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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3841.1.1" ID-GBIF-Dataset="07cca84d-e413-47d9-9ea5-5d24be4dce1b" ID-ISSN="1175-5326" ID-Zenodo-Dep="286746" ID-ZooBank="5FAADB3D-2359-453F-8BF5-1C2A33C6D178" checkinTime="1460460685218" checkinUser="plazi" docAuthor="Gerecke, Reinhard" docDate="2014" docId="03AD87B0FFA3147BFF46BAFEFAC38136" docLanguage="en" docName="zt03841p046.pdf" docOrigin="Zootaxa 3841 (1)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@4b05a9b1" docStyleName="zootaxa.2013.journal_article" docTitle="Aturus protzi Piersig 1901" docType="treatment" docVersion="6" lastPageNumber="19" masterDocId="FF94FFC8FFB31469FFD1BC4BFFE2830B" masterDocTitle="Studies on European species of the water mite family Aturidae Thor (Acari: Hydrachnidia)" masterLastPageNumber="46" masterPageNumber="1" pageNumber="17" updateTime="1677072350355" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Studies on European species of the water mite family Aturidae Thor (Acari: Hydrachnidia)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gerecke, Reinhard</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>3841</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>46</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3841.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">07cca84d-e413-47d9-9ea5-5d24be4dce1b</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">286746</mods:identifier>
<mods:identifier type="ZooBank">5FAADB3D-2359-453F-8BF5-1C2A33C6D178</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6144759" ID-GBIF-Taxon="127669134" ID-Zenodo-Dep="6144759" LSID="urn:lsid:plazi:treatment:03AD87B0FFA3147BFF46BAFEFAC38136" httpUri="http://treatment.plazi.org/id/03AD87B0FFA3147BFF46BAFEFAC38136" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection box="[151,477,1717,1743]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph blockId="16.[151,477,1717,1777]" box="[151,477,1717,1743]" pageId="16" pageNumber="17">
<heading bold="true" box="[151,477,1717,1743]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName authority="Piersig, 1901" authorityName="Piersig" authorityYear="1901" box="[151,477,1717,1743]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="protzi">
<emphasis bold="true" box="[151,477,1717,1743]" pageId="16" pageNumber="17">
<emphasis bold="true" box="[151,309,1717,1743]" italics="true" pageId="16" pageNumber="17">Aturus protzi</emphasis>
<bibRefCitation author="Piersig" box="[317,477,1717,1743]" pageId="16" pageNumber="17" refString="Piersig, R. (1901) Eine neue Aturus-Art aus dem Bohmisch-Bayerischen Walde. Zoologischer Anzeiger, 25 (660), 33 - 35." type="journal article" year="1901">
Piersig,
<number box="[418,477,1717,1743]" pageId="16" pageNumber="17" value="1901.0">1901</number>
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,230,1752,1777]" pageId="16" pageNumber="17" type="description">
<paragraph blockId="16.[151,477,1717,1777]" box="[151,230,1752,1777]" pageId="16" pageNumber="17">
<figureCitation box="[151,230,1752,1777]" captionStart="FIGURE 12" captionStartId="17.[151,250,2001,2023]" captionTargetBox="[421,1174,1344,1987]" captionTargetId="figure@17.[416,1188,1340,1990]" captionTargetPageId="17" captionText="FIGURE 12. Aturus protzi, male. A. IV-L- 4 / 5 posterior (right: IV-L- 4, left: IV-L- 5); B. III-L- 4 / 5 posterior. Scale bar = 100 µm." httpUri="https://zenodo.org/record/286757/files/figure.png" pageId="16" pageNumber="17">
Fig.
<number box="[202,230,1753,1777]" pageId="16" pageNumber="17" value="12.0">12</number>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="17" type="materials_examined">
<paragraph blockId="16.[151,1436,1824,2029]" pageId="16" pageNumber="17">
<emphasis bold="true" box="[151,387,1824,1849]" pageId="16" pageNumber="17">Material examined.</emphasis>
<materialsCitation collectionCode="NHMB" country="Czech Republic" location="Czech Republic" pageId="16" pageNumber="17" specimenCode="NHMB 4579, NHMB 4580" specimenCount="2" specimenCount-female="2">
<collectingCountry box="[396,583,1824,1849]" name="Czech Republic" pageId="16" pageNumber="17">Czech Republic</collectingCountry>
:
<taxonomicName box="[595,743,1825,1848]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[595,743,1825,1848]" italics="true" pageId="16" pageNumber="17">
<originalDetermination box="[595,743,1825,1848]" pageId="16" pageNumber="17">Aturus protzi</originalDetermination>
</emphasis>
</taxonomicName>
,
<specimenCode box="[761,913,1824,1849]" pageId="16" pageNumber="17">NHMB 4579</specimenCode>
,
<specimenCount box="[927,994,1825,1849]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
, one dissected, palps on
<specimenCode box="[1286,1436,1824,1849]" pageId="16" pageNumber="17">
NHMB
<number box="[1380,1436,1824,1849]" pageId="16" pageNumber="17" value="4580.0">4580</number>
</specimenCode>
&quot;
<taxonomicName authority="Piersig" authorityName="Piersig" box="[162,402,1860,1885]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[162,312,1861,1884]" italics="true" pageId="16" pageNumber="17">Aturus protzi</emphasis>
Piersig
</taxonomicName>
♀ &quot;Psg. S. Böhmerwald Mauth Bach unt. Bad
<number box="[970,1028,1861,1885]" pageId="16" pageNumber="17" value="1899.0">1899</number>
/
<number box="[1037,1093,1861,1885]" pageId="16" pageNumber="17" value="1901.0">1901</number>
&quot; &quot;Böhmerwald Coll. Piersig&quot; &quot;VIII/51,52&quot;. &quot;Typus&quot; (no type, see below; state: both specimens dried and squeezed);
</materialsCitation>
<materialsCitation collectionCode="NHMB" collectorName="Piers" country="Czech Republic" location="Czech Republic" pageId="16" pageNumber="17" specimenCode="NHMB 4579, NHMB 4580" specimenCount="3" specimenCount-male="3">
<collectionCode box="[1151,1235,1897,1920]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="16" pageNumber="17">NHMB</collectionCode>
&quot;XXIX/9,
<specimenCount box="[1364,1431,1897,1921]" pageId="16" pageNumber="17" type="male">3 ♂♂</specimenCount>
, without locality and date label &quot;coll.
<collectorName box="[576,638,1932,1957]" pageId="16" pageNumber="17">Piers</collectorName>
.&quot;;
</materialsCitation>
<materialsCitation pageId="16" pageNumber="17">
<collectingCountry box="[676,757,1933,1956]" name="France" pageId="16" pageNumber="17">France</collectingCountry>
:
<accessionNumber box="[769,907,1933,1957]" httpUri="http://www.uniprot.org/uniprot/SMF42928" pageId="16" pageNumber="17">
<collectionCode LSID="urn:lsid:biocol.org:col:34838" box="[769,825,1933,1957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="16" pageNumber="17">SMF</collectionCode>
42928
</accessionNumber>
,
<specimenCount box="[920,966,1933,1957]" pageId="16" pageNumber="17" type="generic">1 ♂</specimenCount>
,
<number box="[980,994,1933,1957]" pageId="16" pageNumber="17" value="1.0">1</number>
♀, &quot;
<collectingCountry box="[1048,1175,1932,1957]" name="France" pageId="16" pageNumber="17">Frankreich</collectingCountry>
, La puoade,
<date box="[1324,1436,1932,1957]" pageId="16" pageNumber="17" value="1948-08-16">
<number box="[1324,1374,1932,1957]" pageId="16" pageNumber="17" value="16.8">16.8</number>
.
<number box="[1378,1436,1932,1957]" pageId="16" pageNumber="17" value="1948.0">1948</number>
</date>
Angelier coll.
<number box="[315,377,1969,1993]" pageId="16" pageNumber="17" value="7040.0">7040</number>
&quot;;
</materialsCitation>
<materialsCitation pageId="16" pageNumber="17">
<collectingCountry box="[398,466,1968,1993]" name="Spain" pageId="16" pageNumber="17">Spain</collectingCountry>
:
<collectionCode box="[478,559,1969,1993]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="16" pageNumber="17">SMNH</collectionCode>
<collectionCode box="[567,718,1969,1993]" pageId="16" pageNumber="17">NHRS-GULI</collectionCode>
0 0 0 0 0 2775, 2 ♂♂ &quot;
<number box="[949,1008,1968,1993]" pageId="16" pageNumber="17" value="4515.0">4515</number>
<taxonomicName authority="Piers." authorityName="Piers." box="[1016,1236,1968,1993]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[1016,1164,1969,1992]" italics="true" pageId="16" pageNumber="17">Aturus protzi</emphasis>
Piers.
</taxonomicName>
♂
<collectingCountry box="[1273,1366,1968,1993]" name="Spain" pageId="16" pageNumber="17">Spanien</collectingCountry>
, Bäck på Sierra Guadarrama
<date box="[404,516,2004,2029]" pageId="16" pageNumber="17" value="1935-10-09">
<number box="[404,455,2004,2029]" pageId="16" pageNumber="17" value="10.9">10.9</number>
.
<number box="[458,516,2004,2029]" pageId="16" pageNumber="17" value="1935.0">1935</number>
</date>
Leg.
<collectingCountry box="[580,603,2005,2029]" name="Iceland" pageId="16" pageNumber="17">O</collectingCountry>
. Lundblad.&quot;.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="discussion">
<paragraph blockId="17.[151,1437,151,1286]" pageId="17" pageNumber="18">
<emphasis bold="true" box="[199,315,151,176]" pageId="17" pageNumber="18">Remarks.</emphasis>
This species was first described from an unclear number of males and we cannot exclude that the specimens on slide
<collectionCode box="[379,463,187,210]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="17" pageNumber="18">NHMB</collectionCode>
<collectionCode box="[473,545,187,211]" pageId="17" pageNumber="18">XXIX</collectionCode>
/
<number box="[554,579,187,211]" pageId="17" pageNumber="18" value="19.0">19</number>
represent
<typeStatus box="[702,800,187,211]" pageId="17" pageNumber="18">syntypes</typeStatus>
of
<taxonomicName box="[843,941,187,210]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[843,941,187,210]" italics="true" pageId="17" pageNumber="18">A. protzi</emphasis>
</taxonomicName>
. Instead, the two females on slide
<collectionCode box="[1352,1436,187,210]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="17" pageNumber="18">NHMB</collectionCode>
<number box="[151,207,221,246]" pageId="17" pageNumber="18" value="4579.0">4579</number>
derive from the area of the
<typeStatus box="[532,580,222,246]" pageId="17" pageNumber="18">type</typeStatus>
locality, but the &quot;
<typeStatus box="[784,857,222,246]" pageId="17" pageNumber="18">Typus</typeStatus>
&quot; label is obviously incorrect. Interestingly, in over hundred years since the first description, no attempt has been made to describe the female sex. The species was in most cases found together with other species of the genus, and no character states are known in both sexes that could be indicators of conspecifity.
</paragraph>
<paragraph blockId="17.[151,1437,151,1286]" pageId="17" pageNumber="18">
Following the original description (and omitting some details without significance for species definition in
<taxonomicName box="[151,226,395,418]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis box="[151,226,395,418]" italics="true" pageId="17" pageNumber="18">Aturus</emphasis>
</taxonomicName>
), male
<taxonomicName box="[317,415,395,418]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[317,415,395,418]" italics="true" pageId="17" pageNumber="18">A. protzi</emphasis>
</taxonomicName>
are characterized by the following important features: Idiosoma with subparallel lateral margins, posteriorly equally rounded, without projections flanking the gonopore. Posterior part of dorsal shield flattened, in the transition area to ventral field on each side an anteriorly-directed bowed line of fine, denselyarranged setae embracing a subtriangular field covered by fine hair-like setae. Dgl-
<number box="[1098,1107,498,523]" pageId="17" pageNumber="18" value="1.0">1</number>
with strong bifurcated setae, all other Dgl with simple, long setae. Lgl-
<number box="[627,666,533,558]" pageId="17" pageNumber="18" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
close to each other, but not in touch, in the dorsal furrow, Lgl-
<number box="[1379,1388,533,558]" pageId="17" pageNumber="18" value="1.0">1</number>
and –
<number box="[166,179,568,592]" pageId="17" pageNumber="18" value="2.0">2</number>
with strong, bifurcated setae, Lgl–
<number box="[578,590,567,592]" pageId="17" pageNumber="18" value="3.0">3</number>
with a long, but simple seta. Posterolateral and posterior idiosoma margin with numerous long and fine setae, none of these with &quot;onion-like&quot; basal inflation; the gonopore flanked by two pairs of enlarged, club-shaped setae. Genital field with ca.
<number box="[816,830,638,662]" pageId="17" pageNumber="18" value="9.0">9</number>
pairs of Ac, three of them flanking the gonopore and directed ventrocaudally. Leg claws with dorsal and ventral clawlet. III-L-
<number box="[988,1000,671,696]" pageId="17" pageNumber="18" value="5.0">5</number>
with numerous long ventral setae and one particularly elongated, distally spiralled &quot;whip seta&quot; in terminal position. IV-L-
<number box="[1106,1119,707,731]" pageId="17" pageNumber="18" value="4.0">4</number>
distally strongly thickened, bearing here three long and strongly enlarged sword setae, four long, slender, pointed setae and, slightly set off from distal margin, two curved setae with a distally denticulate margin. IV-L-
<number box="[1024,1036,775,800]" pageId="17" pageNumber="18" value="5.0">5</number>
curved, proximally with an obtuseangled projection surrounded by
<number box="[528,569,811,835]" pageId="17" pageNumber="18" value="3.5" valueMax="4.0" valueMin="3.0">3–4</number>
strong, but short &quot;elk horn&quot; setae, which are distally truncated and strongly denticulate, and a line of equally-distanced slender and simple setae; distal margin bearing, in addition to several simple setae, one irregularly curved long and flattened seta branched into several fine tips.
</paragraph>
<paragraph blockId="17.[151,1437,151,1286]" pageId="17" pageNumber="18">
The specimens from
<collectionCode box="[433,514,915,939]" httpUri="http://grbio.org/cool/8z6v-f8is" name="Saskatchewan Museum of Natural History" pageId="17" pageNumber="18">SMNH</collectionCode>
agree well with this description and allow some additional observations. Idiosoma L/W
<number box="[212,310,949,974]" pageId="17" pageNumber="18" value="355.0" valueMax="360.0" valueMin="350.0">350–360</number>
/
<number box="[319,418,949,974]" pageId="17" pageNumber="18" value="295.0" valueMax="300.0" valueMin="290.0">290–300</number>
, dorsal shield L/W about
<number box="[719,762,950,974]" pageId="17" pageNumber="18" value="270.0">270</number>
/
<number box="[771,814,950,974]" pageId="17" pageNumber="18" value="270.0">270</number>
. The fact that
<bibRefCitation author="Piersig" box="[983,1144,949,974]" pageId="17" pageNumber="18" refString="Piersig, R. (1901) Eine neue Aturus-Art aus dem Bohmisch-Bayerischen Walde. Zoologischer Anzeiger, 25 (660), 33 - 35." type="journal article" year="1901">
Piersig (
<number box="[1078,1136,950,974]" pageId="17" pageNumber="18" value="1901.0">1901</number>
)
</bibRefCitation>
did not mention Lgl-
<number box="[1394,1408,949,974]" pageId="17" pageNumber="18" value="4.0">4</number>
is probably explained by the fusion Lgl-
<number box="[585,597,983,1008]" pageId="17" pageNumber="18" value="3.0">3</number>
+
<number box="[614,629,983,1008]" pageId="17" pageNumber="18" value="4.0">4</number>
, escaped to his attention. The fusion is documented by the presence of two gland pores and, in addition to the well-developed seta Lgl-
<number box="[891,903,1018,1043]" pageId="17" pageNumber="18" value="3.0">3</number>
a very fine hair like seta on the posteriormost platelet in the dorsal furrow. The setae arranged along the posterolateral and posterior margins are particularly fine and dense, anteriorly they extend to the upwardly-bent flanks of ventral shield which narrows the dorsal furrow posterior to Lgl-
<number box="[342,354,1122,1147]" pageId="17" pageNumber="18" value="3.0">3</number>
+
<number box="[371,386,1122,1147]" pageId="17" pageNumber="18" value="4.0">4</number>
, at the posterior margin they are gradually shortened from the lateral edge to the gonopore area. III-L-
<number box="[275,288,1158,1182]" pageId="17" pageNumber="18" value="4.0">4</number>
bears no particularly elongated distal setae. The setae number on III-L-
<number box="[1090,1102,1157,1182]" pageId="17" pageNumber="18" value="5.0">5</number>
is about
<number box="[1203,1234,1157,1182]" pageId="17" pageNumber="18" value="25.0">25</number>
. Most of them are strong, but simple and arranged in a dense longish group over the distal
<number box="[954,968,1192,1216]" pageId="17" pageNumber="18" value="2.0">2</number>
/
<number box="[978,990,1192,1216]" pageId="17" pageNumber="18" value="3.0">3</number>
of the ventral margin, but in addition to the particularly elongated distal whip seta, three further particular setae are found in the proximal part of the segment (arranged together on a flat posteroventral projection, slightly thickened and distally curved).
</paragraph>
<caption box="[151,1436,2001,2023]" httpUri="https://zenodo.org/record/286757/files/figure.png" pageId="17" pageNumber="18" targetBox="[421,1174,1344,1987]" targetPageId="17">
<paragraph blockId="17.[151,1436,2001,2023]" box="[151,1436,2001,2023]" pageId="17" pageNumber="18">
<emphasis bold="true" box="[151,287,2001,2023]" pageId="17" pageNumber="18">FIGURE 12.</emphasis>
<taxonomicName box="[292,424,2002,2023]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[292,424,2002,2023]" italics="true" pageId="17" pageNumber="18">Aturus protzi</emphasis>
</taxonomicName>
, male. A. IV-L-4/5 posterior (right: IV-L-4, left: IV-L-5); B. III-L-4/5 posterior. Scale bar = 100 µm.
</paragraph>
</caption>
<paragraph blockId="18.[151,1436,151,573]" pageId="18" pageNumber="19">
<collectingCountry box="[199,230,152,176]" name="American Samoa" pageId="18" pageNumber="19">As</collectingCountry>
documented by the widely agreeing descriptions,
<taxonomicName box="[851,954,152,175]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[851,954,152,175]" italics="true" pageId="18" pageNumber="19">A. protzi</emphasis>
</taxonomicName>
is very similar to
<taxonomicName box="[1198,1370,152,176]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis box="[1198,1370,152,176]" italics="true" pageId="18" pageNumber="19">A. intermedius</emphasis>
</taxonomicName>
. Our observations on the latter (see above) show that several previously used character states are variable and thus of restricted taxonomic significance for defining
<taxonomicName box="[668,763,224,247]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[668,763,224,247]" italics="true" pageId="18" pageNumber="19">A. protzi</emphasis>
</taxonomicName>
. This regards mostly the relative distance of Lgl-
<number box="[1319,1328,223,248]" pageId="18" pageNumber="19" value="1.0">1</number>
/-
<number box="[1346,1361,223,248]" pageId="18" pageNumber="19" value="2.0">2</number>
, shape of proximoventral margin of IV-L-
<number box="[547,559,260,285]" pageId="18" pageNumber="19" value="5.0">5</number>
(appearing bluntly pointed or equally convex, depending on little changes of its position) and number of whip-setae on III-L-
<number box="[691,703,295,320]" pageId="18" pageNumber="19" value="5.0">5</number>
(occasionally the second one is shortened could be interpreted as a normal seta); but possibly also the bi-/trifurcation of setae Dgl-
<number box="[935,944,332,357]" pageId="18" pageNumber="19" value="1.0">1</number>
and even the character most stressed for distinguishing the two sister species, the shape of the basal setae on IV-L-
<number box="[1031,1043,367,392]" pageId="18" pageNumber="19" value="5.0">5</number>
(shortened &quot;leaf-like&quot; setae in
<taxonomicName class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis italics="true" pageId="18" pageNumber="19">A. intermedius</emphasis>
</taxonomicName>
may take the &quot;elk horn&quot; aspect typical for
<taxonomicName box="[762,857,404,428]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[762,857,404,428]" italics="true" pageId="18" pageNumber="19">A. protzi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph blockId="18.[151,1436,151,573]" pageId="18" pageNumber="19">
The following, often not easily observed, details allow to distinguish
<taxonomicName box="[1039,1140,440,463]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="protzi">
<emphasis box="[1039,1140,440,463]" italics="true" pageId="18" pageNumber="19">A. protzi</emphasis>
</taxonomicName>
from
<taxonomicName box="[1222,1391,440,464]" class="Arachnida" family="Aturidae" genus="Aturus" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis box="[1222,1391,440,464]" italics="true" pageId="18" pageNumber="19">A. intermedius</emphasis>
</taxonomicName>
(in parentheses): (
<number box="[318,330,476,500]" pageId="18" pageNumber="19" value="1.0">1</number>
) seta Lgl-
<number box="[453,465,476,501]" pageId="18" pageNumber="19" value="3.0">3</number>
as long as setae of Dgl-
<number box="[749,763,476,501]" pageId="18" pageNumber="19" value="2.0">2</number>
/
<number box="[771,783,476,501]" pageId="18" pageNumber="19" value="3.0">3</number>
(short, hair-like); (
<number box="[1003,1018,476,500]" pageId="18" pageNumber="19" value="2.0">2</number>
) III-L-
<number box="[1101,1114,476,500]" pageId="18" pageNumber="19" value="4.0">4</number>
without (with) strong distal seta; (
<number box="[219,233,512,536]" pageId="18" pageNumber="19" value="3.0">3</number>
) IV-L-
<number box="[315,329,512,536]" pageId="18" pageNumber="19" value="4.0">4</number>
, apart from the sword setae, with two or three curved, branched setae (with simple setae only); (
<number box="[1412,1427,512,536]" pageId="18" pageNumber="19" value="4.0">4</number>
) IV-L-
<number box="[216,228,548,573]" pageId="18" pageNumber="19" value="5.0">5</number>
basal setae stout, &quot;elk-horn-shaped&quot; (longish, &quot;leaf-like&quot;, intermediate character states possible).
</paragraph>
</subSubSection>
</treatment>
</document>
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