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<document id="E0123B88FED5D35282F367F9356CF5A8" ID-CLB-Dataset="295759" ID-DOI="10.3906/zoo-1712-1" ID-GBIF-Dataset="b9acd017-c3c4-4b64-b9b6-9d9518b91eba" ID-ISSN="1303-6114" ID-Zenodo-Dep="10973843" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1713182411332" checkinUser="felipe" docAuthor="Lucena, Rudá Amorim &amp; Christoffersen, Martin Lindsey" docDate="2018" docId="03C4812FFFA13A7AFCE65910E53FDFA0" docLanguage="en" docName="TurkJZool.42.4.372-388.pdf" docOrigin="Turkish Journal of Zoology 42 (4)" docSource="http://dx.doi.org/10.3906/zoo-1712-1" docStyle="DocumentStyle:8169CCCB0FD041ECF72354F5C265AA0F.2:TurkJZool.2013-.journal_article.1cover" docStyleId="8169CCCB0FD041ECF72354F5C265AA0F" docStyleName="TurkJZool.2013-.journal_article.1cover" docStyleVersion="2" docTitle="Anoplodactylus ricardoi Lucena &amp; Christoffersen 2018, sp. nov." docType="treatment" docVersion="3" lastPageNumber="383" masterDocId="FFFDF957FFAB3A76FFC75C04E671DE18" masterDocTitle="Anoplodactylus (Pycnogonida: Phoxichilidiidae) from Brazil, new records and two new species" masterLastPageNumber="388" masterPageNumber="372" pageNumber="381" updateTime="1715099430494" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B3CC8ABCC0E0F8F6DE2558F0135AE648">Anoplodactylus (Pycnogonida: Phoxichilidiidae) from Brazil, new records and two new species</mods:title>
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<mods:namePart id="411C777EBCEE6975ED007CC6F3B6E891">Lucena, Rudá Amorim</mods:namePart>
<mods:affiliation id="C99159F9C6C48AD9DCD4FB274A93ED7A">B, Martin Lindsey CHRISTOFFERSEN &amp; B &amp; Postgraduate Program in Biological Sciences</mods:affiliation>
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<mods:namePart id="9F3220CD9433F3DCC0BEE701D95F8D02">Christoffersen, Martin Lindsey</mods:namePart>
<mods:affiliation id="7A77113AD69ADDFD2A18EE8316EED0B2">B, Martin Lindsey CHRISTOFFERSEN &amp; B &amp; Postgraduate Program in Biological Sciences &amp; B &amp; Noninsect Invertebrate Biodiversity Laboratory, Federal University of Paraíba, João Pessoa, Paraíba, Brazil</mods:affiliation>
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<treatment id="03C4812FFFA13A7AFCE65910E53FDFA0" ID-DOI="http://doi.org/10.5281/zenodo.11127868" ID-Zenodo-Dep="11127868" LSID="urn:lsid:plazi:treatment:03C4812FFFA13A7AFCE65910E53FDFA0" httpUri="http://treatment.plazi.org/id/03C4812FFFA13A7AFCE65910E53FDFA0" lastPageId="12" lastPageNumber="383" pageId="10" pageNumber="381">
<subSubSection id="C37763B2FFA13A7CFCE65910E35EDB35" box="[801,1327,1300,1325]" pageId="10" pageNumber="381" type="nomenclature">
<paragraph id="8BD23039FFA13A7CFCE65910E35EDB35" blockId="10.[763,1380,191,1896]" box="[801,1327,1300,1325]" pageId="10" pageNumber="381">
<taxonomicName id="4C6D4BBAFFA13A7CFCE65910E25BDB34" ID-CoL="B4KH4" authority="Lucena &amp; Christoffersen, 2018" authorityName="Lucena &amp; Christoffersen" authorityYear="2018" box="[801,1066,1300,1324]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="ricardoi" status="sp. nov.">
<emphasis id="B919EC2BFFA13A7CFCE65910E25BDB34" bold="true" box="[801,1066,1300,1324]" italics="true" pageId="10" pageNumber="381">Anoplodactylus ricardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A5150FFA13A7CFBF75911E2F1DB34" box="[1072,1152,1301,1324]" pageId="10" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13562CBCFFA13A7CFB4B5911E29EDB35" box="[1164,1263,1301,1325]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="10" pageNumber="381">Figures 3</figureCitation>
a3h)
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA13A7CFCE65930E511DBD3" pageId="10" pageNumber="381" type="materials_examined">
<paragraph id="8BD23039FFA13A7CFCE65930E511DBD3" blockId="10.[763,1380,191,1896]" pageId="10" pageNumber="381">
<emphasis id="B919EC2BFFA13A7CFCE65930E58CDB54" bold="true" box="[801,1021,1332,1356]" pageId="10" pageNumber="381">Material examined:</emphasis>
<materialsCitation id="3B053A64FFA13A7CFBC25930E2ACDB94" collectingDate="2014-07-14" collectionCode="UFPB, R" collectorName="de Pipa &amp; L. M. Falkenberg" location="Rio Grande do Norte" pageId="10" pageNumber="381" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<collectingRegion id="49A9FEDBFFA13A7CFBC25930E285DB54" box="[1029,1268,1332,1356]" country="Brazil" name="Rio Grande do Norte" pageId="10" pageNumber="381">Rio Grande do Norte</collectingRegion>
:
<typeStatus id="54D68E9BFFA13A7CFAC75930E312DB54" box="[1280,1379,1332,1356]" pageId="10" pageNumber="381" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED7CA8FCFFA13A7CFCDE5950E52DDB74" box="[793,860,1364,1388]" country="Brazil" httpUri="http://grbio.org/cool/pjvz-tkai" name="Departamento de Sistematica e Ecologia" pageId="10" pageNumber="381">UFPB</collectionCode>
.PYC245)
<specimenCount id="9D6BFBB0FFA13A7CFC1F5950E273DB76" box="[984,1026,1364,1390]" count="1" pageId="10" pageNumber="381" type="male">1 ♂</specimenCount>
, Praia
<collectorName id="269855EFFFA13A7CFB965950E2D7DB74" box="[1105,1190,1364,1388]" pageId="10" pageNumber="381">de Pipa</collectorName>
, intertidal,
<date id="FFD316F9FFA13A7CFAE35950E541DB94" pageId="10" pageNumber="381" value="2014-07-14">
<collectingDate id="EF97EF11FFA13A7CFAE35950E541DB94" pageId="10" pageNumber="381" value="2014-07-14">14 Jul 2014</collectingDate>
</date>
, coll.
<collectionCode id="ED7CA8FCFFA13A7CFCB75970E5F5DB94" box="[880,900,1396,1420]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="381">R</collectionCode>
.A.
<collectingRegion id="49A9FEDBFFA13A7CFC605970E585DB93" box="[935,1012,1396,1419]" country="Philippines" name="Lucena" pageId="10" pageNumber="381">Lucena</collectingRegion>
and
<collectorName id="269855EFFFA13A7CFBED5970E2ACDB94" box="[1066,1245,1396,1420]" pageId="10" pageNumber="381">L.M. Falkenberg</collectorName>
</materialsCitation>
.
<materialsCitation id="3B053A64FFA13A7CFB205970E52ADBD3" collectingDate="2014-07-14" collectionCode="UFPB" collectorName="de Pipa &amp; L. M. Falkenberg" location="Rio Grande do Norte" pageId="10" pageNumber="381" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Rio Grande do Norte" typeStatus="paratype">
<typeStatus id="54D68E9BFFA13A7CFB205970E33FDB94" box="[1255,1358,1396,1420]" pageId="10" pageNumber="381" type="paratype">Paratypes</typeStatus>
(
<collectionCode id="ED7CA8FCFFA13A7CFCC45997E536DBB3" box="[771,839,1427,1451]" country="Brazil" httpUri="http://grbio.org/cool/pjvz-tkai" name="Departamento de Sistematica e Ecologia" pageId="10" pageNumber="381">UFPB</collectionCode>
.PYC134)
<specimenCount id="9D6BFBB0FFA13A7CFC075997E58CDBB6" box="[960,1021,1427,1454]" count="2" pageId="10" pageNumber="381" type="male">2 ♂♂</specimenCount>
and
<specimenCount id="9D6BFBB0FFA13A7CFBF35997E22CDBB6" box="[1076,1117,1427,1454]" count="1" pageId="10" pageNumber="381" type="female">1 ♀</specimenCount>
, same collection data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA13A7CFCE659D7E279D930" pageId="10" pageNumber="381" type="diagnosis">
<paragraph id="8BD23039FFA13A7CFCE659D7E279D930" blockId="10.[763,1380,191,1896]" pageId="10" pageNumber="381">
<emphasis id="B919EC2BFFA13A7CFCE659D7E5E9DBF3" bold="true" box="[801,920,1491,1515]" pageId="10" pageNumber="381">Diagnosis:</emphasis>
Segmentation incomplete, absent between segments 3 and 4. Lateral processes separated by half of diameter; elongate and with many small setae on dorsal surface. Fingers elongate, with setae at base of fixed finger; movable finger with setae in median region. Palp reduced, formed by a single cylindrical article in female. Propodus with eight spines on heel, 2 large at apex and 6 directed towards sole. Sole with eight small spines along its extension, marginalized by small setae. Female with 2 alar ornamentations, formed by small teen of distal margin, at the base of the proboscis.
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA13A7AFCE65B2BE4D0DAB5" lastPageId="12" lastPageNumber="383" pageId="10" pageNumber="381" type="description">
<paragraph id="8BD23039FFA13A7DFCE65B2BE208D97F" blockId="10.[763,1380,191,1896]" lastBlockId="11.[763,1380,1806,1896]" lastPageId="11" lastPageNumber="382" pageId="10" pageNumber="381">
<emphasis id="B919EC2BFFA13A7CFCE65B2BE275D95F" bold="true" box="[801,1028,1839,1863]" pageId="10" pageNumber="381">Description (Male):</emphasis>
Trunk rounded dorsally, with incomplete segmentation, absent between segments 3 and 4 (
<figureCitation id="13562CBCFFA03A7DFFBE5B0AE6AAD93E" box="[121,219,1806,1830]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="11" pageNumber="382">Figures 3</figureCitation>
a3b). Proboscis cylindrical. Lateral processes separated by half diameter, elongate and with many small setae on dorsal surface (
<figureCitation id="13562CBCFFA03A7DFE425B54E788D970" box="[389,505,1872,1896]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="11" pageNumber="382">Figure 3d</figureCitation>
). Ocular tubercle conical, short; eyes well developed, strongly pigmented. Abdomen cylindrical, slightly larger than ocular tubercle, curved backwards, with distal setae.
</paragraph>
<caption id="DF1260B1FFA03A7DFF995A69E6E4D8C3" ID-DOI="http://doi.org/10.5281/zenodo.11127816" ID-Zenodo-Dep="11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="11" pageNumber="382" startId="11.[94,159,1645,1667]" targetBox="[301,1172,189,1626]" targetPageId="11" targetType="figure">
<paragraph id="8BD23039FFA03A7DFF995A69E6E4D8C3" blockId="11.[94,1380,1645,1755]" pageId="11" pageNumber="382">
<emphasis id="B919EC2BFFA03A7DFF995A69E6C6D89A" bold="true" box="[94,183,1645,1667]" pageId="11" pageNumber="382">Figure 3.</emphasis>
<taxonomicName id="4C6D4BBAFFA03A7DFF7B5A6AE7EDD89B" authority="Lucena &amp; Christoffersen, 2018" authorityName="Lucena &amp; Christoffersen" authorityYear="2018" box="[188,412,1646,1667]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="382" phylum="Arthropoda" rank="species" species="ricardoi" status="sp. nov.">
<emphasis id="B919EC2BFFA03A7DFF7B5A6AE7EDD89B" box="[188,412,1646,1667]" italics="true" pageId="11" pageNumber="382">Anoplodactylus ricardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A5150FFA03A7DFE665A6AE798D89B" box="[417,489,1646,1667]" pageId="11" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
(UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm).
</paragraph>
</caption>
<paragraph id="8BD23039FFA73A7AFF435CBAE760DFEB" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
Cheliphores with scapus uniarticulate, with sparse setae, more concentrated on distal margin. Palm of chela smaller than fingers, with many setae (
<figureCitation id="13562CBCFFA73A7AFE3B5CFAE414DF0E" box="[508,613,254,278]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="12" pageNumber="383">Figure 3e</figureCitation>
). Fingers elongate, with setae at base of fixed finger; movable finger with setae in median region. Oviger with 6 articles (
<figureCitation id="13562CBCFFA73A7AFD475D39E609DF6D" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="12" pageNumber="383">Figure 3f</figureCitation>
). First article very dilated, without setae. Setae beginning on second article. Second and third articles almost of the same size, the third being slightly bigger. Fourth and fifth articles dilated. Sixth article ovalate. Fifth and sixth articles with many setae.
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435DF8E442DD68" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
Legs elongate with many setae (
<figureCitation id="13562CBCFFA73A7AFE285DFFE42FDC0B" box="[495,606,507,531]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="12" pageNumber="383">
Figure
<quantity id="4C959DDCFFA73A7AFD855DFFE42FDC0B" box="[578,606,507,531]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="12" pageNumber="383" unit="g" value="3.0">3g</quantity>
</figureCitation>
). Coxa 1 smaller than 3, together both smaller than coxa 2. Femur and tibia robust, with an elongate setae on dorso-distal region. Tube of cement gland short, directed upwards, forming an almost square rectangle with the femur. Tibia 1 smaller than tibia 2. Tibia 2 smaller than femur. Tarsum rectangular, larger than length; with many setae on ventral surface. Propodus with 8 spines on heel, 2 large ones at apex and 6 directed towards the sole (
<figureCitation id="13562CBCFFA73A7AFDD55EFDE4F3DD09" box="[530,642,761,785]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="12" pageNumber="383">Figure 3h</figureCitation>
). Sole with eight small spines along its extension, marginalized by small setae. Main claw with two-thirds of length of propodus. Auxiliary claws strongly reduced.
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435F73E47EDDF7" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF435F73E709DD97" bold="true" box="[132,376,887,911]" pageId="12" pageNumber="383">Description (Female):</emphasis>
Almost equal to male, differing by owning 2 alar ornamentations, formed by small teeth on distal margin, at base of proboscis. Palp reduced, formed by a single cylindrical article (
<figureCitation id="13562CBCFFA73A7AFE5C5FD3E473DDF7" box="[411,514,983,1007]" captionStart="Figure 3" captionStartId="11.[94,159,1645,1667]" captionTargetBox="[301,1172,189,1626]" captionTargetId="figure-54@11.[299,1176,186,1625]" captionTargetPageId="11" captionText="Figure 3. Anoplodactylus ricardoi sp. nov. (UFPB.PYC245). Adult male. Habitus: a, dorsal view (scale bar: 1 mm); b, lateral view (scale bar: 1 mm); d, lateral process (scale bar: 0.05 mm); e, chela (scale bar: 0.05 mm); f, oviger (scale bar: 0.1 mm); g, leg 3 (scale bar: 0.25 mm; detail of g: 0.025 mm); h, propodus (scale bar: 0.05 mm). (UFPB.PYC134) Adult female. c, proboscis ventral view (scale bar: 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.11127816" httpUri="https://zenodo.org/record/11127816/files/figure.png" pageId="12" pageNumber="383">Figure 3c</figureCitation>
).
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435FF3E4D0DAB5" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF435FF3E460DA16" bold="true" box="[132,529,1014,1039]" pageId="12" pageNumber="383">
Measurements (
<typeStatus id="54D68E9BFFA73A7AFEF35FF2E7EFDA16" box="[308,414,1014,1038]" pageId="12" pageNumber="383" type="holotype">Holotype</typeStatus>
in mm):
</emphasis>
Length of trunk, 1.66; width of trunk (above segment 2), 1.20; length of abdomen, 0.38; length of proboscis, 0.74; length of cheliphore, 0.97; third leg v coxa 1, 0.26; coxa 2, 0.61; coxa 3, 0.39; femur, 1.03; tibia 1, 0.81; tibia 2, 1.00; tarsum, 0.13; propodum, 0.67; main claw, 0.38; auxiliary claws, 0.05.
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA73A7AFF4358B1E42DDAD5" box="[132,604,1205,1229]" pageId="12" pageNumber="383" type="distribution">
<paragraph id="8BD23039FFA73A7AFF4358B1E42DDAD5" blockId="12.[94,711,190,1895]" box="[132,604,1205,1229]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF4358B1E769DAD5" bold="true" box="[132,280,1205,1229]" pageId="12" pageNumber="383">Distribution:</emphasis>
<collectingCountry id="F37A70A9FFA73A7AFEDA58B1E72ADAD5" box="[285,347,1205,1229]" name="Brazil" pageId="12" pageNumber="383">Brazil</collectingCountry>
(
<collectingRegion id="49A9FEDBFFA73A7AFEAE58B1E421DAD5" box="[361,592,1205,1229]" country="Brazil" name="Rio Grande do Norte" pageId="12" pageNumber="383">Rio Grande do Norte</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA73A7AFF4358D1E6D5DB14" pageId="12" pageNumber="383" type="materials_examined">
<paragraph id="8BD23039FFA73A7AFF4358D1E6D5DB14" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
<materialsCitation id="3B053A64FFA73A7AFF4358D1E6EEDB14" collectorName="de Pipa" country="Brazil" location="Praia de Pipa" pageId="12" pageNumber="383" specimenCount="1" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<emphasis id="B919EC2BFFA73A7AFF4358D1E751DAF4" bold="true" box="[132,288,1236,1261]" pageId="12" pageNumber="383">
<typeStatus id="54D68E9BFFA73A7AFF4358D1E6CBDAF5" box="[132,186,1237,1261]" pageId="12" pageNumber="383">Type</typeStatus>
locality:
</emphasis>
<location id="8EB266E2FFA73A7AFEED58D0E7B3DAF4" LSID="urn:lsid:plazi:treatment:03C4812FFFA13A7AFCE65910E53FDFA0:8EB266E2FFA73A7AFEED58D0E7B3DAF4" box="[298,450,1236,1260]" country="Brazil" name="Praia de Pipa" pageId="12" pageNumber="383" stateProvince="Rio Grande do Norte">
Praia
<collectorName id="269855EFFFA73A7AFEAB58D0E7B3DAF4" box="[364,450,1236,1260]" pageId="12" pageNumber="383">de Pipa</collectorName>
</location>
,
<collectingRegion id="49A9FEDBFFA73A7AFE1658D0E4B2DAF4" box="[465,707,1236,1260]" country="Brazil" name="Rio Grande do Norte" pageId="12" pageNumber="383">Rio Grande do Norte</collectingRegion>
,
<collectingCountry id="F37A70A9FFA73A7AFF9958F0E6EEDB14" box="[94,159,1268,1292]" name="Brazil" pageId="12" pageNumber="383">Brazil</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA73A7AFF435910E72EDB93" pageId="12" pageNumber="383" type="etymology">
<paragraph id="8BD23039FFA73A7AFF435910E734DB73" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF435910E774DB34" bold="true" box="[132,261,1300,1324]" pageId="12" pageNumber="383">Etymology:</emphasis>
The specific epithet
<taxonomicName id="4C6D4BBAFFA73A7AFE1A5910E448DB34" authorityName="Lucena &amp; Christoffersen" authorityYear="2018" box="[477,569,1300,1324]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="ricardoi">
<emphasis id="B919EC2BFFA73A7AFE205910E448DB34" box="[487,569,1300,1324]" italics="true" pageId="12" pageNumber="383">ricardoi</emphasis>
</taxonomicName>
” is a homage to the professor Doctor Ricardo
<collectingRegion id="49A9FEDBFFA73A7AFE105930E457DB53" box="[471,550,1332,1355]" country="Philippines" name="Lucena" pageId="12" pageNumber="383">Lucena</collectingRegion>
, from Federal University of
<collectingRegion id="49A9FEDBFFA73A7AFF295957E730DB73" box="[238,321,1363,1387]" country="Brazil" name="Paraiba" pageId="12" pageNumber="383">Paraíba</collectingRegion>
.
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435977E72EDB93" blockId="12.[94,711,190,1895]" box="[132,351,1395,1419]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF435977E6A0DB93" bold="true" box="[132,209,1395,1419]" pageId="12" pageNumber="383">Depth:</emphasis>
Mesolittoral.
</paragraph>
</subSubSection>
<subSubSection id="C37763B2FFA73A7AFF435997E53FDFA0" pageId="12" pageNumber="383" type="discussion">
<paragraph id="8BD23039FFA73A7AFF435997E72AD832" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
<emphasis id="B919EC2BFFA73A7AFF435997E69CDBB3" bold="true" box="[132,237,1427,1451]" pageId="12" pageNumber="383">Remarks:</emphasis>
This species belongs to the group
<taxonomicName id="4C6D4BBAFFA73A7AFD685997E748DBD2" baseAuthorityName="Arango and Maxmen" baseAuthorityYear="2006" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="californicus-digitatus">
<emphasis id="B919EC2BFFA73A7AFD685997E4B0DBB2" box="[687,705,1427,1450]" italics="true" pageId="12" pageNumber="383">A</emphasis>
.
<emphasis id="B919EC2BFFA73A7AFF9959B6E748DBD2" box="[94,313,1458,1482]" italics="true" pageId="12" pageNumber="383">californicus-digitatus</emphasis>
</taxonomicName>
, proposed by
<bibRefCitation id="EFFC4DC8FFA73A7AFE1C59B7E6D6DBF2" author="Arango CP &amp; Maxmen A" pageId="12" pageNumber="383" pagination="51 - 64" refId="ref12650" refString="Arango CP, Maxmen A (2006). Proboscis ornamentation as a diagnostic character for the Anoplodactylus californicus-digitatus complex (Arthropoda: Pycnogonida) with an example from the Anoplodactylus eroticus female. Zootaxa 1311: 51 - 64." type="journal article" year="2006">Arango and Maxmen (2006)</bibRefCitation>
. The group consists of 14 species that have one or more pairs of tubercles in the ventral region of the proboscis of the female.
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435A35E4FBD93F" blockId="12.[94,711,190,1895]" pageId="12" pageNumber="383">
Of the 14 known species, the new species is very similar to
<taxonomicName id="4C6D4BBAFFA73A7AFFBB5A55E762D871" authorityName="Hall" authorityYear="1912" box="[124,275,1617,1641]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B919EC2BFFA73A7AFFBB5A55E6FFD870" box="[124,142,1617,1640]" italics="true" pageId="12" pageNumber="383">A</emphasis>
.
<emphasis id="B919EC2BFFA73A7AFF5C5A55E762D871" box="[155,275,1617,1641]" italics="true" pageId="12" pageNumber="383">californicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6D4BBAFFA73A7AFE8D5A55E7DBD871" authorityName="Marcus" authorityYear="1940" box="[330,426,1617,1641]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="stictus">
<emphasis id="B919EC2BFFA73A7AFE8D5A55E72DD870" box="[330,348,1617,1640]" italics="true" pageId="12" pageNumber="383">A</emphasis>
.
<emphasis id="B919EC2BFFA73A7AFEAE5A56E7DBD871" box="[361,426,1618,1641]" italics="true" pageId="12" pageNumber="383">stictus</emphasis>
</taxonomicName>
, mainly due to the pair of alar processes on the proboscis of the female, the tube of the cement gland, and its body pattern. The absence of dorsal tubercles on the lateral processes, the presence of dentate alar processes on the proboscis of the female, and the large quantity of setae on the appendages and body are the main characters that distinguish the two species.
</paragraph>
<paragraph id="8BD23039FFA73A7AFF435B2BE53FDFA0" blockId="12.[94,711,190,1895]" lastBlockId="12.[763,1380,190,1896]" pageId="12" pageNumber="383">
As for
<taxonomicName id="4C6D4BBAFFA73A7AFF1D5B2BE709D95F" authorityName="Hall" authorityYear="1912" box="[218,376,1839,1863]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="californicus">
<emphasis id="B919EC2BFFA73A7AFF1D5B2BE69DD95E" box="[218,236,1839,1862]" italics="true" pageId="12" pageNumber="383">A</emphasis>
.
<emphasis id="B919EC2BFFA73A7AFEC75B2BE709D95F" box="[256,376,1839,1863]" italics="true" pageId="12" pageNumber="383">californicus</emphasis>
</taxonomicName>
, the closest species, we may distinguish it from
<taxonomicName id="4C6D4BBAFFA73A7AFEF95B4BE44FD97F" authority="Lucena &amp; Christoffersen, 2018" authorityName="Lucena &amp; Christoffersen" authorityYear="2018" box="[318,574,1871,1895]" class="Pycnogonida" family="Phoxichilidiidae" genus="Anoplodactylus" kingdom="Animalia" order="Pantopoda" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="ricardoi" status="sp. nov.">
<emphasis id="B919EC2BFFA73A7AFEF95B4BE44FD97F" box="[318,574,1871,1895]" italics="true" pageId="12" pageNumber="383">Anoplodactylus ricardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A5150FFA73A7AFD8C5B54E4D0D97F" box="[587,673,1872,1895]" pageId="12" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
by the latter being more robust, with lateral processes closer to each other and larger in the distal margin, and with many small dorsal setae; ocular tubercle and abdomen more robust; ocular tubercle closer to the insertion of the cheliphores, which are conical and occupy all the lateral region of the neck; auxiliary claws almost imperceptible; presence of a small tubercle above the insertion of the ovigers.
</paragraph>
</subSubSection>
</treatment>
</document>