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<mods:title id="6047F5E5CE929B2FCA70B063EF121763">New species and records of Caenidae Newman, 1853 (Insecta: Ephemeroptera) from Rondônia State, Northern Brazil</mods:title>
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<mods:namePart id="A78B7865AACEFEA0DB074ECD33A9CE13">Nascimento, Stênio R. S.</mods:namePart>
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<mods:affiliation id="6C1AA3A16B857D7A15267B74EF1BACF9">Universidade Federal de Rondônia (UNIR), Departamento de Biologia, Laboratório de Biodiversidade e Conservação, Rondônia, Brazil &amp; pvilelacruz @ gmail. com; https: // orcid. org / 0000 - 0003 - 2426 - 4628</mods:affiliation>
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<emphasis id="B970EAF7FFF2FFDB3391F8D3424E7647" bold="true" box="[151,315,1846,1872]" italics="true" pageId="2" pageNumber="239">Caenis urupa</emphasis>
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(
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<figureCitation id="133F2A60FFF2FFDB314FF8D3412C7647" box="[585,601,1846,1872]" captionStart="FIGURE 3" captionStartId="5.[152,255,1293,1317]" captionTargetBox="[190,1397,181,1269]" captionTargetId="figure-20@5.[190,1397,181,1269]" captionTargetPageId="5" captionText="FIGURE 3. Caenis urupa sp. nov. male imago, genitalia: A, ventral view (Scale = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309022" httpUri="https://zenodo.org/record/8309022/files/figure.png" pageId="2" pageNumber="239">3</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB3160F8D041DA7658" box="[614,687,1845,1871]" captionStart="FIGURE 4" captionStartId="6.[152,255,1952,1976]" captionTargetBox="[151,1435,489,1928]" captionTargetId="figure-159@6.[151,1435,489,1928]" captionTargetPageId="6" captionText="FIGURE 4. Caenis urupa sp. nov. female imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view (Scale: Figs. 4A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309024" httpUri="https://zenodo.org/record/8309024/files/figure.png" pageId="2" pageNumber="239">4A–C</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB31BAF8D040717658" box="[700,772,1845,1871]" captionStart="FIGURE 5" captionStartId="7.[152,255,1629,1653]" captionTargetBox="[151,1435,181,1606]" captionTargetId="figure-20@7.[151,1435,181,1606]" captionTargetPageId="7" captionText="FIGURE 5. Caenis urupa sp. nov. egg: A, geral view; B, structure of chorion; C, detail of micropyle; D–E, detail of polar cap. (Scale: Figs. 5A, D = 20 μm; 5B = 5 μm; Figs. 5C, E = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.8309026" httpUri="https://zenodo.org/record/8309026/files/figure.png" pageId="2" pageNumber="239">5A–E</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB3017F8D0402E7658" box="[785,859,1845,1871]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="2" pageNumber="239">6A–C</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB306EF8D040DE7658" box="[872,939,1845,1871]" captionStart="FIGURE 7" captionStartId="9.[152,255,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@9.[151,1435,181,1678]" captionTargetPageId="9" captionText="FIGURE 7. Nymph of Caenis urupa sp. nov. A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="2" pageNumber="239">7A–F</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB30BEF8D0408E7658" box="[952,1019,1845,1871]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="2" pageNumber="239">8A–F</figureCitation>
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<figureCitation id="133F2A60FFF2FFDB370EF8D047247658" box="[1032,1105,1845,1871]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="2" pageNumber="239">9A–D</figureCitation>
)
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♁ imago in alcohol (genitalia on slide) from 
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<collectionCode id="ED15AE20FFF2FFDB37F5F847467276AB" box="[1267,1287,1954,1980]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="239">T</collectionCode>
. cols. (
<specimenCode id="DBA29E9EFFF2FFDB3654F847425076F7" collectionCode="INPA-EPH" pageId="2" pageNumber="239">INPA-EPH000042</specimenCode>
)
</materialsCitation>
. 
<materialsCitation id="3B6C3CB8FFF2FFDB323FF823463276F7" box="[313,1351,1990,2018]" collectingDate="2022-08-23" collectionCode="R, V, T" collectorName="Nascimento, S. &amp; Cruz, P. &amp; Lima, C." country="Brazil" latitude="-11.035501" location="Urupa River" longLatPrecision="1" longitude="-62.144863" municipality="Ji-Parana" pageId="2" pageNumber="239" specimenCode="INPA-EPH000043" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
<emphasis id="B970EAF7FFF2FFDB323FF82342CF76F7" bold="true" box="[313,442,1990,2016]" pageId="2" pageNumber="239">
<typeStatus id="54BF8847FFF2FFDB323FF82342C076F7" box="[313,437,1990,2016]" pageId="2" pageNumber="239">Paratypes</typeStatus>
:
</emphasis>
15 ♁ imagos in alcohol (
<collectingMethod id="52454EF2FFF2FFDB31DAF823403F76F7" box="[732,842,1990,2016]" pageId="2" pageNumber="239">light trap</collectingMethod>
), same data as holotype (
<specimenCode id="DBA29E9EFFF2FFDB3761F823464A76F7" box="[1127,1343,1990,2016]" collectionCode="INPA-EPH" pageId="2" pageNumber="239">INPA-EPH000043</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6C3CB8FFF2FFDA3652F823409471A6" collectingDate="2022-08-23" collectionCode="R, V, T" collectorName="Nascimento, S. &amp; Cruz, P. &amp; Lima, C." country="Brazil" lastPageId="3" lastPageNumber="240" latitude="-11.035501" location="Urupa River" longLatPrecision="1" longitude="-62.144863" municipality="Ji-Parana" pageId="2" pageNumber="239" specimenCode="INPA-EPH000045" specimenCount="1" specimenCount-female="1" stateProvince="Rondonia" typeStatus="paratype">
<specimenCount id="9D02FD6CFFF2FFDB3652F82346E976F7" box="[1364,1436,1990,2016]" pageId="2" pageNumber="239" type="female">five ♀</specimenCount>
imagos in alcohol (
<collectingMethod id="52454EF2FFF3FFDA3274FF72429571A6" box="[370,480,151,177]" pageId="3" pageNumber="240">light trap</collectingMethod>
), same data as holotype (
<specimenCode id="DBA29E9EFFF3FFDA3007FF7340AF71A6" box="[769,986,150,177]" collectionCode="INPA-EPH" pageId="3" pageNumber="240">INPA-EPH000045</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6C3CB8FFF3FFDA30E8FF72408371C2" collectingDate="2022-08-23" collectionCode="R, V, T" collectorName="Nascimento, S. &amp; Cruz, P. &amp; Lima, C." country="Brazil" latitude="-11.035501" location="Urupa River" longLatPrecision="1" longitude="-62.144863" municipality="Ji-Parana" pageId="3" pageNumber="240" specimenCode="INPA-EPH000044" specimenCount="1" stateProvince="Rondonia" typeStatus="paratype">
one ♁ nymph in alcohol (mouthparts, legs and opercular gills on slide), same data as holotype (
<specimenCode id="DBA29E9EFFF3FFDA3011FF5E409A71C2" box="[791,1007,187,213]" collectionCode="INPA-EPH" pageId="3" pageNumber="240">INPA-EPH000044</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31E656EFFF3FFDA33C1FF3B429970C6" pageId="3" pageNumber="240" type="diagnosis">
<paragraph id="8BBB36E5FFF3FFDA33C1FF3B429970C6" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FF3B42FA71EE" bold="true" box="[199,399,222,249]" pageId="3" pageNumber="240">Diagnosis. Male:</emphasis>
1) forceps margins slightly sinuous; 2) penis cylindrical (with rectangular outline in ventral view), apically rounded and not laterally projected. Eggs: 1) micropyle linear with narrow and of moderate length micropylar canal, preceded by an scarcely visible oval sperm guide; 2) two polar caps forming a spiral of fine threads with about 13 terminal knobs. Nymph: 1) lateral margins of the pronotum rounded and broader posteriorly; 2) middle and hind coxa semicircular, without projection; 3) fore tarsal claws a row of small denticles basally, middle tarsal claws with six small denticles basally, hind tarsal claws with several small denticles; 4) sternum IX with posterior margin straight.
</paragraph>
</subSubSection>
<subSubSection id="C31E656EFFF3FFDE33C1FE3F43BD763D" lastPageId="7" lastPageNumber="244" pageId="3" pageNumber="240" type="description">
<paragraph id="8BBB36E5FFF3FFDA33C1FE3F4137730E" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FE3F428670E3" bold="true" box="[199,499,474,501]" pageId="3" pageNumber="240">Description. Male imago.</emphasis>
Body length: 2.5–3.0 mm (n= 4); forewing: 
<quantity id="4CFC9B00FFF3FFDA30E2FE3E471B70E2" box="[996,1134,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="3" pageNumber="240" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
; foreleg: 
<quantity id="4CFC9B00FFF3FFDA37D0FE3F461570E2" box="[1238,1376,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="3" pageNumber="240" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
; hind leg: 
<quantity id="4CFC9B00FFF3FFDA33CEFE1B4227730E" box="[200,338,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="3" pageNumber="240" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
; cercus: 
<quantity id="4CFC9B00FFF3FFDA32B2FE1A414B730E" box="[436,574,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="7.949999999999999" metricValueMax="8.2" metricValueMin="7.7" pageId="3" pageNumber="240" unit="mm" value="7.949999999999999" valueMax="8.2" valueMin="7.7">7.7–8.2 mm</quantity>
.
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1FDC746767392" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FDC74266732B" bold="true" box="[199,275,546,572]" pageId="3" pageNumber="240">Ratios</emphasis>
. Foreleg 1.6–1.8× the length of hind leg. Ratio of first segment of the foretarsi length to 2nd:3rd:4th:5th = 0.1–0.2:0.3–0.4:0.3–0.5:0.5–0.8. 
<emphasis id="B970EAF7FFF3FFDA3124FDA241F97376" box="[546,652,583,609]" italics="true" pageId="3" pageNumber="240">Genitalia</emphasis>
: styliger plate length 2.4–2.8× width; forceps length 7.2–8.5× width at half length; distance between extreme lateral points of forceps bases to forceps length = 
<quantity id="4CFC9B00FFF3FFDA3773FD8E478A7392" box="[1141,1279,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="3" pageNumber="240" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 mm</quantity>
.
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1FD6B477E72FA" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FD6B423A73BF" bold="true" box="[199,335,654,680]" pageId="3" pageNumber="240">Coloration.</emphasis>
<emphasis id="B970EAF7FFF3FFDA325CFD6A42E273BE" box="[346,407,655,681]" italics="true" pageId="3" pageNumber="240">Head</emphasis>
: light brown shaded with dark in dorsal view (
<figureCitation id="133F2A60FFF3FFDA30C0FD6A475173BE" box="[966,1060,655,681]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2A</figureCitation>
); venter of head yellowish (
<figureCitation id="133F2A60FFF3FFDA3669FD6A43C873DA" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2D</figureCitation>
). 
<emphasis id="B970EAF7FFF3FFDA33D4FD56425A73DA" box="[210,303,691,717]" italics="true" pageId="3" pageNumber="240">Antenna</emphasis>
: scape and pedicel pale, flagellum grayish (
<figureCitation id="133F2A60FFF3FFDA3022FD5640DE73DA" box="[804,939,691,717]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2A–C</figureCitation>
). 
<emphasis id="B970EAF7FFF3FFDA30B9FD56476573DA" box="[959,1040,691,717]" italics="true" pageId="3" pageNumber="240">Thorax</emphasis>
. Prothorax light brown translucent shaded with black on lateral margin (
<figureCitation id="133F2A60FFF3FFDA3130FD3241C973E6" box="[566,700,727,753]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2A–C</figureCitation>
). Meso- and metanotum light brown shaded with gray on carinae; mesonotum light brown; pleurae and sterna light brown. 
<emphasis id="B970EAF7FFF3FFDA3027FD1E402E7203" box="[801,859,763,788]" italics="true" pageId="3" pageNumber="240">Wing</emphasis>
. forewings hyaline, veins translucent except C, Sc and Rs grayish. 
<emphasis id="B970EAF7FFF3FFDA324CFCFA420A722F" box="[330,383,799,824]" italics="true" pageId="3" pageNumber="240">Legs</emphasis>
. Foreleg dark brown shaded with black, middle and hind femur with subapical pale marks and apically blackish. 
<emphasis id="B970EAF7FFF3FFDA3266FCA642BF724A" box="[352,458,835,861]" italics="true" pageId="3" pageNumber="240">Abdomen</emphasis>
. Terga pale with abdominal segments I–II with a grayish band on medial area and darker laterally, segments III–VI grayish medially, segments VII–VIII yellowish medially and darker laterally, segment IX slightly grayish medially, segment X yellowish (
<figureCitation id="133F2A60FFF3FFDA31BBFC6E406D72B2" box="[701,792,907,933]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2A</figureCitation>
). Sterna pale with segments VII–IX yellowish (
<figureCitation id="133F2A60FFF3FFDA3630FC6E46FA72B2" box="[1334,1423,907,933]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2B</figureCitation>
). Pleura pale (
<figureCitation id="133F2A60FFF3FFDA322FFC4A42F172DE" box="[297,388,943,969]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2B</figureCitation>
). 
<emphasis id="B970EAF7FFF3FFDA329CFC4A417172DE" box="[410,516,943,969]" italics="true" pageId="3" pageNumber="240">Genitalia</emphasis>
. Sternum IX with strongly colored sclerites (
<figureCitation id="133F2A60FFF3FFDA3716FC4A47EF72DE" box="[1040,1178,943,969]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2B, C</figureCitation>
). Forceps dark brown, penis yellowish (
<figureCitation id="133F2A60FFF3FFDA3253FC3642AD72FA" box="[341,472,979,1005]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2C, E</figureCitation>
). Caudal filaments pale translucent. (
<figureCitation id="133F2A60FFF3FFDA307EFC36408A72FA" box="[888,1023,979,1005]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2A–C</figureCitation>
).
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1FC1247777426" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FC1242147506" bold="true" box="[199,353,1015,1041]" pageId="3" pageNumber="240">Morphology.</emphasis>
<emphasis id="B970EAF7FFF3FFDA326AFC1242BC7506" box="[364,457,1015,1041]" italics="true" pageId="3" pageNumber="240">Antenna</emphasis>
: Base of antennal flagellum dilated. 
<emphasis id="B970EAF7FFF3FFDA3082FC1240A07506" box="[900,981,1015,1041]" italics="true" pageId="3" pageNumber="240">Thorax</emphasis>
. Pronotum with lateral margins of the pronotum rounded and broader posteriorly (
<figureCitation id="133F2A60FFF3FFDA319FFBFE418D7522" box="[665,760,1051,1077]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2A</figureCitation>
); prosternal longitudinal ridges forming a triangle shaped structure, closed anteriorly and with straight lateral margins (
<figureCitation id="133F2A60FFF3FFDA3066FBDA40CB754E" box="[864,958,1087,1113]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Fig. 2D</figureCitation>
). 
<emphasis id="B970EAF7FFF3FFDA30D3FBDA474A754E" box="[981,1087,1087,1113]" italics="true" pageId="3" pageNumber="240">Abdomen</emphasis>
. Lateral filaments and fingerlike process on tergum II lacking. 
<emphasis id="B970EAF7FFF3FFDA3128FB8641ED756A" box="[558,664,1123,1149]" italics="true" pageId="3" pageNumber="240">Genitalia</emphasis>
. Styliger plate sclerotized, length 2.4–2.8× width, anterior margin almost straight and posterior margin straight, covering the basal 1/3 of penis; central sclerite elongate dome-shaped; apophyses long and thin, basolateral sclerites weakly discernible, lateral sclerites slender. Forceps long and slender, progressively tapering towards the apex, with margins slightly sinuosity and not fused to lateral margins of styliger plate. Penis cylindrical and apically rounded (with rectangular outline in ventral view), completely fused and not laterally projected; ventral surface with tubercles and groove (
<figureCitation id="133F2A60FFF3FFDA3049FAF240DB7426" box="[847,942,1303,1329]" captionStart="FIGURE 2" captionStartId="4.[152,255,1808,1832]" captionTargetBox="[151,1435,181,1785]" captionTargetId="figure-24@4.[151,1435,181,1785]" captionTargetPageId="4" captionText="FIGURE 2. Caenis urupa sp. nov. male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="3" pageNumber="240">Figs. 2E</figureCitation>
and 
<figureCitation id="133F2A60FFF3FFDA30E2FAF240817426" box="[996,1012,1303,1329]" captionStart="FIGURE 3" captionStartId="5.[152,255,1293,1317]" captionTargetBox="[190,1397,181,1269]" captionTargetId="figure-20@5.[190,1397,181,1269]" captionTargetPageId="5" captionText="FIGURE 3. Caenis urupa sp. nov. male imago, genitalia: A, ventral view (Scale = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309022" httpUri="https://zenodo.org/record/8309022/files/figure.png" pageId="3" pageNumber="240">3</figureCitation>
).
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1FADE40EA74F2" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FADE42037443" bold="true" box="[199,374,1338,1365]" pageId="3" pageNumber="240">Female imago.</emphasis>
Body length: 
<quantity id="4CFC9B00FFF3FFDA3110FADE41E87442" box="[534,669,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="3" pageNumber="240" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7–4.2 mm</quantity>
(n= 5); forewing: 3.0– 
<quantity id="4CFC9B00FFF3FFDA3099FADE408D7442" box="[927,1016,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="240" unit="mm" value="3.2">3.2 mm</quantity>
; foreleg: 
<quantity id="4CFC9B00FFF3FFDA3764FADE47997442" box="[1122,1260,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="3" pageNumber="240" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
; cercus: broken off. Thorax: same as males except mesonotum with more evident dark brown sutures, anterior margin brownish and grayish medially (
<figureCitation id="133F2A60FFF3FFDA3265FA6642CB748A" box="[355,446,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[152,255,1952,1976]" captionTargetBox="[151,1435,489,1928]" captionTargetId="figure-159@6.[151,1435,489,1928]" captionTargetPageId="6" captionText="FIGURE 4. Caenis urupa sp. nov. female imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view (Scale: Figs. 4A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309024" httpUri="https://zenodo.org/record/8309024/files/figure.png" pageId="3" pageNumber="240">Fig. 4A</figureCitation>
); katepisternum shaded with gray while mesopleural sclerites are light brown (
<figureCitation id="133F2A60FFF3FFDA3630FA6646FA748A" box="[1334,1423,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[152,255,1952,1976]" captionTargetBox="[151,1435,489,1928]" captionTargetId="figure-159@6.[151,1435,489,1928]" captionTargetPageId="6" captionText="FIGURE 4. Caenis urupa sp. nov. female imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view (Scale: Figs. 4A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309024" httpUri="https://zenodo.org/record/8309024/files/figure.png" pageId="3" pageNumber="240">Fig. 4B</figureCitation>
). Abdomen: same as males except abdomen with segments I grayish medially; abdominal sternum IX with posterior margin straight, not projected (
<figureCitation id="133F2A60FFF3FFDA32F7FA2E410274F2" box="[497,631,1483,1509]" captionStart="FIGURE 4" captionStartId="6.[152,255,1952,1976]" captionTargetBox="[151,1435,489,1928]" captionTargetId="figure-159@6.[151,1435,489,1928]" captionTargetPageId="6" captionText="FIGURE 4. Caenis urupa sp. nov. female imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view (Scale: Figs. 4A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309024" httpUri="https://zenodo.org/record/8309024/files/figure.png" pageId="3" pageNumber="240">Figs. 4A, C</figureCitation>
). Lateral filaments absent.
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1FA0A411E7762" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1FA0A4388771E" bold="true" box="[199,253,1519,1545]" pageId="3" pageNumber="240">Egg.</emphasis>
Length: 104–119 μm; width: 79–88 μm. Coloration yellowish. Oval-shaped with length 1.2–1.4 × width. Chorion surface with small pores (
<figureCitation id="133F2A60FFF3FFDA3111F9F6411A773A" box="[535,623,1554,1581]" captionStart="FIGURE 5" captionStartId="7.[152,255,1629,1653]" captionTargetBox="[151,1435,181,1606]" captionTargetId="figure-20@7.[151,1435,181,1606]" captionTargetPageId="7" captionText="FIGURE 5. Caenis urupa sp. nov. egg: A, geral view; B, structure of chorion; C, detail of micropyle; D–E, detail of polar cap. (Scale: Figs. 5A, D = 20 μm; 5B = 5 μm; Figs. 5C, E = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.8309026" httpUri="https://zenodo.org/record/8309026/files/figure.png" pageId="3" pageNumber="240">Fig. 5B</figureCitation>
). Micropyle linear with narrow and of moderate length micropylar canal, preceded by a scarcely visible oval sperm guide (
<figureCitation id="133F2A60FFF3FFDA31B8F9D240367746" box="[702,835,1590,1617]" captionStart="FIGURE 5" captionStartId="7.[152,255,1629,1653]" captionTargetBox="[151,1435,181,1606]" captionTargetId="figure-20@7.[151,1435,181,1606]" captionTargetPageId="7" captionText="FIGURE 5. Caenis urupa sp. nov. egg: A, geral view; B, structure of chorion; C, detail of micropyle; D–E, detail of polar cap. (Scale: Figs. 5A, D = 20 μm; 5B = 5 μm; Figs. 5C, E = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.8309026" httpUri="https://zenodo.org/record/8309026/files/figure.png" pageId="3" pageNumber="240">Figs. 5A, C</figureCitation>
). Two polar caps forming a spiral of fine threads with about 13 terminal knobs (
<figureCitation id="133F2A60FFF3FFDA32B0F9BE412B7762" box="[438,606,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[152,255,1629,1653]" captionTargetBox="[151,1435,181,1606]" captionTargetId="figure-20@7.[151,1435,181,1606]" captionTargetPageId="7" captionText="FIGURE 5. Caenis urupa sp. nov. egg: A, geral view; B, structure of chorion; C, detail of micropyle; D–E, detail of polar cap. (Scale: Figs. 5A, D = 20 μm; 5B = 5 μm; Figs. 5C, E = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.8309026" httpUri="https://zenodo.org/record/8309026/files/figure.png" pageId="3" pageNumber="240">Figs. 5A, C, D</figureCitation>
).
</paragraph>
<paragraph id="8BBB36E5FFF3FFDA33C1F99B41EB7612" blockId="3.[151,1437,150,1797]" pageId="3" pageNumber="240">
<emphasis id="B970EAF7FFF3FFDA33C1F99B412A778E" bold="true" box="[199,607,1662,1689]" pageId="3" pageNumber="240">Description. Mature male nymph.</emphasis>
Length: Body, 
<quantity id="4CFC9B00FFF3FFDA3016F99A401C778E" box="[784,873,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="240" unit="mm" value="3.7">3.7 mm</quantity>
, cercus, broken off. Ratios: Mouthparts. Width of maxillary palp segment I 1.7× width of segment II; length of maxillary palp segment I 1.6× length of segment II; length of maxillary palp segment I 0.9× length of segment III; length of labrum 0.4× its maximum width. Foreleg. Length of forefemur 3.1× its maximum width.
</paragraph>
<caption id="DF7B666DFFF4FFDD339EF8F5425E7666" ID-DOI="http://doi.org/10.5281/zenodo.8309018" ID-Zenodo-Dep="8309018" httpUri="https://zenodo.org/record/8309018/files/figure.png" pageId="4" pageNumber="241" startId="4.[152,255,1808,1832]" targetBox="[151,1435,181,1785]" targetPageId="4" targetType="figure">
<paragraph id="8BBB36E5FFF4FFDD339EF8F5425E7666" blockId="4.[151,1437,1808,1905]" pageId="4" pageNumber="241">
<emphasis id="B970EAF7FFF4FFDD339EF8F5426C763F" bold="true" box="[152,281,1808,1832]" pageId="4" pageNumber="241">FIGURE 2.</emphasis>
<taxonomicName id="4C044D66FFF4FFDD3226F8F442DF763F" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[288,426,1809,1832]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="241" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF4FFDD3226F8F442DF763F" box="[288,426,1809,1832]" italics="true" pageId="4" pageNumber="241">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFF4FFDD32B7F8F44177763E" bold="true" box="[433,514,1809,1833]" pageId="4" pageNumber="241">
<taxonomicNameLabel id="A243578CFFF4FFDD32B7F8F44177763E" box="[433,514,1809,1833]" pageId="4" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, detail of prosternum (ventral view); E, detail of genitalia (ventral view). (Scale: Figs. 2A–C = 1 mm; Fig. 2D = 0.5 mm; Fig. 2E = 0.1 mm).
</paragraph>
</caption>
<caption id="DF7B666DFFF5FFDC339EFAE847327432" ID-DOI="http://doi.org/10.5281/zenodo.8309022" ID-Zenodo-Dep="8309022" box="[152,1095,1293,1317]" httpUri="https://zenodo.org/record/8309022/files/figure.png" pageId="5" pageNumber="242" startId="5.[152,255,1293,1317]" targetBox="[190,1397,181,1269]" targetPageId="5" targetType="figure">
<paragraph id="8BBB36E5FFF5FFDC339EFAE847327432" blockId="5.[152,1095,1293,1317]" box="[152,1095,1293,1317]" pageId="5" pageNumber="242">
<emphasis id="B970EAF7FFF5FFDC339EFAE8426D7432" bold="true" box="[152,280,1293,1317]" pageId="5" pageNumber="242">FIGURE 3.</emphasis>
<taxonomicName id="4C044D66FFF5FFDC3219FAE842DD7432" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[287,424,1293,1317]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="242" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF5FFDC3219FAE842DD7432" box="[287,424,1293,1317]" italics="true" pageId="5" pageNumber="242">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFF5FFDC32A8FAE8428A7432" bold="true" box="[430,511,1293,1317]" pageId="5" pageNumber="242">
<taxonomicNameLabel id="A243578CFFF5FFDC32A8FAE8428A7432" box="[430,511,1293,1317]" pageId="5" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male imago, genitalia: A, ventral view (Scale = 0.1 mm).
</paragraph>
</caption>
<paragraph id="8BBB36E5FFF5FFDF33C1FA9241E270C6" blockId="5.[151,1437,1399,2038]" lastBlockId="6.[151,1437,151,465]" lastPageId="6" lastPageNumber="243" pageId="5" pageNumber="242">
<emphasis id="B970EAF7FFF5FFDC33C1FA9241697486" bold="true" box="[199,540,1399,1425]" pageId="5" pageNumber="242">Coloration and Morphology.</emphasis>
<emphasis id="B970EAF7FFF5FFDC3124FA92412A7486" box="[546,607,1399,1425]" italics="true" pageId="5" pageNumber="242">Head</emphasis>
: Brownish; brownish with black markings on posterior margin of occiput; also shaded black behind eyes; mouthparts light brown. 
<emphasis id="B970EAF7FFF5FFDC3008FA79401E74A1" box="[782,875,1436,1462]" italics="true" pageId="5" pageNumber="242">Antenna</emphasis>
: Flagellum brownish, pedicel paler (
<figureCitation id="133F2A60FFF5FFDC360FFA7E46E574A2" box="[1289,1424,1435,1461]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="5" pageNumber="242">Figs. 6A–C</figureCitation>
). Posterior margin of head with setae. Mouthparts: Labrum with lateral margins rounded, with long subapical setae over the dorsal surface (
<figureCitation id="133F2A60FFF5FFDC32A3FA06428A74EA" box="[421,511,1507,1533]" captionStart="FIGURE 7" captionStartId="9.[152,255,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@9.[151,1435,181,1678]" captionTargetPageId="9" captionText="FIGURE 7. Nymph of Caenis urupa sp. nov. A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="5" pageNumber="242">Fig. 7A</figureCitation>
); linguae of hypopharynx medially excavated (
<figureCitation id="133F2A60FFF5FFDC3709FA06471274EA" box="[1039,1127,1507,1533]" captionStart="FIGURE 7" captionStartId="9.[152,255,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@9.[151,1435,181,1678]" captionTargetPageId="9" captionText="FIGURE 7. Nymph of Caenis urupa sp. nov. A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="5" pageNumber="242">Fig. 7B</figureCitation>
); Mandibles with subapical setae varying in size on inner margin (
<figureCitation id="133F2A60FFF5FFDC3159F9E241927736" box="[607,743,1543,1570]" captionStart="FIGURE 7" captionStartId="9.[152,255,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@9.[151,1435,181,1678]" captionTargetPageId="9" captionText="FIGURE 7. Nymph of Caenis urupa sp. nov. A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="5" pageNumber="242">Figs. 7E, F</figureCitation>
); right mandible with outer margin with a dorsolateral row of long setae (
<figureCitation id="133F2A60FFF5FFDC323AF9CE42E67752" box="[316,403,1579,1605]" captionStart="FIGURE 7" captionStartId="9.[152,255,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@9.[151,1435,181,1678]" captionTargetPageId="9" captionText="FIGURE 7. Nymph of Caenis urupa sp. nov. A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="5" pageNumber="242">Fig. 7F</figureCitation>
). 
<emphasis id="B970EAF7FFF5FFDC32AFF9CE428F7752" box="[425,506,1579,1605]" italics="true" pageId="5" pageNumber="242">Thorax</emphasis>
: Nota brownish with brown sutures. Pronotum with anterior margin light brown, lateral zones shaded with black (
<figureCitation id="133F2A60FFF5FFDC3100F9AA4114777E" box="[518,609,1615,1641]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="5" pageNumber="242">Fig. 6A</figureCitation>
). Mesonotum with brownish macula anterior to wing bud bases, and with medial blackish lines (
<figureCitation id="133F2A60FFF5FFDC329EF9964286779A" box="[408,499,1651,1677]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="5" pageNumber="242">Fig. 6A</figureCitation>
). Thoracic sterna brownish (
<figureCitation id="133F2A60FFF5FFDC303FF99640E6779A" box="[825,915,1651,1677]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="5" pageNumber="242">Fig. 6C</figureCitation>
). Legs brownish, femora with blackish marks on subapical zones and paler basally on median and hind femora; tibiae and tarsi blackish on medial zones (
<figureCitation id="133F2A60FFF5FFDC3662F97243A877C2" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="5" pageNumber="242">Figs. 6A–C</figureCitation>
, 
<figureCitation id="133F2A60FFF5FFDC33EEF95E425A77C2" box="[232,303,1723,1749]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">8A–C</figureCitation>
). Fore coxa well-developed without projection, middle and hind coxa semicircular without projection (
<figureCitation id="133F2A60FFF5FFDC3399F93A425377EE" box="[159,294,1759,1785]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Figs. 8A–C</figureCitation>
). Fore coxa with long setae on outer margin, dorsal margin of fore femora with setae robust, varying in size, mostly on apical half, outer margin with simple and robust setae varying in size, setae simple varying in size on basal half of inner margin (
<figureCitation id="133F2A60FFF5FFDC32CEF8C241577656" box="[456,546,1831,1857]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Fig. 8A</figureCitation>
); dorsal margin of the median femora with robust setae, varying in size, mostly on middle region, outer margin with simple and robust setae varying in size on apical half (
<figureCitation id="133F2A60FFF5FFDC3793F8AE47997672" box="[1173,1260,1867,1893]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Fig. 8B</figureCitation>
); dorsal margin of the hind femora with robust setae, varying in size, mostly on middle region, outer margin with long and robust setae on apical half (
<figureCitation id="133F2A60FFF5FFDC327AF87642A176BA" box="[380,468,1939,1965]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Fig. 8C</figureCitation>
). Tibiae with robust setae on inner margins, varying in size (
<figureCitation id="133F2A60FFF5FFDC3775F876478C76BA" box="[1139,1273,1939,1965]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Figs. 8A–C</figureCitation>
). All tarsi with one row of robust setae on inner margin (
<figureCitation id="133F2A60FFF5FFDC3177F852418F76C6" box="[625,762,1975,2001]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Figs. 8A–C</figureCitation>
). Fore tarsal claws a row of small denticles basally, middle tarsal claws with six small denticles basally, hind tarsal claws with several small denticles (
<figureCitation id="133F2A60FFF5FFDC3792F83E466276E2" box="[1172,1303,2011,2037]" captionStart="FIGURE 8" captionStartId="10.[152,255,1245,1269]" captionTargetBox="[151,1435,181,1221]" captionTargetId="figure-24@10.[151,1435,181,1221]" captionTargetPageId="10" captionText="FIGURE 8. Nymph of Caenis urupa sp. nov. A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="5" pageNumber="242">Figs. 8D–F</figureCitation>
). 
<emphasis id="B970EAF7FFF5FFDC362DF83E46E076E2" box="[1323,1429,2011,2037]" italics="true" pageId="5" pageNumber="242">Abdomen</emphasis>
: Terga brownish; segments I–II with a grayish band on medial area and darker laterally; segments III–VI grayish medially, segments VII–VIII brownish medially and darker laterally, segment IX slightly grayish medially, segment X brownish (
<figureCitation id="133F2A60FFF6FFDF322BFF3A42FC71EE" box="[301,393,223,249]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">Fig. 9A</figureCitation>
). Sterna light brown, with only small grayish medial and lateral marks (
<figureCitation id="133F2A60FFF6FFDF37C4FF3A466971EE" box="[1218,1308,223,249]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="6" pageNumber="243">Fig. 6C</figureCitation>
), tergum II with small median projection on posterior margin, triangular with rounded tip. Operculate gills brownish, almost completely shaded with black and laterally with two blackish spots (
<figureCitation id="133F2A60FFF6FFDF3085FEC240937056" box="[899,998,295,321]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="6" pageNumber="243">Figs. 6A</figureCitation>
, 
<figureCitation id="133F2A60FFF6FFDF30F6FEC247617056" box="[1008,1044,295,321]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">9C</figureCitation>
). Operculate gills with submarginal ventral, microtrichia elongated shape and fringed apically, medial Y-ridge complete and well developed; dorsal surface smooth with long, simple setae (
<figureCitation id="133F2A60FFF6FFDF3152FE8A41D9709E" box="[596,684,367,393]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">Fig. 9C</figureCitation>
). Posterolateral projections on abdominal segments III–VI long and pointed (
<figureCitation id="133F2A60FFF6FFDF33FCFE76422170BA" box="[250,340,403,429]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">Fig. 9A</figureCitation>
). Sternum IX with posterior margin straight, laterally posteriorly with long simple setae (
<figureCitation id="133F2A60FFF6FFDF3631FE7646E570BA" box="[1335,1424,403,429]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">Fig. 9B</figureCitation>
). Caudal filaments brownish (
<figureCitation id="133F2A60FFF6FFDF32D2FE52412C70C6" box="[468,601,439,465]" captionStart="FIGURE 6" captionStartId="8.[152,255,1878,1902]" captionTargetBox="[151,1435,181,1854]" captionTargetId="figure-24@8.[151,1435,181,1854]" captionTargetPageId="8" captionText="FIGURE 6. Nymph of Caenis urupa sp. nov. A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="6" pageNumber="243">Figs. 6A–C</figureCitation>
, 
<figureCitation id="133F2A60FFF6FFDF3162FE5241FF70C6" box="[612,650,439,465]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="6" pageNumber="243">9A</figureCitation>
).
</paragraph>
<caption id="DF7B666DFFF6FFDF339EF84542CB76CB" ID-DOI="http://doi.org/10.5281/zenodo.8309024" ID-Zenodo-Dep="8309024" httpUri="https://zenodo.org/record/8309024/files/figure.png" pageId="6" pageNumber="243" startId="6.[152,255,1952,1976]" targetBox="[151,1435,489,1928]" targetPageId="6" targetType="figure">
<paragraph id="8BBB36E5FFF6FFDF339EF84542CB76CB" blockId="6.[151,1436,1952,2012]" pageId="6" pageNumber="243">
<emphasis id="B970EAF7FFF6FFDF339EF845426F76AF" bold="true" box="[152,282,1952,1976]" pageId="6" pageNumber="243">FIGURE 4.</emphasis>
<taxonomicName id="4C044D66FFF6FFDF3224F84542D876AF" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[290,429,1952,1976]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="243" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF6FFDF3224F84542D876AF" box="[290,429,1952,1976]" italics="true" pageId="6" pageNumber="243">Caenis urupa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A243578CFFF6FFDF32B3F845417D76A0" box="[437,520,1952,1976]" pageId="6" pageNumber="243" rank="species">
<emphasis id="B970EAF7FFF6FFDF32B3F845417776AF" bold="true" box="[437,514,1952,1976]" pageId="6" pageNumber="243">sp. nov</emphasis>
.
</taxonomicNameLabel>
female imago: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view (Scale: Figs. 4A–C = 1 mm).
</paragraph>
</caption>
<caption id="DF7B666DFFF7FFDE339EF9B8406E778D" ID-DOI="http://doi.org/10.5281/zenodo.8309026" ID-Zenodo-Dep="8309026" httpUri="https://zenodo.org/record/8309026/files/figure.png" pageId="7" pageNumber="244" startId="7.[152,255,1629,1653]" targetBox="[151,1435,181,1606]" targetPageId="7" targetType="figure">
<paragraph id="8BBB36E5FFF7FFDE339EF9B8406E778D" blockId="7.[151,1437,1629,1690]" pageId="7" pageNumber="244">
<emphasis id="B970EAF7FFF7FFDE339EF9B8426D7761" bold="true" box="[152,280,1629,1654]" pageId="7" pageNumber="244">FIGURE 5.</emphasis>
<taxonomicName id="4C044D66FFF7FFDE3218F9BB42D27762" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[286,423,1630,1653]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="244" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF7FFDE3218F9BB42D27762" box="[286,423,1630,1653]" italics="true" pageId="7" pageNumber="244">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFF7FFDE32ABF9BB42887761" bold="true" box="[429,509,1630,1654]" pageId="7" pageNumber="244">
<taxonomicNameLabel id="A243578CFFF7FFDE32ABF9BB42887761" box="[429,509,1630,1654]" pageId="7" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
egg: A, geral view; B, structure of chorion; C, detail of micropyle; D–E, detail of polar cap. (Scale: Figs. 5A, D = 20 μm; 5B = 5 μm; Figs. 5C, E = 10 μm).
</paragraph>
</caption>
<paragraph id="8BBB36E5FFF7FFDE33C1F90946E87611" blockId="7.[151,1437,1772,1834]" box="[199,1437,1772,1798]" pageId="7" pageNumber="244">
<emphasis id="B970EAF7FFF7FFDE33C1F90942BF7611" bold="true" box="[199,458,1772,1798]" pageId="7" pageNumber="244">Life cycle association.</emphasis>
Male genitalia extracted from mature nymph and compared with the male imagos (
<figureCitation id="133F2A60FFF7FFDE3669F90946E87611" box="[1391,1437,1772,1798]" captionStart="FIGURE 9" captionStartId="11.[152,255,1876,1900]" captionTargetBox="[151,1435,181,1852]" captionTargetId="figure-20@11.[151,1435,181,1852]" captionTargetPageId="11" captionText="FIGURE 9. Nymph of Caenis urupa sp. nov. A, detail of abdomen (dorsal view). B, sternum VIII–IX of mature male nymph, ventral view. C, opercular gill; D, gill IV. (Scale: Fig. 9A = 1 mm; Fig. 9B = 0.2 mm; Figs. 9C–D = 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.8309036" httpUri="https://zenodo.org/record/8309036/files/figure.png" pageId="7" pageNumber="244">Fig.</figureCitation>
</paragraph>
<paragraph id="8BBB36E5FFF7FFDE3391F8F543BD763D" blockId="7.[151,1437,1772,1834]" box="[151,200,1808,1834]" pageId="7" pageNumber="244">9B).</paragraph>
</subSubSection>
<caption id="DF7B666DFFF8FFD1339EF8B342047684" ID-DOI="http://doi.org/10.5281/zenodo.8309030" ID-Zenodo-Dep="8309030" httpUri="https://zenodo.org/record/8309030/files/figure.png" pageId="8" pageNumber="245" startId="8.[152,255,1878,1902]" targetBox="[151,1435,181,1854]" targetPageId="8" targetType="figure">
<paragraph id="8BBB36E5FFF8FFD1339EF8B342047684" blockId="8.[151,1437,1878,1939]" pageId="8" pageNumber="245">
<emphasis id="B970EAF7FFF8FFD1339EF8B3426D7679" bold="true" box="[152,280,1878,1902]" pageId="8" pageNumber="245">FIGURE 6.</emphasis>
Nymph of 
<taxonomicName id="4C044D66FFF8FFD1328DF8B241617679" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[395,532,1879,1902]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="245" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF8FFD1328DF8B241617679" box="[395,532,1879,1902]" italics="true" pageId="8" pageNumber="245">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFF8FFD1311CF8B2411F7678" bold="true" box="[538,618,1879,1903]" pageId="8" pageNumber="245">
<taxonomicNameLabel id="A243578CFFF8FFD1311CF8B2411F7678" box="[538,618,1879,1903]" pageId="8" pageNumber="245" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view. (Scale: Figs. 6A–C = 1 mm).
</paragraph>
</caption>
<caption id="DF7B666DFFF9FFD0339EF943464877F4" ID-DOI="http://doi.org/10.5281/zenodo.8309032" ID-Zenodo-Dep="8309032" httpUri="https://zenodo.org/record/8309032/files/figure.png" pageId="9" pageNumber="246" startId="9.[152,255,1702,1726]" targetBox="[151,1435,181,1678]" targetPageId="9" targetType="figure">
<paragraph id="8BBB36E5FFF9FFD0339EF943464877F4" blockId="9.[151,1436,1702,1763]" pageId="9" pageNumber="246">
<emphasis id="B970EAF7FFF9FFD0339EF943426D77A8" bold="true" box="[152,280,1702,1727]" pageId="9" pageNumber="246">FIGURE 7.</emphasis>
Nymph of 
<taxonomicName id="4C044D66FFF9FFD0328BF942416277A9" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[397,535,1703,1726]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="246" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFF9FFD0328BF942416277A9" box="[397,535,1703,1726]" italics="true" pageId="9" pageNumber="246">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFF9FFD0311BF942411B77A8" bold="true" box="[541,622,1703,1727]" pageId="9" pageNumber="246">
<taxonomicNameLabel id="A243578CFFF9FFD0311BF942411B77A8" box="[541,622,1703,1727]" pageId="9" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, labrum, dorsal view; B, hypopharynx, ventral view; C, labium, ventral view; D, maxilla, ventral view; E, left mandible, dorsal view; F, right mandible, dorsal view. (Scale: Figs. 7A–F = 0.1 mm).
</paragraph>
</caption>
<caption id="DF7B666DFFFAFFD3339EFB3840FC740D" ID-DOI="http://doi.org/10.5281/zenodo.8309034" ID-Zenodo-Dep="8309034" httpUri="https://zenodo.org/record/8309034/files/figure.png" pageId="10" pageNumber="247" startId="10.[152,255,1245,1269]" targetBox="[151,1435,181,1221]" targetPageId="10" targetType="figure">
<paragraph id="8BBB36E5FFFAFFD3339EFB3840FC740D" blockId="10.[151,1436,1245,1306]" pageId="10" pageNumber="247">
<emphasis id="B970EAF7FFFAFFD3339EFB38426D75E2" bold="true" box="[152,280,1245,1269]" pageId="10" pageNumber="247">FIGURE 8.</emphasis>
Nymph of 
<taxonomicName id="4C044D66FFFAFFD3328AFB3B416375E2" authority="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha, 2023" authorityName="Nascimento &amp; Cruz &amp; Lima &amp; Lima &amp; Hamada &amp; Ale-Rocha" authorityYear="2023" box="[396,534,1246,1269]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="247" phylum="Arthropoda" rank="species" species="urupa" status="sp. nov.">
<emphasis id="B970EAF7FFFAFFD3328AFB3B416375E2" box="[396,534,1246,1269]" italics="true" pageId="10" pageNumber="247">Caenis urupa</emphasis>
</taxonomicName>
<emphasis id="B970EAF7FFFAFFD3311DFB38411975E2" bold="true" box="[539,620,1245,1269]" pageId="10" pageNumber="247">
<taxonomicNameLabel id="A243578CFFFAFFD3311DFB38411975E2" box="[539,620,1245,1269]" pageId="10" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, foreleg; B, middle leg; C, hind leg; D, fore tarsal claw; E, middle tarsal claw; F, hind tarsal claw. (Scale: Figs. 8A–C = 0.5 mm; Figs. 8D–F = 0.05 mm).
</paragraph>
</caption>
<subSubSection id="C31E656EFFFAFFD333C1FA89421A7702" pageId="10" pageNumber="247" type="etymology">
<paragraph id="8BBB36E5FFFAFFD333C1FA89421A7702" blockId="10.[151,1436,1387,1558]" pageId="10" pageNumber="247">
<emphasis id="B970EAF7FFFAFFD333C1FA89423B7491" bold="true" box="[199,334,1388,1414]" pageId="10" pageNumber="247">Etymology.</emphasis>
The name of the species is in memory of the extinct native Urupá population that lived on the banks of the river formerly called Ji-Paraná. It has been considered extinct since the beginning of the 20th century, due to territorial disputes with other tribes, but above all to the strong colonizing presence that led to slave labor and the spread of various diseases, with their descendants being culturally eradicated around 1918. The specific epithet is a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>