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<document id="E1F2CC3BE35C5555AC2E6422A10D9A05" ID-DOI="10.37828/em.2020.38.7" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232632" ID-ZooBank="urn:lsid:zoobank.org:pub:4B85DA98-957E-4788-BC8B-25526047DC8B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722914106896" checkinUser="felipe" docAuthor="Saldaitis, Aidas &amp; Volynkin, Anton V." docDate="2020" docId="03CBF90BFF8B4B67C7B09B2A122F5E52" docLanguage="en" docName="EcolMontenegrina.38.51-62.pdf" docOrigin="Ecologica Montenegrina 38" docSource="http://dx.doi.org/10.37828/em.2020.38.7" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Chelis kashmirica" docType="treatment" docVersion="3" lastPageNumber="61" masterDocId="FFF28173FF8E4B6DC7279A2D16565C62" masterDocTitle="On the taxonomy of the Chelis glaphyra (Eversmann, 1843) species-group, with description of a new subspecies of Chelis gratiosa (Grum-Grshimailo, 1890) from Kyrgyzstan, brachipterous females of Chelis kashmirica (Ferguson, 1985) and Chelis golbecki (Dubatolov, 1996), and the checklist of the species-group (Lepidoptera: Erebidae: Arctiinae: Arctiini)" masterLastPageNumber="62" masterPageNumber="51" pageNumber="56" updateTime="1722917624881" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1884D64AD38C680BAB805D4C0D1E4421">On the taxonomy of the Chelis glaphyra (Eversmann, 1843) species-group, with description of a new subspecies of Chelis gratiosa (Grum-Grshimailo, 1890) from Kyrgyzstan, brachipterous females of Chelis kashmirica (Ferguson, 1985) and Chelis golbecki (Dubatolov, 1996), and the checklist of the species-group (Lepidoptera: Erebidae: Arctiinae: Arctiini)</mods:title>
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<mods:namePart id="97538A9217DCF360F0C59247BE5B8C5D">Saldaitis, Aidas</mods:namePart>
<mods:affiliation id="96563624C2A9C208F57A8E41CA0092B2">Nature Research Centre, Akademijos str., 2, LT- 08412, Vilnius- 21, Lithuania. E-mail: saldrasa @ gmail. com &amp; Corresponding author</mods:affiliation>
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<mods:namePart id="29D7FC7F4C48AA6D898F3150B362A5B7">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="A791B7BC96844099A2CA6E0AB105D151">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia. E-mails: monstruncusarctia @ gmail. com; volynkin _ a @ mail. ru &amp; National Research Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia</mods:affiliation>
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<treatment id="03CBF90BFF8B4B67C7B09B2A122F5E52" ID-DOI="http://doi.org/10.5281/zenodo.13232908" ID-Zenodo-Dep="13232908" LSID="urn:lsid:plazi:treatment:03CBF90BFF8B4B67C7B09B2A122F5E52" httpUri="http://treatment.plazi.org/id/03CBF90BFF8B4B67C7B09B2A122F5E52" lastPageId="10" lastPageNumber="61" pageId="5" pageNumber="56">
<subSubSection id="C3781B96FF8B4B68C7B09B2A15605D43" box="[151,822,263,290]" pageId="5" pageNumber="56" type="nomenclature">
<paragraph id="8BDD481DFF8B4B68C7B09B2A15605D43" blockId="5.[151,822,263,324]" box="[151,822,263,290]" pageId="5" pageNumber="56">
<heading id="D095FF71FF8B4B68C7B09B2A15605D43" box="[151,822,263,290]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="56" reason="2">
<taxonomicName id="4C62339EFF8B4B68C7B09B2A14115D40" authority="(Ferguson, 1985)" baseAuthorityName="Ferguson" baseAuthorityYear="1985" box="[151,583,263,290]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF8B4B68C7B09B2A17245D43" bold="true" box="[151,370,263,289]" italics="true" pageId="5" pageNumber="56">Chelis kashmirica</emphasis>
(
<bibRefCitation id="EFF335ECFF8B4B68C6A59B2A14685D40" author="Ferguson, D. C." box="[386,574,263,290]" pageId="5" pageNumber="56" pagination="181 - 275" refId="ref4007" refString="Ferguson, D. C. (1985) Contributions toward reclassification of the world genera of the tribe Arctiini, Part 1 - Introductrion and a revision of the Neoarctia-Grammia group (Lepidoptera: Arctiidae; Arctiinae). The Entomography. An Annual Review for Biosystematics, 3, 181 - 275." type="journal article" year="1985">Ferguson, 1985</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B916940FFF8B4B68C5749B2A15605D43" bold="true" box="[595,822,263,289]" pageId="5" pageNumber="56">stat. &amp; comb. nov.</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF8B4B68C7B09B0417D05D26" box="[151,390,297,324]" pageId="5" pageNumber="56" type="description">
<paragraph id="8BDD481DFF8B4B68C7B09B0417D05D26" blockId="5.[151,822,263,324]" box="[151,390,297,324]" pageId="5" pageNumber="56">
(
<figureCitation id="13595498FF8B4B68C7B89B0417715D26" box="[159,295,297,324]" captionStart="Figures 1016" captionStartId="5.[151,236,1989,2013]" captionTargetBox="[160,1422,480,1948]" captionTargetId="figure-51@5.[151,1434,480,1971]" captionTargetPageId="5" captionText="Figures 1016. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232640" httpUri="https://zenodo.org/record/13232640/files/figure.png" pageId="5" pageNumber="56">Figs 1012</figureCitation>
,
<figureCitation id="13595498FF8B4B68C6149B0417055D26" box="[307,339,297,324]" captionStart="Figures 2123" captionStartId="7.[151,236,1994,2018]" captionTargetBox="[237,1328,196,1963]" captionTargetId="figure-13@7.[151,1406,189,1975]" captionTargetPageId="7" captionText="Figures 2123. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232646" httpUri="https://zenodo.org/record/13232646/files/figure.png" pageId="5" pageNumber="56">21</figureCitation>
,
<figureCitation id="13595498FF8B4B68C6789B0417285D26" box="[351,382,297,324]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="5" pageNumber="56">26</figureCitation>
)
</paragraph>
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<subSubSection id="C3781B96FF8B4B68C7B09B4617DE5DC0" pageId="5" pageNumber="56" type="reference_group">
<paragraph id="8BDD481DFF8B4B68C7B09B4617DE5DC0" blockId="5.[151,1436,363,418]" pageId="5" pageNumber="56">
<taxonomicName id="4C62339EFF8B4B68C7B09B4614105DE1" ID-CoL="4C42W" authority="Ferguson, 1985" authorityName="Ferguson" authorityYear="1985" box="[151,582,363,388]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF8B4B68C7B09B4617DC5DE1" box="[151,394,363,387]" italics="true" pageId="5" pageNumber="56">Palearctia kashmirica</emphasis>
<materialsCitation id="3B0A4240FF8B4B68C6BF9B4114105DE1" box="[408,582,363,388]" country="Pakistan" location="Azad Kashmir" municipality="Baltistan" pageId="5" pageNumber="56" specimenCount="1">
<bibRefCitation id="EFF335ECFF8B4B68C6BF9B4114105DE1" author="Ferguson, D. C." box="[408,582,363,388]" pageId="5" pageNumber="56" pagination="181 - 275" refId="ref4007" refString="Ferguson, D. C. (1985) Contributions toward reclassification of the world genera of the tribe Arctiini, Part 1 - Introductrion and a revision of the Neoarctia-Grammia group (Lepidoptera: Arctiidae; Arctiinae). The Entomography. An Annual Review for Biosystematics, 3, 181 - 275." type="journal article" year="1985">Ferguson, 1985</bibRefCitation>
</materialsCitation>
</taxonomicName>
,
<emphasis id="B916940FFF8B4B68C57F9B4614A15DE1" box="[600,759,363,387]" italics="true" pageId="5" pageNumber="56">Entomography</emphasis>
,
<emphasis id="B916940FFF8B4B68C42D9B4115415DE6" bold="true" box="[778,791,364,388]" pageId="5" pageNumber="56">3</emphasis>
: 222, figs 44, 101, 102 (
<typeStatus id="54D9F6BFFF8B4B68C3649B46122A5DE1" box="[1091,1148,363,387]" pageId="5" pageNumber="56">Type</typeStatus>
locality: [
<collectingCountry id="F375088DFF8B4B68C3D49B4613045DE1" box="[1267,1362,363,387]" name="Pakistan" pageId="5" pageNumber="56">Pakistan</collectingCountry>
,
<location id="8EBD1EC6FF8B4B68C2439B4616A35DC0" LSID="urn:lsid:plazi:treatment:03CBF90BFF8B4B67C7B09B2A122F5E52:8EBD1EC6FF8B4B68C2439B4616A35DC0" country="Pakistan" municipality="Baltistan" name="Azad Kashmir" pageId="5" pageNumber="56">Azad Kashmir</location>
] “
<collectingMunicipality id="6BB9D267FF8B4B68C62A9BA717255DC0" box="[269,371,394,418]" pageId="5" pageNumber="56">Baltistan</collectingMunicipality>
”).
</paragraph>
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<caption id="DF1D1895FF8B4B68C7B09DE815AE5BBF" ID-DOI="http://doi.org/10.5281/zenodo.13232640" ID-Zenodo-Dep="13232640" box="[151,1016,1989,2013]" httpUri="https://zenodo.org/record/13232640/files/figure.png" pageId="5" pageNumber="56" startId="5.[151,236,1989,2013]" targetBox="[160,1422,480,1948]" targetPageId="5" targetType="figure">
<paragraph id="8BDD481DFF8B4B68C7B09DE815AE5BBF" blockId="5.[151,1016,1989,2013]" box="[151,1016,1989,2013]" pageId="5" pageNumber="56">
<emphasis id="B916940FFF8B4B68C7B09DE8176B5BBF" bold="true" box="[151,317,1989,2013]" pageId="5" pageNumber="56">Figures 1016.</emphasis>
<taxonomicName id="4C62339EFF8B4B68C6639DE817E85BBF" box="[324,446,1989,2013]" class="Insecta" family="Erebidae" genus="Chelis" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916940FFF8B4B68C6639DE817DE5BBF" box="[324,392,1989,2013]" italics="true" pageId="5" pageNumber="56">Chelis</emphasis>
spp.
</taxonomicName>
: adults. The specimens are deposited in MWM/ZSM.
</paragraph>
</caption>
<caption id="DF1D1895FF884B6BC7B09DFF13DE5B8B" ID-DOI="http://doi.org/10.5281/zenodo.13232642" ID-Zenodo-Dep="13232642" box="[151,1416,2001,2026]" httpUri="https://zenodo.org/record/13232642/files/figure.png" pageId="6" pageNumber="57" startId="6.[151,236,2002,2026]" targetBox="[163,1426,222,1936]" targetPageId="6" targetType="figure">
<paragraph id="8BDD481DFF884B6BC7B09DFF13DE5B8B" blockId="6.[151,1416,2001,2026]" box="[151,1416,2001,2026]" pageId="6" pageNumber="57">
<emphasis id="B916940FFF884B6BC7B09DFF176B5B88" bold="true" box="[151,317,2002,2026]" pageId="6" pageNumber="57">Figures 1720.</emphasis>
<taxonomicName id="4C62339EFF884B6BC6639DFC17BE5B8B" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[324,488,2001,2025]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF884B6BC6639DFC17BE5B8B" box="[324,488,2001,2025]" italics="true" pageId="6" pageNumber="57">Chelis gratiosa</emphasis>
</taxonomicName>
sspp.: adults. Depositories of the specimens: 17 in ASV/WIGJ; 1820 in MWM/ZSM.
</paragraph>
</caption>
<caption id="DF1D1895FF894B6AC7B09DE715AE5B83" ID-DOI="http://doi.org/10.5281/zenodo.13232646" ID-Zenodo-Dep="13232646" box="[151,1016,1993,2018]" httpUri="https://zenodo.org/record/13232646/files/figure.png" pageId="7" pageNumber="58" startId="7.[151,236,1994,2018]" targetBox="[237,1328,196,1963]" targetPageId="7" targetType="figure">
<paragraph id="8BDD481DFF894B6AC7B09DE715AE5B83" blockId="7.[151,1016,1993,2018]" box="[151,1016,1993,2018]" pageId="7" pageNumber="58">
<emphasis id="B916940FFF894B6AC7B09DE7176B5B80" bold="true" box="[151,317,1994,2018]" pageId="7" pageNumber="58">Figures 2123.</emphasis>
<taxonomicName id="4C62339EFF894B6AC6639DE417E85B83" box="[324,446,1993,2017]" class="Insecta" family="Erebidae" genus="Chelis" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916940FFF894B6AC6639DE417DE5B83" box="[324,392,1993,2017]" italics="true" pageId="7" pageNumber="58">Chelis</emphasis>
spp.
</taxonomicName>
: adults. The specimens are deposited in MWM/ZSM.
</paragraph>
</caption>
<caption id="DF1D1895FF864B65C7B09D5812815BEE" ID-DOI="http://doi.org/10.5281/zenodo.13232648" ID-Zenodo-Dep="13232648" box="[151,1239,1908,1933]" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="8" pageNumber="59" startId="8.[151,236,1909,1933]" targetBox="[155,1436,204,1841]" targetPageId="8" targetType="figure">
<paragraph id="8BDD481DFF864B65C7B09D5812815BEE" blockId="8.[151,1239,1908,1933]" box="[151,1239,1908,1933]" pageId="8" pageNumber="59">
<emphasis id="B916940FFF864B65C7B09D58176B5BEF" bold="true" box="[151,317,1909,1933]" pageId="8" pageNumber="59">Figures 2428.</emphasis>
<taxonomicName id="4C62339EFF864B65C6639D5917E85BEE" box="[324,446,1908,1932]" class="Insecta" family="Erebidae" genus="Chelis" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916940FFF864B65C6639D5917DE5BEE" box="[324,392,1908,1932]" italics="true" pageId="8" pageNumber="59">Chelis</emphasis>
spp.
</taxonomicName>
: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM.
</paragraph>
</caption>
<subSubSection id="C3781B96FF874B64C7B09AE9151F5D40" pageId="9" pageNumber="60" type="materials_examined">
<paragraph id="8BDD481DFF874B64C7B09AE9151F5D40" blockId="9.[151,1437,195,290]" pageId="9" pageNumber="60">
<emphasis id="B916940FFF874B64C7B09AE917D05CBC" bold="true" box="[151,390,196,222]" pageId="9" pageNumber="60">Material examined</emphasis>
.
<materialsCitation id="3B0A4240FF874B64C6B19AEE15B95D62" collectingDate="2000-07-05" collectionCode="ZSM" collectorName="Z. Varga &amp; G. Ronkay &amp; Nos." country="Pakistan" county="Kashmir Deosai" elevation="4525" location="Kashmir" municipality="Himalaya Mts." pageId="9" pageNumber="60" specimenCount="28" specimenCount-female="2" specimenCount-male="26">
<specimenCount id="9D648394FF874B64C6B19AEE14525CBC" box="[406,516,195,222]" count="26" pageId="9" pageNumber="60" type="male">26 males</specimenCount>
,
<specimenCount id="9D648394FF874B64C5359AEE14D15CBC" box="[530,647,195,222]" count="2" pageId="9" pageNumber="60" type="female">2 females</specimenCount>
,
<collectingCountry id="F375088DFF874B64C5B29AEE14AD5CBC" box="[661,763,195,222]" name="Pakistan" pageId="9" pageNumber="60">Pakistan</collectingCountry>
,
<location id="8EBD1EC6FF874B64C42F9AEE15265CBC" LSID="urn:lsid:plazi:treatment:03CBF90BFF8B4B67C7B09B2A122F5E52:8EBD1EC6FF874B64C42F9AEE15265CBC" box="[776,880,195,222]" country="Pakistan" county="Kashmir Deosai" municipality="Himalaya Mts." name="Kashmir" pageId="9" pageNumber="60">Kashmir</location>
,
<collectingMunicipality id="6BB9D267FF874B64C4599AEE12665CBC" box="[894,1072,195,222]" pageId="9" pageNumber="60">Himalaya Mts.</collectingMunicipality>
,
<collectingCounty id="62BC3091FF874B64C3199AEE12A85CBC" box="[1086,1278,195,222]" pageId="9" pageNumber="60">Kashmir Deosai</collectingCounty>
pass.,
<quantity id="4C9AE5F8FF874B64C2749AEE16A35D62" metricMagnitude="3" metricUnit="m" metricValue="4.525" metricValueMax="4.8" metricValueMin="4.25" pageId="9" pageNumber="60" unit="m" value="4525.0" valueMax="4800.0" valueMin="4250.0">
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,
<date id="FFDC6EDDFF874B64C6249AC817D05D62" box="[259,390,229,256]" pageId="9" pageNumber="60" value="2000-07-05">
<collectingDate id="EF989735FF874B64C6249AC817D05D62" box="[259,390,229,256]" pageId="9" pageNumber="60" value="2000-07-05">5.VII.2000</collectingDate>
</date>
, leg.
<collectorName id="26972DCBFF874B64C6EE9ACB14645D62" box="[457,562,230,256]" pageId="9" pageNumber="60">Z. Varga</collectorName>
&amp;
<collectorName id="26972DCBFF874B64C57B9AC814B75D62" box="[604,737,229,256]" pageId="9" pageNumber="60">G. Ronkay</collectorName>
, gen. slide
<collectorName id="26972DCBFF874B64C44D9ACB15F35D62" box="[874,933,230,256]" pageId="9" pageNumber="60">Nos.</collectorName>
:
<collectionCode id="ED73D0D8FF874B64C4949AC815B95D62" box="[947,1007,229,256]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="60" type="Museum">ZSM</collectionCode>
</materialsCitation>
Arct. 2019915 (male),
<collectionCode id="ED73D0D8FF874B64C23C9AC813015D62" box="[1307,1367,229,256]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="60" type="Museum">ZSM</collectionCode>
Arct. 2019916 (female) (prepared by Volynkin) (MWM/
<collectionCode id="ED73D0D8FF874B64C5D89B2A156B5D40" box="[767,829,263,290]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="60" type="Museum">ZSM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF874B64C7B09B6617665EBA" pageId="9" pageNumber="60" type="diagnosis">
<paragraph id="8BDD481DFF874B64C7B09B6617665EBA" blockId="9.[151,1437,330,728]" pageId="9" pageNumber="60">
<emphasis id="B916940FFF874B64C7B09B6617465D07" bold="true" box="[151,272,331,357]" pageId="9" pageNumber="60">Diagnosis</emphasis>
. Male of the species is smaller than those of subspecies of
<taxonomicName id="4C62339EFF874B64C3219B6712CB5D06" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[1030,1181,330,357]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C3219B6712CB5D06" box="[1030,1181,330,357]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
. In the male genital capsule of
<taxonomicName id="4C62339EFF874B64C63B9B4117845DE5" authorityName="Ferguson" authorityYear="1985" box="[284,466,364,391]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C63B9B4117845DE5" box="[284,466,364,391]" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
, the uncus is narrower medially, the vinculum is rather V-shaped (while that of
<taxonomicName id="4C62339EFF874B64C7B09BA3177C5DCA" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[151,298,398,425]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C7B09BA3177C5DCA" box="[151,298,398,425]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
is U-shaped), the juxta is conspicuously shorter and narrower, the valva is less elongate and the cucullus is noticeably smaller than those structures of
<taxonomicName id="4C62339EFF874B64C43F9B9D15FD5DA8" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[792,939,432,459]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C43F9B9D15FD5DA8" box="[792,939,432,459]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
. The aedeagus of
<taxonomicName id="4C62339EFF874B64C3A39B9D13615DA9" authorityName="Ferguson" authorityYear="1985" box="[1156,1335,432,459]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C3A39B9D13615DA9" box="[1156,1335,432,459]" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
is much shorter than in
<taxonomicName id="4C62339EFF874B64C6749BFC17BD5D89" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[339,491,465,492]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C6749BFC17BD5D89" box="[339,491,465,492]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
. In the aedeagus vesica of
<taxonomicName id="4C62339EFF874B64C4189BFC15A05D8E" authorityName="Ferguson" authorityYear="1985" box="[831,1014,465,492]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C4189BFC15A05D8E" box="[831,1014,465,492]" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
, the medial diverticulum is C-like curved (in
<taxonomicName id="4C62339EFF874B64C6069BDE17EC5E6F" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[289,442,499,526]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C6069BDE17EC5E6F" box="[289,442,499,526]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
, it is globular), and the distal section is conspicuously shorter than that of
<taxonomicName id="4C62339EFF874B64C2549BDE16AF5E4D" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C2549BDE16AF5E4D" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
. Female of
<taxonomicName id="4C62339EFF874B64C6B39838141D5E52" authorityName="Ferguson" authorityYear="1985" box="[404,587,533,560]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C6B39838141D5E52" box="[404,587,533,560]" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
is brachipterous and has very short and narrow wings with strongly elongate and pointed apex, while all the subspecies of
<taxonomicName id="4C62339EFF874B64C418981B15805E33" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[831,982,566,593]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C418981B15805E33" box="[831,982,566,593]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
have normal winged females. In the externally similar
<taxonomicName id="4C62339EFF874B64C65F987514455E11" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Dubatolov" baseAuthorityYear="1996" box="[376,531,600,627]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="golbecki">
<emphasis id="B916940FFF874B64C65F987514455E11" box="[376,531,600,627]" italics="true" pageId="9" pageNumber="60">Ch. golbecki</emphasis>
</taxonomicName>
, the both female wings are much longer and broader than those of
<taxonomicName id="4C62339EFF874B64C2549875174F5EF7" authorityName="Ferguson" authorityYear="1985" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C2549875174F5EF7" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
and the forewing apex is more rounded. The female genitalia of
<taxonomicName id="4C62339EFF874B64C33E9857129A5EF7" authorityName="Ferguson" authorityYear="1985" box="[1049,1228,634,661]" class="Insecta" family="Erebidae" genus="Palearctia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="kashmirica">
<emphasis id="B916940FFF874B64C33E9857129A5EF7" box="[1049,1228,634,661]" italics="true" pageId="9" pageNumber="60">Ch. kashmirica</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4C62339EFF874B64C79E98B117185ED4" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" baseAuthorityName="Saldaitis &amp; Volynkin" baseAuthorityYear="1890" box="[185,334,668,695]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="gratiosa">
<emphasis id="B916940FFF874B64C79E98B117185ED4" box="[185,334,668,695]" italics="true" pageId="9" pageNumber="60">Ch. gratiosa</emphasis>
</taxonomicName>
by the much narrower posterior section of the corpus bursae and the conspicuously smaller signa bursae.
</paragraph>
</subSubSection>
<caption id="DF1D1895FF874B64C7B09C9117635A90" ID-DOI="http://doi.org/10.5281/zenodo.13232650" ID-Zenodo-Dep="13232650" httpUri="https://zenodo.org/record/13232650/files/figure.png" pageId="9" pageNumber="60" startId="9.[151,226,1724,1748]" targetBox="[151,1436,741,1705]" targetPageId="9" targetType="figure">
<paragraph id="8BDD481DFF874B64C7B09C9117635A90" blockId="9.[151,1436,1723,1778]" pageId="9" pageNumber="60">
<emphasis id="B916940FFF874B64C7B09C91175D5AB6" bold="true" box="[151,267,1724,1748]" pageId="9" pageNumber="60">Figure 29.</emphasis>
Kyrgyzstan, Transalai Range, Aram-Kungei valley, the type locality of
<taxonomicName id="4C62339EFF874B64C3369C96137A5AB1" authorityName="Saldaitis &amp; Volynkin" authorityYear="2020" box="[1041,1324,1723,1747]" class="Insecta" family="Erebidae" genus="Chelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="gratiosa" subSpecies="kartashovi">
<emphasis id="B916940FFF874B64C3369C96137A5AB1" box="[1041,1324,1723,1747]" italics="true" pageId="9" pageNumber="60">Chelis gratiosa kartashovi</emphasis>
</taxonomicName>
(photo by V. Kartashov).
</paragraph>
</caption>
<subSubSection id="C3781B96FF874B67C7B09D1015DD5DA9" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60" type="description">
<paragraph id="8BDD481DFF874B67C7B09D1015DD5DA9" blockId="9.[151,1436,1853,2048]" lastBlockId="10.[151,1437,195,459]" lastPageId="10" lastPageNumber="61" pageId="9" pageNumber="60">
<emphasis id="B916940FFF874B64C7B09D1017F15B35" bold="true" box="[151,423,1853,1879]" pageId="9" pageNumber="60">Description of female</emphasis>
.
<emphasis id="B916940FFF874B64C69F9D1014905B35" bold="true" box="[440,710,1853,1879]" pageId="9" pageNumber="60">External morphology</emphasis>
(
<figureCitation id="13595498FF874B64C5FF9D1015305B3A" box="[728,870,1853,1880]" captionStart="Figures 1016" captionStartId="5.[151,236,1989,2013]" captionTargetBox="[160,1422,480,1948]" captionTargetId="figure-51@5.[151,1434,480,1971]" captionTargetPageId="5" captionText="Figures 1016. Chelis spp.: adults. The specimens are deposited in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232640" httpUri="https://zenodo.org/record/13232640/files/figure.png" pageId="9" pageNumber="60">Figs 11, 12</figureCitation>
). Forewing length
<quantity id="4C9AE5F8FF874B64C36D9D1012C25B3A" box="[1098,1172,1853,1880]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="60" unit="mm" value="8.0">8 mm</quantity>
. Antenna short, thick, shortly serrate whitish basally and medially and blackish apically. Body robust, brown. Forewing narrow, elongate, with nearly parallel costal and anal margins and pointed apex. Forewing ground color and cilia pale brown. Pattern reduced to two elongate blackish brown spots in the cell connected to each other by thin and diffuse line, longitudinal blackish stroke in subbasal area, intense brown suffusion along the anal margin, and two small diffuse dots in subterminal area. Hindwing pale brown with intense dark brown suffusion basally and along the anal margin, with three diffuse dark brown spots in terminal area. Cilia brown.
<emphasis id="B916940FFF844B67C2679AE917545D62" bold="true" pageId="10" pageNumber="61">Female genitalia</emphasis>
(
<figureCitation id="13595498FF844B67C63F9AC817205D62" box="[280,374,229,256]" captionStart="Figures 2428" captionStartId="8.[151,236,1909,1933]" captionTargetBox="[155,1436,204,1841]" captionTargetId="figure-18@8.[151,1436,189,1872]" captionTargetPageId="8" captionText="Figures 2428. Chelis spp.: adults. Depositories of the specimens: 24 in ASV; 2528 in MWM/ZSM." figureDoi="http://doi.org/10.5281/zenodo.13232648" httpUri="https://zenodo.org/record/13232648/files/figure.png" pageId="10" pageNumber="61">Fig. 26</figureCitation>
). Papillae anales wide, trapezoid with rounded corners, setose. Apophyses posteriores elongate, narrow. Apophyses anteriores fully reduced. Postvaginal area of 8
<superScript id="7C17E555FF844B67C3179B2812695D74" attach="left" box="[1072,1087,261,278]" fontSize="7" pageId="10" pageNumber="61">th</superScript>
sternite sclerotized laterally, with narrow and short medial pocket-like incision. Ostium bursae broad. Ductus bursae long, narrow, dorso-ventrally flattened, narrowed subposteriorly and dilated around ostium, sclerotized medially and posteriorly and membranous anteriorly. Corpus bursae ovoid, membranous, with weak spinulose scobination at the base of appendix bursae and two smalls round signa bursae medially. Appendix bursae broadly conical, membranous, situated postero-ventrally, directed to the left side.
</paragraph>
</subSubSection>
<subSubSection id="C3781B96FF844B67C7B09BD9122F5E52" pageId="10" pageNumber="61" type="distribution">
<paragraph id="8BDD481DFF844B67C7B09BD9122F5E52" blockId="10.[151,1435,499,560]" pageId="10" pageNumber="61">
<emphasis id="B916940FFF844B67C7B09BD917665E6C" bold="true" box="[151,304,500,526]" pageId="10" pageNumber="61">Distribution</emphasis>
. The species is known from
<collectingCountry id="F375088DFF844B67C5969BDE15425E6C" box="[689,788,499,526]" name="Pakistan" pageId="10" pageNumber="61">Pakistan</collectingCountry>
(Azad Kashmir) (
<bibRefCitation id="EFF335ECFF844B67C4D89BDE12EC5E6C" author="Ferguson, D. C." box="[1023,1210,499,526]" pageId="10" pageNumber="61" pagination="181 - 275" refId="ref4007" refString="Ferguson, D. C. (1985) Contributions toward reclassification of the world genera of the tribe Arctiini, Part 1 - Introductrion and a revision of the Neoarctia-Grammia group (Lepidoptera: Arctiidae; Arctiinae). The Entomography. An Annual Review for Biosystematics, 3, 181 - 275." type="journal article" year="1985">Ferguson 1985</bibRefCitation>
) and North
<collectingCountry id="F375088DFF844B67C2789BDE13CD5E6C" box="[1375,1435,499,526]" name="India" pageId="10" pageNumber="61">India</collectingCountry>
(
<collectingRegion id="49A686FFFF844B67C7B89838172F5E52" box="[159,377,533,560]" country="India" name="Himachal Pradesh" pageId="10" pageNumber="61">Himachal Pradesh</collectingRegion>
) (
<bibRefCitation id="EFF335ECFF844B67C6A89838144D5E52" author="Reich, P." box="[399,539,533,560]" pageId="10" pageNumber="61" pagination="233 - 238" refId="ref4062" refString="Reich, P. (1932) Neue Arctiiden. Internationale entomologische Zeitschrift, 26 (22), 233 - 238. [in German]" type="journal article" year="1932">Reich 1932</bibRefCitation>
, as
<taxonomicName id="4C62339EFF844B67C560983B15615E52" box="[583,823,533,560]" class="Insecta" family="Erebidae" genus="Micrarctia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="glaphyra">
<emphasis id="B916940FFF844B67C560983B15615E52" box="[583,823,533,560]" italics="true" pageId="10" pageNumber="61">Micrarctia glaphyra</emphasis>
</taxonomicName>
ab.
<emphasis id="B916940FFF844B67C44E983815845E52" box="[873,978,533,560]" italics="true" pageId="10" pageNumber="61">nebulosa</emphasis>
<bibRefCitation id="EFF335ECFF844B67C4FD9838123B5E52" author="Reich, P." box="[986,1133,533,560]" pageId="10" pageNumber="61" pagination="233 - 238" refId="ref4062" refString="Reich, P. (1932) Neue Arctiiden. Internationale entomologische Zeitschrift, 26 (22), 233 - 238. [in German]" type="journal article" year="1932">Reich, 1932</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>