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<document id="CE996719258E62A20D55361085F1A1FE" ID-DOI="10.5852/ejt.2018.464" ID-GBIF-Dataset="2fc5fa89-0438-4096-b785-78d14b6bfb76" ID-ISSN="2118-9773" ID-Zenodo-Dep="1469475" ID-ZooBank="urn:lsid:zoobank.org:pub:3C501DE6-AC85-4199-8CDC-D28E42753050" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1540291417600" checkinUser="plazi" docAuthor="Zallot, Enrico, Fehér, Zoltán, Bamberger, Sonja &amp; Gittenberger, Edmund" docDate="2018" docId="03C6575CFFFAFFE7C10CE673FB91FDC9" docLanguage="en" docName="ejt-464_zallot_feher_bamberger_gittenberger.pdf.imf" docOrigin="European Journal of Taxonomy 464" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Lovcenia) tropojanum Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger 2018, sp. nov." docType="treatment" docUuid="F986C8B8-F64C-41FE-9422-5B0FF2D749FB" docUuidSource="ZooBank" docVersion="17" lastPageNumber="19" masterDocId="FFFF2F24FFEAFFF5C052E55DFF9AFFC3" masterDocTitle="Cochlostoma revised: the subgenus Lovcenia Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="17" updateTime="1736338903792" updateUser="guilherme">
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<mods:title id="B590891241EB4921100A697F19FDAD62">Cochlostoma revised: the subgenus Lovcenia Zallot et al., 2015 (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="CA1F3E67A85E5CE7E97B6DB1533B77F3">Zallot, Enrico</mods:namePart>
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<mods:namePart id="11FB0F2E5D2D9E55E865430A1B9F8CE3">Fehér, Zoltán</mods:namePart>
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<mods:namePart id="6D036B8C96271AEFF34ADB9C8C526E39">Bamberger, Sonja</mods:namePart>
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<mods:namePart id="FE5AC6725A64829DFC0D28169431CA83">Gittenberger, Edmund</mods:namePart>
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<mods:title id="B69B58E824BE2505BB3F85B50077B5D0">European Journal of Taxonomy</mods:title>
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<mods:number id="DF0B5257EF5EA3E67C7369247E56E7AF">2018-10-02</mods:number>
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<treatment id="03C6575CFFFAFFE7C10CE673FB91FDC9" ID-DOI="http://doi.org/10.5281/zenodo.5631643" ID-GBIF-Taxon="148967380" ID-Zenodo-Dep="5631643" LSID="urn:lsid:zoobank.org:act:F986C8B8-F64C-41FE-9422-5B0FF2D749FB" httpUri="http://treatment.plazi.org/id/03C6575CFFFAFFE7C10CE673FB91FDC9" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" scope_family="Cochlostomatidae" scope_order="Architaenioglossa" scope_subClass="Orthogastropoda" scope_subGenus="Lovcenia" scope_superFamily="Cyclophoroidea" scope_superOrder="Caenogastropoda">
<subSubSection id="C375B5C1FFFAFFE5C10CE673FB4FFC8A" box="[350,1237,814,841]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BD0E64AFFFAFFE5C10CE673FB4FFC8A" blockId="16.[350,1237,814,912]" box="[350,1237,814,841]" pageId="16" pageNumber="17">
<taxonomicName id="4C6F9DC9FFFAFFE5C10CE673FBEEFC8B" ID-CoL="WNPJ" authority="Zallot, Feher &amp; Gittenberger" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[350,1140,814,841]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="tropojanum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C10CE673FC86FC8B" bold="true" box="[350,796,814,841]" italics="true" pageId="16" pageNumber="17">Cochlostoma (Lovcenia) tropojanum</emphasis>
Zallot, Fehér &amp; Gittenberger
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFAFFE5C429E672FB4FFC8A" box="[1147,1237,815,841]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C375B5C1FFFAFFE5C121E60CFCECFC4C" pageId="16" pageNumber="17" type="description">
<paragraph id="8BD0E64AFFFAFFE5C121E60CFB5BFCAF" blockId="16.[350,1237,814,912]" box="[371,1217,849,876]" pageId="16" pageNumber="17">urn:lsid:zoobank.org:act:F986C8B8-F64C-41FE-9422-5B0FF2D749FB</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C2ECE628FCECFC4C" blockId="16.[350,1237,814,912]" box="[702,886,885,912]" pageId="16" pageNumber="17">
<figureCitation id="1354FACFFFFAFFE5C2ECE628FC86FC53" box="[702,796,885,912]" captionStart="Fig. 5" captionStartId="10.[189,232,1859,1885]" captionTargetBox="[189,1398,625,1829]" captionTargetId="figure@10.[189,1398,625,1829]" captionTargetPageId="10" captionText="Fig. 5. Female specimens of Cochlostoma (Lovcenia) Zallot et al., 2015 used for anatomy (shells were destroyed before dissection). A. C. (Lovcenia) erika (A.J. Wagner, 1906) (HNHM 86387). B. C. (Lovcenia) dalmatinum (L. Pfeiffer, 1863) (NHMW 110430/MN/0162). C. C. (Lovcenia) tropojanum sp. nov. (NHMW 111248). D. C. (Lovcenia) jakschae sp. nov. (NHMW 111249). E. C. (Lovcenia) lanatum sp. nov. (HNHM 99861)." figureDoi="http://doi.org/10.5281/zenodo.1469485" httpUri="https://zenodo.org/record/1469485/files/figure.png" pageId="16" pageNumber="17">Figs 5C</figureCitation>
,
<figureCitation id="1354FACFFFFAFFE5C37AE628FCD6FC4C" box="[808,844,885,911]" captionStart="Fig. 6" captionStartId="11.[189,232,1824,1850]" captionTargetBox="[191,1395,878,1778]" captionTargetId="figure@11.[191,1474,844,1819]" captionTargetPageId="11" captionText="Fig. 6. Female genital morphology. See Table 1 for localities and Table 3 for measurements. A. Cochlostoma (Lovcenia) erika (A.J. Wagner, 1906) (HNHM 86387). B. Cochlostoma (Lovcenia) dalmatinum (L. Pfeiffer, 1863) (NHMW 110430/MN/0162). C. Cochlostoma (Lovcenia) tropojanum sp. nov. (NHMW 111248). D1. Cochlostoma (Lovcenia) jakschae sp. nov., Bugarska Peak population (NHMW 111249). D2. Cochlostoma (Lovcenia) jakschae sp. nov., Pejë Pass population (HNHM 97221). E. Cochlostoma (Lovcenia) lanatum sp. nov. (HNHM 99861)." figureDoi="http://doi.org/10.5281/zenodo.1469487" httpUri="https://zenodo.org/record/1469487/files/figure.png" pageId="16" pageNumber="17">6C</figureCitation>
,
<figureCitation id="1354FACFFFFAFFE5C30BE628FCECFC4C" box="[857,886,885,911]" captionStart="Fig. 10" captionStartId="17.[189,232,1927,1953]" captionTargetBox="[189,1398,1138,1893]" captionTargetId="figure@17.[189,1398,1138,1893]" captionTargetPageId="17" captionText="Fig. 10. Cochlostoma (Lovcenia) tropojanum Zallot, Fehér &amp; Gittenberger sp. nov., holotype, Tropojë Valley, Albania (NHMW 111638)." figureDoi="http://doi.org/10.5281/zenodo.1469495" httpUri="https://zenodo.org/record/1469495/files/figure.png" pageId="16" pageNumber="17">10</figureCitation>
</paragraph>
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<subSubSection id="C375B5C1FFFAFFE5C0EFE6EBFED3FADB" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BD0E64AFFFAFFE5C0EFE6EBFEACFC13" blockId="16.[189,310,950,976]" box="[189,310,950,976]" pageId="16" pageNumber="17">
<heading id="D0985126FFFAFFE5C0EFE6EBFEACFC13" bold="true" box="[189,310,950,976]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91B3A58FFFAFFE5C0EFE6EBFEACFC13" bold="true" box="[189,310,950,976]" pageId="16" pageNumber="17">Diagnosis</emphasis>
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<paragraph id="8BD0E64AFFFAFFE5C0EFE6B8FED3FADB" blockId="16.[189,1399,997,1304]" pageId="16" pageNumber="17">
Rather small, with a very slender, spotless and regularly ribbed shell, which has a poorly developed lip, inclined towards the front and broadening at the columellar side, before curving back and covering the umbilicus. In
<taxonomicName id="4C6F9DC9FFFAFFE5C133E171FE6FFB86" baseAuthorityName="A. J. Wagner" baseAuthorityYear="1906" box="[353,501,1067,1094]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="erika" subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C133E171FE6FFB86" box="[353,501,1067,1094]" italics="true" pageId="16" pageNumber="17">C. (L.) erika</emphasis>
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and
<taxonomicName id="4C6F9DC9FFFAFFE5C260E171FC8CFB86" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1863" box="[562,790,1067,1094]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="dalmatinum" subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C260E171FC8CFB86" box="[562,790,1067,1094]" italics="true" pageId="16" pageNumber="17">C. (L.) dalmatinum</emphasis>
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, which are both larger, the columellar lobe differs; the latter species may also be distinguished by its prominent, irregular ribs, which are more uniform in
<taxonomicName id="4C6F9DC9FFFAFFE5C0EFE12FFE0DFB48" authority="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger, 2018" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[189,407,1138,1164]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="tropojanum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C0EFE12FFE0DFB48" box="[189,407,1138,1164]" italics="true" pageId="16" pageNumber="17">C. (L.) tropojanum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFAFFE5C1CFE12FFE6CFB4F" box="[413,502,1138,1164]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C6F9DC9FFFAFFE5C1AEE12FFCB4FB4F" authority="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger, 2018" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[508,814,1138,1164]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="lanatum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C1AEE12FFCB4FB4F" box="[508,814,1138,1164]" italics="true" pageId="16" pageNumber="17">Cochlostoma (L.) lanatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFAFFE5C366E12FFC17FB4F" box="[820,909,1138,1164]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
differs by its regularly spaced, prominent, sharp ribs on the upper whorls. The geographically separate
<taxonomicName id="4C6F9DC9FFFAFFE5C32CE1C8FBA3FB6C" authority="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger, 2018" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[894,1081,1173,1199]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="jakschae" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C32CE1C8FBA3FB6C" box="[894,1081,1173,1199]" italics="true" pageId="16" pageNumber="17">C. (L.) jakschae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFAFFE5C412E1C8FB00FB6C" box="[1088,1178,1173,1199]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
is conchologically similar but less slender, with an even narrower lip. It also differs in the female genitalia, i.e., by the less convoluted visceral oviduct. Moreover,
<taxonomicName id="4C6F9DC9FFFAFFE5C2C3E181FCD4FB36" authority="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger, 2018" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[657,846,1243,1270]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="jakschae" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFAFFE5C2C3E181FCD4FB36" box="[657,846,1243,1270]" italics="true" pageId="16" pageNumber="17">C. (L.) jakschae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFAFFE5C307E181FC2AFB35" box="[853,944,1244,1270]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
has, relative to its shell size, a smaller protoconch.
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFAFFE5C0EFE01DFC50F8FA" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BD0E64AFFFAFFE5C0EFE01DFE30FA99" blockId="16.[189,426,1344,1370]" box="[189,426,1344,1370]" pageId="16" pageNumber="17">
<heading id="D0985126FFFAFFE5C0EFE01DFE30FA99" bold="true" box="[189,426,1344,1370]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91B3A58FFFAFFE5C0EFE01DFE30FA99" bold="true" box="[189,426,1344,1370]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
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<emphasis id="B91B3A58FFFAFFE5C0B8E032FEC6FA4A" bold="true" box="[234,348,1391,1417]" pageId="16" pageNumber="17">Holotype</emphasis>
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:
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(in ethanol),
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gorge of Lumi i Tropojës, ca
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N of Tropojë
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a.s.l.
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,
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<collectingDate id="EF953962FFFAFFE5C279E0E8FD5CFA0C" box="[555,710,1461,1487]" pageId="16" pageNumber="17" value="2014-06-26">26 Jun. 2014</collectingDate>
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<collectorName id="269A839CFFFAFFE5C29DE0E8FCB2FA0C" box="[719,808,1461,1487]" pageId="16" pageNumber="17">Angyal</collectorName>
,
<collectorName id="269A839CFFFAFFE5C366E0E8FCEDFA0C" box="[820,887,1461,1487]" pageId="16" pageNumber="17">Fehér</collectorName>
and
<collectorName id="269A839CFFFAFFE5C3E2E0E8FC63FA0C" box="[944,1017,1461,1487]" pageId="16" pageNumber="17">Grego</collectorName>
leg. (
<specimenCode id="DBC94E31FFFAFFE5C469E0E8FB64FA13" box="[1083,1278,1461,1488]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17">NHMW 111638</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C0B8E0B2FBE7F9B0" blockId="16.[189,1399,1519,1651]" pageId="16" pageNumber="17">
<materialsCitation id="3B07EC17FFFAFFE5C0B8E0B2FE46F993" ID-GBIF-Occurrence="1931541355" collectingDate="2014-06-26" collectionCode="NHMW, HNHM" collectorName="Angyal &amp; Feher &amp; Grego" country="Albania" county="Tropoje District" elevation="925" latitude="42.4692" location="ALBANIA" longLatPrecision="6" longitude="20.1554" municipality="Lumi" pageId="16" pageNumber="17" specimenCode="NHMW 111248/8a+2ja, HNHM 99243/10a+3ja, GR/5a" specimenCount="28" typeStatus="paratype">
<typeStatus id="54D458E8FFFAFFE5C0B8E0B2FEFDF9CA" box="[234,359,1519,1545]" pageId="16" pageNumber="17" type="paratype">
<emphasis id="B91B3A58FFFAFFE5C0B8E0B2FEFDF9CA" bold="true" box="[234,359,1519,1545]" pageId="16" pageNumber="17">Paratypes</emphasis>
</typeStatus>
<collectingCountry id="F378A6DAFFFAFFE5C0EFE34FFEDCF9EF" box="[189,326,1554,1580]" name="Albania" pageId="16" pageNumber="17">ALBANIA</collectingCountry>
:
<specimenCount id="9D692DC3FFFAFFE5C10EE34FFE51F9EF" box="[348,459,1554,1580]" count="28" pageId="16" pageNumber="17" type="generic">28 spec.</specimenCount>
, same collection data as for holotype (
<specimenCode id="DBC94E31FFFAFFE5C38DE34FFA9FF9EF" box="[991,1285,1554,1580]" pageId="16" pageNumber="17">NHMW 111248/8a+2ja</specimenCode>
,
<specimenCode id="DBC94E31FFFAFFE5C54FE34FFEEBF98C" pageId="16" pageNumber="17">HNHM 99243/10a+3ja</specimenCode>
;
<specimenCode id="DBC94E31FFFAFFE5C12FE368FE54F993" box="[381,462,1589,1616]" pageId="16" pageNumber="17">GR/5a</specimenCode>
);
</materialsCitation>
<materialsCitation id="3B07EC17FFFAFFE5C1B1E368FBE7F9B0" ID-GBIF-Occurrence="1931541267" collectingDate="2014-06-26" collectionCode="NHMW, HNHM" collectorName="Angyal &amp; Feher &amp; Grego" country="Albania" elevation="925" latitude="42.4692" location="ca 13 km N of Tropoje" longLatPrecision="6" longitude="20.1554" pageId="16" pageNumber="17" specimenCode="111247/2ja, 99241/1a+1ja" specimenCount="4">
<specimenCount id="9D692DC3FFFAFFE5C1B1E368FDA2F993" box="[483,568,1589,1616]" count="4" pageId="16" pageNumber="17" type="generic">4 spec.</specimenCount>
,
<location id="8EB0B091FFFAFFE5C216E36BFCC8F98C" LSID="urn:lsid:plazi:treatment:03C6575CFFFAFFE7C10CE673FB91FDC9:8EB0B091FFFAFFE5C216E36BFCC8F98C" box="[580,850,1589,1616]" country="Albania" latitude="42.4692" longLatPrecision="6" longitude="20.1554" name="ca 13 km N of Tropoje" pageId="16" pageNumber="17">
ca
<quantity id="4C974BAFFFFAFFE5C234E368FD2AF98C" box="[614,688,1589,1615]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="16" pageNumber="17" unit="km" value="13.0">13 km</quantity>
N of Tropojë
</location>
,
<elevation id="00420179FFFAFFE5C30CE368FC73F98C" box="[862,1001,1589,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="16" pageNumber="17" unit="m" value="925.0">
<quantity id="4C974BAFFFFAFFE5C30CE368FC33F993" box="[862,937,1589,1616]" metricMagnitude="2" metricUnit="m" metricValue="9.25" pageId="16" pageNumber="17" unit="m" value="925.0">925 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE5B808DFFFAFFE5C3A4E368FB1BF993" box="[1014,1153,1589,1616]" degrees="42.4692" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="42.4692">42.4692° N</geoCoordinate>
,
<geoCoordinate id="EE5B808DFFFAFFE5C4DFE368FA8EF993" box="[1165,1300,1589,1616]" degrees="20.1554" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="20.1554">20.1554° E</geoCoordinate>
,
<collectingDate id="EF953962FFFAFFE5C573E368FF61F9B1" pageId="16" pageNumber="17" value="2014-06-26">
<date id="FFD1C08AFFFAFFE5C573E368FF61F9B1" pageId="16" pageNumber="17" value="2014-06-26">26 Jun. 2014</date>
</collectingDate>
,
<collectorName id="269A839CFFFAFFE5C157E305FEC4F9B1" box="[261,350,1624,1650]" pageId="16" pageNumber="17">Angyal</collectorName>
,
<collectorName id="269A839CFFFAFFE5C138E305FE37F9B1" box="[362,429,1624,1650]" pageId="16" pageNumber="17">Fehér</collectorName>
and
<collectorName id="269A839CFFFAFFE5C1B4E305FDB5F9B1" box="[486,559,1624,1650]" pageId="16" pageNumber="17">Grego</collectorName>
leg. (
<collectionCode id="ED7E7E8FFFFAFFE5C222E305FD48F9B1" box="[624,722,1624,1650]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="16" pageNumber="17">NHMW</collectionCode>
<specimenCode id="DBC94E31FFFAFFE5C28BE305FCC5F9B1" box="[729,863,1624,1650]" pageId="16" pageNumber="17">111247/2ja</specimenCode>
,
<collectionCode id="ED7E7E8FFFFAFFE5C338E304FC5EF9B0" box="[874,964,1625,1651]" collectionName="Hungary, Budapest, Hungarian Natural History Museum" pageId="16" pageNumber="17">HNHM</collectionCode>
<specimenCode id="DBC94E31FFFAFFE5C398E305FBEAF9B0" box="[970,1136,1624,1651]" pageId="16" pageNumber="17">99241/1a+1ja</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C0B8E3CFFE3CF96F" blockId="16.[189,1399,1682,1850]" box="[234,422,1682,1708]" pageId="16" pageNumber="17">
<emphasis id="B91B3A58FFFAFFE5C0B8E3CFFE3CF96F" bold="true" box="[234,422,1682,1708]" pageId="16" pageNumber="17">Other material</emphasis>
</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C0EFE3EBFC50F8FA" blockId="16.[189,1399,1682,1850]" pageId="16" pageNumber="17">
<materialsCitation id="3B07EC17FFFAFFE5C0EFE3EBFAEDF930" ID-GBIF-Occurrence="1931541268" collectingDate="2016-07-01" collectingDateMax="2016-07-01" collectingDateMin="2006-06-06" collectionCode="NHMW, HNHM" collectorName="Eross &amp; Feher &amp; Szekeres &amp; Grego" country="Albania" county="Tropoje District" elevation="390" latitude="42.4632" location="gorge of Lumi i Gashit, SW of Dretove" longLatPrecision="6" longitude="20.1258" municipality="Szekeres" pageId="16" pageNumber="17" specimenCode="NHMW 111654, HNHM 99865" specimenCount="1">
<collectingCountry id="F378A6DAFFFAFFE5C0EFE3EBFEDCF913" box="[189,326,1718,1744]" name="Albania" pageId="16" pageNumber="17">ALBANIA</collectingCountry>
:
<collectingCounty id="62B19EC6FFFAFFE5C104E3E8FD80F90C" box="[342,538,1717,1743]" pageId="16" pageNumber="17">Tropojë District</collectingCounty>
,
<location id="8EB0B091FFFAFFE5C27EE3EBFBBDF913" LSID="urn:lsid:plazi:treatment:03C6575CFFFAFFE7C10CE673FB91FDC9:8EB0B091FFFAFFE5C27EE3EBFBBDF913" box="[556,1063,1717,1744]" country="Albania" county="Tropoje District" latitude="42.4632" longLatPrecision="6" longitude="20.1258" municipality="Szekeres" name="gorge of Lumi i Gashit, SW of Dretove" pageId="16" pageNumber="17">gorge of Lumi i Gashit, SW of Dretovë</location>
,
<elevation id="00420179FFFAFFE5C46AE3E8FB54F90C" box="[1080,1230,1717,1744]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="17" unit="m" value="390.0">
<quantity id="4C974BAFFFFAFFE5C46AE3E8FB13F913" box="[1080,1161,1717,1744]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="16" pageNumber="17" unit="m" value="390.0">390 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE5B808DFFFAFFE5C4B3E3E8FAE8F913" box="[1249,1394,1717,1744]" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="42.4009">42.4009° N</geoCoordinate>
,
<geoCoordinate id="EE5B808DFFFAFFE5C0EFE384FED2F930" box="[189,328,1753,1779]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="20.0879">20.0879° E</geoCoordinate>
,
<date id="FFD1C08AFFFAFFE5C10BE384FE72F931" box="[345,488,1752,1779]" pageId="16" pageNumber="17" value="2016-07-01">
<collectingDate id="EF953962FFFAFFE5C10BE384FE72F931" box="[345,488,1752,1779]" pageId="16" pageNumber="17" value="2016-07-01">1 Jul. 2016</collectingDate>
</date>
,
<collectorName id="269A839CFFFAFFE5C1A4E384FDA0F930" box="[502,570,1753,1779]" pageId="16" pageNumber="17">Erőss</collectorName>
,
<collectorName id="269A839CFFFAFFE5C219E385FD0AF931" box="[587,656,1752,1778]" pageId="16" pageNumber="17">Fehér</collectorName>
,
<collectorName id="269A839CFFFAFFE5C2F2E385FC92F931" box="[672,776,1752,1778]" pageId="16" pageNumber="17">Szekeres</collectorName>
and
<collectorName id="269A839CFFFAFFE5C318E384FC09F930" box="[842,915,1753,1779]" pageId="16" pageNumber="17">Grego</collectorName>
leg. (
<specimenCode id="DBC94E31FFFAFFE5C38DE384FB3FF930" box="[991,1189,1752,1779]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17">NHMW 111654</specimenCode>
,
<specimenCode id="DBC94E31FFFAFFE5C4E4E384FAF3F930" box="[1206,1385,1752,1779]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="17">HNHM 99865</specimenCode>
);
</materialsCitation>
<materialsCitation id="3B07EC17FFFAFFE5C0EFE3A1FC50F8FA" ID-GBIF-Occurrence="1931541266" collectedFrom="in rocky grassland" collectingDate="2006-06-06" collectingDateMax="2016-07-01" collectingDateMin="2006-06-06" collectionCode="NHMW, HNHM" collectorName="Barina" country="Albania" county="Tropoje District" elevation="2020" latitude="42.4632" location="above Tropoje, at N foot of Mt Shkelzen" longLatPrecision="6" longitude="20.1258" municipality="Szekeres" pageId="16" pageNumber="17" specimenCode="HNHM 99871" specimenCount="1">
<collectingCounty id="62B19EC6FFFAFFE5C0EFE3A1FE1EF8D5" box="[189,388,1788,1814]" pageId="16" pageNumber="17">Tropojë District</collectingCounty>
,
<location id="8EB0B091FFFAFFE5C1CAE3A1FC2EF8D5" LSID="urn:lsid:plazi:treatment:03C6575CFFFAFFE7C10CE673FB91FDC9:8EB0B091FFFAFFE5C1CAE3A1FC2EF8D5" box="[408,948,1788,1814]" country="Albania" county="Tropoje District" latitude="42.4632" longLatPrecision="6" longitude="20.1258" municipality="Szekeres" name="above Tropoje, at N foot of Mt Shkelzen" pageId="16" pageNumber="17">above Tropojë, at N foot of Mt Shkëlzen</location>
,
<collectedFrom id="3873A5B7FFFAFFE5C39AE3A1FB29F8D5" box="[968,1203,1788,1814]" pageId="16" pageNumber="17">in rocky grassland</collectedFrom>
,
<elevation id="00420179FFFAFFE5C495E3A1FAEAF8D5" box="[1223,1392,1788,1814]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="16" pageNumber="17" unit="m" value="2020.0">
<quantity id="4C974BAFFFFAFFE5C495E3A1FAB3F8D5" box="[1223,1321,1788,1814]" metricMagnitude="3" metricUnit="m" metricValue="2.02" pageId="16" pageNumber="17" unit="m" value="2020.0">2020 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE5B808DFFFAFFE5C0EFE242FEDDF8FA" box="[189,327,1823,1849]" degrees="42.4632" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="42.4632">42.4632° N</geoCoordinate>
,
<geoCoordinate id="EE5B808DFFFAFFE5C101E242FE40F8FA" box="[339,474,1823,1850]" degrees="20.1258" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="20.1258">20.1258° E</geoCoordinate>
,
<date id="FFD1C08AFFFAFFE5C1B4E242FDE8F8FA" box="[486,626,1823,1849]" pageId="16" pageNumber="17" value="2006-06-06">
<collectingDate id="EF953962FFFAFFE5C1B4E242FDE8F8FA" box="[486,626,1823,1849]" pageId="16" pageNumber="17" value="2006-06-06">6 Jun. 2006</collectingDate>
</date>
,
<collectorName id="269A839CFFFAFFE5C22FE242FD51F8FA" box="[637,715,1823,1849]" pageId="16" pageNumber="17">Barina</collectorName>
leg. (
<specimenCode id="DBC94E31FFFAFFE5C35FE242FC27F8FA" box="[781,957,1823,1849]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="17">HNHM 99871</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFAFFE4C0EFE23CFBE6FC00" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8BD0E64AFFFAFFE5C0EFE23CFEEFF8B8" blockId="16.[189,373,1889,1915]" box="[189,373,1889,1915]" pageId="16" pageNumber="17">
<heading id="D0985126FFFAFFE5C0EFE23CFEEFF8B8" bold="true" box="[189,373,1889,1915]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B91B3A58FFFAFFE5C0EFE23CFEEFF8B8" bold="true" box="[189,373,1889,1915]" pageId="16" pageNumber="17">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C0EFE2D2FB8DF86A" blockId="16.[189,1047,1935,1961]" box="[189,1047,1935,1961]" pageId="16" pageNumber="17">
<typeStatus id="54D458E8FFFAFFE5C0EFE2D2FEB5F86A" box="[189,303,1935,1961]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: H
<quantity id="4C974BAFFFFAFFE5C104E2D2FE29F86A" box="[342,435,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="17" unit="mm" value="9.3">9.3 mm</quantity>
, W
<quantity id="4C974BAFFFFAFFE5C1B2E2D2FDA7F86A" box="[480,573,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="17" unit="mm" value="3.8">3.8 mm</quantity>
, Wbw
<quantity id="4C974BAFFFFAFFE5C2DDE2D2FD76F86A" box="[655,748,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="17" unit="mm" value="3.3">3.3 mm</quantity>
, Ha
<quantity id="4C974BAFFFFAFFE5C373E2D2FCE4F86A" box="[801,894,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="17" unit="mm" value="2.7">2.7 mm</quantity>
, Wa
<quantity id="4C974BAFFFFAFFE5C3E4E2D2FB89F86A" box="[950,1043,1935,1961]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="17" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
<paragraph id="8BD0E64AFFFAFFE5C0EFE28CFAEDF828" blockId="16.[189,1399,2000,2027]" box="[189,1399,2000,2027]" pageId="16" pageNumber="17">
<typeStatus id="54D458E8FFFAFFE5C0EFE28CFEB5F828" box="[189,303,2001,2027]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
(N= 12): H
<quantity id="4C974BAFFFFAFFE5C1ECE28DFDD4F828" box="[446,590,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.3" metricValueMin="7.7" pageId="16" pageNumber="17" unit="mm" value="8.5" valueMax="9.3" valueMin="7.7">7.7–9.3 mm</quantity>
, W 3.4–4.0 mm, Wbw
<quantity id="4C974BAFFFFAFFE5C308E28DFC73F828" box="[858,1001,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="16" pageNumber="17" unit="mm" value="3.0" valueMax="3.2" valueMin="2.8">2.8–3.2 mm</quantity>
, Ha
<quantity id="4C974BAFFFFAFFE5C44EE28DFB36F828" box="[1052,1196,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="16" pageNumber="17" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2–2.8 mm</quantity>
, Wa
<quantity id="4C974BAFFFFAFFE5C4B1E28DFAE9F828" box="[1251,1395,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="16" pageNumber="17" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1–2.9 mm</quantity>
.
</paragraph>
<paragraph id="8BD0E64AFFFBFFE4C0EFE456FED4FEE6" blockId="17.[189,1399,267,587]" box="[189,334,267,293]" pageId="17" pageNumber="18">
<heading id="D0985126FFFBFFE4C0EFE456FED4FEE6" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B91B3A58FFFBFFE4C0EFE456FED4FEE6" bold="true" box="[189,334,267,293]" pageId="17" pageNumber="18">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFBFFE4C0EFE467FCB0FD89" blockId="17.[189,1399,267,587]" pageId="17" pageNumber="18">SHELL. Small, with 8½ whorls, H/W ratio 2.8. The initial 2.4–2.5 whorls form the protoconch, which is smooth for only the initial 0.3–0.5 whorl and finely ribbed with narrowly spaced riblets further on. The spotless shell is light yellowish. The teleoconch whorls are convex (Rr =1.24) with whitish, rounded, moderately developed ribs. The ribs are regularly spaced, without any variation between the upper whorls and the body whorl. The ribbing is still present on the final part of the body whorl, approaching the aperture. The aperture is relatively small (Ha/H =0.19–0.20), with a poorly developed lip, inclined towards the front. The lip is broadened at the columellar side before curving back and covering the umbilicus. The body whorl has a small external lobe.</paragraph>
<paragraph id="8BD0E64AFFFBFFE4C0EFE72FFED4FCFF" blockId="17.[189,1399,626,828]" pageId="17" pageNumber="18">FEMALE GENITALIA. Bursa copulatrix large, with a pedunculus connected proximally; its proximal lobe clearly smaller than the distal one. Short seminal receptacle, confined to the ventral side of the body and connected with the pedunculus without a distal oviduct. The 3–4 wide and small loops of the visceral oviduct are situated over the apex of the seminal receptacle. The junction of the uterine gland and the copulatory duct is moderately far from the connection between the seminal receptacle and the pedunculus of the bursa.</paragraph>
<paragraph id="8BD0E64AFFFBFFE4C0EFE639FBE6FC00" blockId="17.[189,1399,867,964]" pageId="17" pageNumber="18">MALE GENITALIA. The penis is longer than the body and thin, with a tapering tip. The spermiduct is narrowly twisted. The body spermiduct is straight. There is a sharp and long groove, ending in a wellformed sperm pocket, delimited on the frontal side by an ascending sperm funnel.</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFFBFFE4C0EFE6B6FC3AFBF0" box="[189,928,1003,1076]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BD0E64AFFFBFFE4C0EFE6B6FEDEFBC6" blockId="17.[189,928,1003,1076]" box="[189,324,1003,1029]" pageId="17" pageNumber="18">
<heading id="D0985126FFFBFFE4C0EFE6B6FEDEFBC6" bold="true" box="[189,324,1003,1029]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="B91B3A58FFFBFFE4C0EFE6B6FEDEFBC6" bold="true" box="[189,324,1003,1029]" pageId="17" pageNumber="18">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFFBFFE4C0EFE144FC3AFBF0" blockId="17.[189,928,1003,1076]" box="[189,928,1049,1076]" pageId="17" pageNumber="18">
This species is named after the Tropojë Valley, its
<typeStatus id="54D458E8FFFBFFE4C358E147FCA7FBF7" box="[778,829,1050,1076]" pageId="17" pageNumber="18">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF10B6C2FFFBFFE4C0EFE2DAFDD5F806" ID-DOI="http://doi.org/10.5281/zenodo.1469495" httpUri="https://zenodo.org/record/1469495/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1927,1953]" targetBox="[189,1398,1138,1893]" targetPageId="17">
<paragraph id="8BD0E64AFFFBFFE4C0EFE2DAFDD5F806" blockId="17.[189,1399,1927,1989]" pageId="17" pageNumber="18">
<emphasis id="B91B3A58FFFBFFE4C0EFE2DAFE80F862" bold="true" box="[189,282,1927,1953]" pageId="17" pageNumber="18">Fig. 10.</emphasis>
<taxonomicName id="4C6F9DC9FFFBFFE4C171E2DAFBA8F862" authority="Zallot, Feher &amp; Gittenberger" authorityName="Zallot &amp; Fehér &amp; Bamberger &amp; Gittenberger" authorityYear="2018" box="[291,1074,1927,1954]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="tropojanum" status="sp. nov." subGenus="Lovcenia">
<emphasis id="B91B3A58FFFBFFE4C171E2DAFD4FF862" box="[291,725,1927,1954]" italics="true" pageId="17" pageNumber="18">Cochlostoma (Lovcenia) tropojanum</emphasis>
Zallot, Fehér &amp; Gittenberger
</taxonomicName>
<taxonomicNameLabel id="A2288723FFFBFFE4C468E2D5FB02F861" box="[1082,1176,1928,1954]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Tropojë Valley, Albania (NHMW 111638).
</paragraph>
</caption>
<subSubSection id="C375B5C1FFF8FFE7C0EFE456FEB5FEB4" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BD0E64AFFF8FFE7C0EFE456FECCFEE6" blockId="18.[189,342,267,293]" box="[189,342,267,293]" pageId="18" pageNumber="19">
<heading id="D0985126FFF8FFE7C0EFE456FECCFEE6" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B91B3A58FFF8FFE7C0EFE456FECCFEE6" bold="true" box="[189,342,267,293]" pageId="18" pageNumber="19">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFF8FFE7C0EFE467FEB5FEB4" blockId="18.[189,1399,314,375]" pageId="18" pageNumber="19">
Mount Shkëlzen in the eastern part of the Prokletije Mts in NE
<collectingCountry id="F378A6DAFFF8FFE7C386E467FBAFFE97" box="[980,1077,314,340]" name="Albania" pageId="18" pageNumber="19">Albania</collectingCountry>
. Known from four nearby locations.
</paragraph>
</subSubSection>
<subSubSection id="C375B5C1FFF8FFE7C0EFE4C3FB91FDC9" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BD0E64AFFF8FFE7C0EFE4C3FE32FE7B" blockId="18.[189,424,414,440]" box="[189,424,414,440]" pageId="18" pageNumber="19">
<heading id="D0985126FFF8FFE7C0EFE4C3FE32FE7B" bold="true" box="[189,424,414,440]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B91B3A58FFF8FFE7C0EFE4C3FE32FE7B" bold="true" box="[189,424,414,440]" pageId="18" pageNumber="19">Habitat preference</emphasis>
</heading>
</paragraph>
<paragraph id="8BD0E64AFFF8FFE7C0EFE490FB91FDC9" blockId="18.[189,1399,461,523]" pageId="18" pageNumber="19">
Like most species of
<taxonomicName id="4C6F9DC9FFF8FFE7C191E490FDC4FE24" box="[451,606,461,487]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="B91B3A58FFF8FFE7C191E490FDC4FE24" box="[451,606,461,487]" italics="true" pageId="18" pageNumber="19">Cochlostoma</emphasis>
</taxonomicName>
, these are obligate rock-dwelling snails that can be found on the surface of limestone cliffs or under boulders on rocky alpine grasslands.
</paragraph>
</subSubSection>
</treatment>
</document>
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