344 lines
40 KiB
XML
344 lines
40 KiB
XML
<document id="83AD018ABC4AB41C3262AC0F2CFF97F6" ID-DOI="10.5281/zenodo.281568" ID-GBIF-Dataset="eb4d0071-03e7-4670-b730-43e20478afa1" ID-ISSN="1175-5326" ID-Zenodo-Dep="281568" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460492821005" checkinUser="plazi" docAuthor="Saigusa, Toyohei" docDate="2012" docId="03C68C2E8070FF8192C99944388A784B" docLanguage="en" docName="zt03353p068.pdf" docOrigin="Zootaxa 3353" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empis (Planempis) mandarina Frey 1953" docType="treatment" docVersion="8" lastPageNumber="60" masterDocId="FFFFF4568072FF84925E9E2B3B1A7E66" masterDocTitle="Revision of the type series of Empis (Planempis) mandarina Frey, the type species of Planempis (Diptera: Empididae)" masterLastPageNumber="68" masterPageNumber="55" pageNumber="57" updateTime="1698310423591" updateUser="plazi">
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<mods:title id="8EBFE591B8C65679DED3D23E480D6CD7">Revision of the type series of Empis (Planempis) mandarina Frey, the type species of Planempis (Diptera: Empididae)</mods:title>
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<mods:namePart id="046DECD056CAD5A246B95AFAFC43365F">Saigusa, Toyohei</mods:namePart>
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<mods:title id="D0726C9321DC89D39189B3B1AA40EE22">Zootaxa</mods:title>
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<mods:date id="1EC2928F37329A81284334E0016AB5B1">2012</mods:date>
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<mods:number id="C9E242D9CA9D57E9888FBCDCF6AAFE38">3353</mods:number>
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<mods:identifier id="65AB21C9B24A4676E4A961DDF6D58F0B" type="DOI">10.5281/zenodo.281568</mods:identifier>
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<treatment id="03C68C2E8070FF8192C99944388A784B" ID-DOI="http://doi.org/10.5281/zenodo.6180350" ID-GBIF-Taxon="119519897" ID-Zenodo-Dep="6180350" LSID="urn:lsid:plazi:treatment:03C68C2E8070FF8192C99944388A784B" httpUri="http://treatment.plazi.org/id/03C68C2E8070FF8192C99944388A784B" lastPageId="5" lastPageNumber="60" pageId="2" pageNumber="57">
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<subSubSection id="C3756EB38070FF8692C999443AD079CF" pageId="2" pageNumber="57" type="nomenclature">
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<paragraph id="8BD03D388070FF8692C99944398879EF" blockId="2.[151,658,1903,1961]" box="[151,658,1903,1929]" pageId="2" pageNumber="57">
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<heading id="D0988A548070FF8692C99944398879EF" bold="true" box="[151,658,1903,1929]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
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<taxonomicName id="4C6F46BB8070FF8692C99944398879EF" ID-CoL="8BZLZ" authority="Frey, 1953" authorityName="Frey" authorityYear="1953" box="[151,658,1903,1929]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
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<emphasis id="B91BE12A8070FF8692C99944398879EF" bold="true" box="[151,658,1903,1929]" pageId="2" pageNumber="57">
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<emphasis id="B91BE12A8070FF8692C999443BFE79EF" bold="true" box="[151,228,1903,1929]" italics="true" pageId="2" pageNumber="57">Empis</emphasis>
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(
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||
<emphasis id="B91BE12A8070FF8692AA99443A6979EF" bold="true" box="[244,371,1903,1929]" italics="true" pageId="2" pageNumber="57">Planempis</emphasis>
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)
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<emphasis id="B91BE12A8070FF8693DA9944391379EF" bold="true" box="[388,521,1903,1929]" italics="true" pageId="2" pageNumber="57">mandarina</emphasis>
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<bibRefCitation id="EFFE40C98070FF86904F9944398879EF" author="Frey" box="[529,658,1903,1929]" pageId="2" pageNumber="57" refString="Frey, R. (1953) Studien uber ostasiatiche Dipteren. I. Die Gattung Empis L. Notulae Entomologicae, 33, 29 - 57." type="journal article" year="1953">Frey, 1953</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD03D388070FF8692C999BA3AD079CF" blockId="2.[151,658,1903,1961]" box="[151,458,1937,1961]" pageId="2" pageNumber="57">
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(
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<figureCitation id="135421BD8070FF8692C199BA3BF279CF" box="[159,232,1937,1961]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="2" pageNumber="57">Figs. 1</figureCitation>
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A, D, 2, 3A, 5A, B)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756EB38070FF8792C999F039C97EAA" lastPageId="3" lastPageNumber="58" pageId="2" pageNumber="57" type="reference_group">
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<paragraph id="8BD03D388070FF8692C999F03E337996" blockId="2.[151,1321,2010,2032]" box="[151,1321,2010,2032]" pageId="2" pageNumber="57">
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<treatmentCitationGroup id="AB7F1A168070FF8692C999F03E337996" box="[151,1321,2010,2032]" pageId="2" pageNumber="57">
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<taxonomicName id="4C6F46BB8070FF8692C999F039737989" ID-CoL="8BZLZ" authority="Frey, 1953: 40" authorityName="Frey" authorityPageNumber="40" authorityYear="1953" box="[151,617,2010,2032]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
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<emphasis id="B91BE12A8070FF8692C999F03BCD7996" box="[151,215,2011,2032]" italics="true" pageId="2" pageNumber="57">Empis</emphasis>
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(
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<emphasis id="B91BE12A8070FF8692BB99F13A4A7996" box="[229,336,2010,2032]" italics="true" pageId="2" pageNumber="57">Planempis</emphasis>
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)
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<emphasis id="B91BE12A8070FF86930099F13AD77996" box="[350,461,2010,2032]" italics="true" pageId="2" pageNumber="57">mandarina</emphasis>
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<treatmentCitation id="0ACE1B298070FF86938A99F039737989" author="Frey" box="[468,617,2010,2032]" page="40" pageId="2" pageNumber="57" year="1953">
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<bibRefCitation id="EFFE40C98070FF86938A99F0395E7996" author="Frey" box="[468,580,2010,2032]" pageId="2" pageNumber="57" refString="Frey, R. (1953) Studien uber ostasiatiche Dipteren. I. Die Gattung Empis L. Notulae Entomologicae, 33, 29 - 57." type="journal article" year="1953">Frey, 1953</bibRefCitation>
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: 40
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</treatmentCitation>
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</taxonomicName>
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. Type-localities:
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<collectingCountry id="F3787DA88070FF86914999F1384D7996" box="[791,855,2010,2032]" name="China" pageId="2" pageNumber="57">China</collectingCountry>
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: Fukien, Kuatun [after
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<typeStatus id="54D4839A8070FF86961F99F13F847996" box="[1089,1182,2010,2032]" pageId="2" pageNumber="57" type="lectotype">lectotype</typeStatus>
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designation].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD03D388071FF8792C99EB239457EC8" blockId="3.[151,723,152,205]" box="[151,607,152,174]" pageId="3" pageNumber="58">
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<treatmentCitationGroup id="AB7F1A168071FF8792C99EB239457EC8" box="[151,607,152,174]" pageId="3" pageNumber="58">
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<taxonomicName id="4C6F46BB8071FF8792C99EB23AE27ECB" ID-CoL="8BZLZ" authority="Frey, 1954: 417" authorityName="Frey" authorityPageNumber="417" authorityYear="1954" box="[151,504,152,174]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="mandarina">
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<emphasis id="B91BE12A8071FF8792C99EB23A517EC8" box="[151,331,152,174]" italics="true" pageId="3" pageNumber="58">Empis mandarina</emphasis>
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:
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<treatmentCitation id="0ACE1B298071FF8793079EB23AE27ECB" author="Frey" box="[345,504,152,174]" page="417" pageId="3" pageNumber="58" year="1954">
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<bibRefCitation id="EFFE40C98071FF8793079EB23AD27EC8" author="Frey" box="[345,456,152,174]" pageId="3" pageNumber="58" refString="Frey, R. (1954) 28. Empididae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region IV, 4. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 400 - 432, pl. 35 - 36." type="journal article" year="1954">Frey, 1954</bibRefCitation>
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: 417
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</treatmentCitation>
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</taxonomicName>
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(partim.).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD03D388071FF8792C99E9339C97EAA" blockId="3.[151,723,152,205]" box="[151,723,182,205]" pageId="3" pageNumber="58">
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||
<treatmentCitationGroup id="AB7F1A168071FF8792C99E9339C97EAA" box="[151,723,182,205]" pageId="3" pageNumber="58">
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||
<taxonomicName id="4C6F46BB8071FF8792C99E9339777EAA" ID-CoL="8BZLZ" authority="Frey, 1955: 11" authorityName="Frey" authorityPageNumber="11" authorityYear="1955" box="[151,621,182,205]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
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<emphasis id="B91BE12A8071FF8792C99E933BCD7EAB" box="[151,215,184,205]" italics="true" pageId="3" pageNumber="58">Empis</emphasis>
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(
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<emphasis id="B91BE12A8071FF8792BB9E9C3A4A7EAB" box="[229,336,183,205]" italics="true" pageId="3" pageNumber="58">Planempis</emphasis>
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||
)
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<emphasis id="B91BE12A8071FF8793009E9C3AD77EAB" box="[350,461,183,205]" italics="true" pageId="3" pageNumber="58">mandarina</emphasis>
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||
:
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<treatmentCitation id="0ACE1B298071FF8793859E9C39777EAA" author="Frey" box="[475,621,182,204]" page="11" pageId="3" pageNumber="58" year="1955">
|
||
<bibRefCitation id="EFFE40C98071FF8793859E9C39517EAA" author="Frey" box="[475,587,182,204]" pageId="3" pageNumber="58" refString="Frey, R. (1955) Studien uber ostasiatische Dipteren IV. Hilara Meig. (Suppl.), Empis L. (Suppl.), Platypalpus Macq. Notulae Entomologicae, 35, 1 - 14." type="journal article" year="1955">Frey, 1955</bibRefCitation>
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: 11
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</treatmentCitation>
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</taxonomicName>
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(partim.).
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||
</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3756EB38071FF8792C99ED83E3E7F49" pageId="3" pageNumber="58" type="diagnosis">
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||
<paragraph id="8BD03D388071FF8792C99ED83E3E7F49" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">
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<emphasis id="B91BE12A8071FF8792C99ED83A107F6A" bold="true" box="[151,266,243,268]" pageId="3" pageNumber="58">Diagnosis</emphasis>
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. Medium-sized blackish species of
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<emphasis id="B91BE12A8071FF8790EB9EDE39DC7F6A" box="[693,710,245,268]" italics="true" pageId="3" pageNumber="58">E</emphasis>
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. (
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<taxonomicName id="4C6F46BB8071FF8790BD9EDF38437F6A" box="[739,857,244,268]" class="Insecta" family="Empididae" genus="Planempis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B91BE12A8071FF8790BD9EDF38437F6A" box="[739,857,244,268]" italics="true" pageId="3" pageNumber="58">Planempis</emphasis>
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</taxonomicName>
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) with holoptic male eyes, brownish simple legs, entirely infuscated wings, dull blackish mesonotum without acrostichals, and flat-topped male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3756EB38071FF8192979F103BE57A1B" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58" type="description">
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<paragraph id="8BD03D388071FF8792979F103F7F7CF1" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">
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<emphasis id="B91BE12A8071FF8792979F103A4E7F32" bold="true" box="[201,340,315,340]" pageId="3" pageNumber="58">Description</emphasis>
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.
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<emphasis id="B91BE12A8071FF87933F9F17390A7F32" bold="true" box="[353,528,316,340]" pageId="3" pageNumber="58">
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Male
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<typeStatus id="54D4839A8071FF8793FB9F17390A7F32" box="[421,528,316,340]" pageId="3" pageNumber="58" type="lectotype">lectotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="135421BD8071FF87907E9F1739457F32" box="[544,607,316,340]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
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A). Head shrivelled, antennal stylus broken, abdominal terga more or less greasy. Head black, compound eyes touching for 0.25 distance on frons, with upper facets slightly enlarged, 1.5X as large as lower facets; frons and face light grey pollinose, clypeus polished blackish brown; ocellar tubercle with pair of short black setae; occiput densely grey pollinose, with postocular ciliation of rather short black bristles, irregularly biserial rows of ca. 10 fairly long black occipital bristles, and on ventral surface clothed with many long yellow hairs, and similar but shorter ones close to lower postocular area. Antenna except for stylus slightly shorter than shrivelled head length, two basal segments dark brown, bearing a few short black setae; basiflagellomere black, slightly longer than
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<date id="FFD11BF88071FF87939E9C1C3AF87C29" box="[448,482,567,591]" pageId="3" pageNumber="58">3X</date>
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thickness; relative lengths of 3 antennal segments 5:3:10; stylus missing in
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<typeStatus id="54D4839A8071FF8797709C1C3E827C29" box="[1326,1432,567,591]" pageId="3" pageNumber="58" type="lectotype">lectotype</typeStatus>
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. Palpus tawny basally, darkened towards apex, bearing 3 long black setae 0.75X as long as palpus below, and several shorter setae above; labrum reddish brown, 1.4X as long as shrivelled head height.
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</paragraph>
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<paragraph id="8BD03D388071FF8792979C8F3F1A7A66" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">
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Thorax blackish brown (
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<figureCitation id="135421BD8071FF8793AC9C8F39237CDA" box="[498,569,676,700]" captionStart="FIGURE 1" captionStartId="2.[151,250,1769,1792]" captionTargetBox="[340,1246,591,1738]" captionTargetId="figure@2.[340,1247,591,1745]" captionTargetPageId="2" captionText="FIGURE 1. Empis (Planempis) spp., male; A – C: habitus, lateral aspect; D-F: anterior part of body, dorsal aspect. A, D: E. (P.) mandarina Frey, lectotype male; B, E: E. (P.) lindebergi sp. nov., holotype male; C, F: E. (P.) janssoni sp. nov." httpUri="https://zenodo.org/record/281569/files/figure.png" pageId="3" pageNumber="58">Fig. 1</figureCitation>
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D); mesonotum densely pollinose, dull black, subdorsal stripes between acrostichal and dorsocentral stripes dark grey and subequal in width to latter two stripes; margin of humeral callus and notopleural depression chestnut brown; scutellum, postnotum and pleura rather densely dark grey pollinose; spiracles yellowish brown. Chaetotaxy: Most of mesonotal bristles black; acrostichal setae unrecognizable; dorsocentrals uniserial, consisting of at most 15 short fine yellowish setae 0.33—0.5X as long as diameter of prothoracic spiracle, ending in longish prescutellar bristle; humerus with 3—4 black setae and more yellowish hairs; posthumeral to notopleural areas sparsely yellow-haired; 1 short posthumeral, 3 strong notopleurals, 1 weak postalar bristle; scutellum with 3 pairs of bristles, subapical ones longest; anterior pronotum with a few black bristles and on lateral part scattered yellow hairs; proepisternum with several yellow hairs, prosternum bare, laterotergal bristles yellow to yellowish brown arising from oblique oval area.
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</paragraph>
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<paragraph id="8BD03D388071FF8792979A26380E7BEA" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">
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Legs moderately slender and simple, reddish brown to brown and black setose. Coxae as in pleura in colour, clothed with yellow setae and bearing black bristles along distal and posterior margins. Femora very short-setose dorsally, black at extreme tip; f1 with av and pv rows of very short setae, of which apical 2—3 are longer, 0.5X as long as f1 thickness; f2 and f3 (
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<figureCitation id="135421BD8071FF8793A29A5C39257AE9" box="[508,575,1143,1167]" captionStart="FIGURE 5" captionStartId="8.[151,250,1180,1203]" captionTargetBox="[340,1247,447,1151]" captionTargetId="figure@8.[340,1247,447,1155]" captionTargetPageId="8" captionText="FIGURE 5. Hind legs of Empis (Planempis) spp., male; A-D: left hind leg excluding coxae; B: middle part of hind tibia showing ventral erect pile; E: hind femur showing pale posteroventral hairs; A, B: E. (P.) mandarina Frey, lectotype; C: E. (P.) lindebergi sp. nov., holotype; D, E: E. (P.) janssoni sp. nov., holotype." httpUri="https://zenodo.org/record/281573/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
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A) with av and pv row of short black bristles 0.33—0.
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<date id="FFD11BF88071FF8796C49A533FB37AF6" box="[1178,1193,1144,1168]" pageId="3" pageNumber="58">5</date>
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X as long as f2 thickness. Tibiae reddish brown, t3 infuscated to brown except for subbasal part; t1 very short-setose above and beneath; t2 short setose above, densely covered with minute pile beneath, and with 4—6 ad and a few av weak bristles; t3 gradually dilated apically, with subapical part nearly
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<date id="FFD11BF88071FF8790B29ACF38147A9A" box="[748,782,1252,1276]" pageId="3" pageNumber="58">2X</date>
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as thick as subbasal part, densely covered with erect pilelike minute hairs on ventral to pv surfaces (
|
||
<figureCitation id="135421BD8071FF8790D99B2C39D07B79" box="[647,714,1287,1311]" captionStart="FIGURE 5" captionStartId="8.[151,250,1180,1203]" captionTargetBox="[340,1247,447,1151]" captionTargetId="figure@8.[340,1247,447,1155]" captionTargetPageId="8" captionText="FIGURE 5. Hind legs of Empis (Planempis) spp., male; A-D: left hind leg excluding coxae; B: middle part of hind tibia showing ventral erect pile; E: hind femur showing pale posteroventral hairs; A, B: E. (P.) mandarina Frey, lectotype; C: E. (P.) lindebergi sp. nov., holotype; D, E: E. (P.) janssoni sp. nov., holotype." httpUri="https://zenodo.org/record/281573/files/figure.png" pageId="3" pageNumber="58">Fig. 5</figureCitation>
|
||
B), moderately long setae above and bearing av and pv rows of longish bristles 0.75—1.0X as long as t3 thickness. Fore and mid tarsi slender, short setose above, metatarsi with longish bristles on ventral surface; hind tarsus black, metatarsus slightly thicker than subbasal part of t3, short setose and bearing a few longish dorsal and ventral bristles.
|
||
</paragraph>
|
||
<paragraph id="8BD03D388071FF8792979BBC3A8D782E" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">
|
||
Wing (
|
||
<figureCitation id="135421BD8071FF8793499BBC3A407BC9" box="[279,346,1431,1455]" captionStart="FIGURE 3" captionStartId="5.[151,250,786,809]" captionTargetBox="[340,1242,207,757]" captionTargetId="figure@5.[340,1247,193,765]" captionTargetPageId="5" captionText="FIGURE 3. Wings of Empis (Planempis) spp. A: E. (P.) mandarina Frey, lectotype male; B: E. (P.) lindebergi sp. nov., holotype male; C: E. (P.) janssoni sp. nov., holotype male; D: ditto, paratype female 1; E: ditto, paratype female 2; F: E. (P.) sp. female syntype of E. (P.) mandarina sensu Frey from Kuatun." httpUri="https://zenodo.org/record/281571/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
|
||
A) as illustrated, almost
|
||
<date id="FFD11BF88071FF87902B9BBC398D7BC9" box="[629,663,1431,1455]" pageId="3" pageNumber="58">3X</date>
|
||
as long as wide, rather broad, only slightly tapered towards tip and distinctly brownish, pterostigma and veins dark brown; M1, M2, M4 and CuA+CuP not weakened towards wing margin; R4 forming ca. 90˚ to R5 basally, then distinctly curved posteriorly at basal 0.33; outer margin of cell r2+3 about 0.38X as long as cell r4; apex of cell cua not so sharp as in following two species. Halter blackish brown with yellowish brown shaft.
|
||
</paragraph>
|
||
<paragraph id="8BD03D388071FF879297987F3F9078BE" blockId="3.[151,1437,243,2004]" pageId="3" pageNumber="58">Abdomen moderately slender, blackish brown, densely dark grey pollinose, clothed with short yellow hairs, those on lateral parts of abdominal segments 1—3 very long. Abdominal tergum 8 short, 0.2X as long as tergum 7, fused with narrow lateral areas of sternum, and bearing 2—3 minute setae on each side; abdominal sternum 8 0.66X as long as sternum 7, and bearing short posterior marginal setae 0.33X as long as sternum.</paragraph>
|
||
<paragraph id="8BD03D388071FF80929798CF380F7F02" blockId="3.[151,1437,243,2004]" lastBlockId="4.[151,1436,151,356]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
|
||
Male genitalia (
|
||
<figureCitation id="135421BD8071FF87932098CF3AD9789A" box="[382,451,1764,1788]" captionStart="FIGURE 2" captionStartId="4.[151,250,1969,1992]" captionTargetBox="[173,1423,429,1926]" captionTargetId="figure@4.[151,1436,411,1945]" captionTargetPageId="4" captionText="FIGURE 2. Empis (Planempis) mandarina Frey, lectotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281570/files/figure.png" pageId="3" pageNumber="58">Fig. 2</figureCitation>
|
||
A): Flat-topped
|
||
<typeStatus id="54D4839A8071FF87902298CE39AA789B" box="[636,688,1765,1789]" pageId="3" pageNumber="58">type</typeStatus>
|
||
. Cercus dark reddish brown, clothed with short yellow hairs apically; epandrial lobe blackish brown, brownish distally, dark grey pollinose and clothed with black setae. Cerci (
|
||
<figureCitation id="135421BD8071FF8792C199073BEA7922" box="[159,240,1836,1860]" captionStart="FIGURE 2" captionStartId="4.[151,250,1969,1992]" captionTargetBox="[173,1423,429,1926]" captionTargetId="figure@4.[151,1436,411,1945]" captionTargetPageId="4" captionText="FIGURE 2. Empis (Planempis) mandarina Frey, lectotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281570/files/figure.png" pageId="3" pageNumber="58">Figs. 2</figureCitation>
|
||
B, C) with dorsal surface flattened, combined cerci viewed from above subquadrate with lateral margins parallel with each other on basal 0.66, with distal margin only slightly produced, surrounding subquadrate fenestrula, of which posterior open space is 0.5X as long as wide; dorsal process of cercus strongly curved inwardly, simple, only slightly produced ventrally at tip and with single concavity at tip; middle swelling of cercus moderately large, ventral process of cercus short, only slightly extending posteriorly beyond middle swelling, weakly widened distally, produced into short, narrow dorsodistal projection; supra-anal area without pigmentation or sclerotization, subanal area with weak oval sclerotization; epandrial lobe with distal margin more or less straight, oblique and pointed dorsodistally, bearing stiff spine-like bristles in addition to ordinary bristles; hypandrium narrow and separated ventrally; phallus outside from coelom (
|
||
<figureCitation id="135421BD8076FF8091729F2F38607F7A" box="[812,890,260,284]" captionStart="FIGURE 2" captionStartId="4.[151,250,1969,1992]" captionTargetBox="[173,1423,429,1926]" captionTargetId="figure@4.[151,1436,411,1945]" captionTargetPageId="4" captionText="FIGURE 2. Empis (Planempis) mandarina Frey, lectotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect." httpUri="https://zenodo.org/record/281570/files/figure.png" pageId="4" pageNumber="59">Figs. 2</figureCitation>
|
||
D, E) long, middle 0.33 swollen, with pair of triangular lamellate lateral expansions, dorsal keel triangularly produced at middle and irregular transverse wrinkles, its apical 0.33 slender and evenly curved semi-circularly.
|
||
</paragraph>
|
||
<caption id="DF106DB08076FF8092C9999A3F507980" httpUri="https://zenodo.org/record/281570/files/figure.png" pageId="4" pageNumber="59" targetBox="[173,1423,429,1926]" targetPageId="4">
|
||
<paragraph id="8BD03D388076FF8092C9999A3F507980" blockId="4.[151,1436,1969,2022]" pageId="4" pageNumber="59">
|
||
<emphasis id="B91BE12A8076FF8092C9999A3A0E79A0" bold="true" box="[151,276,1969,1992]" pageId="4" pageNumber="59">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6F46BB8076FF8093459999399679A1" authority="Frey" authorityName="Frey" box="[283,652,1969,1991]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
|
||
<emphasis id="B91BE12A8076FF80934599993A4079A1" box="[283,346,1970,1991]" italics="true" pageId="4" pageNumber="59">Empis</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8076FF809335999A3ACC79A1" box="[363,470,1969,1991]" italics="true" pageId="4" pageNumber="59">Planempis</emphasis>
|
||
)
|
||
<emphasis id="B91BE12A8076FF8093BB999A394E79A1" box="[485,596,1969,1991]" italics="true" pageId="4" pageNumber="59">mandarina</emphasis>
|
||
Frey
|
||
</taxonomicName>
|
||
, lectotype male; A; genitalia, lateral aspect; B: cerci, dorsal aspect; C: ditto, posterior aspect; D: phallus (excluding part in coelom), lateral aspect; E. ditto, ventral aspect.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF106DB08077FF8192C99D3938117D00" httpUri="https://zenodo.org/record/281571/files/figure.png" pageId="5" pageNumber="60" targetBox="[340,1242,207,757]" targetPageId="5">
|
||
<paragraph id="8BD03D388077FF8192C99D3938117D00" blockId="5.[151,1436,786,870]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192C99D393A097D4E" bold="true" box="[151,275,786,809]" pageId="5" pageNumber="60">FIGURE 3.</emphasis>
|
||
Wings of
|
||
<taxonomicName id="4C6F46BB8077FF8193209D3F395B7D4F" authority="(Planempis)" baseAuthorityName="Planempis" box="[382,577,787,809]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91BE12A8077FF8193209D3F3AA47D4F" box="[382,446,788,809]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8193939D3839227D4F" box="[461,568,787,809]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
spp. A:
|
||
<taxonomicName id="4C6F46BB8077FF8190C69D3F389F7D4F" authority="Frey" authorityName="Frey" box="[664,901,787,809]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
|
||
<emphasis id="B91BE12A8077FF8190C69D3F39B57D4F" box="[664,687,788,809]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8190E09D3F39D57D4F" box="[702,719,788,809]" italics="true" pageId="5" pageNumber="60">P.</emphasis>
|
||
)
|
||
<emphasis id="B91BE12A8077FF8190BE9D3838557D4F" box="[736,847,787,809]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
|
||
Frey
|
||
</taxonomicName>
|
||
, lectotype male; B:
|
||
<taxonomicName id="4C6F46BB8077FF81960C9D3F3E1A7D4F" box="[1106,1280,787,809]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="lindebergi" status="sp. nov." subGenus="Planempis">
|
||
<emphasis id="B91BE12A8077FF81960C9D3F3F737D4F" box="[1106,1129,788,809]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8196299D3F3F927D4F" box="[1143,1160,788,809]" italics="true" pageId="5" pageNumber="60">P.</emphasis>
|
||
)
|
||
<emphasis id="B91BE12A8077FF8196C79D383E1A7D4F" box="[1177,1280,787,809]" italics="true" pageId="5" pageNumber="60">lindebergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2285C518077FF8197599D383E437D4E" box="[1287,1369,787,808]" pageId="5" pageNumber="60" rank="species">
|
||
<emphasis id="B91BE12A8077FF8197599D383E437D4E" bold="true" box="[1287,1369,787,808]" pageId="5" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male; C:
|
||
<taxonomicName id="4C6F46BB8077FF8193749D193AD27D21" box="[298,456,818,839]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="janssoni" status="sp. nov." subGenus="Planempis">
|
||
<emphasis id="B91BE12A8077FF8193749D193A5B7D21" box="[298,321,818,839]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF81930E9D193A787D21" box="[336,354,818,839]" italics="true" pageId="5" pageNumber="60">P.</emphasis>
|
||
)
|
||
<emphasis id="B91BE12A8077FF81932D9D193AD27D21" box="[371,456,818,839]" italics="true" pageId="5" pageNumber="60">janssoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2285C518077FF81938E9D19393B7D21" box="[464,545,818,839]" pageId="5" pageNumber="60" rank="species">
|
||
<emphasis id="B91BE12A8077FF81938E9D19393B7D21" bold="true" box="[464,545,818,839]" pageId="5" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male; D: ditto, paratype female 1; E: ditto, paratype female 2; F:
|
||
<emphasis id="B91BE12A8077FF8197689D193E577D21" box="[1334,1357,818,839]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8197029D193E777D21" box="[1372,1389,818,839]" italics="true" pageId="5" pageNumber="60">P.</emphasis>
|
||
) sp. female syntype of
|
||
<taxonomicName id="4C6F46BB8077FF81930D9D7A39607D00" authority="Frey" authorityName="Frey" box="[339,634,848,870]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina" subGenus="Planempis">
|
||
<emphasis id="B91BE12A8077FF81930D9D7A3A737D00" box="[339,361,849,870]" italics="true" pageId="5" pageNumber="60">E.</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8193299D7A3A927D00" box="[375,392,849,870]" italics="true" pageId="5" pageNumber="60">P.</emphasis>
|
||
)
|
||
<emphasis id="B91BE12A8077FF8193C69D7B391D7D00" box="[408,519,848,870]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
|
||
sensu Frey
|
||
</taxonomicName>
|
||
from Kuatun.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD03D388077FF8192979DA639717DC3" blockId="5.[151,1437,909,1581]" box="[201,619,909,933]" pageId="5" pageNumber="60">
|
||
Length: Body
|
||
<quantity id="4C9790DD8077FF8193349DA63AD97DC3" box="[362,451,909,933]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="5" pageNumber="60" unit="mm" value="7.4">7.4 mm</quantity>
|
||
; wing
|
||
<quantity id="4C9790DD8077FF8190509DA6397D7DC3" box="[526,615,909,933]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="5" pageNumber="60" unit="mm" value="8.1">8.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD03D388077FF8192979D9A3BE57A1B" blockId="5.[151,1437,909,1581]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192979D9A3AD87DAC" bold="true" box="[201,450,945,970]" pageId="5" pageNumber="60">Specimens examined</emphasis>
|
||
.
|
||
<emphasis id="B91BE12A8077FF81938F9D9A39677DAC" bold="true" box="[465,637,945,970]" pageId="5" pageNumber="60">
|
||
<typeStatus id="54D4839A8077FF81938F9D9A39677DAC" box="[465,637,945,970]" pageId="5" pageNumber="60" type="lectotype">LECTOTYPE</typeStatus>
|
||
</emphasis>
|
||
ɗ (here designated), labelled: Kuatun (
|
||
<quantity id="4C9790DD8077FF8196629D9A3F947DAF" box="[1084,1166,945,969]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="60" unit="m" value="2300.0">2300m</quantity>
|
||
) 27,
|
||
<specimenCount id="9D69F6B18077FF8196E09D9A3FE37DAF" box="[1214,1273,945,969]" pageId="5" pageNumber="60" type="generic">40 n.</specimenCount>
|
||
Br. /117,40 ö. L.J. Klapperich/
|
||
<date id="FFD11BF88077FF81933B9DFE3ADD7D8B" box="[357,455,981,1005]" pageId="5" pageNumber="60" value="1938-04-02">2.4.1938</date>
|
||
(Fukien) // Spec. typ. //
|
||
<taxonomicName id="4C6F46BB8077FF81915E9DFE3F717D8B" authority="Frey" authorityName="Frey" box="[768,1131,981,1005]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">Rhamphomyia mandarina Frey</taxonomicName>
|
||
// [
|
||
<typeStatus id="54D4839A8077FF8196CC9DFE3E227D8B" box="[1170,1336,981,1005]" pageId="5" pageNumber="60" type="lectotype">LECTOTYPE</typeStatus>
|
||
] /
|
||
<taxonomicName id="4C6F46BB8077FF8197089DFC3A057A77" authority="(Planempis)" baseAuthorityName="Planempis" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91BE12A8077FF8197089DFC3E867D88" box="[1366,1436,983,1006]" italics="true" pageId="5" pageNumber="60">Empis</emphasis>
|
||
(
|
||
<emphasis id="B91BE12A8077FF8192FE9DD23A0C7A77" box="[160,278,1017,1041]" italics="true" pageId="5" pageNumber="60">Planempis</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C6F46BB8077FF81936A9DD239377A77" authority="Frey, 1953" authorityName="Frey" authorityYear="1953" box="[308,557,1017,1042]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">
|
||
<emphasis id="B91BE12A8077FF81936A9DD23AB57A77" box="[308,431,1017,1041]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
|
||
<bibRefCitation id="EFFE40C98077FF8193E89DD139377A77" author="Frey" box="[438,557,1017,1042]" pageId="5" pageNumber="60" refString="Frey, R. (1953) Studien uber ostasiatiche Dipteren. I. Die Gattung Empis L. Notulae Entomologicae, 33, 29 - 57." type="journal article" year="1953">Frey, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
/designated by/
|
||
<bibRefCitation id="EFFE40C98077FF8190BA9DD138A57A77" author="SAIGUSA" box="[740,959,1017,1041]" pageId="5" pageNumber="60" refString="Liu, X., Saigusa, T. & Yang, D. (2012) Two new species of Empis (Planempis) from Oriental China, with an updated key to species of China (Diptera: Empidoidea). Zootaxa, 3239, 51 - 57." type="journal article" year="2012">T. SAIGUSA, 2012</bibRefCitation>
|
||
(male genitalia and posterior part of abdomen macerated and preserved in a vial with glycerol, and a wing detached from body and mounted between cover slips with
|
||
<collectingCountry id="F3787DA88077FF81934A9A6A3A727A3F" box="[276,360,1089,1113]" name="Canada" pageId="5" pageNumber="60">Canada</collectingCountry>
|
||
balsam; left hind leg detached from body and mounted on minuten, attached to main pin) (ZFMK).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756EB38077FF8192979AA23BC97A8F" pageId="5" pageNumber="60" type="materials_examined">
|
||
<paragraph id="8BD03D388077FF8192979AA23E597AC7" blockId="5.[151,1437,909,1581]" box="[201,1347,1161,1186]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192979AA239347AC4" bold="true" box="[201,558,1161,1186]" pageId="5" pageNumber="60">Additional material examined</emphasis>
|
||
.
|
||
<specimenCount id="9D69F6B18077FF8190629AA239917AC7" box="[572,651,1161,1185]" pageId="5" pageNumber="60" type="male">1 male</specimenCount>
|
||
, FUKIEN, S.
|
||
<collectingCountry id="F3787DA88077FF8191769AA238777AC7" box="[808,877,1161,1185]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
|
||
/Shaowu: T. C. Maa//
|
||
<date id="FFD11BF88077FF81963C9AA23FCE7AC7" box="[1122,1236,1161,1185]" pageId="5" pageNumber="60" value="1945-03-15">15.iii.1945</date>
|
||
(
|
||
<collectionCode id="ED7EA5FD8077FF8196BD9AA23E2D7AC7" LSID="urn:lsid:biocol.org:col:34697" box="[1251,1335,1161,1185]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="5" pageNumber="60">BPBM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD03D388077FF8192979A853BC97A8F" blockId="5.[151,1437,909,1581]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192979A853A7A7AA0" bold="true" box="[201,352,1197,1222]" pageId="5" pageNumber="60">
|
||
<typeStatus id="54D4839A8077FF8192979A853A187AA0" box="[201,258,1198,1222]" pageId="5" pageNumber="60">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F3787DA88077FF8193309A863AAF7AA3" box="[366,437,1197,1221]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
|
||
: Fukien, Kuatun (
|
||
<quantity id="4C9790DD8077FF8190229A8639CC7AA3" box="[636,726,1197,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="60" unit="m" value="2300.0">2300 m</quantity>
|
||
), 2740ʹ N, 11740ʹ E". This locality is located in the Wui Mountains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756EB38077FF8192979ADE390F7B6B" box="[201,533,1269,1294]" pageId="5" pageNumber="60" type="distribution">
|
||
<paragraph id="8BD03D388077FF8192979ADE390F7B6B" blockId="5.[151,1437,909,1581]" box="[201,533,1269,1294]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192979ADE3A417B68" bold="true" box="[201,347,1269,1294]" pageId="5" pageNumber="60">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3787DA88077FF8193379ADE3AB67B6B" box="[361,428,1269,1293]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
|
||
(Fujian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3756EB38077FF8192979B31388A784B" pageId="5" pageNumber="60" type="discussion">
|
||
<paragraph id="8BD03D388077FF8192979B313A867BFB" blockId="5.[151,1437,909,1581]" pageId="5" pageNumber="60">
|
||
<emphasis id="B91BE12A8077FF8192979B313A2C7B54" bold="true" box="[201,310,1306,1330]" pageId="5" pageNumber="60">Remarks</emphasis>
|
||
. As stated in the above section, the male from Kuatun is selected as the
|
||
<typeStatus id="54D4839A8077FF81962C9B323FC27B57" box="[1138,1240,1305,1329]" pageId="5" pageNumber="60" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6F46BB8077FF81975F9B313A587B33" authority="Frey. Although" authorityName="Frey. Although" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">
|
||
<emphasis id="B91BE12A8077FF81975F9B313E867B57" box="[1281,1436,1305,1329]" italics="true" pageId="5" pageNumber="60">E. mandarina</emphasis>
|
||
Frey. Although
|
||
</taxonomicName>
|
||
the specimen is different from the original description in the absence of acrostichal setae, and its male genitalia were not illustrated in Die Fliegen, the author selected the male from
|
||
<collectingCountry id="F3787DA88077FF8196179B4A3F967B1F" box="[1097,1164,1377,1401]" name="China" pageId="5" pageNumber="60">China</collectingCountry>
|
||
in accordance with specific name,
|
||
<taxonomicName id="4C6F46BB8077FF8193449BAD3A8F7BF8" box="[282,405,1414,1438]" class="Insecta" family="Chironomidae" genus="Rhamphomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">
|
||
<emphasis id="B91BE12A8077FF8193449BAD3A8F7BF8" box="[282,405,1414,1438]" italics="true" pageId="5" pageNumber="60">mandarina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD03D388077FF8192979B823F707B83" blockId="5.[151,1437,909,1581]" pageId="5" pageNumber="60">The additional male has a fine, short yellowish seta on the dorsomedial line on the middle of mesoscutum, therefore this species seems to have a potential to bear a few uniserial acrostichal setae.</paragraph>
|
||
<paragraph id="8BD03D388077FF8192979BD9388A784B" blockId="5.[151,1437,909,1581]" pageId="5" pageNumber="60">
|
||
<taxonomicName id="4C6F46BB8077FF8192979BD93A8F786F" box="[201,405,1521,1545]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="mandarina">
|
||
<emphasis id="B91BE12A8077FF8192979BD93A8F786F" box="[201,405,1521,1545]" italics="true" pageId="5" pageNumber="60">Empis mandarina</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from the following two new species by the tawny legs, broader wings, dorsal process of the cercus only slightly produced ventrally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |