treatments-xml/data/03/CE/F9/03CEF92012742941F628FC25FD29F8A5.xml

<document id="D6480BCC3F695037F0DA04BE220EC679" ID-CLB-Dataset="3698" ID-DOI="10.35929/RSZ.0048" ID-GBIF-Dataset="880cb736-92eb-42c0-b026-54b62378344f" ID-ISSN="0035-418" ID-Zenodo-Dep="5639911" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635892246867" checkinUser="felipe" docAuthor="Lienhard, Charles" docDate="2021" docId="03CEF92012742941F628FC25FD29F8A5" docLanguage="en" docName="RevSuisseZool.128.2.227-235.pdf" docOrigin="Revue suisse de Zoologie 128 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Prionoglaris kapralovi Lienhard 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="233" masterDocId="FFF7815812762946F734FFE6FF95FFCA" masterDocTitle="A new species of Prionoglaris Enderlein (Psocodea: ‘ Psocoptera’: Prionoglarididae) from an Armenian cave, with an account of the distribution of the genus" masterLastPageNumber="235" masterPageNumber="227" pageNumber="228" updateTime="1699467450576" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="03E34C1AAC274BFCB77A6140E1C24DB3">A new species of Prionoglaris Enderlein (Psocodea: ‘ Psocoptera’: Prionoglarididae) from an Armenian cave, with an account of the distribution of the genus</mods:title>
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<mods:namePart id="FDFA6AAC8FD57F55A913ABFA7448FDB8">Lienhard, Charles</mods:namePart>
<mods:affiliation id="F7B48ED75ADF184206EF700042D08AB9">Muséum d’histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
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<treatment id="03CEF92012742941F628FC25FD29F8A5" ID-DOI="http://doi.org/10.5281/zenodo.5646277" ID-GBIF-Taxon="189990954" ID-Zenodo-Dep="5646277" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5" httpUri="http://treatment.plazi.org/id/03CEF92012742941F628FC25FD29F8A5" lastPageId="7" lastPageNumber="233" pageId="2" pageNumber="228">
<subSubSection id="C37D1BBD12742944F628FC25FDE5FC10" box="[284,624,962,986]" pageId="2" pageNumber="228" type="nomenclature">
<paragraph id="8BD8483612742944F628FC25FDE5FC10" blockId="2.[284,624,962,1017]" box="[284,624,962,986]" pageId="2" pageNumber="228">
<heading id="D090FF5A12742944F628FC25FDE5FC10" bold="true" box="[284,624,962,986]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="228" reason="2">
<emphasis id="B913942412742944F628FC25FDE5FC10" bold="true" box="[284,624,962,986]" pageId="2" pageNumber="228">
<taxonomicName id="4C6733B512742944F628FC25FD81FC10" authority="Lienhard, 2021" authorityName="Lienhard" authorityYear="2021" box="[284,532,963,986]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="228" phylum="Arthropoda" rank="species" species="kapralovi" status="sp. nov.">
<emphasis id="B913942412742944F628FC25FD81FC10" bold="true" box="[284,532,963,986]" italics="true" pageId="2" pageNumber="228">Prionoglaris kapralovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220295F12742944F52EFC24FDE5FC10" box="[538,624,962,986]" pageId="2" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C37D1BBD12742944F6ADFC07FE64FC33" box="[409,497,993,1017]" pageId="2" pageNumber="228" type="description">
<paragraph id="8BD8483612742944F6ADFC07FE64FC33" blockId="2.[284,624,962,1017]" box="[409,497,993,1017]" pageId="2" pageNumber="228">
<figureCitation id="135C54B312742944F6ADFC07FE64FC33" box="[409,497,993,1017]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[151,186,1997,2019]" captionStartId-1="4.[151,186,1465,1487]" captionTargetBox-0="[151,1436,709,1956]" captionTargetBox-1="[158,1427,263,1417]" captionTargetId-0="figure-237@3.[151,1436,709,1956]" captionTargetId-1="figure-268@4.[151,1436,254,1434]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." captionText-1="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5639915" figureDoi-1="http://doi.org/10.5281/zenodo.5639917" httpUri-0="https://zenodo.org/record/5639915/files/figure.png" httpUri-1="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Figs 1-2</figureCitation>
</paragraph>
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<subSubSection id="C37D1BBD12742944F7A3FBF6FE3AFB1E" pageId="2" pageNumber="228" type="materials_examined">
<paragraph id="8BD8483612742944F7A3FBF6FD76FB4C" blockId="2.[151,756,1040,1158]" pageId="2" pageNumber="228">
<materialsCitation id="3B0F426B12742944F7A3FBF6FD76FB4C" ID-GBIF-Occurrence="3398241315" collectedFrom="under stones near cave entrance" collectingDate="2018-08-14" collectionCode="MHNG" collectorName="S. A. Kapralov." country="Armenia" elevation="1540" latitude="39.71258" location="Mozrov Cave" longLatPrecision="1" longitude="45.25953" pageId="2" pageNumber="228" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B913942412742944F7A3FBF6FE9CFBE2" bold="true" box="[151,265,1040,1064]" pageId="2" pageNumber="228">
<typeStatus id="54DCF69412742944F7A3FBF6FE91FBE2" box="[151,260,1040,1064]" pageId="2" pageNumber="228" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED76D0F312742944F621FBF6FEFFFBE2" box="[277,362,1040,1064]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="228" type="Museum">MHNG</collectionCode>
; 
<specimenCount id="9D6183BF12742944F64FFBF6FE3AFBE2" box="[379,431,1040,1064]" pageId="2" pageNumber="228" type="male">male</specimenCount>
(some parts slide-mounted); 
<collectingCountry id="F37008A612742944F7A3FBC9FF62FB8D" box="[151,247,1071,1095]" name="Armenia" pageId="2" pageNumber="228">Armenia</collectingCountry>
, 
<locationDeviation id="9A6A511412742944F631FBC9FDDCFB8D" box="[261,585,1071,1095]" location="Yeghegnadzor" pageId="2" pageNumber="228">
<quantity id="4C9FE5D312742944F631FBC9FEC6FB8D" box="[261,339,1071,1095]" metricMagnitude="3" metricUnit="m" metricValue="8.3" pageId="2" pageNumber="228" unit="km" value="8.3">8.3 km</quantity>
SW of Yeghegnadzor
</locationDeviation>
, 
<location id="8EB81EED12742944F563FBD6FD7AFB8D" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12742944F563FBD6FD7AFB8D" box="[599,751,1071,1095]" country="Armenia" latitude="39.71258" longLatPrecision="1" longitude="45.25953" name="Mozrov Cave" pageId="2" pageNumber="228">Mozrov Cave</location>
; WGS84 
<geoCoordinate id="EE532EF112742944F7C0FBA9FECEFBAD" box="[244,347,1103,1127]" degrees="39.71258" direction="north" orientation="latitude" pageId="2" pageNumber="228" precision="1" value="39.71258">39.71258</geoCoordinate>
, 
<geoCoordinate id="EE532EF112742944F651FBA9FE59FBAD" box="[357,460,1103,1127]" degrees="45.25953" direction="east" orientation="longitude" pageId="2" pageNumber="228" precision="1" value="45.25953">45.25953</geoCoordinate>
; 
<elevation id="004AAF0512742944F6E3FBA9FDF4FBAD" box="[471,609,1103,1127]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="228" unit="m" value="1540.0">
<quantity id="4C9FE5D312742944F6E3FBA9FDBDFBAD" box="[471,552,1103,1127]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="228" unit="m" value="1540.0">1540 m</quantity>
a.s.l.
</elevation>
, 
<collectedFrom id="387B0BCB12742944F55AFBA9FEF4FB4C" pageId="2" pageNumber="228">under stones near cave entrance</collectedFrom>
; 
<date id="FFD96EF612742944F658FB88FD93FB4C" box="[364,518,1134,1158]" pageId="2" pageNumber="228" value="2018-08-14">
<collectingDate id="EF9D971E12742944F658FB88FD93FB4C" box="[364,518,1134,1158]" pageId="2" pageNumber="228" value="2018-08-14">14.AUG.2018</collectingDate>
</date>
; leg. 
<collectorName id="26922DE012742944F50BFB88FD76FB4C" box="[575,739,1134,1158]" pageId="2" pageNumber="228">S. A. Kapralov.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BD8483612742944F7A3FB7BFE3AFB1E" blockId="2.[151,756,1181,1236]" pageId="2" pageNumber="228">
<materialsCitation id="3B0F426B12742944F7A3FB7BFE3AFB1E" ID-GBIF-Occurrence="3398241328" collectedFrom="under stones near cave entrance" collectingDate="2018-08-14" collectionCode="MHNG" collectorName="S. A. Kapralov." country="Armenia" elevation="1540" latitude="39.71258" location="Mozrov Cave" longLatPrecision="1" longitude="45.25953" pageId="2" pageNumber="228" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B913942412742944F7A3FB7BFE9DFB7F" bold="true" box="[151,264,1181,1205]" pageId="2" pageNumber="228">
<typeStatus id="54DCF69412742944F7A3FB7BFE96FB7F" box="[151,259,1181,1205]" pageId="2" pageNumber="228" type="paratype">Paratype</typeStatus>
:
</emphasis>
<collectionCode id="ED76D0F312742944F625FB7BFEF3FB7F" box="[273,358,1181,1205]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="228" type="Museum">MHNG</collectionCode>
; female (some parts slide-mounted); same data as for holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12742944F7A3FB0DFE9EFA6A" pageId="2" pageNumber="228" type="diagnosis">
<paragraph id="8BD8483612742944F7A3FB0DFE9EFA6A" blockId="2.[151,757,1259,1440]" pageId="2" pageNumber="228">
<emphasis id="B913942412742944F7A3FB0DFE85FAC9" bold="true" box="[151,272,1259,1283]" pageId="2" pageNumber="228">Diagnosis:</emphasis>
This new species can be distinguished from all known species of the genus 
<taxonomicName id="4C6733B512742944F570FAECFD58FAE8" authorityName="Enderlein" authorityYear="1909" box="[580,717,1290,1314]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412742944F570FAECFD58FAE8" box="[580,717,1290,1314]" italics="true" pageId="2" pageNumber="228">Prionoglaris</emphasis>
</taxonomicName>
by the absence of an internal membranous vesicle on the anterior claw and by the absence of a basal tooth on the posterior claw of each pretarsus (
<figureCitation id="135C54B312742944F539FA8FFDC4FA4B" box="[525,593,1385,1409]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2</figureCitation>
C-D). See also key below.
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12742944F7A3FA51FDC2FA25" pageId="2" pageNumber="228" type="etymology">
<paragraph id="8BD8483612742944F7A3FA51FDC2FA25" blockId="2.[151,756,1463,1520]" pageId="2" pageNumber="228">
<emphasis id="B913942412742944F7A3FA51FE88FA05" bold="true" box="[151,285,1463,1487]" pageId="2" pageNumber="228">Etymology:</emphasis>
The species is dedicated to Sergei A. Kapralov who collected the 
<typeStatus id="54DCF69412742944F6F0FA3EFE67FA3A" box="[452,498,1496,1520]" pageId="2" pageNumber="228">type</typeStatus>
material.
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12742945F40BFEE7FD4CFEAA" lastPageId="3" lastPageNumber="229" pageId="2" pageNumber="228" type="description">
<paragraph id="8BD8483612742944F40BFEE7FC56FED3" blockId="2.[831,1437,257,1517]" box="[831,963,257,281]" pageId="2" pageNumber="228">
<emphasis id="B913942412742944F40BFEE7FC56FED3" bold="true" box="[831,963,257,281]" pageId="2" pageNumber="228">Description</emphasis>
</paragraph>
<paragraph id="8BD8483612742944F40BFEC7FBBCFDB4" blockId="2.[831,1437,257,1517]" pageId="2" pageNumber="228">
<emphasis id="B913942412742944F40BFEC7FB9AFEF3" box="[831,1039,289,313]" italics="true" pageId="2" pageNumber="228">General characters</emphasis>
: Head and thorax (including antenna, maxillary palp and legs) yellowish to light or medium brown. Vertex, frons and genae with some small brown patches. Compound eye uniformly dark grey to black (after 2 years in alcohol). Wings hyaline, veins medium brown, pterostigma colourless but slightly opaque (
<figureCitation id="135C54B312742944F473FE03FC09FE37" box="[839,924,485,509]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1B</figureCitation>
). Abdomen whitish, dorsally and laterally with numerous patches of dark hypodermal pigment forming irregular transversal bands (
<figureCitation id="135C54B312742944F358FDC0FB3DFDF4" box="[1132,1192,550,574]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1</figureCitation>
A-B). Terminalia light to medium brown, ventro-median process of phallosome dark brown (
<figureCitation id="135C54B312742944F4FCFD80FB89FDB4" box="[968,1052,614,638]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="8BD8483612742944F40BFD61FB56FAA1" blockId="2.[831,1437,257,1517]" pageId="2" pageNumber="228">
Mouthparts typical for the genus; lacinial rudiment with an acuminate tip (
<figureCitation id="135C54B312742944F33DFD41FBF5FD75" box="[1033,1120,679,703]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2E</figureCitation>
). Antennae long and slender (
<figureCitation id="135C54B312742944F473FD2EFC0BFD2A" box="[839,926,712,736]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1A</figureCitation>
), both damaged in male and female examined (highest number of flagellomeres observed is 7, in right antenna of 
<typeStatus id="54DCF69412742944F4C2FCEFFBC3FCEB" box="[1014,1110,777,801]" pageId="2" pageNumber="228" type="holotype">holotype</typeStatus>
, 7th flagellomere broken near base). Anterior claw of each pretarsus (
<figureCitation id="135C54B312742944F3C6FCCCFAD2FC88" box="[1266,1351,810,834]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2C</figureCitation>
) with a long basal seta, lacking internal membranous vesicle and preapical denticle. Posterior claw of each pretarsus simple (
<figureCitation id="135C54B312742944F4AAFC6DFC69FC69" box="[926,1020,907,931]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2D</figureCitation>
), lacking basal tooth and preapical denticle. Forewing broadly rounded, clearly longer than twice its width (FW/FWw about 2.4); venation typical for the genus (
<figureCitation id="135C54B312742944F4DEFC0BFBD3FBCF" box="[1002,1094,1005,1029]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2A</figureCitation>
), but AP somewhat shorter in male than in female (in male its marginal length about same as its height, see 
<figureCitation id="135C54B312742944F31AFBC8FB17FB8C" box="[1070,1154,1070,1094]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1A</figureCitation>
; in female marginal length slightly exceeding height, see 
<figureCitation id="135C54B312742944F394FBA9FB73FBAD" box="[1184,1254,1103,1127]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2</figureCitation>
A-B). Hindwing as in 
<taxonomicName id="4C6733B512742944F44BFB96FC49FB4D" authorityName="Enderlein" authorityYear="1909" box="[895,988,1136,1159]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="228" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412742944F44BFB96FC49FB4D" box="[895,988,1136,1159]" italics="true" pageId="2" pageNumber="228">P. stygia</emphasis>
</taxonomicName>
. Vein aberrations observed in forewings and hindwings: right forewing of female with a crossvein between 
<specimenCode id="DBC1E04D12742944F4E1FB56FC61FB02" box="[981,1012,1200,1224]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="228">R1</specimenCode>
and 
<specimenCode id="DBC1E04D12742944F31DFB56FBDEFB02" box="[1065,1099,1200,1224]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="228">R2</specimenCode>
+3, and M1 bifurcate near wing margin (
<figureCitation id="135C54B312742944F4A9FB37FC60FB23" box="[925,1013,1233,1257]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2B</figureCitation>
); right forewing of male with a short spurvein on 
<specimenCode id="DBC1E04D12742944F4FDFB17FC7FFAC3" box="[969,1002,1265,1289]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="228">R1</specimenCode>
, just basally to point where Sc reaches 
<specimenCode id="DBC1E04D12742944F40BFAF4FCCAFAE0" box="[831,863,1298,1322]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="228">R1</specimenCode>
(
<figureCitation id="135C54B312742944F444FAF4FC5CFAE0" box="[880,969,1298,1322]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1A</figureCitation>
); left hindwing of male with a transversal vein between 
<specimenCode id="DBC1E04D12742944F4E3FAD4FC62FA80" box="[983,1015,1330,1354]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="228">R1</specimenCode>
and Rs (
<figureCitation id="135C54B312742944F36EFAD4FB25FA80" box="[1114,1200,1330,1354]" captionStart="Fig" captionStartId="3.[151,186,1997,2019]" captionTargetBox="[151,1436,709,1956]" captionTargetId="figure-237@3.[151,1436,709,1956]" captionTargetPageId="3" captionText="Fig. 1. Prionoglaris kapralovi sp. nov. (A) Habitus of male, dorsal view of holotype, in alcohol (body length 3.3 mm). (B) Habitus of female, lateral view of paratype, in alcohol (body length 4.1 mm). (C) Male terminalia, ventral view of holotype, in alcohol. Photographs by S. A. Kapralov." figureDoi="http://doi.org/10.5281/zenodo.5639915" httpUri="https://zenodo.org/record/5639915/files/figure.png" pageId="2" pageNumber="228">Fig. 1A</figureCitation>
), and both hindwings of female with slightly aberrant M2.
</paragraph>
<paragraph id="8BD8483612742945F40BFA95FD4CFEAA" blockId="2.[831,1437,257,1517]" lastBlockId="3.[151,756,257,633]" lastPageId="3" lastPageNumber="229" pageId="2" pageNumber="228">
<emphasis id="B913942412742944F40BFA95FC75FA41" box="[831,992,1395,1419]" italics="true" pageId="2" pageNumber="228">Male genitalia</emphasis>
: Sac-like proximal body of phallosome relatively short (
<figureCitation id="135C54B312742944F4CEFA72FBABFA66" box="[1018,1086,1428,1452]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2</figureCitation>
H-I), about half of total length of phallosome; ventro-median process strongly curved in lateral view (
<figureCitation id="135C54B312742944F4C3FA33FBC7FA27" box="[1015,1106,1493,1517]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="2" pageNumber="228">Fig. 2H</figureCitation>
); dorso-lateral appendages of about same width between base and apex, only weakly narrowing towards apex (
<figureCitation id="135C54B312752945F6ECFEC2FDA6FEF6" box="[472,563,292,316]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="3" pageNumber="229">Fig. 2I</figureCitation>
); medio-internal appendage with one fine hair on each side (
<figureCitation id="135C54B312752945F550FEAEFD31FEAA" box="[612,676,328,352]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="3" pageNumber="229">Fig. 2</figureCitation>
H-I).
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12752945F7A3FE8DFB8EFE18" pageId="3" pageNumber="229" type="description">
<paragraph id="8BD8483612752945F7A3FE8DFA08FEF1" blockId="3.[151,756,257,633]" lastBlockId="3.[831,1437,257,315]" pageId="3" pageNumber="229">
<emphasis id="B913942412752945F7A3FE8DFEC1FE49" box="[151,340,363,387]" italics="true" pageId="3" pageNumber="229">Female genitalia</emphasis>
: Subgenital plate and gonapophyses typical for the genus, only external valvula well developed, bearing 6 apically hooked setae near broadly rounded apex. Spermapore sclerite a thick and heavily sclerotized plate, proximally broadly rounded, distally narrowly rounded (
<figureCitation id="135C54B312752945F659FDFCFE56FDF8" box="[365,451,538,562]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="3" pageNumber="229">Fig. 2F</figureCitation>
); spermathecal duct curved inside the sclerite (
<figureCitation id="135C54B312752945F6B6FDD8FE74FD9C" box="[386,481,574,598]" captionStart="Fig" captionStartId="4.[151,186,1465,1487]" captionTargetBox="[158,1427,263,1417]" captionTargetId="figure-268@4.[151,1436,254,1434]" captionTargetPageId="4" captionText="Fig. 2. Prionoglaris kapralovi sp. nov., female (A-G), male (H-I). (A) Left forewing. (B) Right forewing. (C) Anterior claw of hind pretarsus. (D) Posterior claw of hind pretarsus. (E) Lacinial rudiment (length 0.2 mm). (F) Spermapore sclerite and distal part of spermathecal duct, ventral view. (G) Idem, optical longitudinal section, lateral view. (H) Phallosome, lateral view. (I) Idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5639917" httpUri="https://zenodo.org/record/5639917/files/figure.png" pageId="3" pageNumber="229">Fig. 2G</figureCitation>
) and widened towards spermathecal vesicle shortly after the spermapore sclerite (length of the narrow portion of the duct, before widening, about 1.5 times length of spermapore sclerite).
</paragraph>
<paragraph id="8BD8483612752945F40BFEB2FB8EFE18" blockId="3.[831,1436,340,466]" pageId="3" pageNumber="229">
<emphasis id="B913942412752945F40BFEB2FC64FEA6" bold="true" box="[831,1009,340,364]" pageId="3" pageNumber="229">Measurements:</emphasis>
<emphasis id="B913942412752945F4CFFEB2FBA7FEA6" box="[1019,1074,340,364]" italics="true" pageId="3" pageNumber="229">Male</emphasis>
: BL = 
<quantity id="4C9FE5D312752945F3BEFEB2FB71FEA6" box="[1162,1252,340,364]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="229" unit="mm" value="3.3">3.3 mm</quantity>
; FW = 
<quantity id="4C9FE5D312752945F20BFEB2FA0DFEA6" box="[1343,1432,340,364]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="229" unit="mm" value="3.7">3.7 mm</quantity>
; FWw = 
<quantity id="4C9FE5D312752945F4AEFE90FC65FE44" box="[922,1008,374,398]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="229" unit="mm" value="1.5">1.5 mm</quantity>
; HW = 
<quantity id="4C9FE5D312752945F373FE90FB0BFE44" box="[1095,1182,374,398]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="229" unit="mm" value="2.5">2.5 mm</quantity>
; IO/D = 1.8. – 
<emphasis id="B913942412752945F271FE90FA00FE44" box="[1349,1429,374,398]" italics="true" pageId="3" pageNumber="229">Female</emphasis>
: BL = 
<quantity id="4C9FE5D312752945F4B6FE7EFC4FFE7A" box="[898,986,408,432]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="229" unit="mm" value="4.1">4.1 mm</quantity>
; FW = 
<quantity id="4C9FE5D312752945F305FE7EFB1DFE7A" box="[1073,1160,408,432]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="3" pageNumber="229" unit="mm" value="4.1">4.1 mm</quantity>
; FWw = 
<quantity id="4C9FE5D312752945F3C7FE7EFADEFE7A" box="[1267,1355,408,432]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="229" unit="mm" value="1.7">1.7 mm</quantity>
; HW = 
<quantity id="4C9FE5D312752945F40BFE5CFC01FE1B" box="[831,916,442,466]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="229" unit="mm" value="2.7">2.7 mm</quantity>
; IO/D = 1.9.
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12752945F40BFE0DFC31FDEF" pageId="3" pageNumber="229" type="distribution">
<paragraph id="8BD8483612752945F40BFE0DFC31FDEF" blockId="3.[831,1436,491,549]" pageId="3" pageNumber="229">
<emphasis id="B913942412752945F40BFE0DFC42FDC9" bold="true" box="[831,983,491,515]" pageId="3" pageNumber="229">Distribution:</emphasis>
Only known from 
<typeStatus id="54DCF69412752945F3E9FE0DFA99FDC9" box="[1245,1292,491,515]" pageId="3" pageNumber="229">type</typeStatus>
locality in 
<collectingCountry id="F37008A612752945F40BFDEBFC35FDEF" box="[831,928,525,549]" name="Armenia" pageId="3" pageNumber="229">Armenia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12752942F40BFDD8FF45F969" lastPageId="4" lastPageNumber="230" pageId="3" pageNumber="229" type="distribution">
<paragraph id="8BD8483612752942F40BFDD8FF45F969" blockId="3.[831,1437,574,631]" lastBlockId="4.[151,756,1642,1699]" lastPageId="4" lastPageNumber="230" pageId="3" pageNumber="229">
<emphasis id="B913942412752945F40BFDD8FC6DFD9C" bold="true" box="[831,1016,574,598]" pageId="3" pageNumber="229">Type locality:</emphasis>
<collectingCountry id="F37008A612752945F323FDD8FBE2FD9C" box="[1047,1143,574,598]" name="Armenia" pageId="3" pageNumber="229">Armenia</collectingCountry>
, Mozrov Cave near Yeghegnadzor, 
<quantity id="4C9FE5D312752945F4D1FDB9FBA2FDBD" box="[997,1079,607,631]" metricMagnitude="3" metricUnit="m" metricValue="1.54" pageId="3" pageNumber="229" unit="m" value="1540.0">1540 m</quantity>
a.s.l. This cave is situated on the rolling steppe-like plateau on the left bank of the Arpi river.
</paragraph>
</subSubSection>
<subSubSection id="C37D1BBD12722943F7A3F95AFB76FDB4" lastPageId="5" lastPageNumber="231" pageId="4" pageNumber="230" type="discussion">
<paragraph id="8BD8483612722942F7A3F95AFC69F7D7" blockId="4.[151,757,1723,2077]" lastBlockId="4.[831,1437,1642,2077]" pageId="4" pageNumber="230">
<emphasis id="B913942412722942F7A3F95AFE9EF91E" bold="true" box="[151,267,1724,1748]" pageId="4" pageNumber="230">Remarks:</emphasis>
Judging from most morphological characters, especially the male genitalia, this species clearly belongs to the genus 
<taxonomicName id="4C6733B512722942F6D9F91BFDE2F8DF" authorityName="Enderlein" authorityYear="1909" box="[493,631,1789,1813]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412722942F6D9F91BFDE2F8DF" box="[493,631,1789,1813]" italics="true" pageId="4" pageNumber="230">Prionoglaris</emphasis>
</taxonomicName>
. However, the asymmetry of the claws of each pretarsus is less pronounced in the new species than in the other species of this genus, especially due to the absence of the membranous vesicle on the anterior claw. This membranous extension is always well developed in the other species of 
<taxonomicName id="4C6733B512722942F6B3F825FD84F811" authorityName="Enderlein" authorityYear="1909" box="[391,529,1987,2011]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412722942F6B3F825FD84F811" box="[391,529,1987,2011]" italics="true" pageId="4" pageNumber="230">Prionoglaris</emphasis>
</taxonomicName>
. The presence of a membranous extension on the anterior claw has been considered as an autapomorphy of the subfamily 
<taxonomicName id="4C6733B512722942F474F98CFB5EF948" authority="(Lienhard, 2004)" baseAuthorityName="Lienhard" baseAuthorityYear="2004" box="[832,1227,1642,1666]" class="Insecta" family="Prionoglarididae" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="subFamily" subFamily="Prionoglaridinae">
Prionoglaridinae (
<bibRefCitation id="EFF635C712722942F339F98CFB51F948" author="Lienhard C." box="[1037,1220,1642,1666]" pageId="4" pageNumber="230" pagination="865 - 875" refId="ref6497" refString="Lienhard C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
)
</taxonomicName>
. No intra-generic variability concerning this character has previously been observed in this subfamily. The membranous extension is particularly well developed in the 
<collectingRegion id="49A386D412722942F271F937FA09F923" box="[1349,1436,1745,1769]" country="Morocco" name="Oriental" pageId="4" pageNumber="230">Oriental</collectingRegion>
genus 
<taxonomicName id="4C6733B512722942F4B8F915FB5FF8C1" authority="Lienhard, 2004" authorityName="Lienhard" authorityYear="2004" box="[908,1226,1779,1803]" class="Insecta" family="Prionoglarididae" genus="Siamoglaris" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412722942F4B8F915FB9BF8C1" box="[908,1038,1779,1803]" italics="true" pageId="4" pageNumber="230">Siamoglaris</emphasis>
<bibRefCitation id="EFF635C712722942F328F915FB5FF8C1" author="Lienhard C." box="[1052,1226,1779,1803]" pageId="4" pageNumber="230" pagination="865 - 875" refId="ref6497" refString="Lienhard C. 2004. Siamoglaris zebrina gen. n., sp. n., the first representative of Prionoglarididae from the Oriental Region (Insecta: Psocoptera). Revue suisse de Zoologie 111 (4): 865 - 875." type="journal article" year="2004">Lienhard, 2004</bibRefCitation>
</taxonomicName>
, and in the fossil genus 
<taxonomicName id="4C6733B512722942F4B2F8F3FABFF8E7" authority="Azar, Huang &amp; Nel" authorityName="Azar, Huang &amp; Nel" authorityYear="2017" box="[902,1322,1813,1838]" class="Insecta" family="Prionoglarididae" genus="Palaeosiamoglaris" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412722942F4B2F8F3FBDAF8E7" box="[902,1103,1813,1837]" italics="true" pageId="4" pageNumber="230">Palaeosiamoglaris</emphasis>
Azar, Huang &amp; Nel
</taxonomicName>
in 
<bibRefCitation id="EFF635C712722942F279F8F3FC36F89A" author="Azar D. &amp; Huang Diying &amp; El-Hajj L. &amp; Cai Chenyang &amp; Nel A. &amp; Maksoud S." pageId="4" pageNumber="230" pagination="146 - 156" refId="ref5968" refString="Azar D., Huang Diying, El-Hajj L., Cai Chenyang, Nel A., Maksoud S. 2017. New Prionoglarididae from Burmese amber (Psocodea: Trogiomorpha: Prionoglaridetae). Cretaceous Research 75: 146 - 156." type="journal article" year="2017">
Azar 
<emphasis id="B913942412722942F2BDF8F0FCC0F89A" italics="true" pageId="4" pageNumber="230">et al</emphasis>
., 2017
</bibRefCitation>
from Burmese amber, but weakly developed in the fourth genus of this subfamily, the troglobitic 
<taxonomicName id="4C6733B512722942F474F89AFBADF85E" authority="Lienhard" authorityName="Lienhard" authorityYear="2010" box="[832,1080,1916,1940]" class="Insecta" family="Prionoglarididae" genus="Speleopsocus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="230" phylum="Arthropoda" rank="genus">
<emphasis id="B913942412722942F474F89AFC45F85E" box="[832,976,1916,1940]" italics="true" pageId="4" pageNumber="230">Speleopsocus</emphasis>
Lienhard
</taxonomicName>
in 
<bibRefCitation id="EFF635C712722942F354F89AFAC7F85E" author="Lienhard C. &amp; Holusa O. &amp; Grafitti G." box="[1120,1362,1916,1940]" pageId="4" pageNumber="230" pagination="185 - 197" refId="ref6642" refString="Lienhard C., Holusa O., Grafitti G. 2010 a. Two new cavedwelling Prionoglarididae from Venezuela and Namibia (Psocodea: ' Psocoptera': Trogiomorpha). Revue suisse de Zoologie 117 (2): 185 - 197." type="journal article" year="2010">
Lienhard 
<emphasis id="B913942412722942F3FDF89BFB6FF85E" box="[1225,1274,1916,1940]" italics="true" pageId="4" pageNumber="230">et al</emphasis>
., 2010a
</bibRefCitation>
which is only known from a single Venezuelan cave. Therefore the absence of this structure in the new species has to be interpreted as an apomorphic reduction. See the discussion below.
</paragraph>
<paragraph id="8BD8483612732943F7A3FEE7FDB6FED3" blockId="5.[151,757,257,2078]" box="[151,547,257,281]" pageId="5" pageNumber="231">
<emphasis id="B913942412732943F7A3FEE7FDB6FED3" bold="true" box="[151,547,257,281]" pageId="5" pageNumber="231">Remarks on a Moroccan sample of</emphasis>
</paragraph>
<paragraph id="8BD8483612732943F7A3FEC7FEFDFEF2" blockId="5.[151,757,257,2078]" box="[151,360,289,312]" pageId="5" pageNumber="231">
<taxonomicName id="4C6733B512732943F7A3FEC7FEFDFEF2" authorityName="Enderlein" authorityYear="1909" box="[151,360,289,312]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F7A3FEC7FEFDFEF2" bold="true" box="[151,360,289,312]" italics="true" pageId="5" pageNumber="231">Prionoglaris stygia</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BD8483612732943F7A3FEA7FEBDFA51" blockId="5.[151,757,257,2078]" pageId="5" pageNumber="231">
<location id="8EB81EED12732943F7A3FEA7FF55FE93" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12732943F7A3FEA7FF55FE93" box="[151,192,321,345]" name="The" pageId="5" pageNumber="231">The</location>
only morphological character that distinguishes the closely related species 
<taxonomicName id="4C6733B512732943F6BAFE84FE7CFEB3" authorityName="Enderlein" authorityYear="1909" box="[398,489,354,377]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F6BAFE84FE7CFEB3" box="[398,489,354,377]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6733B512732943F52AFE84FD27FEB3" authorityName="Lienhard" authorityYear="1988" box="[542,690,353,377]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="dactyloides">
<emphasis id="B913942412732943F52AFE84FD27FEB3" box="[542,690,353,377]" italics="true" pageId="5" pageNumber="231">P. dactyloides</emphasis>
</taxonomicName>
is the shape of the dorso-lateral appendages of the phallosome. Specimens from a Moroccan sample assigned to 
<taxonomicName id="4C6733B512732943F5A1FE44FD7BFE73" authorityName="Enderlein" authorityYear="1909" box="[661,750,418,441]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F5A1FE44FD7BFE73" box="[661,750,418,441]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
, collected in the 
<location id="8EB81EED12732943F677FE27FE4EFE13" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12732943F677FE27FE4EFE13" box="[323,475,449,473]" name="Friouato Cave" pageId="5" pageNumber="231">Friouato Cave</location>
(see 
<collectorName id="26922DE012732943F520FE27FD0DFE13" box="[532,664,449,473]" pageId="5" pageNumber="231">Distribution</collectorName>
, below), slightly differ from 
<collectorName id="26922DE012732943F647FE04FE4CFE33" box="[371,473,482,505]" pageId="5" pageNumber="231">European</collectorName>
<taxonomicName id="4C6733B512732943F6D6FE04FDD5FE33" authorityName="Enderlein" authorityYear="1909" box="[482,576,482,505]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F6D6FE04FDD5FE33" box="[482,576,482,505]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
by the shape of these appendages, and by the presence of a preapical denticle on the posterior claw of each pretarsus (
<figureCitation id="135C54B312732943F5A1FDC7FD7DFDF3" box="[661,744,545,569]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3C</figureCitation>
). 
<location id="8EB81EED12732943F7A3FDA7FF55FD93" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12732943F7A3FDA7FF55FD93" box="[151,192,577,601]" name="The" pageId="5" pageNumber="231">The</location>
dorso-lateral appendages of the phallosome are apically slightly narrowing in the 
<collectorName id="26922DE012732943F53DFD84FDE7FDB3" box="[521,626,610,633]" pageId="5" pageNumber="231">Morrocan</collectorName>
males (
<figureCitation id="135C54B312732943F5FDFD87FF37FD50" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3</figureCitation>
E-F and 
<bibRefCitation id="EFF635C712732943F634FD64FE33FD50" author="Lienhard C." box="[256,422,642,666]" pageId="5" pageNumber="231" pagination="87 - 151" refId="ref6425" refString="Lienhard C. 1996. Psocopteres nouveaux ou peu connus de quelques iles atlantiques (Canaries, Madere, Acores, Ascension) et de l'Afrique du Nord (Insecta: Psocoptera). Boletim do Museu Municipal do Funchal (Historia Natural) 48 (267): 87 - 151." type="journal article" year="1996">Lienhard, 1996</bibRefCitation>
: figs 4-6, 1998: fig. 41b), while they are broadly rounded in 
<taxonomicName id="4C6733B512732943F6EBFD45FDABFD70" authorityName="Enderlein" authorityYear="1909" box="[479,574,675,698]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F6EBFD45FDABFD70" box="[479,574,675,698]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
males from the type locality of this species (
<figureCitation id="135C54B312732943F6E1FD24FDB4FD10" box="[469,545,706,730]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3I</figureCitation>
) and in most other European males. The variability of the phallosome in several European populations of 
<taxonomicName id="4C6733B512732943F6CDFCE5FDC6FCD0" authorityName="Enderlein" authorityYear="1909" box="[505,595,771,794]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F6CDFCE5FDC6FCD0" box="[505,595,771,794]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
and in the two known Greek populations of 
<taxonomicName id="4C6733B512732943F6DEFCC5FD11FCF0" authorityName="Lienhard" authorityYear="1988" box="[490,644,802,826]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="dactyloides">
<emphasis id="B913942412732943F6DEFCC5FD11FCF0" box="[490,644,802,826]" italics="true" pageId="5" pageNumber="231">P. dactyloides</emphasis>
</taxonomicName>
from the 
<location id="8EB81EED12732943F7A3FCA4FE8BFC90" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12732943F7A3FCA4FE8BFC90" box="[151,286,834,858]" country="Switzerland" county="These" municipality="Eastern Crete" name="Peloponnese" pageId="5" pageNumber="231">Peloponnese</location>
(male 
<typeStatus id="54DCF69412732943F641FCA4FE41FC90" box="[373,468,834,858]" pageId="5" pageNumber="231" type="holotype">holotype</typeStatus>
) and from 
<collectingMunicipality id="6BBCD24C12732943F569FCA5FD61FC90" box="[605,756,834,858]" pageId="5" pageNumber="231">Eastern Crete</collectingMunicipality>
(male 
<typeStatus id="54DCF69412732943F7E9FC85FEAEFCB1" box="[221,315,867,891]" pageId="5" pageNumber="231" type="paratype">paratype</typeStatus>
) was illustrated by 
<bibRefCitation id="EFF635C712732943F521FC84FD54FCB1" author="Lienhard, C." box="[533,705,866,891]" pageId="5" pageNumber="231" pagination="89 - 108" refId="ref6387" refString="Lienhard, C. 1988. Vorarbeiten zu einer Psocopteren - Fauna der Westpalaarktis. IV. Die Gattung Prionoglaris Enderlein (Psocoptera: Prionoglarididae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 61: 89 - 108." type="journal article" year="1988">Lienhard (1988</bibRefCitation>
: figs 17-36). 
<collectingCounty id="62B930BA12732943F7DBFC64FEBAFC50" box="[239,303,898,922]" pageId="5" pageNumber="231">These</collectingCounty>
figures also show the high variability of the number of fine lateral hairs of the medio-internal appendage of the phallosome (1-13 hairs on each side, but often not the same number on both sides of the appendage). 
<collectorName id="26922DE012732943F614FBE5FEC1FBD0" box="[288,340,1027,1050]" pageId="5" pageNumber="231">Here</collectorName>
figures of the dorso-lateral and mediointernal appendages of two additional 
<collectorName id="26922DE012732943F512FBC5FD19FBF0" box="[550,652,1059,1082]" pageId="5" pageNumber="231">European</collectorName>
males are presented, one from 
<collectingCountry id="F37008A612732943F659FBA5FE79FB91" box="[365,492,1091,1115]" name="Switzerland" pageId="5" pageNumber="231">Switzerland</collectingCountry>
(
<figureCitation id="135C54B312732943F6CEFBA5FDD8FB91" box="[506,589,1091,1115]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3G</figureCitation>
), the other from 
<collectingCountry id="F37008A612732943F7A3FB85FF41FBB1" box="[151,212,1123,1147]" name="Spain" pageId="5" pageNumber="231">Spain</collectingCountry>
(
<figureCitation id="135C54B312732943F7D0FB85FEAEFBB1" box="[228,315,1123,1147]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3H</figureCitation>
). 
<collectorName id="26922DE012732943F664FB85FE11FBB1" box="[336,388,1123,1147]" pageId="5" pageNumber="231">Both</collectorName>
males are somewhat intermediate between 
<taxonomicName id="4C6733B512732943F7CDFB62FEC0FB51" authorityName="Enderlein" authorityYear="1909" box="[249,341,1156,1179]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F7CDFB62FEC0FB51" box="[249,341,1156,1179]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
males from the type locality (
<figureCitation id="135C54B312732943F595FB65FD79FB51" box="[673,748,1155,1179]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3I</figureCitation>
) and the 
<location id="8EB81EED12732943F7C9FB42FEFCFB71" LSID="urn:lsid:plazi:treatment:03CEF92012742941F628FC25FD29F8A5:8EB81EED12732943F7C9FB42FEFCFB71" box="[253,361,1188,1211]" country="Switzerland" municipality="However" name="Moroccan" pageId="5" pageNumber="231">Moroccan</location>
males (
<figureCitation id="135C54B312732943F6FAFB45FD81FB71" box="[462,532,1187,1211]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3</figureCitation>
E-F). 
<collectingMunicipality id="6BBCD24C12732943F569FB42FD55FB71" box="[605,704,1188,1211]" pageId="5" pageNumber="231">However</collectingMunicipality>
, the preapical denticle on the posterior claw, always present in the 
<collectorName id="26922DE012732943F7EBFB02FEDEFB31" box="[223,331,1252,1275]" pageId="5" pageNumber="231">Moroccan</collectorName>
specimens (though sometimes smaller than in 
<figureCitation id="135C54B312732943F7D2FAE5FEA2FAD1" box="[230,311,1283,1307]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3C</figureCitation>
), is completely absent in these 
<specimenCount id="9D6183BF12732943F5B5FAE2FD65FAD1" box="[641,752,1283,1307]" pageId="5" pageNumber="231" type="male">two males</specimenCount>
, as it is generally the case in the 
<collectorName id="26922DE012732943F6CEFAC2FDF5FAF1" box="[506,608,1316,1339]" pageId="5" pageNumber="231">European</collectorName>
<taxonomicName id="4C6733B512732943F55DFAC2FD50FAF1" authorityName="Enderlein" authorityYear="1909" box="[617,709,1316,1339]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F55DFAC2FD50FAF1" box="[617,709,1316,1339]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
and in 
<taxonomicName id="4C6733B512732943F78DFAA2FEC6FA91" authorityName="Lienhard" authorityYear="1988" box="[185,339,1347,1371]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="dactyloides">
<emphasis id="B913942412732943F78DFAA2FEC6FA91" box="[185,339,1347,1371]" italics="true" pageId="5" pageNumber="231">P. dactyloides</emphasis>
</taxonomicName>
(only in one mesothoracic leg of a 
<typeStatus id="54DCF69412732943F7A3FA82FF66FAB6" box="[151,243,1380,1404]" pageId="5" pageNumber="231" type="paratype">paratype</typeStatus>
of the latter could a minute preapical denticle be observed).
</paragraph>
<paragraph id="8BD8483612732943F7A3FA45FE4CF8D6" blockId="5.[151,757,257,2078]" pageId="5" pageNumber="231">
When looking for other potentially diagnostic characters which could allow the Moroccan specimens to be distinguished from the European 
<taxonomicName id="4C6733B512732943F537FA03FDF5FA36" authorityName="Enderlein" authorityYear="1909" box="[515,608,1509,1532]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F537FA03FDF5FA36" box="[515,608,1509,1532]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
, I discovered some differences in the ratio between forewing length and width (FW/FWw) and in the micromorphology of the spermapore sclerite. In both sexes the Moroccan specimens have particularly broad and slightly shortened wings (
<figureCitation id="135C54B312732943F7D2F962FEA9F956" box="[230,316,1668,1692]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3D</figureCitation>
), with FW/FWw varying from 2.0 to 2.1 (N = 2). In the European 
<taxonomicName id="4C6733B512732943F699F943FD9DF976" authorityName="Enderlein" authorityYear="1909" box="[429,520,1701,1724]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F699F943FD9DF976" box="[429,520,1701,1724]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
forewings are clearly longer than twice their width, FW/FWw varying from 2.2 to 2.5 (N = 6; the mean is 2.4). In 
<taxonomicName id="4C6733B512732943F6C9F903FD05F936" authorityName="Lienhard" authorityYear="1988" box="[509,656,1764,1788]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="dactyloides">
<emphasis id="B913942412732943F6C9F903FD05F936" box="[509,656,1764,1788]" italics="true" pageId="5" pageNumber="231">P. dactyloides</emphasis>
</taxonomicName>
this ratio varies from 2.1 to 2.3 (N = 2).
</paragraph>
<paragraph id="8BD8483612732943F7A3F8C3FBC2FE90" blockId="5.[151,757,257,2078]" lastBlockId="5.[832,1437,257,638]" pageId="5" pageNumber="231">
In the only Moroccan female available for this study the spermapore sclerite is proximally broadly rounded and distally slightly elongated and beak-shaped (
<figureCitation id="135C54B312732943F5B9F883FD7DF8B7" box="[653,744,1893,1917]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3K</figureCitation>
). In the European 
<taxonomicName id="4C6733B512732943F67FF860FE33F857" authorityName="Enderlein" authorityYear="1909" box="[331,422,1926,1949]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F67FF860FE33F857" box="[331,422,1926,1949]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
this sclerite is similar in shape to that of 
<taxonomicName id="4C6733B512732943F638F840FE04F877" authority="Lienhard, 2021" authorityName="Lienhard" authorityYear="2021" box="[268,401,1957,1981]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="kapralovi" status="sp. nov.">
<emphasis id="B913942412732943F638F840FE04F877" box="[268,401,1957,1981]" italics="true" pageId="5" pageNumber="231">P. kapralovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220295F12732943F6A8F840FE63F877" box="[412,502,1958,1981]" pageId="5" pageNumber="231" rank="species">sp. nov.</taxonomicNameLabel>
, i.e. narrowly rounded distally but not elongated and beak-shaped (see figures in 
<bibRefCitation id="EFF635C712732943F7A3F803FEA8F837" author="Lienhard, C." box="[151,317,2021,2045]" pageId="5" pageNumber="231" pagination="89 - 108" refId="ref6387" refString="Lienhard, C. 1988. Vorarbeiten zu einer Psocopteren - Fauna der Westpalaarktis. IV. Die Gattung Prionoglaris Enderlein (Psocoptera: Prionoglarididae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 61: 89 - 108." type="journal article" year="1988">Lienhard, 1988</bibRefCitation>
, 
<bibRefCitation id="EFF635C712732943F673F803FE15F837" author="Lienhard C." box="[327,384,2021,2045]" pageId="5" pageNumber="231" pagination="1 - 517" refId="ref6478" refString="Lienhard C. 1998. Psocopteres euro-mediterraneens. Faune de France 83: I-XX, 1 - 517." type="book chapter" year="1998">1998</bibRefCitation>
). However, in a female of 
<taxonomicName id="4C6733B512732943F5AEF800FD61F837" authorityName="Enderlein" authorityYear="1909" box="[666,756,2022,2045]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F5AEF800FD61F837" box="[666,756,2022,2045]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
from the 
<typeStatus id="54DCF69412732943F7C1F7E0FEB6F7D4" box="[245,291,2054,2078]" pageId="5" pageNumber="231">type</typeStatus>
locality (French Pyrenees) a distally weakly elongated spermapore sclerite was observed (
<figureCitation id="135C54B312732943F207FEE7FA1AFED3" box="[1331,1423,257,281]" captionStart="Fig" captionStartId="6.[151,186,1412,1434]" captionTargetBox="[162,1422,261,1370]" captionTargetId="figure-209@6.[151,1436,257,1376]" captionTargetPageId="6" captionText="Fig. 3. Prionoglaris spp.: P. stygia males from Friouato Cave, Morocco (A-F), P. stygia male from Pierre à Perret Cave, Switzerland (G), P.stygia male fromW of Atzeneta del Maestrat, Spain (H), P.stygia male from the type locality, Compagnaga Cave, French Pyrenees (I), P. dactyloides, male holotype (J), P. stygia female from Friouato Cave, Morocco (K-L), P. stygia female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="5" pageNumber="231">Fig. 3M</figureCitation>
), about intermediate in shape between the other 
<taxonomicName id="4C6733B512732943F274FEC5FA08FEF0" authorityName="Enderlein" authorityYear="1909" box="[1344,1437,291,314]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F274FEC5FA08FEF0" box="[1344,1437,291,314]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
and the Moroccan female.
</paragraph>
<paragraph id="8BD8483612732943F474FE85FB76FDB4" blockId="5.[832,1437,257,638]" pageId="5" pageNumber="231">
In view of the considerable variability of the above mentioned characters, and of the low numbers of specimens examined for this study, the slight differences observed between the Moroccan population and the European 
<taxonomicName id="4C6733B512732943F499FE03FB9DFE36" authorityName="Enderlein" authorityYear="1909" box="[941,1032,485,508]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="231" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412732943F499FE03FB9DFE36" box="[941,1032,485,508]" italics="true" pageId="5" pageNumber="231">P. stygia</emphasis>
</taxonomicName>
are not sufficient to warrant a formal separation of the Moroccan form as a new species. The only character which can be considered as diagnostic for the Moroccan form is the presence of a preapical denticle on the posterior claw of each pretarsus.
</paragraph>
</subSubSection>
<caption id="DF1818BE12702940F7A3FA62FD56F98C" ID-DOI="http://doi.org/10.5281/zenodo.5639921" ID-Zenodo-Dep="5639921" httpUri="https://zenodo.org/record/5639921/files/figure.png" pageId="6" pageNumber="232" startId="6.[151,186,1412,1434]" targetBox="[162,1422,261,1370]" targetPageId="6">
<paragraph id="8BD8483612702940F7A3FA62FD56F98C" blockId="6.[151,1437,1412,1606]" pageId="6" pageNumber="232">
Fig. 3. 
<taxonomicName id="4C6733B512702940F7D7FA62FE1AFA53" box="[227,399,1412,1434]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B913942412702940F7D7FA62FECBFA50" box="[227,350,1412,1434]" italics="true" pageId="6" pageNumber="232">Prionoglaris</emphasis>
spp.
</taxonomicName>
: 
<taxonomicName id="4C6733B512702940F6ACFA63FE7CFA50" authorityName="Enderlein" authorityYear="1909" box="[408,489,1413,1434]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F6ACFA63FE7CFA50" box="[408,489,1413,1434]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
males from Friouato Cave, Morocco (A-F), 
<taxonomicName id="4C6733B512702940F4ADFA63FC7FFA50" authorityName="Enderlein" authorityYear="1909" box="[921,1002,1413,1434]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F4ADFA63FC7FFA50" box="[921,1002,1413,1434]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
male from Pierre à Perret Cave, Switzerland (G), 
<taxonomicName id="4C6733B512702940F63BFA47FECBFA7C" authorityName="Enderlein" authorityYear="1909" box="[271,350,1441,1462]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F63BFA47FECBFA7C" box="[271,350,1441,1462]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
male from W of Atzeneta del Maestrat, Spain (H), 
<taxonomicName id="4C6733B512702940F40AFA47FC18FA7C" authorityName="Enderlein" authorityYear="1909" box="[830,909,1441,1462]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F40AFA47FC18FA7C" box="[830,909,1441,1462]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
male from the type locality, Compagnaga Cave, French Pyrenees (I), 
<taxonomicName id="4C6733B512702940F657FA58FE72FA19" authorityName="Lienhard" authorityYear="1988" box="[355,487,1469,1491]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="dactyloides">
<emphasis id="B913942412702940F657FA58FE72FA19" box="[355,487,1469,1491]" italics="true" pageId="6" pageNumber="232">P. dactyloides</emphasis>
</taxonomicName>
, male holotype (J), 
<taxonomicName id="4C6733B512702940F593FA58FD62FA19" authorityName="Enderlein" authorityYear="1909" box="[679,759,1470,1491]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F593FA58FD62FA19" box="[679,759,1470,1491]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
female from Friouato Cave, Morocco (K-L), 
<taxonomicName id="4C6733B512702940F399FA58FB68FA19" authorityName="Enderlein" authorityYear="1909" box="[1197,1277,1470,1491]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="232" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="B913942412702940F399FA58FB68FA19" box="[1197,1277,1470,1491]" italics="true" pageId="6" pageNumber="232">P. stygia</emphasis>
</taxonomicName>
female from the type locality, Compagnaga Cave, French Pyrenees (M). – (A) Phallosome, ventral view. (B) Anterior claw of hind pretarsus. (C) Posterior claw of hind pretarsus. (D) Left forewing. (E-J) Dorso-lateral appendages and medio-internal appendage of phallosome, not in situ (E same male as in A). (K-M) Spermapore sclerite and distal part of spermathecal duct (K and M ventral view; L optical longitudinal section, lateral view).
</paragraph>
</caption>
<subSubSection id="C37D1BBD12712941F628F8CEFD29F8A5" pageId="7" pageNumber="233" type="distribution">
<paragraph id="8BD8483612712941F628F8CEFDE5F88A" blockId="7.[284,624,1832,1856]" box="[284,624,1832,1856]" pageId="7" pageNumber="233">
<heading id="D090FF5A12712941F628F8CEFDE5F88A" bold="true" box="[284,624,1832,1856]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="233" reason="2">
<emphasis id="B913942412712941F628F8CEFDE5F88A" bold="true" box="[284,624,1832,1856]" pageId="7" pageNumber="233">
<taxonomicName id="4C6733B512712941F628F8CEFD81F8F5" authority="Lienhard, 2021" authorityName="Lienhard" authorityYear="2021" box="[284,532,1832,1855]" class="Insecta" family="Prionoglarididae" genus="Prionoglaris" kingdom="Animalia" order="Psocodea" pageId="7" pageNumber="233" phylum="Arthropoda" rank="species" species="kapralovi" status="sp. nov.">
<emphasis id="B913942412712941F628F8CEFD81F8F5" bold="true" box="[284,532,1832,1855]" italics="true" pageId="7" pageNumber="233">Prionoglaris kapralovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220295F12712941F52EF8CEFDE5F88A" box="[538,624,1832,1856]" pageId="7" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD8483612712941F7A3F8B1FD29F8A5" blockId="7.[151,700,1879,1903]" box="[151,700,1879,1903]" pageId="7" pageNumber="233">
<emphasis id="B913942412712941F7A3F8B1FECAF8A5" bold="true" box="[151,351,1879,1903]" pageId="7" pageNumber="233">Previous records:</emphasis>
<collectingCountry id="F37008A612712941F650F8B1FE51F8A5" box="[356,452,1879,1903]" name="Armenia" pageId="7" pageNumber="233">Armenia</collectingCountry>
, see description above.
</paragraph>
</subSubSection>
</treatment>
</document>