284 lines
33 KiB
XML
284 lines
33 KiB
XML
<document id="81E08448A63229B4260F3ADEE71D44E3" ID-DOI="10.11646/zootaxa.3670.1.5" ID-GBIF-Dataset="09317f41-bc49-448b-a019-0b78e079ba4a" ID-ISSN="1175-5326" ID-Zenodo-Dep="283933" ID-ZooBank="C6F33726-B9EC-4000-8957-B24158E98AC2" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633519470" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto & Novelo-Gutiérrez, Rodolfo" docDate="2013" docId="03B13135FFFBFFA7FF50FC79FC28FE42" docLanguage="en" docName="zt03670p070.pdf" docOrigin="Zootaxa 3670 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psephenops shepardi Arce-Pérez & Novelo-Gutiérrez, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="69" masterDocId="FF88494DFFF8FFA1FFC7FFD6FFCEFFB6" masterDocTitle="Two new species of Psephenops Grouvelle from Belize (Coleoptera: Byrrhoidea: Psephenidae), with a key to the known species from Mexico and Central America" masterLastPageNumber="70" masterPageNumber="63" pageNumber="66" updateTime="1698334777422" updateUser="plazi">
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<mods:mods id="D8AFAA365A0CAC33943CDA5E71B79AC3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="07359DA4FF5128905784032BB2E31681">
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<mods:title id="A703E50E7C81277668C6B508627CCA35">Two new species of Psephenops Grouvelle from Belize (Coleoptera: Byrrhoidea: Psephenidae), with a key to the known species from Mexico and Central America</mods:title>
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</mods:titleInfo>
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<mods:name id="36B3B6291A058BBE1A761721E2FCA2AF" type="personal">
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<mods:roleTerm id="6B4B95E1CFF1D6F929A2F7C0E9A61A36">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4C362AD855FC0979A9515F5458C07636">Arce-Pérez, Roberto</mods:namePart>
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</mods:name>
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<mods:name id="94CE75EC3D20E552A79B297721D20DAE" type="personal">
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<mods:role id="E5373756ADB17AD0E848EB60D8C0B471">
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<mods:roleTerm id="1FBB0F47FCC4CF13C0D95CE7D0F2B482">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="41F4B71BE876E09BA1CED96B53A13BF6">Novelo-Gutiérrez, Rodolfo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B2EA2CDC61F64E852CD98B286FF63D16">text</mods:typeOfResource>
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<mods:relatedItem id="918B02679133111B7F12F573A5713779" type="host">
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<mods:titleInfo id="DFFE19D0915E249D7B52EE91707EB719">
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<mods:title id="A1C5F8C660A581596E34BE19302834B7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="23B6851AA0D83E895A81010AF34268EF">
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<mods:date id="AE2B6CAA11D8478399F129187A8F11DD">2013</mods:date>
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<mods:detail id="D30AB35722079D264385E8D724ACA147" type="volume">
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<mods:number id="6271E1C18C14B7B2F5F2A2B487D04789">3670</mods:number>
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</mods:detail>
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<mods:detail id="D22717ECBCDF3F5339D4FCF534A455D5" type="issue">
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<mods:number id="8D35D718A745FC18ACA611327CBE0077">1</mods:number>
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</mods:detail>
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<mods:extent id="322EAE63BF22902E8ABC49EE1E93202C" unit="page">
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<mods:start id="401315FD0FCD4985FBAFE55467D4AFCC">63</mods:start>
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<mods:end id="12DBAD11C5D41A35BA014038B69C69BF">70</mods:end>
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</mods:extent>
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<mods:classification id="5A0BBEA30ED3915E4BE880D940E3170D">journal article</mods:classification>
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<mods:identifier id="64ED69FEE59B80470887F517B106D804" type="DOI">10.11646/zootaxa.3670.1.5</mods:identifier>
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<mods:identifier id="B93B0B8C3C02D1F291FD7C027577ECB1" type="GBIF-Dataset">09317f41-bc49-448b-a019-0b78e079ba4a</mods:identifier>
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<mods:identifier id="6A6F8B12F71DFD8942189F0498D449BC" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="90EF04BE5DBA662F4496931496F0945B" type="Zenodo-Dep">283933</mods:identifier>
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<mods:identifier id="11D3DA85EC4CED36BF373694EAA0F662" type="ZooBank">C6F33726-B9EC-4000-8957-B24158E98AC2</mods:identifier>
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</mods:mods>
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<treatment id="03B13135FFFBFFA7FF50FC79FC28FE42" ID-DOI="http://doi.org/10.5281/zenodo.6148889" ID-GBIF-Taxon="127664369" ID-Zenodo-Dep="6148889" LSID="urn:lsid:plazi:treatment:03B13135FFFBFFA7FF50FC79FC28FE42" httpUri="http://treatment.plazi.org/id/03B13135FFFBFFA7FF50FC79FC28FE42" lastPageId="6" lastPageNumber="69" pageId="3" pageNumber="66">
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<subSubSection id="C302D3A8FFFBFFA2FF50FC79FEF5FC5C" pageId="3" pageNumber="66" type="nomenclature">
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<paragraph id="8BA78023FFFBFFA2FF50FC79FD43FC7F" blockId="3.[151,653,943,1002]" box="[151,653,943,969]" pageId="3" pageNumber="66">
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<heading id="D0EF374FFFFBFFA2FF50FC79FD43FC7F" bold="true" box="[151,653,943,969]" fontSize="11" level="1" pageId="3" pageNumber="66" reason="1">
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<emphasis id="B96C5C31FFFBFFA2FF50FC79FD43FC7F" bold="true" box="[151,653,943,969]" pageId="3" pageNumber="66">
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<taxonomicName id="4C18FBA0FFFBFFA2FF50FC79FE5BFC7F" box="[151,405,943,969]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
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<emphasis id="B96C5C31FFFBFFA2FF50FC79FE5BFC7F" bold="true" box="[151,405,943,969]" italics="true" pageId="3" pageNumber="66">Psephenops shepardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25FE14AFFFBFFA2FE5AFC79FE34FC7F" box="[413,506,943,969]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
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Arce-Pérez
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA78023FFFBFFA2FF50FC04FEF5FC5C" blockId="3.[151,653,943,1002]" box="[151,315,978,1002]" pageId="3" pageNumber="66">
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(
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<figureCitation id="13239CA6FFFBFFA2FF58FC04FF24FC5C" box="[159,234,978,1002]" captionStart="FIGURE 1" captionStartId="1.[151,255,1314,1338]" captionTargetBox="[159,1436,265,1275]" captionTargetId="figure@1.[151,1436,264,1293]" captionTargetPageId="1" captionText="FIGURE 1. Hind tarsi of Psephenops, dorsolateral view: a) P. spiniparameri sp. nov.; b) P. shepardi sp. nov." httpUri="https://zenodo.org/record/283934/files/figure.png" pageId="3" pageNumber="66">Figs. 1</figureCitation>
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b, 5–7)
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</paragraph>
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</subSubSection>
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<subSubSection id="C302D3A8FFFBFFA5FF50FBCCFE9BFCA2" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66" type="description">
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<paragraph id="8BA78023FFFBFFA5FF50FBCCFA44FDE9" blockId="3.[151,1437,1050,2010]" lastBlockId="4.[151,1436,151,968]" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66">
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<emphasis id="B96C5C31FFFBFFA2FF50FBCCFE31FB85" bold="true" box="[151,511,1050,1075]" pageId="3" pageNumber="66">
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Description.
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<typeStatus id="54A33E81FFFBFFA2FEFCFBCDFE66FB85" box="[315,424,1051,1075]" pageId="3" pageNumber="66" type="holotype">Holotype</typeStatus>
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Male:
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</emphasis>
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Body oval, depressed; integument yellowish-red, with yellow iridescence, completely covered with iridescent, reddish-yellow, short setae. Head, antennal flagellum, scutellum and elytra dark yellowish-red; pronotum and legs light yellowish-red. Venter dark yellowish-red, vestiture short, fine and dense, gray-yellowish with reflections yellow.
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<emphasis id="B96C5C31FFFBFFA2FD70FB50FD3DFB28" box="[695,755,1158,1182]" italics="true" pageId="3" pageNumber="66">Head</emphasis>
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: short, transverse; clypeus subrectangular, declivous at an angle of less than 90 degrees from the plane of the frons, distal margin widely emarginate; fronto-clypeal surface with stout, dark reddish setae; vertex minutely punctate, with minute setae and very shallow longitudinal furrow in middle. Antennae moniliform, 11-segmented, scape yellow, larger than other segments, pedicel yellow, subglobose, half as long as scape; flagellum with 8 small segments, apical segment acuminate. Eyes lateral, rounded, very prominent, black, with a yellow post-ocular area. Labrum similar to clypeus but shorter. Maxillary palpi 4-segmented, covered with short setae, segment 1 smallest, apical segment largest, relative proportion of segments: 0.4, 0.9, 0.7, 1.0; basal segments 1–3 subconical, segment 4 oval, inflated from the base and gradually narrowing to tip (acuminate). Labial palpi very short, 3-segmented, segments 1–2 rounded and wide, segment 3 subconical, slightly shorter than preceding ones.
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<emphasis id="B96C5C31FFFBFFA2FD0AFA1DFCD0FA55" box="[717,798,1483,1507]" italics="true" pageId="3" pageNumber="66">Thorax</emphasis>
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: Pronotum light reddish-brown (dark reddish-brown in some
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<typeStatus id="54A33E81FFFBFFA2FF1AFA38FE83F9B0" box="[221,333,1518,1542]" pageId="3" pageNumber="66" type="paratype">paratypes</typeStatus>
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), subtrapezoidal; anterior margin arcuate, posterior margin bisinuate, almost as wide as elytral base; lateral margins straight along anterior half and expanded laterally on posterior half; posterolateral angles subacute; all margins narrowly rimmed. Pronotal disc with median longitudinal carina well developed on distal half, and wide, shallow, circular depression at each side of carina; in profile view, pronotal disc (except for posterolateral expansions) convex; pronotal surface without punctation, covered with short, grayish-yellow setae, shorter than those of head. Scutellum short, with apex rounded.
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<emphasis id="B96C5C31FFFBFFA2FC4AF975FC1DF90D" box="[909,979,1699,1723]" italics="true" pageId="3" pageNumber="66">Elytra</emphasis>
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: with short reddish-yellow setae; sides parallel in basal 0.75, then gradually converging rearward; base lightly depressed on internal side of humeri. Prosternum short and slightly carinate, with short, lanceolate, slightly carinate prosternal process, not reaching anterior half of mesoventrite. Mesoventrite short, with wide groove for reception of prosternal process. Metaventrite large and bulky.
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<emphasis id="B96C5C31FFFBFFA2FE28F8E2FDEAF8FD" box="[495,548,1844,1867]" italics="true" pageId="3" pageNumber="66">Legs</emphasis>
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: procoxae globose, mesocoxae conical, metacoxae transverse; femora robust; tibiae slender, metatibiae longer than femora, protibiae with distinct, apicolateral denticle. Tarsi 5-segmented (
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<figureCitation id="13239CA6FFFBFFA2FAA8F880FF6CF824" captionStart="FIGURE 1" captionStartId="1.[151,255,1314,1338]" captionTargetBox="[159,1436,265,1275]" captionTargetId="figure@1.[151,1436,264,1293]" captionTargetPageId="1" captionText="FIGURE 1. Hind tarsi of Psephenops, dorsolateral view: a) P. spiniparameri sp. nov.; b) P. shepardi sp. nov." httpUri="https://zenodo.org/record/283934/files/figure.png" pageId="3" pageNumber="66">Fig. 1</figureCitation>
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b), tarsomeres 1 and 2 lobed ventrally, with a thick sponge-like vestiture; ventral lobe of tarsomere 1 extending distally over basal 0.25 of tarsomere 2; tarsomere 2 the longest, twice as long as tarsomere 1, extending distally and completely covering tarsomeres 3, 4, and basal 0.70 of tarsomere 5; proportional length of tarsomeres: 0.50, 1.0, 0.2, 0.2, 0.57; a pair of long, curved, slender apical claws, with small basal tooth.
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<emphasis id="B96C5C31FFFCFFA5FBF8FF4EFB67FF06" box="[1087,1193,152,176]" italics="true" pageId="4" pageNumber="67">Abdomen</emphasis>
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: vestiture short, fine, dense, with yellow iridescence and intermingled dark reddish setae. Seven visible sterna (ventrites) in relaxed specimens; usually ventrite VI hidden under ventrite V; ventrite II the largest. Posterior margin of ventrite I deeply concave at middle, sinuate at sides; posterior margin of ventrite II as in ventrite I but concavity shallow medially; posterior margin of ventrites III and IV straight, ventrite V with wide, shallow concavity, ventrite VI very short, posterior margin widely concave. Pygidial plate oval, long, narrow and densely covered with a long, stiff, dark reddish setae.
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<emphasis id="B96C5C31FFFCFFA5FEFBFEA6FE68FE3E" box="[316,422,368,392]" italics="true" pageId="4" pageNumber="67">Genitalia</emphasis>
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: short, wide, subrectagular, trilobate (
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<figureCitation id="13239CA6FFFCFFA5FC98FEB9FC07FE31" box="[863,969,367,391]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Figs 5–7</figureCitation>
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). Parameres robust, slightly shorter than median lobe, in dorsal view (
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<figureCitation id="13239CA6FFFCFFA5FE25FE42FDE8FE1A" box="[482,550,404,428]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 5</figureCitation>
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) with internal margins fused at basal 0.66, apices broadly separate, U-shaped, external margin abruptly and strongly convergent with the internal margin at 0.80 of parameres length, ending in a sharp point, with lateroapical membrane (
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<figureCitation id="13239CA6FFFCFFA5FDB0FE0AFD73FE42" box="[631,701,476,500]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 5</figureCitation>
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); in ventral view with small tooth on internal margin near apex (
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<figureCitation id="13239CA6FFFCFFA5FF58FE29FF2BFDA1" box="[159,229,511,535]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 6</figureCitation>
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). Penis, in ventral view, subrectangular, wider and longer than parameres, external margins concave, apical margin strongly convex with apical membrane (
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<figureCitation id="13239CA6FFFCFFA5FD70FDF2FD32FD8A" box="[695,764,548,572]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 6</figureCitation>
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), bearing longitudinal, ventral, subtriangular sclerite which is 0.54 as long as penis; dorsal area of phallobase concave (
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<figureCitation id="13239CA6FFFCFFA5FCF6FD91FCBBFDE9" box="[817,885,583,607]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 5</figureCitation>
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); ventral portion bulky, subrectangular (
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<figureCitation id="13239CA6FFFCFFA5FAFFFD91FAB3FDE9" box="[1336,1405,583,607]" captionStart="FIGURES 5 – 7" captionStartId="4.[151,269,1967,1991]" captionTargetBox="[151,1423,1013,1944]" captionTargetId="figure@4.[151,1436,1002,1962]" captionTargetPageId="4" captionText="FIGURES 5 – 7. Male genitalia of Psephenops shepardi sp. nov.: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view." httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67">Fig. 6</figureCitation>
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).
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</paragraph>
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<paragraph id="8BA78023FFFCFFA5FF00FDBAFE9BFCA2" blockId="4.[151,1436,151,968]" pageId="4" pageNumber="67">
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||
<emphasis id="B96C5C31FFFCFFA5FF00FDBAFBAFFD32" bold="true" box="[199,1121,619,644]" pageId="4" pageNumber="67">
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Measurements (in mm) [in square brackets the
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<typeStatus id="54A33E81FFFCFFA5FCD0FDBAFC43FD32" box="[791,909,620,644]" pageId="4" pageNumber="67" type="paratype">paratypes</typeStatus>
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variation N=25]:
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</emphasis>
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Total length 2.65 [2.5-3.0, mean=2.65]; maxillary palp: 0.04, 0.09, 0.07, 0.10. Pronotum: posterior margin 0.98 [0.92-1.02, mean=0.97], lateral margin 0.44 [0.42-0.55, mean=0.47]; elytra: humeral width 1.16 [1.0-1.16, mean=1.10]; length of tarsomeres: 0.14, 0.28, 0.05, 0.05, 0.16. Genitalia: total length 0.46; parameres 0.28, penis 0.31; phallobase: length 0.20, width 0.16.
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</paragraph>
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</subSubSection>
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<subSubSection id="C302D3A8FFFCFFA7FF00FCF6FE22FF62" lastPageId="6" lastPageNumber="69" pageId="4" pageNumber="67" type="materials_examined">
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<paragraph id="8BA78023FFFCFFA5FF00FCF6FB3DFC71" blockId="4.[151,1436,151,968]" pageId="4" pageNumber="67">
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||
<emphasis id="B96C5C31FFFCFFA5FF00FCF6FEB7FC8E" bold="true" box="[199,377,799,824]" pageId="4" pageNumber="67">
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<typeStatus id="54A33E81FFFCFFA5FF00FCF6FECCFC8E" box="[199,258,800,824]" pageId="4" pageNumber="67">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54A33E81FFFCFFA5FE43FCC9FE22FC81" box="[388,492,799,823]" pageId="4" pageNumber="67" type="holotype">Holotype</typeStatus>
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labeled:
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<collectingCountry id="F30FC0B3FFFCFFA5FD9CFCC9FD69FC81" box="[603,679,799,823]" name="Belize" pageId="4" pageNumber="67">Belize</collectingCountry>
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: Cayo Dist,
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<quantity id="4CE02DC6FFFCFFA5FCF4FCC9FCB0FC81" box="[819,894,799,823]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="4" pageNumber="67" unit="mi" value="1.7">1.7 mi</quantity>
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E. O. Top, Camp,
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||
<date id="FFA6A6E3FFFCFFA5FBA1FCC9FB04FC81" box="[1126,1226,799,823]" pageId="4" pageNumber="67" value="1995-01-09">9-I-1995</date>
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||
, trib to Dr/Creek, WDS-A-1127 (EMEC),
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||
<typeStatus id="54A33E81FFFCFFA5FE6EFC93FDD9FCEB" box="[425,535,837,861]" pageId="4" pageNumber="67" type="paratype">paratypes</typeStatus>
|
||
, same data as
|
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<typeStatus id="54A33E81FFFCFFA5FD7FFC92FCD4FCEA" box="[696,794,836,860]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
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||
14 3 [9(IEXA); 5 3 (EMEC)]; Cayo Dist, Over the Top Camp,
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<date id="FFA6A6E3FFFCFFA5FF2EFCB1FE83FCC9" box="[233,333,871,895]" pageId="4" pageNumber="67" value="1995-01-09">9-I-1995</date>
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||
, St Margaret ‘s Cr, WDS-A-1126,1 3 (IEXA); Toledo Dist.,
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<quantity id="4CE02DC6FFFCFFA5FBC6FCB1FB90FCC9" box="[1025,1118,871,895]" metricMagnitude="4" metricUnit="m" metricValue="3.51" pageId="4" pageNumber="67" unit="km" value="35.1">35.1 km</quantity>
|
||
SE Belmopan, 3-Vlll-1993, unnamed stream, WDS-A-985, 4 3 [2 (IEXA); 2 (EMEC)]; Toledo Dist., Hellgate,
|
||
<date id="FFA6A6E3FFFCFFA5FB88FC5AFB0FFC12" box="[1103,1217,908,932]" pageId="4" pageNumber="67" value="1995-01-11">11-I-1995</date>
|
||
, WDS-A-1138, 13 (EMEC);
|
||
<collectingCountry id="F30FC0B3FFFCFFA5FEC1FC79FE83FC71" box="[262,333,943,967]" name="Belize" pageId="4" pageNumber="67">Belize</collectingCountry>
|
||
Dist.,
|
||
<quantity id="4CE02DC6FFFCFFA5FE5FFC79FE05FC71" box="[408,459,943,967]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="4" pageNumber="67" unit="mi" value="3.0">3 mi</quantity>
|
||
W
|
||
<collectingCountry id="F30FC0B3FFFCFFA5FE34FC79FDFDFC71" box="[499,563,943,967]" name="United Kingdom" pageId="4" pageNumber="67">Gales</collectingCountry>
|
||
Point,
|
||
<date id="FFA6A6E3FFFCFFA5FD44FC79FD3BFC71" box="[643,757,943,967]" pageId="4" pageNumber="67" value="1995-01-14">14-I-1995</date>
|
||
, Soldier Creek, WDS-A-
|
||
<date id="FFA6A6E3FFFCFFA5FBD5FC79FBB0FC7E" box="[1042,1150,943,968]" pageId="4" pageNumber="67" value="1143-04-03">1143, 4 3</date>
|
||
(EMEC)..
|
||
</paragraph>
|
||
<caption id="DF67D0ABFFFCFFA5FF50F879FD36F851" httpUri="https://zenodo.org/record/283936/files/figure.png" pageId="4" pageNumber="67" targetBox="[151,1423,1013,1944]" targetPageId="4">
|
||
<paragraph id="8BA78023FFFCFFA5FF50F879FD36F851" blockId="4.[151,1436,1967,2023]" pageId="4" pageNumber="67">
|
||
<emphasis id="B96C5C31FFFCFFA5FF50F879FE88F871" bold="true" box="[151,326,1967,1991]" pageId="4" pageNumber="67">FIGURES 5–7.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C18FBA0FFFCFFA5FDD1F866FD32F871" box="[534,764,1968,1991]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFCFFA5FDD1F866FD32F871" box="[534,764,1968,1991]" italics="true" pageId="4" pageNumber="67">Psephenops shepardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFCFFA5FCC1F879FC91F871" bold="true" box="[774,863,1967,1991]" pageId="4" pageNumber="67">
|
||
<taxonomicNameLabel id="A25FE14AFFFCFFA5FCC1F879FC91F871" box="[774,863,1967,1991]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: 5) phallobase, parameres and median lobe (penis), dorsal view; 6) same, ventral view; 7) same, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF67D0ABFFFDFFA4FF50F8A4FDF0F844" httpUri="https://zenodo.org/record/283937/files/figure.png" pageId="5" pageNumber="68" targetBox="[151,1432,199,1864]" targetPageId="5">
|
||
<paragraph id="8BA78023FFFDFFA4FF50F8A4FDF0F844" blockId="5.[151,1437,1906,2034]" pageId="5" pageNumber="68">
|
||
<emphasis id="B96C5C31FFFDFFA4FF50F8A4FE9EF83C" bold="true" box="[151,336,1906,1930]" pageId="5" pageNumber="68">FIGURES 8–13.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FDD2F8A4FD5AF83F" box="[533,660,1906,1929]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96C5C31FFFDFFA4FDD2F8A4FD5AF83F" box="[533,660,1906,1929]" italics="true" pageId="5" pageNumber="68">Psephenops</emphasis>
|
||
</taxonomicName>
|
||
spp. occurring in Mexico and Central America, dorsal view (scale = 0.1 mm): 8)
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FF30F843FE4AF81A" box="[247,388,1941,1964]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B96C5C31FFFDFFA4FF30F843FE4AF81A" box="[247,388,1941,1964]" italics="true" pageId="5" pageNumber="68">P. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
; 9)
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FE76F843FDC0F81A" box="[433,526,1941,1964]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="smithi">
|
||
<emphasis id="B96C5C31FFFDFFA4FE76F843FDC0F81A" box="[433,526,1941,1964]" italics="true" pageId="5" pageNumber="68">P. smithi</emphasis>
|
||
</taxonomicName>
|
||
; 10)
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FD8EF843FD3DF81A" box="[585,755,1941,1964]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="argentinensis">
|
||
<emphasis id="B96C5C31FFFDFFA4FD8EF843FD3DF81A" box="[585,755,1941,1964]" italics="true" pageId="5" pageNumber="68">P. argentinensis</emphasis>
|
||
</taxonomicName>
|
||
; 11)
|
||
<emphasis id="B96C5C31FFFDFFA4FCEAF843FC4AF81A" box="[813,900,1941,1964]" italics="true" pageId="5" pageNumber="68">P. lu pit a</emphasis>
|
||
; 12)
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FC04F843FBAEF81A" box="[963,1120,1941,1964]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="maculicollis">
|
||
<emphasis id="B96C5C31FFFDFFA4FC04F843FBAEF81A" box="[963,1120,1941,1964]" italics="true" pageId="5" pageNumber="68">P. maculicollis</emphasis>
|
||
</taxonomicName>
|
||
; 13)
|
||
<taxonomicName id="4C18FBA0FFFDFFA4FB5BF843FADCF81A" box="[1180,1298,1941,1964]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="68" phylum="Arthropoda" rank="species" species="prestone">
|
||
<emphasis id="B96C5C31FFFDFFA4FB5BF843FADCF81A" box="[1180,1298,1941,1964]" italics="true" pageId="5" pageNumber="68">P. prestone</emphasis>
|
||
</taxonomicName>
|
||
[Fig. 8 after Arce-Pérez y Novelo-Gutiérrez (2000); Figs 9–10 modified from Delève (1967); Fig. 11 after Arce-Pérez (2002); Figs. 12–13 modified from Spangler (1990)].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA78023FFFEFFA7FF00FF4EFE22FF62" blockId="6.[151,1436,151,500]" pageId="6" pageNumber="69">
|
||
<emphasis id="B96C5C31FFFEFFA7FF00FF4EFEA7FF06" bold="true" box="[199,361,151,176]" pageId="6" pageNumber="69">
|
||
<typeStatus id="54A33E81FFFEFFA7FF00FF4EFECEFF06" box="[199,256,152,176]" pageId="6" pageNumber="69">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F30FC0B3FFFEFFA7FEB1FF41FE0FFF19" box="[374,449,151,175]" name="Belize" pageId="6" pageNumber="69">Belize</collectingCountry>
|
||
; Cayo District,
|
||
<quantity id="4CE02DC6FFFEFFA7FD46FF41FD03FF19" box="[641,717,151,175]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="6" pageNumber="69" unit="mi" value="1.7">1.7 mi</quantity>
|
||
east of Over the Top Camp on the Hummingbird Highway, tributary to Dry Creek,
|
||
<specimenCount id="9D1E4BAAFFFEFFA7FE59FF6AFE22FF62" box="[414,492,188,212]" pageId="6" pageNumber="69" type="generic">
|
||
<quantity id="4CE02DC6FFFEFFA7FE59FF6AFE29FF62" box="[414,487,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="69" unit="m" value="115.0">115 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302D3A8FFFEFFA7FF00FF36FE1DFEAA" pageId="6" pageNumber="69" type="etymology">
|
||
<paragraph id="8BA78023FFFEFFA7FF00FF36FE1DFEAA" blockId="6.[151,1436,151,500]" pageId="6" pageNumber="69">
|
||
<emphasis id="B96C5C31FFFEFFA7FF00FF36FE80FF4E" bold="true" box="[199,334,224,248]" pageId="6" pageNumber="69">Etymology.</emphasis>
|
||
This species is respectfully dedicated to Dr. William D. Shepard for his great contribution to the study of aquatic
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FE96FED2FE1EFEAA" box="[337,464,260,284]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302D3A8FFFEFFA7FF00FEFEFC28FE42" pageId="6" pageNumber="69" type="discussion">
|
||
<paragraph id="8BA78023FFFEFFA7FF00FEFEFC28FE42" blockId="6.[151,1436,151,500]" pageId="6" pageNumber="69">
|
||
<emphasis id="B96C5C31FFFEFFA7FF00FEFEFEF5FEF6" bold="true" box="[199,315,296,320]" pageId="6" pageNumber="69">Remarks.</emphasis>
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FE85FEFEFDFEFEF6" box="[322,560,296,320]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFEFFA7FE85FEFEFDFEFEF6" box="[322,560,296,320]" italics="true" pageId="6" pageNumber="69">Psephenops shepardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFEFFA7FDF0FEFEFD5EFEF6" bold="true" box="[567,656,296,320]" pageId="6" pageNumber="69">
|
||
<taxonomicNameLabel id="A25FE14AFFFEFFA7FDF0FEFEFD5EFEF6" box="[567,656,296,320]" pageId="6" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated easily from other known
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FBB5FEFEFB36FEF6" box="[1138,1272,296,320]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96C5C31FFFEFFA7FBB5FEFEFB36FEF6" box="[1138,1272,296,320]" italics="true" pageId="6" pageNumber="69">Psephenops</emphasis>
|
||
</taxonomicName>
|
||
species by the following features [other species in brackets]: antennae moniliform [filiform,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FBCEFE9AFB84FED2" box="[1033,1098,332,356]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="lupita">
|
||
<emphasis id="B96C5C31FFFEFFA7FBCEFE9AFB84FED2" box="[1033,1098,332,356]" italics="true" pageId="6" pageNumber="69">lupita</emphasis>
|
||
</taxonomicName>
|
||
]; pronotum without tubercles [with 3 small tubercles,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FE6EFEA6FDD7FE3E" box="[425,537,368,392]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="grouvellei">
|
||
<emphasis id="B96C5C31FFFEFFA7FE6EFEA6FDD7FE3E" box="[425,537,368,392]" italics="true" pageId="6" pageNumber="69">grouvellei</emphasis>
|
||
</taxonomicName>
|
||
]; protibiae with denticle [without denticle,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FBC0FEA7FBBAFE3E" box="[1031,1140,369,392]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="prestonae">
|
||
<emphasis id="B96C5C31FFFEFFA7FBC0FEA7FBBAFE3E" box="[1031,1140,369,392]" italics="true" pageId="6" pageNumber="69">prestonae</emphasis>
|
||
</taxonomicName>
|
||
]; tarsal lobes short [tarsal lobes enlarged
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FE8EFE42FE1CFE1A" box="[329,466,404,428]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="maculicollis">
|
||
<emphasis id="B96C5C31FFFEFFA7FE8EFE42FE1CFE1A" box="[329,466,404,428]" italics="true" pageId="6" pageNumber="69">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FE22FE43FD95FE1A" box="[485,603,405,428]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B96C5C31FFFEFFA7FE22FE43FD95FE1A" box="[485,603,405,428]" italics="true" pageId="6" pageNumber="69">mexicanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FDA9FE43FD15FE1A" box="[622,731,405,428]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="prestonae">
|
||
<emphasis id="B96C5C31FFFEFFA7FDA9FE43FD15FE1A" box="[622,731,405,428]" italics="true" pageId="6" pageNumber="69">prestonae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FD29FE43FC42FE1A" box="[750,908,405,428]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="spiniparameri" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFEFFA7FD29FE43FC42FE1A" box="[750,908,405,428]" italics="true" pageId="6" pageNumber="69">spiniparameri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFEFFA7FC5FFE42FC38FE1A" bold="true" box="[920,1014,404,428]" pageId="6" pageNumber="69">
|
||
<taxonomicNameLabel id="A25FE14AFFFEFFA7FC5FFE42FC38FE1A" box="[920,1014,404,428]" pageId="6" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
]. By general morphology and male genitalia
|
||
<emphasis id="B96C5C31FFFEFFA7FEC6FE6FFEB2FE66" box="[257,380,440,464]" italics="true" pageId="6" pageNumber="69">P. s h e p ard i</emphasis>
|
||
<emphasis id="B96C5C31FFFEFFA7FE4EFE6EFE28FE66" bold="true" box="[393,486,440,464]" pageId="6" pageNumber="69">sp. nov.</emphasis>
|
||
appears closely related to
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FCDBFE6FFC0DFE66" box="[796,963,440,464]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="maculicollis">
|
||
<emphasis id="B96C5C31FFFEFFA7FCDBFE6FFC0DFE66" box="[796,963,440,464]" italics="true" pageId="6" pageNumber="69">P. maculicollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B96C5C31FFFEFFA7FC39FE6FFB48FE66" box="[1022,1158,441,464]" italics="true" pageId="6" pageNumber="69">P. p re s t o n a e</emphasis>
|
||
.
|
||
<taxonomicName id="4C18FBA0FFFEFFA7FB5CFE6FFAD4FE66" box="[1179,1306,440,464]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="69" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFEFFA7FB5CFE6FFB67FE66" box="[1179,1193,441,464]" italics="true" pageId="6" pageNumber="69">P</emphasis>
|
||
.
|
||
<emphasis id="B96C5C31FFFEFFA7FB7EFE6EFAD4FE66" box="[1209,1306,440,464]" italics="true" pageId="6" pageNumber="69">shepardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFEFFA7FAE3FE6EFA4EFE66" bold="true" box="[1316,1408,440,464]" pageId="6" pageNumber="69">
|
||
<taxonomicNameLabel id="A25FE14AFFFEFFA7FAE3FE6EFA4EFE66" box="[1316,1408,440,464]" pageId="6" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently known from lotic regions 2 and 3 of
|
||
<collectingCountry id="F30FC0B3FFFEFFA7FD5AFE0AFD2AFE42" box="[669,740,476,500]" name="Belize" pageId="6" pageNumber="69">Belize</collectingCountry>
|
||
(
|
||
<emphasis id="B96C5C31FFFEFFA7FD32FE0BFCFDFE42" box="[757,819,477,500]" italics="true" pageId="6" pageNumber="69">sensu</emphasis>
|
||
Shepard 2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |