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<document id="1444F6D47BE0A0A3B3CA4A5EE995E9E8" ID-DOI="10.15298/rusentj.32.3.09" ID-ISSN="0132-8069" ID-Zenodo-Dep="13179128" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722595318349" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2023" docId="03B3CC7F7868B637FA38A645FD9BFB0C" docLanguage="en" docName="RussEntomolJ.32.3.330-355.pdf" docOrigin="Russian Entomological Journal 32 (3)" docSource="http://dx.doi.org/10.15298/rusentj.32.3.09" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Empis (Anacrostichus) verralli Collin 1927" docType="treatment" docVersion="3" lastPageNumber="350" masterDocId="FF8AB407787CB623FB37A704FFE5FF97" masterDocTitle="Review of the Nearctic species of the subgenus Anacrostichus Bezzi, 1909 (Diptera: Empididae)" masterLastPageNumber="355" masterPageNumber="330" pageNumber="350" updateTime="1722636720815" updateUser="ExternalLinkService">
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<mods:title id="85D54B763D1BA7142967768F26341D38">Review of the Nearctic species of the subgenus Anacrostichus Bezzi, 1909 (Diptera: Empididae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="6643A664BC78FB64028CCFF11A82BE71">Shamshev, I. V.</mods:namePart>
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<mods:typeOfResource id="EF5BE4A09976066FEDF8692299CC218D">text</mods:typeOfResource>
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<mods:title id="455814C8B77522166D2B44930CA1CB22">Russian Entomological Journal</mods:title>
</mods:titleInfo>
<mods:part id="6395631F048127B5A40397407706F4C7">
<mods:date id="238A4867AB16DA399E3E3562786269B4">2023</mods:date>
<mods:detail id="54F90848C6EB67F5BBADBC3819CB3DC2" type="pubDate">
<mods:number id="9B4A75C71CB35C0CA1BB8A890200EB79">2023-09-30</mods:number>
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<mods:number id="96825CBA6D34A9C6D2519306837D0B5B">32</mods:number>
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<mods:number id="A2A11EB1523E68436E3AACD022631AB2">3</mods:number>
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<mods:url id="F48526C4E276391273745B9B14F9B198">http://dx.doi.org/10.15298/rusentj.32.3.09</mods:url>
</mods:location>
<mods:classification id="E61BD344180B7EF15A7267B05E31D35E">journal article</mods:classification>
<mods:identifier id="8B4261E709DFD6C582D15B8A78A9615B" type="DOI">10.15298/rusentj.32.3.09</mods:identifier>
<mods:identifier id="44B2B742F0C1835B66D3597D8D0C0517" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="F9E568FD6B8B6E45387FE4DB03DF3206" type="Zenodo-Dep">13179128</mods:identifier>
</mods:mods>
<treatment id="03B3CC7F7868B637FA38A645FD9BFB0C" ID-DOI="http://doi.org/10.5281/zenodo.13179304" ID-Zenodo-Dep="13179304" LSID="urn:lsid:plazi:treatment:03B3CC7F7868B637FA38A645FD9BFB0C" httpUri="http://treatment.plazi.org/id/03B3CC7F7868B637FA38A645FD9BFB0C" lastPageNumber="350" pageId="20" pageNumber="350">
<subSubSection id="C3002EE27868B637FA38A645FDC6FEE4" pageId="20" pageNumber="350" type="nomenclature">
<paragraph id="8BA57D697868B637FA38A645FD01FECF" blockId="20.[271,740,321,371]" box="[271,740,321,344]" pageId="20" pageNumber="350">
<heading id="D0EDCA057868B637FA38A645FD01FECF" box="[271,740,321,344]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="350" reason="8">
<taxonomicName id="4C1A06EA7868B637FA38A645FD01FECF" ID-CoL="8BZ2X" authority="Collin, 1927" authorityName="Collin" authorityYear="1927" box="[271,740,321,344]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli" subGenus="Anacrostichus">
<emphasis id="B96EA17B7868B637FA38A645FEB7FECF" box="[271,338,321,344]" italics="true" pageId="20" pageNumber="350">Empis</emphasis>
(
<emphasis id="B96EA17B7868B637FA56A645FE1CFECF" box="[353,505,321,344]" italics="true" pageId="20" pageNumber="350">Anacrostichus</emphasis>
)
<emphasis id="B96EA17B7868B637F93FA645FDBDFECF" box="[520,600,321,344]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
Collin, 1927
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA57D697868B637FAE6A658FDC6FEE4" blockId="20.[271,740,321,371]" box="[465,547,348,371]" pageId="20" pageNumber="350">Fig. 36.</paragraph>
</subSubSection>
<subSubSection id="C3002EE27868B637FBC2A69CFE52FE6F" pageId="20" pageNumber="350" type="reference_group">
<paragraph id="8BA57D697868B637FBC2A69CFD0FFE55" blockId="20.[207,806,408,1179]" pageId="20" pageNumber="350">
<taxonomicName id="4C1A06EA7868B637FBC2A69CFD6DFE3F" ID-CoL="8BZ2X" authority="Collin, 1927: 21" authorityName="Collin" authorityPageNumber="21" authorityYear="1927" box="[245,648,408,424]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli" subGenus="Anacrostichus">
<emphasis id="B96EA17B7868B637FBC2A69CFEC0FE3F" box="[245,293,408,424]" italics="true" pageId="20" pageNumber="350">Empis</emphasis>
(
<emphasis id="B96EA17B7868B637FA00A69CFE43FE3F" box="[311,422,408,424]" italics="true" pageId="20" pageNumber="350">Anacrostichus</emphasis>
)
<emphasis id="B96EA17B7868B637FA80A69CFE17FE3F" box="[439,498,408,424]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
<materialsCitation id="3B7277347868B637FACAA69CFD6DFE3F" box="[509,648,408,424]" country="United Kingdom" location="UK" pageId="20" pageNumber="350" specimenCount="2" specimenCount-female="1" specimenCount-male="1">Collin, 1927: 21</materialsCitation>
</taxonomicName>
(
<specimenCount id="9D1CB6E07868B637F9ADA69CFD4DFE3F" box="[666,680,408,424]" count="1" pageId="20" pageNumber="350" type="male">♂</specimenCount>
and
<specimenCount id="9D1CB6E07868B637F9EDA69CFD0DFE3F" box="[730,744,408,424]" count="1" pageId="20" pageNumber="350" type="female">♀</specimenCount>
).
<typeStatus id="54A1C3CB7868B637F9C9A69CFCC1FE3F" box="[766,804,408,424]" pageId="20" pageNumber="350">Type</typeStatus>
localities:
<collectingCountry id="F30D3DF97868B637FA16A6B6FEDBFE55" box="[289,318,434,450]" name="United Kingdom" pageId="20" pageNumber="350">UK</collectingCountry>
, England: Snowdon (Carnarvon), Yorkshire;
<collectingCountry id="F30D3DF97868B637F997A6B6FD02FE55" box="[672,743,434,450]" name="United Kingdom" pageId="20" pageNumber="350">Scotland</collectingCountry>
.
</paragraph>
<paragraph id="8BA57D697868B637FBC2A6C9FE52FE6F" blockId="20.[207,806,408,1179]" pageId="20" pageNumber="350">
<taxonomicName id="4C1A06EA7868B637FBC2A6C9FDB7FE4A" ID-CoL="92RJZ" authority="Frey, 1953: 35" authorityName="Frey" authorityPageNumber="35" authorityYear="1953" box="[245,594,461,477]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="minor" subGenus="Anacrostichus">
<emphasis id="B96EA17B7868B637FBC2A6C9FEC1FE4A" box="[245,292,461,477]" italics="true" pageId="20" pageNumber="350">Empis</emphasis>
(
<emphasis id="B96EA17B7868B637FA05A6C9FE79FE4A" box="[306,412,461,477]" italics="true" pageId="20" pageNumber="350">Anacrostichus</emphasis>
)
<emphasis id="B96EA17B7868B637FA9DA6C9FE32FE4A" box="[426,471,461,477]" italics="true" pageId="20" pageNumber="350">minor</emphasis>
Frey, 1953: 35
</taxonomicName>
(
<specimenCount id="9D1CB6E07868B637F957A6C9FD8BFE4A" box="[608,622,461,477]" count="1" pageId="20" pageNumber="350" type="male">♂</specimenCount>
and
<specimenCount id="9D1CB6E07868B637F9AEA6C9FD42FE4A" box="[665,679,461,477]" count="1" pageId="20" pageNumber="350" type="female">♀</specimenCount>
).
<materialsCitation id="3B7277347868B637F98DA6C9FE56FE6F" country="Russia" location="Ozernaya" pageId="20" pageNumber="350" specimenCount="1" stateProvince="Kamchatka">
<typeStatus id="54A1C3CB7868B637F98DA6C9FD3AFE4A" box="[698,735,461,477]" pageId="20" pageNumber="350">Type</typeStatus>
locality:
<collectingCountry id="F30D3DF97868B637FBF8A6ECFEE6FE6F" box="[207,259,488,504]" name="Russia" pageId="20" pageNumber="350">Russia</collectingCountry>
,
<collectingRegion id="49DEB38B7868B637FA3DA6ECFE87FE6F" box="[266,354,488,504]" country="Russia" name="Kamchatka" pageId="20" pageNumber="350">Kamchatka</collectingRegion>
:
<location id="8EC52BB27868B637FA5EA6ECFE56FE6F" LSID="urn:lsid:plazi:treatment:03B3CC7F7868B637FA38A645FD9BFB0C:8EC52BB27868B637FA5EA6ECFE56FE6F" box="[361,435,488,504]" country="Russia" name="Ozernaya" pageId="20" pageNumber="350" stateProvince="Kamchatka">Ozernaya</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3002EE27868B637FBC2A504FE11FCB5" pageId="20" pageNumber="350" type="materials_examined">
<paragraph id="8BA57D697868B637FBC2A504FE11FCB5" blockId="20.[207,806,408,1179]" pageId="20" pageNumber="350">
<materialsCitation id="3B7277347868B637FBC2A504FDF6FDA7" collectingDate="1961-06-29" collectionCode="MATERIAL, EXAMINED" collectorName="Unalakleet" country="United States of America" latitude="63.866665" location="Alaska" longLatPrecision="1011" longitude="-160.78334" pageId="20" pageNumber="350" specimenCount="1" stateProvince="Alaska">
<collectionCode id="ED0BE5AC7868B637FBC2A504FDE7FD82" box="[245,514,512,533]" pageId="20" pageNumber="350">MATERIAL EXAMINED</collectionCode>
.
<emphasis id="B96EA17B7868B637F921A504FD4EFD82" bold="true" box="[534,683,512,533]" pageId="20" pageNumber="350">
<collectingCountry id="F30D3DF97868B637F921A504FDADFD82" box="[534,584,512,533]" name="United States of America" pageId="20" pageNumber="350">USA</collectingCountry>
.
<collectingRegion id="49DEB38B7868B637F96BA504FD43FD82" box="[604,678,512,533]" country="United States of America" name="Alaska" pageId="20" pageNumber="350">Alaska</collectingRegion>
:
</emphasis>
<collectorName id="26EF18BF7868B637F98BA504FCC1FD82" box="[700,804,512,533]" pageId="20" pageNumber="350">Unalakleet</collectorName>
[
<geoCoordinate id="EE2E1BAE7868B637FBE2A51FFEC2FDA7" box="[213,295,539,560]" degrees="63" direction="north" minutes="52" orientation="latitude" pageId="20" pageNumber="350" precision="925" value="63.866665">63°52′N</geoCoordinate>
<geoCoordinate id="EE2E1BAE7868B637FA06A51FFE72FDA7" box="[305,407,539,560]" degrees="160" direction="west" minutes="47" orientation="longitude" pageId="20" pageNumber="350" precision="925" value="-160.78334">160°47′W</geoCoordinate>
],
<date id="FFA45BA97868B637FA9DA51FFDF6FDA7" box="[426,531,539,560]" pageId="20" pageNumber="350" value="1961-06-29">
<collectingDate id="EFE0A2417868B637FA9DA51FFDF6FDA7" box="[426,531,539,560]" pageId="20" pageNumber="350" value="1961-06-29">29.vi.1961</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B7277347868B637F916A51FFDA8FDDC" collectingDate="1961-06-23" collectionCode="MATERIAL, EXAMINED" collectorName="B. S. Heming" country="United States of America" latitude="63.866665" location="Alaska" longLatPrecision="1011" longitude="-160.78334" pageId="20" pageNumber="350" specimenCount="3" specimenCount-male="3" stateProvince="Alaska">
<collectorName id="26EF18BF7868B637F916A51FFD44FDA7" box="[545,673,539,560]" pageId="20" pageNumber="350">B.S. Heming</collectorName>
(
<specimenCount id="9D1CB6E07868B637F983A51FFD3FFDA7" box="[692,730,539,560]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
); same locality and collector,
<date id="FFA45BA97868B637FAA9A532FDE0FDDC" box="[414,517,566,587]" pageId="20" pageNumber="350" value="1961-06-23">
<collectingDate id="EFE0A2417868B637FAA9A532FDE0FDDC" box="[414,517,566,587]" pageId="20" pageNumber="350" value="1961-06-23">23.vi.1961</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637F927A532FDA1FDDC" box="[528,580,566,587]" count="2" pageId="20" pageNumber="350" type="male">2 ♂♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="3B7277347868B637F961A532FEBEFDF1" collectingDate="1961-06-20" collectionCode="MATERIAL, EXAMINED" collectorName="B. S. Heming" country="United States of America" latitude="63.866665" location="Alaska" longLatPrecision="1011" longitude="-160.78334" pageId="20" pageNumber="350" specimenCount="12" specimenCount-female="5" specimenCount-male="7" stateProvince="Alaska">
same data:
<date id="FFA45BA97868B637F989A532FCC1FDDC" box="[702,804,566,587]" pageId="20" pageNumber="350" value="1961-06-20">
<collectingDate id="EFE0A2417868B637F989A532FCC1FDDC" box="[702,804,566,587]" pageId="20" pageNumber="350" value="1961-06-20">20.vi.1961</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637FBE0A555FEEBFDF1" box="[215,270,593,614]" count="7" pageId="20" pageNumber="350" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="9D1CB6E07868B637FA2CA555FEB7FDF1" box="[283,338,593,614]" count="5" pageId="20" pageNumber="350" type="female">5 ♀♀</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="3B7277347868B637FA50A555FDEFFDF1" box="[359,522,593,614]" collectingDate="1961-06-29" collectionCode="MATERIAL, EXAMINED" collectorName="B. S. Heming" country="United States of America" isTailEnumeration="true" latitude="63.866665" location="Alaska" longLatPrecision="1011" longitude="-160.78334" pageId="20" pageNumber="350" specimenCount="1" specimenCount-female="1" stateProvince="Alaska">
<date id="FFA45BA97868B637FA50A555FE28FDF1" box="[359,461,593,614]" pageId="20" pageNumber="350" value="1961-06-29">
<collectingDate id="EFE0A2417868B637FA50A555FE28FDF1" box="[359,461,593,614]" pageId="20" pageNumber="350" value="1961-06-29">29.vi.1961</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637FAE9A555FDE4FDF1" box="[478,513,593,614]" count="1" pageId="20" pageNumber="350" type="female">1 ♀</specimenCount>
)
</materialsCitation>
,
<date id="FFA45BA97868B637F921A555FD9DFDF1" box="[534,632,593,614]" pageId="20" pageNumber="350" value="1961-07-05">5.vii.1961</date>
(
<specimenCount id="9D1CB6E07868B637F9BFA555FD4EFDF1" box="[648,683,593,614]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D1CB6E07868B637F98FA555FD38FDF1" box="[696,733,593,614]" count="1" pageId="20" pageNumber="350" type="female">1 ♀</specimenCount>
);
<materialsCitation id="3B7277347868B637F9C4A555FE74FD17" collectingDate="1961-06-22" collectionCode="MATERIAL, EXAMINED" collectorName="B. S. Heming" country="United States of America" isTailEnumeration="true" latitude="63.866665" location="Alaska" longLatPrecision="1011" longitude="-160.78334" pageId="20" pageNumber="350" specimenCount="1" stateProvince="Alaska">
same locality,
<date id="FFA45BA97868B637FA1FA56FFE74FD17" box="[296,401,619,640]" pageId="20" pageNumber="350" value="1961-06-22">
<collectingDate id="EFE0A2417868B637FA1FA56FFE74FD17" box="[296,401,619,640]" pageId="20" pageNumber="350" value="1961-06-22">22.vi.1961</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B7277347868B637FA97A56FFEE3FD0C" collectingDate="1961-06-24" collectionCode="R" collectorName="Madge" country="United States of America" pageId="20" pageNumber="350" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Alaska">
<collectionCode id="ED0BE5AC7868B637FA97A56FFE56FD17" box="[416,435,619,640]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="350">R</collectionCode>
.
<collectorName id="26EF18BF7868B637FAF4A56FFDE3FD17" box="[451,518,619,640]" pageId="20" pageNumber="350">Madge</collectorName>
(
<specimenCount id="9D1CB6E07868B637F92EA56FFDB0FD17" box="[537,597,619,640]" count="3" pageId="20" pageNumber="350" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D1CB6E07868B637F950A56FFD46FD17" box="[615,675,619,640]" count="5" pageId="20" pageNumber="350" type="female">5 ♀♀</specimenCount>
),
<date id="FFA45BA97868B637F989A56FFCC1FD17" box="[702,804,619,640]" pageId="20" pageNumber="350" value="1961-06-24">
<collectingDate id="EFE0A2417868B637F989A56FFCC1FD17" box="[702,804,619,640]" pageId="20" pageNumber="350" value="1961-06-24">24.vi.1961</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637FBEEA582FF18FD0C" box="[217,253,646,667]" count="1" pageId="20" pageNumber="350" type="female">1 ♀</specimenCount>
)
</materialsCitation>
; Nome [
<geoCoordinate id="EE2E1BAE7868B637FA6AA582FE4AFD0C" box="[349,431,646,667]" degrees="64" direction="north" minutes="30" orientation="latitude" pageId="20" pageNumber="350" precision="925" value="64.5">64°30′N</geoCoordinate>
<geoCoordinate id="EE2E1BAE7868B637FA80A582FDF8FD0C" box="[439,541,646,667]" degrees="165" direction="west" minutes="24" orientation="longitude" pageId="20" pageNumber="350" precision="925" value="-165.4">165°24′W</geoCoordinate>
],
<date id="FFA45BA97868B637F919A582FD74FD0C" box="[558,657,646,667]" pageId="20" pageNumber="350" value="1951-07-02">2.vii.1951</date>
,
<materialsCitation id="3B7277347868B637F9A9A582FD51FD21" collectingDate="1951-06-20" collectionCode="R" collectorName="D. P. Whillans" country="United States of America" pageId="20" pageNumber="350" specimenCount="2" specimenCount-male="2" stateProvince="Alaska">
<collectorName id="26EF18BF7868B637F9A9A582FCC0FD0C" box="[670,805,646,667]" pageId="20" pageNumber="350">D.P. Whillans</collectorName>
(
<specimenCount id="9D1CB6E07868B637FBEEA5A5FF1FFD21" box="[217,250,673,694]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
); same locality and collector,
<date id="FFA45BA97868B637F922A5A5FD99FD21" box="[533,636,673,694]" pageId="20" pageNumber="350" value="1951-06-20">
<collectingDate id="EFE0A2417868B637F922A5A5FD99FD21" box="[533,636,673,694]" pageId="20" pageNumber="350" value="1951-06-20">20.vi.1951</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637F9BDA5A5FD4EFD21" box="[650,683,673,694]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
)
</materialsCitation>
,
<materialsCitation id="3B7277347868B637F989A5A5FEEFFD46" collectingDate="1951-06-15" collectionCode="R" collectorName="D. P. Whillans" country="United States of America" isTailEnumeration="true" pageId="20" pageNumber="350" specimenCount="1" specimenCount-male="1" stateProvince="Alaska">
<date id="FFA45BA97868B637F989A5A5FCC0FD21" box="[702,805,673,694]" pageId="20" pageNumber="350" value="1951-06-15">
<collectingDate id="EFE0A2417868B637F989A5A5FCC0FD21" box="[702,805,673,694]" pageId="20" pageNumber="350" value="1951-06-15">15.vi.1951</collectingDate>
</date>
(
<specimenCount id="9D1CB6E07868B637FBEEA5B8FEE4FD46" box="[217,257,700,721]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
)
</materialsCitation>
,
<date id="FFA45BA97868B637FA2CA5B8FE90FD46" box="[283,373,700,721]" pageId="20" pageNumber="350" value="1951-06-06">6.vi.1951</date>
(
<specimenCount id="9D1CB6E07868B637FABCA5B8FE56FD46" box="[395,435,700,721]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
),
<date id="FFA45BA97868B637FAFAA5B8FDD6FD46" box="[461,563,700,721]" pageId="20" pageNumber="350" value="1951-06-18">18.vi.1951</date>
(
<specimenCount id="9D1CB6E07868B637F97FA5B8FD61FD46" box="[584,644,700,721]" count="3" pageId="20" pageNumber="350" type="male">3 ♂♂</specimenCount>
),
<date id="FFA45BA97868B637F9A9A5B8FCE1FD46" box="[670,772,700,721]" pageId="20" pageNumber="350" value="1951-06-19">19.vi.1951</date>
(
<specimenCount id="9D1CB6E07868B637F820A5B8FF16FD7B" count="4" pageId="20" pageNumber="350" type="male">4 ♂♂</specimenCount>
),
<date id="FFA45BA97868B637FA3FA5D3FE8BFD7B" box="[264,366,727,748]" pageId="20" pageNumber="350" value="1951-06-12">12.vi.1951</date>
(
<specimenCount id="9D1CB6E07868B637FA4BA5D3FE5AFD7B" box="[380,447,727,748]" count="18" pageId="20" pageNumber="350" type="male">18 ♂♂</specimenCount>
,
<specimenCount id="9D1CB6E07868B637FAFBA5D3FE15FD7B" box="[460,496,727,748]" count="1" pageId="20" pageNumber="350" type="female">1 ♀</specimenCount>
); Naknek [
<geoCoordinate id="EE2E1BAE7868B637F96BA5D3FD4BFD7B" box="[604,686,727,748]" degrees="58" direction="north" minutes="43" orientation="latitude" pageId="20" pageNumber="350" precision="925" value="58.716667">58°43′N</geoCoordinate>
<geoCoordinate id="EE2E1BAE7868B637F982A5D3FCFEFD7B" box="[693,795,727,748]" degrees="157" direction="west" minutes="00" orientation="longitude" pageId="20" pageNumber="350" precision="925" value="-157.0">157°00′W</geoCoordinate>
], on tundra,
<date id="FFA45BA97868B637FA73A5F6FE51FC90" box="[324,436,754,775]" pageId="20" pageNumber="350" value="1952-07-18">18.vii.1952</date>
,
<materialsCitation id="3B7277347868B637FAF3A5F6FED7FCB5" collectingDate="1952-07-03" collectionCode="R" collectorName="J. B. Hartley" country="United States of America" isTailEnumeration="true" pageId="20" pageNumber="350" specimenCount="1" specimenCount-male="1" stateProvince="Alaska">
<collectorName id="26EF18BF7868B637FAF3A5F6FDDAFC90" box="[452,575,754,775]" pageId="20" pageNumber="350">J.B. Hartley</collectorName>
(
<specimenCount id="9D1CB6E07868B637F962A5F6FD9BFC90" box="[597,638,754,775]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
); same locality,
<date id="FFA45BA97868B637FBF8A409FED7FCB5" box="[207,306,781,802]" pageId="20" pageNumber="350" value="1952-07-03">
<collectingDate id="EFE0A2417868B637FBF8A409FED7FCB5" box="[207,306,781,802]" pageId="20" pageNumber="350" value="1952-07-03">3.vii.1952</collectingDate>
</date>
</materialsCitation>
, W.
<materialsCitation id="3B7277347868B637FA6FA409FE15FCB5" box="[344,496,781,802]" collectionCode="R" collectorName="Mason" country="United States of America" pageId="20" pageNumber="350" specimenCount="1" specimenCount-male="1" stateProvince="Alaska">
<collectionCode id="ED0BE5AC7868B637FA6FA409FE8CFCB5" box="[344,361,781,802]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="350">R</collectionCode>
.
<collectorName id="26EF18BF7868B637FA44A409FE51FCB5" box="[371,436,781,802]" pageId="20" pageNumber="350">Mason</collectorName>
(
<specimenCount id="9D1CB6E07868B637FAF4A409FE03FCB5" box="[451,486,781,802]" count="1" pageId="20" pageNumber="350" type="male">1 ♂</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3002EE27868B637FBC2A42CFD4BFB83" pageId="20" pageNumber="350" type="discussion">
<paragraph id="8BA57D697868B637FBC2A42CFD4BFB83" blockId="20.[207,806,408,1179]" pageId="20" pageNumber="350">
REMARKS. It seems that this species was not described by the North American authors. Main differences of
<taxonomicName id="4C1A06EA7868B637F9F5A447FCC1FCCF" authorityName="Collin" authorityYear="1927" box="[706,804,835,856]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli">
<emphasis id="B96EA17B7868B637F9F5A447FD34FCCF" box="[706,721,835,856]" italics="true" pageId="20" pageNumber="350">E</emphasis>
.
<emphasis id="B96EA17B7868B637F9EBA447FCC1FCCF" box="[732,804,835,856]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
</taxonomicName>
from other Nearctic species of
<taxonomicName id="4C1A06EA7868B637FAC9A45AFD63FCE4" authorityName="Bezzi" authorityYear="1909" box="[510,646,862,883]" pageId="20" pageNumber="354" rank="subGenus" subGenus="Anacrostichus">
<emphasis id="B96EA17B7868B637FAC9A45AFD63FCE4" box="[510,646,862,883]" italics="true" pageId="20" pageNumber="350">Anacrostichus</emphasis>
</taxonomicName>
are indicated in the key.
<taxonomicName id="4C1A06EA7868B637FA15A47DFE4BFC19" authorityName="Collin" authorityYear="1927" box="[290,430,889,910]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli">
<emphasis id="B96EA17B7868B637FA15A47DFE4BFC19" box="[290,430,889,910]" italics="true" pageId="20" pageNumber="350">Empis verralli</emphasis>
</taxonomicName>
was redescribed by Collin [1961] and Chvála [1994]. The wing venation of
<taxonomicName id="4C1A06EA7868B637F901A490FD7DFC3E" authorityName="Collin" authorityYear="1927" box="[566,664,916,937]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli">
<emphasis id="B96EA17B7868B637F901A490FDA0FC3E" box="[566,581,916,937]" italics="true" pageId="20" pageNumber="350">E</emphasis>
.
<emphasis id="B96EA17B7868B637F967A490FD7DFC3E" box="[592,664,916,937]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
</taxonomicName>
is unstable (as both these authors have noted), including different degree of a reduction of the radial fork (R
<subScript id="179E7F2C7868B637FAC3A4DCFDEFFC73" attach="left" box="[500,522,984,996]" fontSize="5" pageId="20" pageNumber="350">4+5</subScript>
) and some other veins, extra crossveins, etc. In addition, the frons of the female sometimes is entirely pruinose (usually shiny on the middle).
</paragraph>
</subSubSection>
<subSubSection id="C3002EE27868B637FBC2A31EFD9BFB0C" pageId="20" pageNumber="350" type="distribution">
<paragraph id="8BA57D697868B637FBC2A31EFD9BFB0C" blockId="20.[207,806,408,1179]" pageId="20" pageNumber="350">
DISTRIBUTION. Holarctic. In Eurasia,
<taxonomicName id="4C1A06EA7868B637F9ABA31EFCE3FBB8" authorityName="Collin" authorityYear="1927" box="[668,774,1050,1071]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli">
<emphasis id="B96EA17B7868B637F9ABA31EFD4EFBB8" box="[668,683,1050,1071]" italics="true" pageId="20" pageNumber="350">E</emphasis>
.
<emphasis id="B96EA17B7868B637F989A31EFCE3FBB8" box="[702,774,1050,1071]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
</taxonomicName>
is known from British Isles, extreme north of the European part of
<collectingCountry id="F30D3DF97868B637FBDDA354FEC9FBF2" box="[234,300,1104,1125]" name="Russia" pageId="20" pageNumber="350">Russia</collectingCountry>
, Siberia (including subarctic areas) and the Russian Far East [Shamshev, 2016]. In North America,
<taxonomicName id="4C1A06EA7868B637F991A36FFCEEFB17" authorityName="Collin" authorityYear="1927" box="[678,779,1131,1152]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="350" phylum="Arthropoda" rank="species" species="verralli">
<emphasis id="B96EA17B7868B637F991A36FFD50FB17" box="[678,693,1131,1152]" italics="true" pageId="20" pageNumber="350">E</emphasis>
.
<emphasis id="B96EA17B7868B637F9F4A36FFCEEFB17" box="[707,779,1131,1152]" italics="true" pageId="20" pageNumber="350">verralli</emphasis>
</taxonomicName>
is known only from the western part of Alaska.
</paragraph>
</subSubSection>
</treatment>
</document>
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