId="4" pageNumber="5">7</td>
</tr>
<tr id="3543600B692210F7FF4EADD7FA952642" box="[197,1373,745,801]" gridrow="8" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EADD7FD392642" box="[197,753,745,801]" gridcol="0" gridrow="8" pageId="4" pageNumber="5">Ban Xai Na Pho, Champasak (7044)</th>
<td id="76920977692210F7FCBCADD7FC952642" box="[823,861,745,801]" gridcol="1" gridrow="8" pageId="4" pageNumber="5">3</td>
<td id="76920977692210F7FC13ADD7FBCD2642" box="[920,1029,745,801]" gridcol="2" gridrow="8" pageId="4" pageNumber="5">26.0−43.6 34.8±4.06</td>
<td id="76920977692210F7FB9EADD7FB4A2642" box="[1045,1154,745,801]" gridcol="3" gridrow="8" pageId="4" pageNumber="5">13.1−18.1 15.3±1.20</td>
<td id="76920977692210F7FB12ADD7FB302642" box="[1177,1272,745,801]" gridcol="4" gridrow="8" pageId="4" pageNumber="5">1.8−2.5 2.2±0.14</td>
<td id="76920977692210F7FAABADD7FA952642" box="[1312,1373,745,801]" gridcol="5" gridrow="8" pageId="4" pageNumber="5">7</td>
</tr>
<tr id="3543600B692210F7FF4EAC12FA952627" box="[197,1373,812,836]" gridrow="9" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAC12FA952627" box="[197,1373,812,836]" colspan="6" colspanRight="5" gridcol="0" gridrow="9" pageId="4" pageNumber="5">
<taxonomicName id="4C7319CA6922EF0CFF4EAC12FDF82627" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[197,560,812,836]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="xiengkhaungensis" status="sp. nov.">
<emphasis id="B907BE5B6922EF0CFF4EAC12FDF82627" bold="true" box="[197,560,812,836]" italics="true" pageId="4" pageNumber="5">Amphidromus xiengkhaungensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206922EF0CFDBCAC13FD422627" box="[567,650,813,836]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
</th>
</tr>
<tr id="3543600B692210F7FF4EAC6FFA9526EB" box="[197,1373,849,904]" gridrow="10" pageId="4" pageNumber="5">
<th id="76920977692210F7FF4EAC6FFD3926EB" box="[197,753,849,904]" gridcol="0" gridrow="10" pageId="4" pageNumber="5">Nong Tang, Xieng Khaung (7045, 7046)</th>
<td id="76920977692210F7FCBCAC6FFC9526EB" box="[823,861,849,904]" gridcol="1" gridrow="10" pageId="4" pageNumber="5">4</td>
<td id="76920977692210F7FC13AC6FFBCD26EB" box="[920,1029,849,904]" gridcol="2" gridrow="10" pageId="4" pageNumber="5">21.4−29.8 25.6±3.49</td>
<td id="76920977692210F7FB9EAC6FFB4A26EB" box="[1045,1154,849,904]" gridcol="3" gridrow="10" pageId="4" pageNumber="5">14.8−17.0 15.7±1.04</td>
<td id="76920977692210F7FB12AC6FFB3026EB" box="[1177,1272,849,904]" gridcol="4" gridrow="10" pageId="4" pageNumber="5">1.4−1.7 1.6±0.13</td>
<td id="76920977692210F7FAABAC6FFA9526EB" box="[1312,1373,849,904]" gridcol="5" gridrow="10" pageId="4" pageNumber="5">6</td>
</tr>
</table>
</paragraph>
<caption id="DF0C32C16922EF0CFF36A8F9FB972282" ID-DOI="http://doi.org/10.5281/zenodo.3829746" ID-Zenodo-Dep="3829746" box="[189,1119,1991,2018]" httpUri="https://zenodo.org/record/3829746/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1991,2017]" targetBox="[416,1172,981,1949]" targetPageId="4">
<paragraph id="8BCC62496922EF0CFF36A8F9FB972282" blockId="4.[189,1119,1991,2018]" box="[189,1119,1991,2018]" pageId="4" pageNumber="5">
<emphasis id="B907BE5B6922EF0CFF36A8F9FEC32282" bold="true" box="[189,267,1991,2017]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
Distribution map of 
<taxonomicName id="4C7319CA6922EF0CFD8BA8F9FD6C2282" authorityName="Albers" authorityYear="1850" box="[512,676,1991,2017]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B6922EF0CFD8BA8F9FD6C2282" box="[512,676,1991,2017]" italics="true" pageId="4" pageNumber="5">Amphidromus</emphasis>
</taxonomicName>
spp. in Laos recognized in this study.
</paragraph>
</caption>
<subSubSection id="C36931C26923EF00FF36AE35FC6D2409" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BCC62496923EF0DFF36AE35FE622446" blockId="5.[189,426,267,293]" box="[189,426,267,293]" pageId="5" pageNumber="6">
<heading id="D084D5256923EF0DFF36AE35FE622446" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B907BE5B6923EF0DFF36AE35FE622446" bold="true" box="[189,426,267,293]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC62496923EF0DFF61AE04FE512430" blockId="5.[189,1399,313,411]" box="[234,409,313,340]" pageId="5" pageNumber="6">
<emphasis id="B907BE5B6923EF0DFF61AE04FE512430" bold="true" box="[234,409,313,340]" pageId="5" pageNumber="6">
<typeStatus id="54C8DCEB6923EF0DFF61AE04FEEF2437" box="[234,295,314,340]" pageId="5" pageNumber="6">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BCC62496923EF0DFF36AE63FCD024F8" blockId="5.[189,1399,313,411]" pageId="5" pageNumber="6">
<collectingCountry id="F36422D96923EF0DFF36AE63FE912414" box="[189,345,349,375]" name="Thailand" pageId="5" pageNumber="6">THAILAND</collectingCountry>
: 
<typeStatus id="54C8DCEB6923EF0DFEEEAE63FE152414" box="[357,477,349,375]" pageId="5" pageNumber="6" type="lectotype">Lectotype</typeStatus>
, 
<collectionCode id="ED62FA8C6923EF0DFE61AE63FD912414" box="[490,601,349,375]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="6" type="Museum">NHMUK</collectionCode>
19601462 (
<bibRefCitation id="EFE21FB86923EF0DFD6CAE62FC172414" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[743,991,348,375]" pageId="5" pageNumber="6" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B6923EF0DFCD7AE63FC582414" box="[860,912,349,375]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 13j); 
<typeStatus id="54C8DCEB6923EF0DFBD1AE62FB332415" box="[1114,1275,348,374]" pageId="5" pageNumber="6" type="paralectotype">paralectotype</typeStatus>
, 
<collectionCode id="ED62FA8C6923EF0DFA83AE63FABF2414" box="[1288,1399,349,375]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="5" pageNumber="6" type="Museum">NHMUK</collectionCode>
19601463 (1S shell, 
<bibRefCitation id="EFE21FB86923EF0DFE3BAEBEFD6A24F8" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[432,674,384,411]" pageId="5" pageNumber="6" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B6923EF0DFDAAAEBFFD9D24F9" box="[545,597,384,410]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2015
</bibRefCitation>
: fig. 13k).
</paragraph>
<paragraph id="8BCC62496923EF0DFF61AEFFFE6E24B8" blockId="5.[189,1399,449,1004]" box="[234,422,449,475]" pageId="5" pageNumber="6">
<emphasis id="B907BE5B6923EF0DFF61AEFFFE6E24B8" bold="true" box="[234,422,449,475]" pageId="5" pageNumber="6">Other material</emphasis>
</paragraph>
<paragraph id="8BCC62496923EF0DFF36AEDBFC1C2688" blockId="5.[189,1399,449,1004]" pageId="5" pageNumber="6">
<collectingCountry id="F36422D96923EF0DFF36AEDBFEC6249C" box="[189,270,485,511]" name="Laos" pageId="5" pageNumber="6">LAOS</collectingCountry>
: 5D + 3S shells, 
<collectingRegion id="49B7ACAB6923EF0DFE46AEDBFD84249D" box="[461,588,484,511]" country="Laos" name="Xekong" pageId="5" pageNumber="6">Ban Phone</collectingRegion>
Can, Yommalat District, 
<collectingRegion id="49B7ACAB6923EF0DFCF9AEDAFBC2249D" box="[882,1034,484,510]" country="Laos" name="Khammouan" pageId="5" pageNumber="6">Khammouan</collectingRegion>
(
<collectionCode id="ED62FA8C6923EF0DFB9CAEDBFBA6249C" box="[1047,1134,485,511]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7001); 5D + 1S shells, Wat Paphar, Khamkert District, Bolikhamxay (
<collectionCode id="ED62FA8C6923EF0DFD69AD36FCF12741" box="[738,825,520,546]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7002); 2D + 1S shells, Thad Mouang, Khamkert District, Bolikhamxay (
<collectionCode id="ED62FA8C6923EF0DFE58AD15FDE22726" box="[467,554,555,581]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7003); 4D + 10S specimens, in ethanol (
<figureCitation id="13487ECC6923EF0DFB98AD15FB912726" box="[1043,1113,555,581]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
E–F, 
<collectionCode id="ED62FA8C6923EF0DFB13AD15FB252726" box="[1176,1261,555,581]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7004), 13S specimens in freezer (
<figureCitation id="13487ECC6923EF0DFE44AD70FDF9270B" box="[463,561,590,616]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[189,1399,1098,1849]" captionTargetId="figure@5.[189,1399,1098,1849]" captionTargetPageId="5" captionText="Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
, 
<collectionCode id="ED62FA8C6923EF0DFDC8AD70FD50270B" box="[579,664,590,616]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7005), Tam Mung Korn, Khamkert District, Bolikhamxay; 1D + 1S specimens, in ethanol (
<figureCitation id="13487ECC6923EF0DFDB6AD4FFD4C27E8" box="[573,644,625,651]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
C–D, 
<collectionCode id="ED62FA8C6923EF0DFD40AD4FFCE827E8" box="[715,800,625,651]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7006), 4D + 5S specimens in freezer (
<figureCitation id="13487ECC6923EF0DFB7AAD4FFAFE27E8" box="[1265,1334,625,651]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[214,1373,265,1744]" captionTargetId="figure@6.[214,1373,265,1744]" captionTargetPageId="6" captionText="Fig. 3. Aestivation sites of Amphidromus species. A–B. Amphidromus roseolabiatus Fulton, 1896 (shell height about 34 mm) from NamTurn Bridge, Bolikhamxay, Laos, aestivated. A. Inside sterile fronds of stag horn ferns, Platycerium Desv. B. In a small hole on a Jackfruit tree trunk, Artocarpus heterophyllus Lam., with other snails, Quantula Baker, 1941 and Durgella Blanford, 1863. C. Amphidromus givenchyi Geret, 1912 (shell height about 43 mm), hiding in a small hole of a dipterocarpus tree shell. D. Amphidromus syndromoideus sp. nov. (shell height about 28 mm) from the type locality, hiding in a hole of a tree, visible after removal of clusters of sterile fronds of the basket fern Drynaria (Bory) J. Sm." figureDoi="http://doi.org/10.5281/zenodo.3829750" httpUri="https://zenodo.org/record/3829750/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
A–B, 
<collectionCode id="ED62FA8C6923EF0DFF36ADABFEDA27CC" box="[189,274,661,687]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7007), Nam Turn Bridge, Khamkert District, Bolikhamxay; 8D + 6S shells, Limestone quarry, Bolikhamxay (
<collectionCode id="ED62FA8C6923EF0DFEE0AD86FE0A27B1" box="[363,450,696,722]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
4013); 4D + 3S shells, pink aperture, Km. 70 on the road from Laos to Vietnam border, Yommalat District, 
<collectingRegion id="49B7ACAB6923EF0DFD80ADE5FD6B2796" box="[523,675,731,757]" country="Laos" name="Khammouan" pageId="5" pageNumber="6">Khammouan</collectingRegion>
(
<collectionCode id="ED62FA8C6923EF0DFD3DADE5FCC52796" box="[694,781,731,757]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7008); 8D shells, pink aperture, Tam Narng Lod, Yommalat District, 
<collectingRegion id="49B7ACAB6923EF0DFE2BADC0FDF0267B" box="[416,568,766,792]" country="Laos" name="Khammouan" pageId="5" pageNumber="6">Khammouan</collectingRegion>
(
<collectionCode id="ED62FA8C6923EF0DFDCEADC0FD54267B" box="[581,668,766,792]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7009); 3D + 1S shells, pink aperture, Hauy In River, Yommalat District, 
<collectingRegion id="49B7ACAB6923EF0DFEA9AC1FFE722658" box="[290,442,801,827]" country="Laos" name="Khammouan" pageId="5" pageNumber="6">Khammouan</collectingRegion>
(
<collectionCode id="ED62FA8C6923EF0DFE4CAC1FFDD62658" box="[455,542,801,827]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7010); 12D + 8S specimens, in ethanol, pink aperture (
<figureCitation id="13487ECC6923EF0DFB12AC1FFB152658" box="[1177,1245,801,827]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
A–B, 
<collectionCode id="ED62FA8C6923EF0DFAA9AC1FFABF2658" box="[1314,1399,801,827]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7011), 5D specimens, in ethanol (
<figureCitation id="13487ECC6923EF0DFDD7AC7AFD09263C" box="[604,705,836,863]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[189,1399,1098,1849]" captionTargetId="figure@5.[189,1399,1098,1849]" captionTargetPageId="5" captionText="Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="5" pageNumber="6">Figs 2A</figureCitation>
, 
<figureCitation id="13487ECC6923EF0DFD59AC7AFD28263D" box="[722,736,836,862]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
A–B, 7A−C, 
<collectionCode id="ED62FA8C6923EF0DFC0EAC7BFC12263C" box="[901,986,837,863]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7012), Ban Pha Vong, Yommalat District, 
<collectingRegion id="49B7ACAB6923EF0DFEAFAC56FE7726E1" box="[292,447,872,898]" country="Laos" name="Khammouan" pageId="5" pageNumber="6">Khammouan</collectingRegion>
; 1D + 1S shells, pink aperture (
<collectionCode id="ED62FA8C6923EF0DFCBFAC56FC4326E1" box="[820,907,872,898]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7013), 6D specimens, in ethanol (
<collectionCode id="ED62FA8C6923EF0DFAABAC56FABF26E1" box="[1312,1399,872,898]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7047), 4D specimens, in ethanol (
<collectionCode id="ED62FA8C6923EF0DFDDDACB5FD6526C6" box="[598,685,907,933]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7048), 1D + 1S shells, pink aperture (
<collectionCode id="ED62FA8C6923EF0DFB0FACB5FB1326C6" box="[1156,1243,907,933]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7035), Thad Khaungsy Waterfall, Luang Phrabang District, Luang Phrabang; 6D shells (
<collectionCode id="ED62FA8C6923EF0DFBBDAC90FB4526AB" box="[1078,1165,942,968]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
40107), 10S + 21D shells (
<collectionCode id="ED62FA8C6923EF0DFE84ACEFFEAE2688" box="[271,358,977,1003]" country="0" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="6">CUMZ</collectionCode>
7049), Ban Phahom, Vang Vieng District, 
<collectingRegion id="49B7ACAB6923EF0DFCD0ACEFFC072688" box="[859,975,977,1003]" country="Laos" name="Viangchan" pageId="5" pageNumber="6">Vientiane</collectingRegion>
.
</paragraph>
<caption id="DF0C32C16923EF0DFF36A85AFAEC228B" ID-DOI="http://doi.org/10.5281/zenodo.3829748" ID-Zenodo-Dep="3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1892,1918]" targetBox="[189,1399,1098,1849]" targetPageId="5">
<paragraph id="8BCC62496923EF0DFF36A85AFAEC228B" blockId="5.[189,1399,1892,2025]" pageId="5" pageNumber="6">
<emphasis id="B907BE5B6923EF0DFF36A85AFEC0221D" bold="true" box="[189,264,1892,1918]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
Living snails. 
<emphasis id="B907BE5B6923EF0DFE25A85AFE2D221D" bold="true" box="[430,485,1892,1918]" pageId="5" pageNumber="6">A–B</emphasis>
. 
<taxonomicName id="4C7319CA6923EF0DFE79A85BFC1C221D" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[498,980,1892,1919]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6923EF0DFE79A85BFCF1221C" box="[498,825,1893,1919]" italics="true" pageId="5" pageNumber="6">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB86923EF0DFCB6A85AFC1C221D" author="Fulton H. C." box="[829,980,1892,1918]" pageId="5" pageNumber="6" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B907BE5B6923EF0DFC57A85AFC39221D" bold="true" box="[988,1009,1892,1918]" pageId="5" pageNumber="6">A</emphasis>
. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). 
<emphasis id="B907BE5B6923EF0DFDEAA8B6FDBD22C1" bold="true" box="[609,629,1928,1954]" pageId="5" pageNumber="6">B</emphasis>
. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). 
<emphasis id="B907BE5B6923EF0DFEF8A895FE4022A6" bold="true" box="[371,392,1963,1989]" pageId="5" pageNumber="6">C</emphasis>
. 
<taxonomicName id="4C7319CA6923EF0DFE1DA895FCF722A6" authority="Geret, 1912" authorityName="Geret" authorityYear="1912" box="[406,831,1963,1989]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6923EF0DFE1DA895FD6722A6" box="[406,687,1963,1989]" italics="true" pageId="5" pageNumber="6">Amphidromus givenchyi</emphasis>
<bibRefCitation id="EFE21FB86923EF0DFD3EA895FCF722A6" author="Geret P." box="[693,831,1963,1989]" pageId="5" pageNumber="6" pagination="55 - 56" refId="ref18483" refString="Geret P. 1912. Description d'un nouvel Amphidromus. Journal de Conchyliologie 60: 55 - 56. Available from http: // biodiversitylibrary. org / page / 16358498 [accessed 15 Jul. 2013]." type="journal article" year="1912">Geret, 1912</bibRefCitation>
</taxonomicName>
from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). 
<emphasis id="B907BE5B6923EF0DFE9BA8F0FEED228B" bold="true" box="[272,293,1998,2024]" pageId="5" pageNumber="6">D</emphasis>
. 
<taxonomicName id="4C7319CA6923EF0DFEBFA8F0FD58228B" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[308,656,1998,2024]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B6923EF0DFEBFA8F0FD58228B" box="[308,656,1998,2024]" italics="true" pageId="5" pageNumber="6">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206923EF0DFD1CA8F1FD3C228A" box="[663,756,1999,2025]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype from the type locality (CUMZ 7019).
</paragraph>
</caption>
<caption id="DF0C32C16920EF0EFF36A9C3FABE2288" ID-DOI="http://doi.org/10.5281/zenodo.3829750" ID-Zenodo-Dep="3829750" httpUri="https://zenodo.org/record/3829750/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1789,1815]" targetBox="[214,1373,265,1744]" targetPageId="6">
<paragraph id="8BCC62496920EF0EFF36A9C3FABE2288" blockId="6.[189,1399,1789,2027]" pageId="6" pageNumber="7">
<emphasis id="B907BE5B6920EF0EFF36A9C3FEC22274" bold="true" box="[189,266,1789,1815]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
Aestivation sites of 
<taxonomicName id="4C7319CA6920EF0EFE73A9C3FD542274" authorityName="Albers" authorityYear="1850" box="[504,668,1789,1815]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B6920EF0EFE73A9C3FD542274" box="[504,668,1789,1815]" italics="true" pageId="6" pageNumber="7">Amphidromus</emphasis>
</taxonomicName>
species. 
<emphasis id="B907BE5B6920EF0EFC8EA9C3FCF42274" bold="true" box="[773,828,1789,1815]" pageId="6" pageNumber="7">A–B</emphasis>
. 
<taxonomicName id="4C7319CA6920EF0EFCC1A9C3FAE72274" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[842,1327,1789,1815]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6920EF0EFCC1A9C3FB5B2274" box="[842,1171,1789,1815]" italics="true" pageId="6" pageNumber="7">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB86920EF0EFB12A9C3FAE72274" author="Fulton H. C." box="[1177,1327,1789,1815]" pageId="6" pageNumber="7" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
(shell height about 34 mm) from NamTurn Bridge, Bolikhamxay, Laos, aestivated. 
<emphasis id="B907BE5B6920EF0EFBC2A81EFB962259" bold="true" box="[1097,1118,1824,1850]" pageId="6" pageNumber="7">A</emphasis>
. Inside sterile fronds of stag horn ferns, 
<taxonomicName id="4C7319CA6920EF0EFEF3A87AFD58223D" authority="Desv. B. In" authorityName="Desv. B. In" box="[376,656,1860,1886]" class="Polypodiopsida" family="Polypodiaceae" genus="Platycerium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B907BE5B6920EF0EFEF3A87AFDCF223D" box="[376,519,1860,1886]" italics="true" pageId="6" pageNumber="7">Platycerium</emphasis>
Desv. 
<emphasis id="B907BE5B6920EF0EFDDDA87AFDA2223D" bold="true" box="[598,618,1860,1886]" pageId="6" pageNumber="7">B</emphasis>
. In
</taxonomicName>
a small hole on a Jackfruit tree trunk, 
<taxonomicName id="4C7319CA6920EF0EFBC6A87AFF3522E2" authority="Lam." authorityName="Lam." class="Magnoliopsida" family="Moraceae" genus="Artocarpus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B907BE5B6920EF0EFBC6A87AFABF223D" box="[1101,1399,1860,1886]" italics="true" pageId="6" pageNumber="7">Artocarpus heterophyllus</emphasis>
Lam.
</taxonomicName>
, with other snails, 
<taxonomicName id="4C7319CA6920EF0EFE77A859FCDA22E2" authority="Baker, 1941" authorityName="Baker" authorityYear="1941" box="[508,786,1895,1921]" class="Gastropoda" family="Dyakiidae" genus="Quantula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B6920EF0EFE77A859FDA322E2" box="[508,619,1895,1921]" italics="true" pageId="6" pageNumber="7">Quantula</emphasis>
Baker, 1941
</taxonomicName>
and 
<taxonomicName id="4C7319CA6920EF0EFCD7A859FB4722E2" authority="Blanford, 1863" authorityName="Blanford" authorityYear="1863" box="[860,1167,1895,1921]" class="Gastropoda" family="Helicarionidae" genus="Durgella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B907BE5B6920EF0EFCD7A859FC0D22E2" box="[860,965,1895,1921]" italics="true" pageId="6" pageNumber="7">Durgella</emphasis>
Blanford, 1863
</taxonomicName>
. 
<emphasis id="B907BE5B6920EF0EFB2DA859FB7322E2" bold="true" box="[1190,1211,1895,1921]" pageId="6" pageNumber="7">C</emphasis>
. 
<taxonomicName id="4C7319CA6920EF0EFB58A859FE0922C7" authority="Geret, 1912" authorityName="Geret" authorityYear="1912" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6920EF0EFB58A859FEE522C7" italics="true" pageId="6" pageNumber="7">Amphidromus givenchyi</emphasis>
<bibRefCitation id="EFE21FB86920EF0EFEBDA8B4FE0922C7" author="Geret P." box="[310,449,1930,1956]" pageId="6" pageNumber="7" pagination="55 - 56" refId="ref18483" refString="Geret P. 1912. Description d'un nouvel Amphidromus. Journal de Conchyliologie 60: 55 - 56. Available from http: // biodiversitylibrary. org / page / 16358498 [accessed 15 Jul. 2013]." type="journal article" year="1912">Geret, 1912</bibRefCitation>
</taxonomicName>
(shell height about 43 mm), hiding in a small hole of a dipterocarpus tree shell. 
<emphasis id="B907BE5B6920EF0EFF36A893FF1A22A4" bold="true" box="[189,210,1965,1991]" pageId="6" pageNumber="7">D</emphasis>
. 
<taxonomicName id="4C7319CA6920EF0EFF6AA893FDF522A4" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[225,573,1965,1991]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B6920EF0EFF6AA893FDF522A4" box="[225,573,1965,1991]" italics="true" pageId="6" pageNumber="7">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206920EF0EFDCFA890FD5722AB" box="[580,671,1966,1992]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(shell height about 28 mm) from the type locality, hiding in a hole of a tree, visible after removal of clusters of sterile fronds of the basket fern 
<taxonomicName id="4C7319CA6920EF0EFBEAA8EFFABE2288" authority="(Bory) J. Sm." authorityName="J. Sm." baseAuthorityName="Bory" box="[1121,1398,2000,2027]" class="Polypodiopsida" family="Polypodiaceae" genus="Drynaria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polypodiales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B907BE5B6920EF0EFBEAA8EFFB062289" box="[1121,1230,2001,2026]" italics="true" pageId="6" pageNumber="7">Drynaria</emphasis>
(Bory) J. Sm.
</taxonomicName>
</paragraph>
</caption>
<caption id="DF0C32C16921EF0FFF36A9E6FDCA228A" ID-DOI="http://doi.org/10.5281/zenodo.3829752" ID-Zenodo-Dep="3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1752,1778]" targetBox="[189,1399,375,1710]" targetPageId="7">
<paragraph id="8BCC62496921EF0FFF36A9E6FDCA228A" blockId="7.[189,1399,1752,2026]" pageId="7" pageNumber="8">
<emphasis id="B907BE5B6921EF0FFF36A9E6FEC52390" bold="true" box="[189,269,1752,1779]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
Shells of 
<taxonomicName id="4C7319CA6921EF0FFE07A9E7FD3A2390" authority="(Amphidromus)" box="[396,754,1753,1779]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="subGenus" subGenus="Amphidromus">
<emphasis id="B907BE5B6921EF0FFE07A9E7FDF82390" box="[396,560,1753,1779]" italics="true" pageId="7" pageNumber="8">Amphidromus</emphasis>
(
<emphasis id="B907BE5B6921EF0FFDCFA9E7FD202390" box="[580,744,1753,1779]" italics="true" pageId="7" pageNumber="8">Amphidromus</emphasis>
)
</taxonomicName>
spp. 
<emphasis id="B907BE5B6921EF0FFCBDA9E6FCA62391" bold="true" box="[822,878,1752,1778]" pageId="7" pageNumber="8">A−F</emphasis>
. 
<taxonomicName id="4C7319CA6921EF0FFCF4A9E7FABA2390" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[895,1394,1753,1779]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B6921EF0FFCF4A9E7FB042390" box="[895,1228,1753,1779]" italics="true" pageId="7" pageNumber="8">Amphidromus roseolabiatus</emphasis>
<bibRefCitation id="EFE21FB86921EF0FFB5DA9E7FABA2390" author="Fulton H. C." box="[1238,1394,1753,1779]" pageId="7" pageNumber="8" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B907BE5B6921EF0FFF36A9C2FF3C2275" bold="true" box="[189,244,1788,1814]" pageId="7" pageNumber="8">A–B</emphasis>
. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). 
<emphasis id="B907BE5B6921EF0FFBD6A9C2FB5E2275" bold="true" box="[1117,1174,1788,1814]" pageId="7" pageNumber="8">C–D</emphasis>
. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). 
<emphasis id="B907BE5B6921EF0FFC35A821FC3A225A" bold="true" box="[958,1010,1823,1849]" pageId="7" pageNumber="8">E–F</emphasis>
. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). 
<emphasis id="B907BE5B6921EF0FFD77A87CFCDB223F" bold="true" box="[764,787,1858,1884]" pageId="7" pageNumber="8">G</emphasis>
. 
<taxonomicName id="4C7319CA6921EF0FFCA9A87CFB11223F" authority="Bavay, 1898" authorityName="Bavay" authorityYear="1898" box="[802,1241,1858,1885]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="laosianus">
<emphasis id="B907BE5B6921EF0FFCA9A87CFBF5223F" box="[802,1085,1858,1884]" italics="true" pageId="7" pageNumber="8">Amphidromus laosianus</emphasis>
<bibRefCitation id="EFE21FB86921EF0FFBCDA87DFB11223F" author="Bavay A." box="[1094,1241,1858,1885]" pageId="7" pageNumber="8" pagination="15 - 20" refId="ref18014" refString="Bavay A. 1898. Coquilles nouvelles provenant des recoltes de M. L. Levay dans le Haut-Mekong pendant la campagne du Massie (1893 - 95). Journal de Conchyliologie 46: 15 - 20. Available from http: // biodiversitylibrary. org / page / 15920414 [accessed 17 Jul. 2013]." type="journal article" year="1898">Bavay, 1898</bibRefCitation>
</taxonomicName>
from Khone, Champasak, Laos (RMNH 101049). 
<emphasis id="B907BE5B6921EF0FFDFEA858FD4422E3" bold="true" box="[629,652,1894,1920]" pageId="7" pageNumber="8">H</emphasis>
. 
<taxonomicName id="4C7319CA6921EF0FFD17A85BFACB22E3" authority="Lehmann &amp; Maassen, 2004" authorityName="Lehmann &amp; Maassen" authorityYear="2004" box="[668,1283,1893,1920]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="protania">
<emphasis id="B907BE5B6921EF0FFD17A85BFC65221C" box="[668,941,1893,1919]" italics="true" pageId="7" pageNumber="8">Amphidromus protania</emphasis>
<bibRefCitation id="EFE21FB86921EF0FFC3DA85BFACB22E3" author="Lehmann H. &amp; Maassen W. J. M." box="[950,1283,1893,1920]" pageId="7" pageNumber="8" pagination="17 - 20" refId="ref18701" refString="Lehmann H. &amp; Maassen W. J. M. 2004. A new species of Amphidromus from Laos (Gastropoda, Pulmonata, Camaenidae). Basteria 68: 17 - 20. Available from http: // natuurtijdschriften. nl / download? t ype = document &amp; docid = 597271 [accessed 8 Jun. 2016]." type="journal article" year="2004">Lehmann &amp; Maassen, 2004</bibRefCitation>
</taxonomicName>
, holotype (RMNH 98143). 
<emphasis id="B907BE5B6921EF0FFE0FA8B7FE4722C0" bold="true" box="[388,399,1929,1955]" pageId="7" pageNumber="8">I</emphasis>
. 
<taxonomicName id="4C7319CA6921EF0FFE17A8B7FC8B22C0" authority="Geret, 1912" authorityName="Geret" authorityYear="1912" box="[412,835,1929,1955]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="givenchyi">
<emphasis id="B907BE5B6921EF0FFE17A8B7FD7D22C0" box="[412,693,1929,1955]" italics="true" pageId="7" pageNumber="8">Amphidromus givenchyi</emphasis>
<bibRefCitation id="EFE21FB86921EF0FFD30A8B7FC8B22C0" author="Geret P." box="[699,835,1929,1955]" pageId="7" pageNumber="8" pagination="55 - 56" refId="ref18483" refString="Geret P. 1912. Description d'un nouvel Amphidromus. Journal de Conchyliologie 60: 55 - 56. Available from http: // biodiversitylibrary. org / page / 16358498 [accessed 15 Jul. 2013]." type="journal article" year="1912">Geret, 1912</bibRefCitation>
</taxonomicName>
from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). 
<emphasis id="B907BE5B6921EF0FFE9AA892FE8D22A5" bold="true" box="[273,325,1964,1990]" pageId="7" pageNumber="8">J–K</emphasis>
. 
<taxonomicName id="4C7319CA6921EF0FFEDDA892FD7B22A5" authority="Inkhavilay &amp; Sutcharit &amp; Panha, 2017" authorityName="Inkhavilay &amp; Sutcharit &amp; Panha" authorityYear="2017" box="[342,691,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="syndromoideus" status="sp. nov.">
<emphasis id="B907BE5B6921EF0FFEDDA892FD7B22A5" box="[342,691,1964,1990]" italics="true" pageId="7" pageNumber="8">Amphidromus syndromoideus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23403206921EF0FFD30A892FCDF22A5" box="[699,791,1964,1990]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
from the type locality. 
<emphasis id="B907BE5B6921EF0FFBA4A892FBF622A5" bold="true" box="[1071,1086,1964,1990]" pageId="7" pageNumber="8">J</emphasis>
. Holotype (CUMZ 7019). 
<emphasis id="B907BE5B6921EF0FFF36A8F1FF1C228A" bold="true" box="[189,212,1999,2025]" pageId="7" pageNumber="8">K</emphasis>
. Paratype (CUMZ 7020).
</paragraph>
</caption>
<paragraph id="8BCC6249692EEF00FF36AE32FC2E2445" blockId="8.[189,998,267,294]" box="[189,998,267,294]" pageId="8" pageNumber="9">
<collectingCountry id="F36422D9692EEF00FF36AE32FE812445" box="[189,329,268,294]" name="Vietnam" pageId="8" pageNumber="9">VIETNAM</collectingCountry>
: 2D+3S shells, Phong Nha National Park (
<collectionCode id="ED62FA8C692EEF00FCB7AE35FC5B2446" box="[828,915,267,293]" country="0" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="9">CUMZ</collectionCode>
7053).
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AE71FC6D2409" blockId="8.[189,933,335,362]" box="[189,933,335,362]" pageId="8" pageNumber="9">
<collectingCountry id="F36422D9692EEF00FF36AE71FE91240A" box="[189,345,335,361]" name="Thailand" pageId="8" pageNumber="9">THAILAND</collectingCountry>
: 1D shell, 
<collectingRegion id="49B7ACAB692EEF00FE5EAE6EFDB0240A" box="[469,632,335,362]" country="Thailand" name="Nan" pageId="8" pageNumber="9">Nan Province</collectingRegion>
, Thailand (
<collectionCode id="ED62FA8C692EEF00FD70AE71FC9A240A" box="[763,850,335,361]" country="0" name="Cameroon University, Museum of Zoology" pageId="8" pageNumber="9">CUMZ</collectionCode>
7054).
</paragraph>
</subSubSection>
<subSubSection id="C36931C2692EEF00FF36AEADFB97230F" pageId="8" pageNumber="9" type="description">
<paragraph id="8BCC6249692EEF00FF36AEADFE8624CE" blockId="8.[189,334,403,429]" box="[189,334,403,429]" pageId="8" pageNumber="9">
<heading id="D084D525692EEF00FF36AEADFE8624CE" bold="true" box="[189,334,403,429]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B907BE5B692EEF00FF36AEADFE8624CE" bold="true" box="[189,334,403,429]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AEFAFE432786" blockId="8.[189,1399,452,741]" pageId="8" pageNumber="9">Shell chirally dimorphic, elongate to ovate conical, rather thin and glossy. Spire conical; apex acute, light brown and without black spot on tip. Whorls 6 to 7 nearly smooth; suture wide and depressed; last whorl rounded. Periostracum usually deciduous to yellowish-green radial streaks, more conspicuous on last whorl and faded in previous whorls. Last whorl processes thin, whitish subsutural band, sometimes with one or two reddish-brown spiral bands below periphery but usually not reaching lip; varix absent. Parietal callus thin and transparent. Aperture ovate to elongate; peristome expanded and not reflected; lip usually purplish-pink or white. Columella straight, thickened, purplish-pink or white. Umbilicus narrowly opened.</paragraph>
<paragraph id="8BCC6249692EEF00FF61AC31FE8D264A" blockId="8.[189,1398,783,997]" box="[234,325,783,809]" pageId="8" pageNumber="9">
<emphasis id="B907BE5B692EEF00FF61AC31FE8D264A" bold="true" box="[234,325,783,809]" pageId="8" pageNumber="9">Radula</emphasis>
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AC0AFABE2687" blockId="8.[189,1398,783,997]" pageId="8" pageNumber="9">
Each row contains about 130 (66-(10-6)-1-(6-10)-63) teeth. Central tooth unicuspid and spatulate with curved cusp. Lateral teeth bicuspid. Endocone cylindrical, slightly oblique, with wide notch; ectocone large, with truncated to rounded cusp. Marginal teeth tricuspid, start around tooth number 6 to 10; endocone small; mesocone large, with rounded cusp, ectocone small, with sharp cusp. Outermost teeth with small and sharp cusp on endocone and extocone; mesocone large, with curved cusps (
<figureCitation id="13487ECC692EEF00FB6DACF4FAE32687" box="[1254,1323,970,996]" captionStart="Fig" captionStartId="12.[189,232,1823,1849]" captionTargetBox="[196,1393,710,1775]" captionTargetId="figure@12.[189,1399,704,1781]" captionTargetPageId="12" captionText="Fig. 7. SEM images of the radula. A–C. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos (CUMZ 7012). D–F. Amphidromus syndromoideus sp. nov., holotype (CUMZ 7019). G–I. Amphidromus areolatus (Pfeiffer, 1861) from Thad Fek, Attapue, Laos (CUMZ 7023). A, D, G = central tooth with the first to fifth to eighth lateral teeth; B, E, H = lateral teeth with the tricuspid marginal teeth transition; C, F, I = outermost marginal teeth. Numbers indicate the order of the lateral and marginal teeth. Central tooth indicated by ‘C’." figureDoi="http://doi.org/10.5281/zenodo.3829758" httpUri="https://zenodo.org/record/3829758/files/figure.png" pageId="8" pageNumber="9">Fig. 7</figureCitation>
A−C).
</paragraph>
<paragraph id="8BCC6249692EEF00FF61AB30FE6C214B" blockId="8.[189,1399,1038,1253]" box="[234,420,1038,1064]" pageId="8" pageNumber="9">
<emphasis id="B907BE5B692EEF00FF61AB30FE6C214B" bold="true" box="[234,420,1038,1064]" pageId="8" pageNumber="9">Genital organs</emphasis>
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AB0AFE0E2186" blockId="8.[189,1399,1038,1253]" pageId="8" pageNumber="9">
Atrium (at) rather short (n = 10). Penis (p) long and stout. Epiphallus (e) long, about two times as long as penis; flagellum (fl) longer than epiphallus and terminated in folded coil. Appendix very small, extends from folded coil of flagellum, about same length as epiphallus. Penial retractor muscle (pr) short, thickened and inserted distally on penis. Vas deferens (vd) narrow tube connecting epiphallus and free oviduct (
<figureCitation id="13487ECC692EEF00FED1ABF4FE712186" box="[346,441,1226,1253]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="8" pageNumber="9">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AA30FEBB2017" blockId="8.[189,1399,1294,1396]" pageId="8" pageNumber="9">
Internal penial wall corrugated into series of thick penial pilasters (pp), which form a fringe around penial verge. Penial verge (pv) rather long and curved, conical, with smooth surface and orifice located at tip (
<figureCitation id="13487ECC692EEF00FE82AA67FEAD2017" box="[265,357,1369,1396]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="8" pageNumber="9">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="8BCC6249692EEF00FF36AAA3FAAE2360" blockId="8.[189,1398,1437,1539]" pageId="8" pageNumber="9">
Vagina cylindrical, about two times as long as penis. Gametolytic duct very long, proximal to genital orifice large, coiled; distal to genital orifice tapering to small tube, short, about one-third length of proximal part and connected to enlarged gametolytic sac. Oviduct and albumin gland small (
<figureCitation id="13487ECC692EEF00FB71AAD6FA912360" box="[1274,1369,1512,1539]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="8" pageNumber="9">Fig. 6A</figureCitation>
).
</paragraph>
<paragraph id="8BCC6249692EEF00FF36A915FB97230F" blockId="8.[189,1398,1579,1644]" pageId="8" pageNumber="9">
Internally, vaginal wall performs longitudinal vaginal pilasters (vp); proximal to genital orifice with small corrugated continuous ridges, ridges distally becoming smooth (
<figureCitation id="13487ECC692EEF00FC7EA96FFB99230F" box="[1013,1105,1617,1644]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="8" pageNumber="9">Fig. 6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36931C2692EEF00FF36A9ABFB992266" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BCC6249692EEF00FF36A9ABFE9E23CC" blockId="8.[189,342,1685,1711]" box="[189,342,1685,1711]" pageId="8" pageNumber="9">
<heading id="D084D525692EEF00FF36A9ABFE9E23CC" bold="true" box="[189,342,1685,1711]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B907BE5B692EEF00FF36A9ABFE9E23CC" bold="true" box="[189,342,1685,1711]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692EEF00FF36A9F8FB992266" blockId="8.[189,1398,1734,1798]" pageId="8" pageNumber="9">
This species is distributed widely, from many localities in central and northern 
<collectingCountry id="F36422D9692EEF00FB0BA9F8FB752383" box="[1152,1213,1734,1760]" name="Laos" pageId="8" pageNumber="9">Laos</collectingCountry>
. There are also records from 
<collectingRegion id="49B7ACAB692EEF00FED0A9D2FE362266" box="[347,510,1771,1798]" country="Thailand" name="Nan" pageId="8" pageNumber="9">Nan Province</collectingRegion>
, 
<collectingCountry id="F36422D9692EEF00FD81A9D5FDBA2266" box="[522,626,1771,1797]" name="Thailand" pageId="8" pageNumber="9">Thailand</collectingCountry>
and Phong Nha National Park, 
<collectingCountry id="F36422D9692EEF00FC6DA9D5FB852266" box="[998,1101,1771,1797]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36931C2692EEF01FF36A811FD49272F" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BCC6249692EEF00FF36A811FEE7222A" blockId="8.[189,303,1839,1865]" box="[189,303,1839,1865]" pageId="8" pageNumber="9">
<heading id="D084D525692EEF00FF36A811FEE7222A" bold="true" box="[189,303,1839,1865]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B907BE5B692EEF00FF36A811FEE7222A" bold="true" box="[189,303,1839,1865]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC6249692EEF00FF36A85EFAF32288" blockId="8.[189,1399,1888,2027]" pageId="8" pageNumber="9">
<taxonomicName id="4C7319CA692EEF00FF36A85EFDC02219" authorityName="Fulton" authorityYear="1896" box="[189,520,1888,1914]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B692EEF00FF36A85EFDC02219" box="[189,520,1888,1914]" italics="true" pageId="8" pageNumber="9">Amphidromus roseolabiatus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C7319CA692EEF00FD2CA85EFC792219" authority="Fulton, 1896" authorityName="Fulton" authorityYear="1896" box="[679,945,1888,1914]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="smithi">
<emphasis id="B907BE5B692EEF00FD2CA85EFCD92219" box="[679,785,1888,1914]" italics="true" pageId="8" pageNumber="9">A. smithi</emphasis>
<bibRefCitation id="EFE21FB8692EEF00FC92A85EFC792219" author="Fulton H. C." box="[793,945,1888,1914]" pageId="8" pageNumber="9" pagination="66 - 94" refId="ref18418" refString="Fulton H. C. 1896. A list of the species of Amphidromus, Albers, with critical notes and descriptions of some hitherto undescribed species and varieties. Annals and Magazine of Natural History, Series 6 17: 66 - 94. Available from http: // biodiversitylibrary. org / page / 16006918 [accessed 19 Sept. 2013]." type="journal article" year="1896">Fulton, 1896</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7319CA692EEF00FC67A85EFABF2219" authority="Mollendorff, 1900" authorityName="Mollendorff" authorityYear="1900" box="[1004,1399,1888,1914]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ventrosulus">
<emphasis id="B907BE5B692EEF00FC67A85EFB5C2219" box="[1004,1172,1888,1914]" italics="true" pageId="8" pageNumber="9">A. ventrosulus</emphasis>
Möllendorff, 1900
</taxonomicName>
(
<bibRefCitation id="EFE21FB8692EEF00FF4EA8BBFE8222C3" author="Zilch A." box="[197,330,1925,1952]" pageId="8" pageNumber="9" pagination="131 - 140" refId="ref20321" refString="Zilch A. 1953. Die Typen und Typoide des Natur-Museums Senckenberg 10: Mollusca, Pleurodontidae (l). Archiv fur Molluskenkunde 82: 131 - 140." type="journal article" year="1953">Zilch 1953</bibRefCitation>
; 
<bibRefCitation id="EFE21FB8692EEF00FEDDA8BBFD8E22C3" author="Sutcharit C. &amp; Ablett J. &amp; Tongkerd P. &amp; Naggs F. &amp; Panha S." box="[342,582,1925,1952]" pageId="8" pageNumber="9" pagination="49 - 105" refId="ref19986" refString="Sutcharit C., Ablett J., Tongkerd P., Naggs F. &amp; Panha S. 2015. Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species. ZooKeys 492: 49 - 105. https: // doi. org / 10.3897 / zookeys. 492.8641" type="journal article" year="2015">
Sutcharit 
<emphasis id="B907BE5B692EEF00FE43A8B8FE3322C3" box="[456,507,1926,1952]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2015
</bibRefCitation>
) in having an enantiomorphic shell coiling, purplish-pink lip and fine green streaks. In contrast, 
<taxonomicName id="4C7319CA692EEF00FE73A892FDAA22A6" authorityName="Fulton" authorityYear="1896" box="[504,610,1963,1990]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="smithi">
<emphasis id="B907BE5B692EEF00FE73A892FDAA22A6" box="[504,610,1963,1990]" italics="true" pageId="8" pageNumber="9">A. smithi</emphasis>
</taxonomicName>
has a sinistral shell, brownish lip with dark spot on the apex, and 
<taxonomicName id="4C7319CA692EEF00FF36A8EFFEAC2288" authorityName="Mollendorff" authorityYear="1900" box="[189,356,2001,2027]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="ventrosulus">
<emphasis id="B907BE5B692EEF00FF36A8EFFEAC2288" box="[189,356,2001,2027]" italics="true" pageId="8" pageNumber="9">A. ventrosulus</emphasis>
</taxonomicName>
has a sinistral shell, uniform green color, elongate spire and more depressed suture.
</paragraph>
<paragraph id="8BCC6249692FEF01FF36AE35FD49272F" blockId="9.[189,1399,267,588]" pageId="9" pageNumber="10">
<bibRefCitation id="EFE21FB8692FEF01FF36AE35FE202445" author="Laidlaw F. F. &amp; Solem A." box="[189,488,267,294]" pageId="9" pageNumber="10" pagination="507 - 677" refId="ref18651" refString="Laidlaw F. F. &amp; Solem A. 1961. The land snail genus Amphidromus: A synoptic catalogue. Fieldiana: Zoology 41: 507 - 677. Available from http: // biodiversitylibrary. org / item / 20789 [accessed 18 Jul. 2013]." type="journal article" year="1961">Laidlaw &amp; Solem (1961)</bibRefCitation>
recognized 
<taxonomicName id="4C7319CA692FEF01FDF1AE32FCF32446" authorityName="Fulton" authorityYear="1896" box="[634,827,267,294]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="roseolabiatus">
<emphasis id="B907BE5B692FEF01FDF1AE32FCF32446" box="[634,827,267,294]" italics="true" pageId="9" pageNumber="10">A. roseolabiatus</emphasis>
</taxonomicName>
as belonging to an uncertain subgenus that was probably close to the subgenus 
<taxonomicName id="4C7319CA692FEF01FDA4AE0EFD782429" authorityName="Pilsbry" authorityYear="1900" box="[559,688,304,330]" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="subGenus" subGenus="Syndromus">
<emphasis id="B907BE5B692FEF01FDA4AE0EFD782429" box="[559,688,304,330]" italics="true" pageId="9" pageNumber="10">Syndromus</emphasis>
</taxonomicName>
. However, this species exhibits dimorphic shell coiling with a long flagellum and appendix, which are typical characters of the nominotypical subgenus (
<figureCitation id="13487ECC692FEF01FA82AE6BFAA22413" box="[1289,1386,341,368]" captionStart="Fig" captionStartId="11.[189,232,1894,1920]" captionTargetBox="[189,1399,348,1849]" captionTargetId="figure@11.[189,1399,348,1849]" captionTargetPageId="11" captionText="Fig. 6. Genitalia of Amphidromusspp. A–B. Amphidromus roseolabiatus Fulton, 1896 from Ban Phavong, Khammouan, Laos showing reproductive system and interior structures of penis and vaginal chamber (CUMZ 7012). C–D. Amphidromus syndromoideus sp. nov., holotype, showing general characteristics of the genital system and interior structures of penis and vagina chamber (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829756" httpUri="https://zenodo.org/record/3829756/files/figure.png" pageId="9" pageNumber="10">Fig. 6A</figureCitation>
). Two color forms were observed in the recently collected materials. The typical color form has a greenish radial streaked periostracum and pinkish apertural lip (
<figureCitation id="13487ECC692FEF01FCB0AEA1FC5424DA" box="[827,924,415,441]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[189,1399,1098,1849]" captionTargetId="figure@5.[189,1399,1098,1849]" captionTargetPageId="5" captionText="Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="9" pageNumber="10">Figs 2A</figureCitation>
, 
<figureCitation id="13487ECC692FEF01FC22AEA1FC7024DA" box="[937,952,415,441]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
A–B). There are some specimens with the typical color form showing one or two additional reddish-brown lower, peripheral bands (
<figureCitation id="13487ECC692FEF01FA87AEFAFAA124BD" box="[1292,1385,452,478]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="9" pageNumber="10">Fig. 4B</figureCitation>
). The second, whitish form usually has a greenish periostracum, as in the typical color form, but a white apertural lip (
<figureCitation id="13487ECC692FEF01FEE9AD33FE0E274B" box="[354,454,525,552]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[189,1399,1098,1849]" captionTargetId="figure@5.[189,1399,1098,1849]" captionTargetPageId="5" captionText="Fig. 2. Living snails. A–B. Amphidromus roseolabiatus Fulton, 1896. A. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7012). B. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7005). C. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7018). D. Amphidromus syndromoideus sp. nov., holotype from the type locality (CUMZ 7019)." figureDoi="http://doi.org/10.5281/zenodo.3829748" httpUri="https://zenodo.org/record/3829748/files/figure.png" pageId="9" pageNumber="10">Figs 2B</figureCitation>
, 
<figureCitation id="13487ECC692FEF01FE5EAD30FE2C274B" box="[469,484,526,552]" captionStart="Fig" captionStartId="7.[189,232,1752,1778]" captionTargetBox="[189,1399,375,1710]" captionTargetId="figure@7.[189,1399,375,1710]" captionTargetPageId="7" captionText="Fig. 4. Shells of Amphidromus (Amphidromus) spp. A−F. Amphidromus roseolabiatus Fulton, 1896. A–B. Typical form from Ban Phavong, Khammouan, Laos (CUMZ 7011). C–D. Form with white lip from Nam Turn Bridge, Bolikhamxay, Laos (CUMZ 7006). E–F. Form with white lip from Tam Mung Korn, Bolikhamxay, Laos (CUMZ 7004). G. Amphidromus laosianus Bavay, 1898 from Khone, Champasak, Laos (RMNH 101049). H. Amphidromus protania Lehmann &amp; Maassen, 2004, holotype (RMNH 98143). I. Amphidromus givenchyi Geret, 1912 from Thad Lor Waterfall, Salavan, Laos (CUMZ 7015). J–K. Amphidromus syndromoideus sp. nov. from the type locality. J. Holotype (CUMZ 7019). K. Paratype (CUMZ 7020)." figureDoi="http://doi.org/10.5281/zenodo.3829752" httpUri="https://zenodo.org/record/3829752/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
C–F). However, these two color forms have identical genital structures and sculptures inside the penis and vagina.
</paragraph>
</subSubSection>
</treatment>
</document>