260 lines
30 KiB
XML
260 lines
30 KiB
XML
<document id="03A80EFA680B62421B238ACA6449566F" ID-DOI="10.11646/zootaxa.3670.1.5" ID-GBIF-Dataset="09317f41-bc49-448b-a019-0b78e079ba4a" ID-ISSN="1175-5326" ID-Zenodo-Dep="283933" ID-ZooBank="C6F33726-B9EC-4000-8957-B24158E98AC2" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633519470" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto & Novelo-Gutiérrez, Rodolfo" docDate="2013" docId="03B13135FFF9FFA2FF50FF41FE8CFCEA" docLanguage="en" docName="zt03670p070.pdf" docOrigin="Zootaxa 3670 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Psephenops spiniparameri Arce-Pérez & Novelo-Gutiérrez, 2013, sp. nov." docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="FF88494DFFF8FFA1FFC7FFD6FFCEFFB6" masterDocTitle="Two new species of Psephenops Grouvelle from Belize (Coleoptera: Byrrhoidea: Psephenidae), with a key to the known species from Mexico and Central America" masterLastPageNumber="70" masterPageNumber="63" pageNumber="64" updateTime="1698334777422" updateUser="plazi">
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<mods:mods id="FB32BFB6EB12E0639D1553384C797B0B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="218010673F34346AC5BFF2D0AE6699EB">
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<mods:title id="75DFF4B92C26BBAFE15147430107389E">Two new species of Psephenops Grouvelle from Belize (Coleoptera: Byrrhoidea: Psephenidae), with a key to the known species from Mexico and Central America</mods:title>
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<mods:name id="575098EFFCF42227D1BF0C412203BEFF" type="personal">
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<mods:namePart id="808DBC62F74A71D3E12F41743C219FD9">Arce-Pérez, Roberto</mods:namePart>
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<mods:name id="0C61E7C33D56890DE7C030F8156C76E1" type="personal">
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<mods:namePart id="9629CA8CE34CE7C340F42357C704FA65">Novelo-Gutiérrez, Rodolfo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A77E702D2B58F6495A5D53C3FFDB0C4A">text</mods:typeOfResource>
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<mods:titleInfo id="B5860A158D6608FACA5FC8F2B93D6158">
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<mods:title id="3D0C38B15A22EC363143E62D3F695288">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3FFB4A9D4EEDB4A67C6666490CF9C9FA">
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<mods:date id="13359E8CE0649D23CF21D632D1DE7704">2013</mods:date>
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<mods:detail id="3900CF305CF66BC841FFAAD80BA8FFA9" type="volume">
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<mods:number id="E4CAF26C3DC237FBBC18D94B5B8F464A">3670</mods:number>
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<mods:detail id="93344C8C29BE9103A5370D7B5B4FD5FC" type="issue">
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<mods:number id="E2A935C0600A48ECC8B790F9CA601140">1</mods:number>
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<mods:classification id="9EECB25C2945D8BEF4278051FE02CB38">journal article</mods:classification>
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<mods:identifier id="E6C156B4F16A50263FA11160A996E917" type="DOI">10.11646/zootaxa.3670.1.5</mods:identifier>
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<mods:identifier id="92EBDF1AAA4E945DF559F44AC5549DA8" type="GBIF-Dataset">09317f41-bc49-448b-a019-0b78e079ba4a</mods:identifier>
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<mods:identifier id="84995D03CEC038733C4E3FB90247B12B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0FA6DD86D54E2BD5C229E82D4F3970A5" type="Zenodo-Dep">283933</mods:identifier>
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<mods:identifier id="F29A194460AE41FF7DF34D8EFAB91A77" type="ZooBank">C6F33726-B9EC-4000-8957-B24158E98AC2</mods:identifier>
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</mods:mods>
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<treatment id="03B13135FFF9FFA2FF50FF41FE8CFCEA" ID-DOI="http://doi.org/10.5281/zenodo.6148887" ID-GBIF-Taxon="127664370" ID-Zenodo-Dep="6148887" LSID="urn:lsid:plazi:treatment:03B13135FFF9FFA2FF50FF41FE8CFCEA" httpUri="http://treatment.plazi.org/id/03B13135FFF9FFA2FF50FF41FE8CFCEA" lastPageId="3" lastPageNumber="66" pageId="1" pageNumber="64">
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<subSubSection id="C302D3A8FFF9FFA0FF50FF41FEFDFF64" pageId="1" pageNumber="64" type="nomenclature">
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<paragraph id="8BA78023FFF9FFA0FF50FF41FD00FF07" blockId="1.[151,718,151,210]" box="[151,718,151,177]" pageId="1" pageNumber="64">
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<heading id="D0EF374FFFF9FFA0FF50FF41FD00FF07" bold="true" box="[151,718,151,177]" fontSize="11" level="1" pageId="1" pageNumber="64" reason="1">
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<emphasis id="B96C5C31FFF9FFA0FF50FF41FD00FF07" bold="true" box="[151,718,151,177]" pageId="1" pageNumber="64">
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<taxonomicName id="4C18FBA0FFF9FFA0FF50FF41FE19FF07" box="[151,471,151,177]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="spiniparameri" status="sp. nov.">
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<emphasis id="B96C5C31FFF9FFA0FF50FF41FE19FF07" bold="true" box="[151,471,151,177]" italics="true" pageId="1" pageNumber="64">Psephenops spiniparameri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25FE14AFFF9FFA0FE19FF41FDF5FF07" box="[478,571,151,177]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
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Arce-Pérez
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA78023FFF9FFA0FF50FF6CFEFDFF64" blockId="1.[151,718,151,210]" box="[151,307,186,210]" pageId="1" pageNumber="64">
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(
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<figureCitation id="13239CA6FFF9FFA0FF58FF6CFF2CFF64" box="[159,226,186,210]" captionStart="FIGURE 1" captionStartId="1.[151,255,1314,1338]" captionTargetBox="[159,1436,265,1275]" captionTargetId="figure@1.[151,1436,264,1293]" captionTargetPageId="1" captionText="FIGURE 1. Hind tarsi of Psephenops, dorsolateral view: a) P. spiniparameri sp. nov.; b) P. shepardi sp. nov." httpUri="https://zenodo.org/record/283934/files/figure.png" pageId="1" pageNumber="64">Figs 1</figureCitation>
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a, 2–4)
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</paragraph>
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</subSubSection>
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<caption id="DF67D0ABFFF9FFA0FF50FAF4FAE9FA8C" box="[151,1319,1314,1338]" httpUri="https://zenodo.org/record/283934/files/figure.png" pageId="1" pageNumber="64" targetBox="[159,1436,265,1275]" targetPageId="1">
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<paragraph id="8BA78023FFF9FFA0FF50FAF4FAE9FA8C" blockId="1.[151,1319,1314,1338]" box="[151,1319,1314,1338]" pageId="1" pageNumber="64">
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<emphasis id="B96C5C31FFF9FFA0FF50FAF4FED7FA8C" bold="true" box="[151,281,1314,1338]" pageId="1" pageNumber="64">FIGURE 1.</emphasis>
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Hind tarsi of
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<taxonomicName id="4C18FBA0FFF9FFA0FE6AFAF4FDE2FA8F" box="[429,556,1314,1337]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B96C5C31FFF9FFA0FE6AFAF4FDE2FA8F" box="[429,556,1314,1337]" italics="true" pageId="1" pageNumber="64">Psephenops</emphasis>
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</taxonomicName>
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, dorsolateral view: a)
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<taxonomicName id="4C18FBA0FFF9FFA0FCDBFAF4FC03FA8F" box="[796,973,1314,1337]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="spiniparameri" status="sp. nov.">
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<emphasis id="B96C5C31FFF9FFA0FCDBFAF4FC03FA8F" box="[796,973,1314,1337]" italics="true" pageId="1" pageNumber="64">P. spiniparameri</emphasis>
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</taxonomicName>
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<emphasis id="B96C5C31FFF9FFA0FC13FAF4FBE7FA8C" bold="true" box="[980,1065,1314,1338]" pageId="1" pageNumber="64">
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<taxonomicNameLabel id="A25FE14AFFF9FFA0FC13FAF4FBE7FA8C" box="[980,1065,1314,1338]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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; b)
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<taxonomicName id="4C18FBA0FFF9FFA0FB93FAF4FB04FA8F" box="[1108,1226,1314,1337]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="shepardi" status="sp. nov.">
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<emphasis id="B96C5C31FFF9FFA0FB93FAF4FB04FA8F" box="[1108,1226,1314,1337]" italics="true" pageId="1" pageNumber="64">P. shepardi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25FE14AFFF9FFA0FB16FAF4FAE9FA8C" box="[1233,1319,1314,1338]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</caption>
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<subSubSection id="C302D3A8FFF9FFA2FF50FAB3FE83FE89" lastPageId="3" lastPageNumber="66" pageId="1" pageNumber="64" type="description">
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<paragraph id="8BA78023FFF9FFA3FF50FAB3FB6DFCC9" blockId="1.[151,1436,1381,2018]" lastBlockId="2.[151,1437,151,896]" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
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<emphasis id="B96C5C31FFF9FFA0FF50FAB3FDCFFAC8" bold="true" box="[151,513,1381,1406]" pageId="1" pageNumber="64">
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Description.
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<typeStatus id="54A33E81FFF9FFA0FEFCFAB0FE66FAC8" box="[315,424,1382,1406]" pageId="1" pageNumber="64" type="holotype">Holotype</typeStatus>
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Male:
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</emphasis>
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Body oval, depressed; integument yellowish-red, with yellow reflections, completely covered with iridescent, yellow-grayish, short setae. Head, pronotum and scutellum dark, yellowishred; antennae and elytra light, yellowish-red; legs with coxae, trochanters, and femora light yellowish-red, tibiae and tarsi dark yellowish-red. Venter dark yellowish-red, vestiture short, fine and dense, grayish-yellow, with yellow reflections.
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<emphasis id="B96C5C31FFF9FFA0FEDBFA20FE96F9B8" box="[284,344,1526,1550]" italics="true" pageId="1" pageNumber="64">Head</emphasis>
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: short, transverse; clypeus subrectangular, declivous at an angle of less than 90 degrees from the plane of the frons, distal margin widely emarginate; fronto-clypeal surface with long, stout, dark reddish setae; vertex minutely punctate, with minute setae and a very shallow longitudinal furrow in middle. Antennae moniliform, 11-segmented, scape larger than other segments, pedicel 0.50 length of scape, subglobose; eight flagellar segments small, apical segment acuminate. Eyes lateral, rounded, very prominent, black; with post-ocular area yellow. Labrum similar to clypeus but shorter; maxillary palp 4-segmented, covered with short setae, segment 1 smallest, apical segment largest, relative proportion of segments: 0.4, 0.8, 0.6, 1.0, basal segments 1–3 subconical, segment 4 oval, inflated from the base and gradually narrowing to tip (acuminate). Labial palpi very short, 3-segmented, segments 1–2 globular and wide, segment 3 subconical, slightly longer than preceding ones.
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<emphasis id="B96C5C31FFF9FFA0FF50F8ECFF26F8E4" box="[151,232,1850,1874]" italics="true" pageId="1" pageNumber="64">Thorax</emphasis>
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: Pronotum, subtrapezoidal, anterior margin arcuate, posterior margin bisinuate, almost as wide as elytral base, lateral margins straight along anterior half, expanded laterally on posterior half, posterolateral angles lightly round, all margins narrowly rimmed. Pronotal disc with a median longitudinal carina well developed on distal half and wide, shallow, circular depression at each side of carina; in profile view, pronotal disc (except for posterolateral expansions) convex, pronotal surface with punctation coarser than that of the head, covered with short, grayishyellow setae and long dark reddish setae, setae shorter than those of head. Scutellum short, with apex rounded.
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<emphasis id="B96C5C31FFFAFFA3FF50FF6AFF13FF62" box="[151,221,188,212]" italics="true" pageId="2" pageNumber="65">Elytra</emphasis>
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: with short grayish-yellow setae intermingled with long, dark, reddish setae shorter than those of head; sides parallel in basal 0.75, then gradually converging posteriorly, base lightly depressed on internal side of humeri. Prosternum short, slightly carinate, with long, lanceolate, slightly carinate prosternal process, which reaches half of mesoventrite. Mesoventrite short, with wide groove for reception of prosternal process. Metaventrite large and bulky.
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<emphasis id="B96C5C31FFFAFFA3FF20FE9BFED2FED2" box="[231,284,333,356]" italics="true" pageId="2" pageNumber="65">Legs</emphasis>
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: procoxae globose, mesocoxae conical, metacoxae transverse; femora robust, reddish-yellow (dark reddish-brown in some
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<typeStatus id="54A33E81FFFAFFA3FE59FEB9FDC0FE31" box="[414,526,367,391]" pageId="2" pageNumber="65" type="paratype">paratypes</typeStatus>
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); tibiae slender, metatibiae longer than femora, protibiae with distinct apicolateral denticle. Tarsi 5-segmented (
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<figureCitation id="13239CA6FFFAFFA3FE2EFE42FDE3FE1A" box="[489,557,404,428]" captionStart="FIGURE 1" captionStartId="1.[151,255,1314,1338]" captionTargetBox="[159,1436,265,1275]" captionTargetId="figure@1.[151,1436,264,1293]" captionTargetPageId="1" captionText="FIGURE 1. Hind tarsi of Psephenops, dorsolateral view: a) P. spiniparameri sp. nov.; b) P. shepardi sp. nov." httpUri="https://zenodo.org/record/283934/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
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a), tarsomeres 1 and 2 ventrally lobed, with a thick, sponge-like vestiture, ventral lobe of tarsomere 1 extending distally over basal 0.25 of tarsomere 2, tarsomere 2 twice as long as tarsomere 1, ventral lobe of tarsomere 2 extending distally and completely covering area of tarsomeres 3, 4, and basal 0.90 of tarsomere 5; proportional length of tarsomeres: 0.5, 1.0, 0.1, 0.1, 0.45; a pair of long, curved, slender apical claws, with a small basal tooth.
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<emphasis id="B96C5C31FFFAFFA3FD83FDF2FD63FD8A" box="[580,685,548,572]" italics="true" pageId="2" pageNumber="65">Abdomen</emphasis>
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: vestiture short, fine, dense, with yellow iridescence, and also with long, sparse, dark reddish setae. Seven visible sterna (ventrites) in relaxed specimens, usually ventrite VI hidden under ventrite V, ventrite II the largest; posterior margin of ventrite I deeply concave at middle, sinuate at sides, posterior margin of ventrite II as in ventrite I but concavity shallow medially, posterior margin of ventrites III and IV straight, ventrite V with a wide, shallow concavity; ventrite VI very short, posterior margin widely concave. Pygidial plate oval, long, narrow and densely covered with long, stiff, dark, reddish setae.
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<emphasis id="B96C5C31FFFAFFA3FB7EFD0EFAEDFD46" box="[1209,1315,728,752]" italics="true" pageId="2" pageNumber="65">Genitalia</emphasis>
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: aedeagus trilobate (
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<figureCitation id="13239CA6FFFAFFA3FECEFD2AFE9DFCA2" box="[265,339,764,788]" captionStart="FIGURES 2 – 4" captionStartId="2.[151,269,1969,1993]" captionTargetBox="[151,1428,936,1916]" captionTargetId="figure@2.[151,1436,934,1948]" captionTargetPageId="2" captionText="FIGURES 2 – 4. Male genitalia of Psephenops spinaparameri sp. nov.: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view." httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
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). Parameres robust, subtriangular, strongly divergent in apical third, curved, spine-like, with internal margins fused at basal
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<quantity id="4CE02DC6FFFAFFA3FDC4FCC9FD9BFC81" box="[515,597,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="65" unit="in" value="0.1">0.10 in</quantity>
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dorsal view (
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<figureCitation id="13239CA6FFFAFFA3FD31FCC9FCF1FC81" box="[758,831,799,823]" captionStart="FIGURES 2 – 4" captionStartId="2.[151,269,1969,1993]" captionTargetBox="[151,1428,936,1916]" captionTargetId="figure@2.[151,1436,934,1948]" captionTargetPageId="2" captionText="FIGURES 2 – 4. Male genitalia of Psephenops spinaparameri sp. nov.: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view." httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
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). Penis visibly longer than parameres (
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<figureCitation id="13239CA6FFFAFFA3FAC2FCC9FAA1FC81" box="[1285,1391,799,823]" captionStart="FIGURES 2 – 4" captionStartId="2.[151,269,1969,1993]" captionTargetBox="[151,1428,936,1916]" captionTargetId="figure@2.[151,1436,934,1948]" captionTargetPageId="2" captionText="FIGURES 2 – 4. Male genitalia of Psephenops spinaparameri sp. nov.: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view." httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65">Figs 2–4</figureCitation>
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), in ventral view finger-like, bearing a ventral, longitudinal, long, slender sclerite which is 0.52 as long as penis (
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<figureCitation id="13239CA6FFFAFFA3FAA9FC92FF69FCC9" captionStart="FIGURES 2 – 4" captionStartId="2.[151,269,1969,1993]" captionTargetBox="[151,1428,936,1916]" captionTargetId="figure@2.[151,1436,934,1948]" captionTargetPageId="2" captionText="FIGURES 2 – 4. Male genitalia of Psephenops spinaparameri sp. nov.: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view." httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65">Fig. 3</figureCitation>
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). Phallobase, in dorsal view, concave, in ventral view, bulky and subcylindrical (
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<figureCitation id="13239CA6FFFAFFA3FBF7FCB1FB58FCC9" box="[1072,1174,871,895]" captionStart="FIGURES 2 – 4" captionStartId="2.[151,269,1969,1993]" captionTargetBox="[151,1428,936,1916]" captionTargetId="figure@2.[151,1436,934,1948]" captionTargetPageId="2" captionText="FIGURES 2 – 4. Male genitalia of Psephenops spinaparameri sp. nov.: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view." httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65">Figs 2–3</figureCitation>
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).
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</paragraph>
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<caption id="DF67D0ABFFFAFFA3FF50F867FC81F85A" httpUri="https://zenodo.org/record/283935/files/figure.png" pageId="2" pageNumber="65" targetBox="[151,1428,936,1916]" targetPageId="2">
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<paragraph id="8BA78023FFFAFFA3FF50F867FC81F85A" blockId="2.[151,1436,1969,2028]" pageId="2" pageNumber="65">
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<emphasis id="B96C5C31FFFAFFA3FF50F867FE86F87F" bold="true" box="[151,328,1969,1993]" pageId="2" pageNumber="65">FIGURES 2–4.</emphasis>
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Male genitalia of
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<taxonomicName id="4C18FBA0FFFAFFA3FDE7F864FC86F87F" box="[544,840,1970,1993]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="spinaparameri" status="sp. nov.">
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<emphasis id="B96C5C31FFFAFFA3FDE7F864FC86F87F" box="[544,840,1970,1993]" italics="true" pageId="2" pageNumber="65">Psephenops spinaparameri</emphasis>
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</taxonomicName>
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<emphasis id="B96C5C31FFFAFFA3FC93F867FC60F87F" bold="true" box="[852,942,1969,1993]" pageId="2" pageNumber="65">
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<taxonomicNameLabel id="A25FE14AFFFAFFA3FC93F867FC60F87F" box="[852,942,1969,1993]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: 2) phallobase, parameres and median lobe (penis), dorsal view; 3) same, ventral view; 4) same, lateral view.
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</paragraph>
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</caption>
|
||
<paragraph id="8BA78023FFFBFFA2FF00FF4EFE83FE89" blockId="3.[151,1436,151,860]" pageId="3" pageNumber="66">
|
||
<emphasis id="B96C5C31FFFBFFA2FF00FF4EFB9AFF06" bold="true" box="[199,1108,151,176]" pageId="3" pageNumber="66">
|
||
Measurements (in mm) [in square brackets the
|
||
<typeStatus id="54A33E81FFFBFFA2FCD0FF4EFC43FF06" box="[791,909,152,176]" pageId="3" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
variation N=6]:
|
||
</emphasis>
|
||
Total length 2.75 [2.54-3.0, mean=2.67]; length of segments of maxillary palpi: 0.04, 0.08, 0.06, 0.10. Pronotum: posterior margin 1.02 [1.02- 1.10, mean=1.06], lateral margin 0.5 [0.5-0.62, mean=0.56]; humeral width 1.2 [1.18-1.24, mean=1.20]; length of tarsomeres: 0.20, 0.40, 0.04, 0.04, 0.18. Genitalia: total length 0.54; parameres 0.36, penis 0.38, phallobase: length 0.22, width 0.17
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302D3A8FFFBFFA2FF00FE9AFE22FD8A" pageId="3" pageNumber="66" type="materials_examined">
|
||
<paragraph id="8BA78023FFFBFFA2FF00FE9AFEAFFE42" blockId="3.[151,1436,151,860]" pageId="3" pageNumber="66">
|
||
<emphasis id="B96C5C31FFFBFFA2FF00FE9AFEB8FED2" bold="true" box="[199,374,331,356]" pageId="3" pageNumber="66">
|
||
<typeStatus id="54A33E81FFFBFFA2FF00FE9AFECCFED2" box="[199,258,332,356]" pageId="3" pageNumber="66">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54A33E81FFFBFFA2FEB8FE9AFE29FED2" box="[383,487,332,356]" pageId="3" pageNumber="66" type="holotype">Holotype</typeStatus>
|
||
labeled
|
||
<collectingCountry id="F30FC0B3FFFBFFA2FD8EFE9AFD5BFED2" box="[585,661,332,356]" name="Belize" pageId="3" pageNumber="66">Belize</collectingCountry>
|
||
: Cayo District,
|
||
<quantity id="4CE02DC6FFFBFFA2FC8EFE9AFC5CFED2" box="[841,914,332,356]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="3" pageNumber="66" unit="mi" value="1.7">1.7 mi</quantity>
|
||
E. O. Top. Camp,
|
||
<date id="FFA6A6E3FFFBFFA2FBAAFE9AFB1FFED2" box="[1133,1233,332,356]" pageId="3" pageNumber="66" value="1995-01-09">9-I-1995</date>
|
||
, trib to Dr/Creek, WDS-A-1127 (EMEC); Two male
|
||
<typeStatus id="54A33E81FFFBFFA2FDFBFEA6FD64FE3E" box="[572,682,368,392]" pageId="3" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
, same data as
|
||
<typeStatus id="54A33E81FFFBFFA2FCAFFEB9FC04FE31" box="[872,970,367,391]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
(IEXA); Toledo District,
|
||
<quantity id="4CE02DC6FFFBFFA2FACDFEB9FAA3FE31" box="[1290,1389,367,391]" metricMagnitude="4" metricUnit="m" metricValue="3.51" pageId="3" pageNumber="66" unit="km" value="35.1">35.1 km</quantity>
|
||
SE Belmopan, 3-Vlll-1993, unnamed stream, WDS-A-985, 2 3 (EMEC); Cayo Dist.,
|
||
<quantity id="4CE02DC6FFFBFFA2FBF9FE42FB4EFE1A" box="[1086,1152,404,428]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="3" pageNumber="66" unit="mi" value="20.0">20 mi</quantity>
|
||
SE Belmopan,
|
||
<date id="FFA6A6E3FFFBFFA2FAF4FE42FA59FE1A" box="[1331,1431,404,428]" pageId="3" pageNumber="66" value="1993-01-02">2-I-1993</date>
|
||
, Sibun River, WDS-A-946, 1 3, (EMEC); Cayo Dist.,
|
||
<quantity id="4CE02DC6FFFBFFA2FD3CFE61FC96FE79" box="[763,856,439,463]" metricMagnitude="4" metricUnit="m" metricValue="2.84" pageId="3" pageNumber="66" unit="km" value="28.4">28.4 km</quantity>
|
||
SE Belmopan,
|
||
<date id="FFA6A6E3FFFBFFA2FBCDFE61FB58FE79" box="[1034,1174,439,463]" pageId="3" pageNumber="66" value="1993-08-03">3-VIII-1993</date>
|
||
, Sibun River, WDS-A- 984, 13 (EMEC).
|
||
</paragraph>
|
||
<paragraph id="8BA78023FFFBFFA2FF00FDD6FE22FD8A" blockId="3.[151,1436,151,860]" pageId="3" pageNumber="66">
|
||
<emphasis id="B96C5C31FFFBFFA2FF00FDD6FEA7FDAE" bold="true" box="[199,361,511,536]" pageId="3" pageNumber="66">
|
||
<typeStatus id="54A33E81FFFBFFA2FF00FDD6FECEFDAE" box="[199,256,512,536]" pageId="3" pageNumber="66">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F30FC0B3FFFBFFA2FEB1FE29FE0FFDA1" box="[374,449,511,535]" name="Belize" pageId="3" pageNumber="66">Belize</collectingCountry>
|
||
; Cayo District,
|
||
<quantity id="4CE02DC6FFFBFFA2FD46FE29FD03FDA1" box="[641,717,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.7358848" pageId="3" pageNumber="66" unit="mi" value="1.7">1.7 mi</quantity>
|
||
east of Over the Top Camp on the Hummingbird Highway, tributary to Dry Creek,
|
||
<specimenCount id="9D1E4BAAFFFBFFA2FE59FDF2FE22FD8A" box="[414,492,548,572]" pageId="3" pageNumber="66" type="generic">
|
||
<quantity id="4CE02DC6FFFBFFA2FE59FDF2FE29FD8A" box="[414,487,548,572]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="66" unit="m" value="115.0">115 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302D3A8FFFBFFA2FF00FD9EFBFCFDE9" box="[199,1074,583,608]" pageId="3" pageNumber="66" type="etymology">
|
||
<paragraph id="8BA78023FFFBFFA2FF00FD9EFBFCFDE9" blockId="3.[151,1436,151,860]" box="[199,1074,583,608]" pageId="3" pageNumber="66">
|
||
<emphasis id="B96C5C31FFFBFFA2FF00FD9EFE80FDD6" bold="true" box="[199,334,584,608]" pageId="3" pageNumber="66">Etymology.</emphasis>
|
||
The name
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FE0CFD9FFDA7FDD6" box="[459,617,585,608]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="spiniparameri">
|
||
<emphasis id="B96C5C31FFFBFFA2FE0CFD9FFDA7FDD6" box="[459,617,585,608]" italics="true" pageId="3" pageNumber="66">spiniparameri</emphasis>
|
||
</taxonomicName>
|
||
refers to the parameres being spine-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C302D3A8FFFBFFA2FF00FDBAFE8CFCEA" pageId="3" pageNumber="66" type="discussion">
|
||
<paragraph id="8BA78023FFFBFFA2FF00FDBAFE8CFCEA" blockId="3.[151,1436,151,860]" pageId="3" pageNumber="66">
|
||
<emphasis id="B96C5C31FFFBFFA2FF00FDBAFEF5FD32" bold="true" box="[199,315,620,644]" pageId="3" pageNumber="66">Remarks.</emphasis>
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FE84FDBAFDBEFD32" box="[323,624,620,644]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="spiniparameri" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFBFFA2FE84FDBAFDBEFD32" box="[323,624,620,644]" italics="true" pageId="3" pageNumber="66">Psephenops spiniparameri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFBFFA2FDBFFDBAFD1DFD32" bold="true" box="[632,723,620,644]" pageId="3" pageNumber="66">
|
||
<taxonomicNameLabel id="A25FE14AFFFBFFA2FDBFFDBAFD1DFD32" box="[632,723,620,644]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be separated easily from other known
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FB7AFDBAFA8DFD32" box="[1213,1347,620,644]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96C5C31FFFBFFA2FB7AFDBAFA8DFD32" box="[1213,1347,620,644]" italics="true" pageId="3" pageNumber="66">Psephenops</emphasis>
|
||
</taxonomicName>
|
||
species by the following features [other species in brackets]: antennae moniliform [filiform,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FBB5FD46FB7DFD1E" box="[1138,1203,656,680]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="lupita">
|
||
<emphasis id="B96C5C31FFFBFFA2FBB5FD46FB7DFD1E" box="[1138,1203,656,680]" italics="true" pageId="3" pageNumber="66">lupita</emphasis>
|
||
</taxonomicName>
|
||
]; pronotum without tubercles [with 3 small tubercles,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FDE4FD62FD5AFD7A" box="[547,660,692,716]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="grouvellei">
|
||
<emphasis id="B96C5C31FFFBFFA2FDE4FD62FD5AFD7A" box="[547,660,692,716]" italics="true" pageId="3" pageNumber="66">grouvellei</emphasis>
|
||
</taxonomicName>
|
||
]; protibiae with denticle [without denticle,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FB57FD63FB33FD7A" box="[1168,1277,693,716]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="prestonae">
|
||
<emphasis id="B96C5C31FFFBFFA2FB57FD63FB33FD7A" box="[1168,1277,693,716]" italics="true" pageId="3" pageNumber="66">prestonae</emphasis>
|
||
</taxonomicName>
|
||
]; tarsal lobes enlarged [tarsal lobes short,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FE12FD0EFD8BFD46" box="[469,581,728,752]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="grouvellei">
|
||
<emphasis id="B96C5C31FFFBFFA2FE12FD0EFD8BFD46" box="[469,581,728,752]" italics="true" pageId="3" pageNumber="66">grouvellei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FD93FD0EFD5BFD46" box="[596,661,728,752]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="lupita">
|
||
<emphasis id="B96C5C31FFFBFFA2FD93FD0EFD5BFD46" box="[596,661,728,752]" italics="true" pageId="3" pageNumber="66">lupita</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FD63FD0EFCCBFD46" box="[676,773,728,752]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="shepardi">
|
||
<emphasis id="B96C5C31FFFBFFA2FD63FD0EFCCBFD46" box="[676,773,728,752]" italics="true" pageId="3" pageNumber="66">shepardi</emphasis>
|
||
</taxonomicName>
|
||
]; apices of parameres spine-like, divergent [not spine-like, all other species]. By general morphology and male genitalia
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FC82FD2BFC31FCA2" box="[837,1023,765,788]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="spiniparameri" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFBFFA2FC82FD2BFC31FCA2" box="[837,1023,765,788]" italics="true" pageId="3" pageNumber="66">P. spiniparameri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFBFFA2FBC1FD2AFB91FCA2" bold="true" box="[1030,1119,764,788]" pageId="3" pageNumber="66">
|
||
<taxonomicNameLabel id="A25FE14AFFFBFFA2FBC1FD2AFB91FCA2" box="[1030,1119,764,788]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears closely related to
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FA40FD2BFEC3FC8E" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B96C5C31FFFBFFA2FA40FD2BFEC3FC8E" italics="true" pageId="3" pageNumber="66">P. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C18FBA0FFFBFFA2FEE7FCF6FD98FC8E" box="[288,598,800,824]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="spinaparameri" status="sp. nov.">
|
||
<emphasis id="B96C5C31FFFBFFA2FEE7FCF6FD98FC8E" box="[288,598,800,824]" italics="true" pageId="3" pageNumber="66">Psephenops spinaparameri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C5C31FFFBFFA2FDA5FCF6FD0EFC8E" bold="true" box="[610,704,800,824]" pageId="3" pageNumber="66">
|
||
<taxonomicNameLabel id="A25FE14AFFFBFFA2FDA5FCF6FD0EFC8E" box="[610,704,800,824]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently known only from lotic region 3 of
|
||
<collectingCountry id="F30FC0B3FFFBFFA2FAC6FCC9FA87FC81" box="[1281,1353,799,823]" name="Belize" pageId="3" pageNumber="66">Belize</collectingCountry>
|
||
(
|
||
<emphasis id="B96C5C31FFFBFFA2FA99FCF7FA52FC8E" box="[1374,1436,801,824]" italics="true" pageId="3" pageNumber="66">sensu</emphasis>
|
||
Shepard 2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |