treatments-xml/data/03/AD/87/03AD87A83800FFB51891318AFC66FC61.xml

<document id="D7A6FBDE9215DDC5E5CF429CB8EEDDE7" ID-CLB-Dataset="46061" ID-DOI="10.11646/zootaxa.3785.2.4" ID-GBIF-Dataset="9fd64151-f80d-4835-8771-1178cda25236" ID-ISSN="1175-5326" ID-Zenodo-Dep="251956" ID-ZooBank="B18DBB73-3F51-4FA5-88DC-E88E80B0EB9E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460406043830" checkinUser="plazi" docAuthor="Velasco, Julián A. &amp; Hurtado-Gómez, Juan Pablo" docDate="2014" docId="03AD87A83800FFB51891318AFC66FC61" docLanguage="en" docName="zt03785p216.pdf" docOrigin="Zootaxa 3785 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anolis limon Velasco &amp; Hurtado-Gómez, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="212" masterDocId="FF94FFD03802FFBE1806374DFFACFFE1" masterDocTitle="A new green anole lizard of the &quot; Dactyloa &quot; clade (Squamata: Dactyloidae) from the Magdalena river valley of Colombia" masterLastPageNumber="216" masterPageNumber="201" pageNumber="203" updateTime="1698299808622" updateUser="ExternalLinkService">
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<mods:title id="4A5CF8816255AC5DD90C330B9938E5CB">A new green anole lizard of the &quot; Dactyloa &quot; clade (Squamata: Dactyloidae) from the Magdalena river valley of Colombia</mods:title>
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<mods:namePart id="46D96AAA28A9708F3F03D81A3DFD8CE3">Velasco, Julián A.</mods:namePart>
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<mods:namePart id="825392CE8D2A540D42F2B45C560C3751">Hurtado-Gómez, Juan Pablo</mods:namePart>
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<subSubSection id="C31E65353800FFBC1891318AFEE0F8E5" pageId="2" pageNumber="203" type="nomenclature">
<paragraph id="8BBB36BE3800FFBC1891318AFE20F901" blockId="2.[151,396,1735,1797]" box="[151,396,1735,1760]" pageId="2" pageNumber="203">
<heading id="D0F381D23800FFBC1891318AFE20F901" bold="true" box="[151,396,1735,1760]" fontSize="10" level="2" pageId="2" pageNumber="203" reason="2">
<emphasis id="B970EAAC3800FFBC1891318AFE20F901" bold="true" box="[151,396,1735,1760]" pageId="2" pageNumber="203">
<taxonomicName id="4C044D3D3800FFBC1891318AFE87F901" box="[151,299,1735,1760]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="203" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3800FFBC1891318AFE87F901" bold="true" box="[151,299,1735,1760]" italics="true" pageId="2" pageNumber="203">Anolis limon</emphasis>
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<taxonomicNameLabel id="A24357D73800FFBC1934318AFE20F901" box="[306,396,1735,1760]" pageId="2" pageNumber="203" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BBB36BE3800FFBC189131A1FEE0F8E5" blockId="2.[151,396,1735,1797]" box="[151,332,1772,1797]" pageId="2" pageNumber="203">
(
<figureCitation id="133F2A3B3800FFBC189831A1FF52F8E5" box="[158,254,1772,1797]" captionStart="FIGURE 1" captionStartId="3.[151,250,1513,1535]" captionTargetBox="[159,1426,301,1491]" captionTargetId="figure@3.[159,1427,301,1491]" captionTargetPageId="3" captionText="FIGURE 1. Anolis limon sp. nov. Top: male holotype (MHUA-R 11760); Bottom: female paratype (MHUA-R 10939)." httpUri="https://zenodo.org/record/251957/files/figure.png" pageId="2" pageNumber="203">Figure 1</figureCitation>
and 
<figureCitation id="133F2A3B3800FFBC193231A1FEEFF8E5" box="[308,323,1772,1796]" captionStart="FIGURE 2" captionStartId="4.[151,250,1957,1979]" captionTargetBox="[394,1185,193,1932]" captionTargetId="figure@4.[394,1185,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dewlap of Anolis limon sp. nov. and closest putative relatives. Top: Anolis limon sp. nov. (Photo: JPHG). Middle: Anolis chocorum (Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: Anolis ibanezi (Photo: Steven Poe)." httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="2" pageNumber="203">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31E65353800FFBD18913079FAA6FF19" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203" type="materials_examined">
<paragraph id="8BBB36BE3800FFBC18913079FE02F875" blockId="2.[151,1436,1844,2013]" pageId="2" pageNumber="203">
<emphasis id="B970EAAC3800FFBC18913079FEA7F8AC" bold="true" box="[151,267,1844,1869]" pageId="2" pageNumber="203">
<typeStatus id="54BF881C3800FFBC18913079FEAAF8AC" box="[151,262,1844,1869]" pageId="2" pageNumber="203" type="holotype">Holotype</typeStatus>
.
</emphasis>
MHUA-R 11760; adult male, from department of Antioquia, Gómez Plata municipality, Vereda La Clara, Hacienda Vegas de la Clara; 
<geoCoordinate id="EE3050793800FFBC1A2A301AFD05F891" box="[556,681,1879,1904]" direction="north" orientation="latitude" pageId="2" pageNumber="203" precision="15" value="6.581389">6°34’53”N</geoCoordinate>
, 
<geoCoordinate id="EE3050793800FFBC1AB1301AFCE5F891" box="[695,841,1879,1904]" direction="west" orientation="longitude" pageId="2" pageNumber="203" precision="15" value="-75.19528">75°11’43’’W</geoCoordinate>
(
<quantity id="4CFC9B5B3800FFBC1B5C3015FC1EF891" box="[858,946,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.093" pageId="2" pageNumber="203" unit="m" value="1093.0">1093 m</quantity>
) collected by Wilmar Munera on 
<date id="FFBA107E3800FFBC1D33301AFF63F875" pageId="2" pageNumber="203" value="2008-04-22">April 22, 2008</date>
(
<figureCitation id="133F2A3B3800FFBC18DB3031FE91F875" box="[221,317,1916,1941]" captionStart="FIGURE 1" captionStartId="3.[151,250,1513,1535]" captionTargetBox="[159,1426,301,1491]" captionTargetId="figure@3.[159,1427,301,1491]" captionTargetPageId="3" captionText="FIGURE 1. Anolis limon sp. nov. Top: male holotype (MHUA-R 11760); Bottom: female paratype (MHUA-R 10939)." httpUri="https://zenodo.org/record/251957/files/figure.png" pageId="2" pageNumber="203">Figure 1</figureCitation>
, 
<figureCitation id="133F2A3B3800FFBC194D3031FEF5F875" box="[331,345,1916,1940]" captionStart="FIGURE 2" captionStartId="4.[151,250,1957,1979]" captionTargetBox="[394,1185,193,1932]" captionTargetId="figure@4.[394,1185,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dewlap of Anolis limon sp. nov. and closest putative relatives. Top: Anolis limon sp. nov. (Photo: JPHG). Middle: Anolis chocorum (Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: Anolis ibanezi (Photo: Steven Poe)." httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="2" pageNumber="203">2</figureCitation>
and 
<figureCitation id="133F2A3B3800FFBC19963031FE0CF875" box="[400,416,1916,1940]" captionStart="FIGURE 3" captionStartId="5.[151,250,1074,1096]" captionTargetBox="[151,1435,193,1051]" captionTargetId="figure@5.[151,1436,193,1053]" captionTargetPageId="5" captionText="FIGURE 3. Holotype of Anolis limon sp. nov. (MHUA-R 11760, adult male). A) lateral, B) dorsal, and C) ventral view of head; D) detail of dorsal scales at mid-dorsum of body, arrow indicate the middorsal scales; E) detail of ventral scales at middorsum of body; F) detail of flank scales at mid-dorsum of body; G) ventral view of cloacal region showing enlarged postcloacal scales." httpUri="https://zenodo.org/record/251959/files/figure.png" pageId="2" pageNumber="203">3</figureCitation>
).
</paragraph>
<paragraph id="8BBB36BE3800FFBD18C130EDFAA6FF19" blockId="2.[151,1436,1844,2013]" lastBlockId="3.[151,1436,151,248]" lastPageId="3" lastPageNumber="204" pageId="2" pageNumber="203">
<typeStatus id="54BF881C3800FFBC18C130EDFE93F858" box="[199,319,1952,1977]" pageId="2" pageNumber="203" type="paratype">
<emphasis id="B970EAAC3800FFBC18C130EDFE93F858" bold="true" box="[199,319,1952,1977]" pageId="2" pageNumber="203">Paratypes</emphasis>
</typeStatus>
(all from 
<collectingCountry id="F313762E3800FFBC19B930EDFD9FF858" box="[447,563,1952,1977]" name="Colombia" pageId="2" pageNumber="203">Colombia</collectingCountry>
). MHUA-R 10053, 11267 collected in department of Antioquia, Yolombo municipality, Normandia, 
<geoCoordinate id="EE3050793800FFBC19CC3089FD9FF83C" box="[458,563,1988,2013]" direction="north" orientation="latitude" pageId="2" pageNumber="203" precision="15" value="6.735">6°44'6''N</geoCoordinate>
, 
<geoCoordinate id="EE3050793800FFBC1A433089FD6CF83C" box="[581,704,1988,2013]" direction="west" orientation="longitude" pageId="2" pageNumber="203" precision="15" value="-75.08083">75°4'51''W</geoCoordinate>
(
<quantity id="4CFC9B5B3800FFBC1AD03089FC85F83D" box="[726,809,1988,2013]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="2" pageNumber="203" unit="m" value="935.0">935 m</quantity>
); MHUA-R 10939 department of Antioquia, Amalfi municipality, vereda San Ignacio, 
<geoCoordinate id="EE3050793801FFBD1A2637DAFD3BFF51" box="[544,663,151,176]" direction="north" orientation="latitude" pageId="3" pageNumber="204" precision="15" value="6.9119444">6°54'43''N</geoCoordinate>
, 
<geoCoordinate id="EE3050793801FFBD1AA337DAFC8CFF51" box="[677,800,151,176]" direction="west" orientation="longitude" pageId="3" pageNumber="204" precision="15" value="-75.07833">75°4'42''W</geoCoordinate>
(
<quantity id="4CFC9B5B3801FFBD1B3737D5FC22FF51" box="[817,910,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="m" value="1000.0">1000 m</quantity>
); MHUA-R 11020 collected in department of Caldas, Norcasia municipality, vereda El Tigre, 
<geoCoordinate id="EE3050793801FFBD1AB137F1FCEFFF34" box="[695,835,188,213]" direction="north" orientation="latitude" pageId="3" pageNumber="204" precision="1" value="5.5590835">5°33'32.7''N</geoCoordinate>
, 
<geoCoordinate id="EE3050793801FFBD1B5637F1FC4CFF34" box="[848,992,188,213]" direction="west" orientation="longitude" pageId="3" pageNumber="204" precision="1" value="-74.868225">74°52'5,6''W</geoCoordinate>
(
<quantity id="4CFC9B5B3801FFBD1BE937F1FB90FF35" box="[1007,1084,188,213]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="204" unit="m" value="520.0">520 m</quantity>
); MHUA-R 11248 collected in department of Antioquia, Anor municipality, vereda El Retiro 
<geoCoordinate id="EE3050793801FFBD1B573792FC47FF19" box="[849,1003,223,248]" direction="north" orientation="latitude" pageId="3" pageNumber="204" precision="1" value="6.812961">6°48'46.66''N</geoCoordinate>
, 
<geoCoordinate id="EE3050793801FFBD1BF03792FB38FF19" box="[1014,1172,223,248]" direction="west" orientation="longitude" pageId="3" pageNumber="204" precision="1" value="-75.05954">75°3'34,34''W</geoCoordinate>
(
<quantity id="4CFC9B5B3801FFBD1CA53792FB51FF19" box="[1187,1277,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.538" pageId="3" pageNumber="204" unit="m" value="1538.0">1538 m</quantity>
).
</paragraph>
</subSubSection>
<caption id="DF7B66363801FFBD189132A4FAFDFA1E" box="[151,1361,1513,1535]" httpUri="https://zenodo.org/record/251957/files/figure.png" pageId="3" pageNumber="204" targetBox="[159,1426,301,1491]" targetPageId="3">
<paragraph id="8BBB36BE3801FFBD189132A4FAFDFA1E" blockId="3.[151,1361,1513,1535]" box="[151,1361,1513,1535]" pageId="3" pageNumber="204">
<emphasis id="B970EAAC3801FFBD189132A4FEBFFA1F" bold="true" box="[151,275,1513,1535]" pageId="3" pageNumber="204">FIGURE 1.</emphasis>
<taxonomicName id="4C044D3D3801FFBD191F32A4FE36FA1E" box="[281,410,1513,1535]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3801FFBD191F32A4FE36FA1E" box="[281,410,1513,1535]" italics="true" pageId="3" pageNumber="204">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73801FFBD19A632A4FE5CFA1F" box="[416,496,1513,1534]" pageId="3" pageNumber="204" rank="species">
<emphasis id="B970EAAC3801FFBD19A632A4FE5CFA1F" bold="true" box="[416,496,1513,1534]" pageId="3" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
Top: male holotype (MHUA-R 11760); Bottom: female paratype (MHUA-R 10939).
</paragraph>
</caption>
<subSubSection id="C31E65353801FFBD18C1316BFBB0F94A" pageId="3" pageNumber="204" type="diagnosis">
<paragraph id="8BBB36BE3801FFBD18C1316BFBB0F94A" blockId="3.[151,1436,1574,2031]" pageId="3" pageNumber="204">
<emphasis id="B970EAAC3801FFBD18C1316BFEEDF9DE" bold="true" box="[199,321,1574,1599]" pageId="3" pageNumber="204">Diagnosis.</emphasis>
An 
<taxonomicName id="4C044D3D3801FFBD1972316BFE10F9DF" box="[372,444,1574,1598]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="genus">
<emphasis id="B970EAAC3801FFBD1972316BFE10F9DF" box="[372,444,1574,1598]" italics="true" pageId="3" pageNumber="204">Anolis</emphasis>
</taxonomicName>
of the “ 
<taxonomicName id="4C044D3D3801FFBD1A1E316BFD2EF9DE" box="[536,642,1574,1599]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="3" pageNumber="214" phylum="Chordata" rank="genus">Dactyloa</taxonomicName>
” clade (
<bibRefCitation id="EF954B4F3801FFBD1AE6316AFC14F9DE" author="Savage" box="[736,952,1574,1599]" pageId="3" pageNumber="204" refString="Savage, J. &amp; Guyer, C. (1989) Infrageneric classification and species composition of the anole genera, Anolis, Ctenonotus, Dactyloa, Norops and Semiurus (Sauria: Iguanidae). Amphibia-Reptilia, 10, 105 - 116. http: // dx. doi. org / 10.1163 / 156853889 x 00142" type="journal article" year="1989">
Savage 
<emphasis id="B970EAAC3801FFBD1B3D316AFCD8F9DF" box="[827,884,1574,1598]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
1989
</bibRefCitation>
; Castañeda &amp; de 
<bibRefCitation id="EF954B4F3801FFBD1C87316BFAB1F9DE" author="Queiroz" box="[1153,1309,1574,1599]" pageId="3" pageNumber="204" refString="Castaneda, M. R. &amp; de Queiroz, K. (2011) Phylogenetic relationships of the Dactyloa clade of Anolis lizards based on nuclear and mitochondrial DNA sequence data. Molecular phylogenetics and evolution, 61, 784 - 800. http: // dx. doi. org / 10.1016 / j. ympev. 2011.07.004" type="journal article" year="2011">Queiroz 2011</bibRefCitation>
, 
<bibRefCitation id="EF954B4F3801FFBD1D2C316AFAC8F9DE" author="Queiroz" box="[1322,1380,1575,1599]" pageId="3" pageNumber="204" refString="Castaneda, M. R. &amp; de Queiroz, K. (2013) Phylogeny of the Dactyloa Clade of Anolis Lizards: New Insights from Combining Morphological and Molecular Data. Bulletin of the Museum of Comparative Zoology, 160, 345 - 398. http: // dx. doi. org / 10.3099 / 0027 - 4100 - 160.7.345" type="journal article" year="2013">2013</bibRefCitation>
) and the 
<taxonomicName id="4C044D3D3801FFBD18C33107FE8FF983" box="[197,291,1610,1634]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B970EAAC3801FFBD18C33107FE8FF983" box="[197,291,1610,1634]" italics="true" pageId="3" pageNumber="204">latifrons</emphasis>
</taxonomicName>
series (Etheridge 1959; Williams 1976a) according to the following combination of osteological characters: eight or less septate caudal vertebrae, lack of transverse processes on most or all of the autotomic caudal vertebra and having at least four parasternal chevrons attached to the dorsal ribs.
</paragraph>
</subSubSection>
<subSubSection id="C31E65353801FFBB18C131FBFDB4F9E8" lastPageId="5" lastPageNumber="206" pageId="3" pageNumber="204" type="reference_group">
<paragraph id="8BBB36BE3801FFBD18C131FBFE08F862" blockId="3.[151,1436,1574,2031]" pageId="3" pageNumber="204">
<taxonomicName id="4C044D3D3801FFBD18C131FBFEF7F92F" box="[199,347,1718,1742]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3801FFBD18C131FBFEF7F92F" box="[199,347,1718,1742]" italics="true" pageId="3" pageNumber="204">Anolis limon</emphasis>
</taxonomicName>
differs from most species of the 
<taxonomicName id="4C044D3D3801FFBD1AF531FAFCCCF92F" box="[755,864,1719,1742]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="punctatus">
<emphasis id="B970EAAC3801FFBD1AF531FAFCCCF92F" box="[755,864,1719,1742]" italics="true" pageId="3" pageNumber="204">punctatus</emphasis>
</taxonomicName>
and 
<emphasis id="B970EAAC3801FFBD1BA731FAFC54F92F" box="[929,1016,1719,1742]" italics="true" pageId="3" pageNumber="204">tigrinus</emphasis>
groups (
<bibRefCitation id="EF954B4F3801FFBD1C6331FBFABEF92E" author="Williams" box="[1125,1298,1718,1743]" pageId="3" pageNumber="204" refString="Williams, E. E. (1992) New or problematic Anolis from Colombia. VII. Anolis lamari, a new anole from the Cordillera Oriental of Colombia, with a discussion of tigrinus and punctatus species group boundaries. Breviora, 495, 1 - 24." type="journal article" year="1992">Williams 1992</bibRefCitation>
) by having relatively smaller head scales; from the 
<taxonomicName id="4C044D3D3801FFBD1A513197FD34F913" box="[599,664,1754,1778]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="laevis">
<emphasis id="B970EAAC3801FFBD1A513197FD34F913" box="[599,664,1754,1778]" italics="true" pageId="3" pageNumber="204">laevis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF954B4F3801FFBD1B4E3197FC5CF912" author="Williams" box="[840,1008,1754,1779]" pageId="3" pageNumber="204" refString="Williams, E. E. (1979) South American Anoles: The species groups. 2. The proboscis anoles (Anolis laevis group). Breviora, 449, 1 - 21." type="journal article" year="1979">Williams 1979</bibRefCitation>
) by lacking a soft 
<taxonomicName id="4C044D3D3801FFBD1CC43197FA9DF912" box="[1218,1329,1754,1779]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="proboscis">proboscis</taxonomicName>
; from the 
<emphasis id="B970EAAC3801FFBD189131B2FF73F8F7" box="[151,223,1791,1814]" italics="true" pageId="3" pageNumber="204">roquet</emphasis>
species group (Williams 1976b) by possessing supraorbital semicircles separated from each other, the interparietal separated from the supraorbital semicircles, and a straight posterior border of the mental (
<bibRefCitation id="EF954B4F3801FFBD1D19306EFA3CF8DA" author="Poe" box="[1311,1424,1827,1851]" pageId="3" pageNumber="204" refString="Poe, S. (2004) Phylogeny of Anoles. Herpetological Monographs, 18, 37. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
); and from the 
<taxonomicName id="4C044D3D3801FFBD192A300BFE26F8BF" box="[300,394,1862,1886]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="latifrons">
<emphasis id="B970EAAC3801FFBD192A300BFE26F8BF" box="[300,394,1862,1886]" italics="true" pageId="3" pageNumber="204">latifrons</emphasis>
</taxonomicName>
clade (Castañeda &amp; de Quieroz 2011, 2013) by having a snout-vent length in adults less than 
<quantity id="4CFC9B5B3801FFBD18913026FF58F863" box="[151,244,1899,1923]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="204" unit="mm" value="100.0">100 mm</quantity>
(
<bibRefCitation id="EF954B4F3801FFBD19043026FE34F862" author="Savage" box="[258,408,1899,1923]" pageId="3" pageNumber="204" refString="Savage, J. (2002) The Amphibians and Reptiles of Costa Rica: A Herpetofauna between two Continents, between two Seas. University of Chicago press, 934 pp." type="book" year="2002">Savage 2002</bibRefCitation>
).
</paragraph>
<paragraph id="8BBB36BE3801FFBB18C130C3FDB4F9E8" blockId="3.[151,1436,1574,2031]" lastBlockId="5.[151,1437,1232,2014]" lastPageId="5" lastPageNumber="206" pageId="3" pageNumber="204">
<taxonomicName id="4C044D3D3801FFBD18C130C3FEFBF847" box="[199,343,1934,1958]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3801FFBD18C130C3FEFBF847" box="[199,343,1934,1958]" italics="true" pageId="3" pageNumber="204">Anolis limon</emphasis>
</taxonomicName>
shares intermediate adult body size (
<quantity id="4CFC9B5B3801FFBD1B0730C3FCD1F847" box="[769,893,1934,1959]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="3" pageNumber="204" unit="mm" value="80.0" valueMax="95.0" valueMin="65.0">65–95 mm</quantity>
SVL), smooth ventral scales, and mostly green dorsal coloration with 
<taxonomicName id="4C044D3D3801FFBD199830FEFD9DF82B" box="[414,561,1970,1994]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3801FFBD199830FEFD9DF82B" box="[414,561,1970,1994]" italics="true" pageId="3" pageNumber="204">A. chocorum</emphasis>
</taxonomicName>
and 
<emphasis id="B970EAAC3801FFBD1A6930FEFD45F82B" box="[623,745,1970,1994]" italics="true" pageId="3" pageNumber="204">
<taxonomicName id="4C044D3D3801FFBD1A6930FEFD4AF82B" box="[623,742,1970,1994]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="ibanezi">A. ibanezi</taxonomicName>
,
</emphasis>
(see 
<bibRefCitation id="EF954B4F3801FFBD1B2A30FEFC5CF82A" author="Poe" box="[812,1008,1970,1995]" pageId="3" pageNumber="204" refString="Poe, S., Latella, I., Ryan, M. J. &amp; Schaad, E. W. (2009) A new species of Anolis lizard (Squamata, Iguania) from Panama. Phyllomedusa, 8, 81 - 87." type="journal article" year="2009" yearSuffix="a">
Poe 
<emphasis id="B970EAAC3801FFBD1B6730FEFC30F82B" box="[865,924,1970,1994]" italics="true" pageId="3" pageNumber="204">et al.</emphasis>
2009a
</bibRefCitation>
), but differs of these both species in dewlap and color pattern (
<figureCitation id="133F2A3B3801FFBD19BD309BFDB2F80E" box="[443,542,2006,2031]" captionStart="FIGURE 2" captionStartId="4.[151,250,1957,1979]" captionTargetBox="[394,1185,193,1932]" captionTargetId="figure@4.[394,1185,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dewlap of Anolis limon sp. nov. and closest putative relatives. Top: Anolis limon sp. nov. (Photo: JPHG). Middle: Anolis chocorum (Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: Anolis ibanezi (Photo: Steven Poe)." httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="3" pageNumber="204">Figure 2</figureCitation>
). 
<taxonomicName id="4C044D3D3801FFBD1A34309BFD6DF80F" box="[562,705,2006,2030]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3801FFBD1A34309BFD6DF80F" box="[562,705,2006,2030]" italics="true" pageId="3" pageNumber="204">Anolis limon</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C044D3D3801FFBD1BBD309AFBE6F80F" box="[955,1098,2006,2030]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3801FFBD1BBD309AFBE6F80F" box="[955,1098,2006,2030]" italics="true" pageId="3" pageNumber="204">A. chocorum</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C044D3D3801FFBD1C86309AFB5CF80F" box="[1152,1264,2006,2030]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="204" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC3801FFBD1C86309AFB5CF80F" box="[1152,1264,2006,2030]" italics="true" pageId="3" pageNumber="204">A. ibanezi</emphasis>
</taxonomicName>
by its uniform light tan or yellow near the throat and tan in the distal portion with rows of green scales in males and color pattern. The dewlap in 
<taxonomicName id="4C044D3D3807FFBB194733B8FE7EFAEC" box="[321,466,1269,1293]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB194733B8FE7EFAEC" box="[321,466,1269,1293]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
is orange and white anteriorly with rows of white scales. The dewlap in 
<taxonomicName id="4C044D3D3807FFBB1D1633B8FA2DFAEC" box="[1296,1409,1269,1293]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC3807FFBB1D1633B8FA2DFAEC" box="[1296,1409,1269,1293]" italics="true" pageId="5" pageNumber="206">A. ibanezi</emphasis>
</taxonomicName>
is green and white anteriorly orange with rows of green scales (
<bibRefCitation id="EF954B4F3807FFBB1B4F3254FC55FAD0" author="Poe" box="[841,1017,1305,1329]" pageId="5" pageNumber="206" refString="Poe, S., Latella, I., Ryan, M. J. &amp; Schaad, E. W. (2009) A new species of Anolis lizard (Squamata, Iguania) from Panama. Phyllomedusa, 8, 81 - 87." type="journal article" year="2009">
Poe 
<emphasis id="B970EAAC3807FFBB1B7A3254FC19FAD0" box="[892,949,1305,1329]" italics="true" pageId="5" pageNumber="206">et al.</emphasis>
2009
</bibRefCitation>
). The color pattern in 
<taxonomicName id="4C044D3D3807FFBB1CF23254FAFFFAD0" box="[1268,1363,1305,1329]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3807FFBB1CF23254FAFFFAD0" box="[1268,1363,1305,1329]" italics="true" pageId="5" pageNumber="206">A. limon</emphasis>
</taxonomicName>
males is green with wide black bands on the flanks of males (males of 
<taxonomicName id="4C044D3D3807FFBB1B623270FC5FFAB4" box="[868,1011,1341,1365]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB1B623270FC5FFAB4" box="[868,1011,1341,1365]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
exhibit rows of ocelli and males of 
<taxonomicName id="4C044D3D3807FFBB1D823270FF44FA98" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC3807FFBB1D823270FF44FA98" italics="true" pageId="5" pageNumber="206">A. ibanezi</emphasis>
</taxonomicName>
exhibit thin lines). In females of 
<taxonomicName id="4C044D3D3807FFBB1A64322CFD6EFA98" box="[610,706,1377,1401]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3807FFBB1A64322CFD6EFA98" box="[610,706,1377,1401]" italics="true" pageId="5" pageNumber="206">A. limon</emphasis>
</taxonomicName>
there is diffuse dark green spots on the flanks (there are rows of ocelli in females of 
<taxonomicName id="4C044D3D3807FFBB197132C8FDAAFA7C" box="[375,518,1413,1437]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB197132C8FDAAFA7C" box="[375,518,1413,1437]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
and thin lines in females of 
<taxonomicName id="4C044D3D3807FFBB1B4E32C8FC15FA7C" box="[840,953,1413,1437]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC3807FFBB1B4E32C8FC15FA7C" box="[840,953,1413,1437]" italics="true" pageId="5" pageNumber="206">A. ibanezi</emphasis>
</taxonomicName>
; however, we do not known whether these spots would be homologous with the ocelli present in 
<taxonomicName id="4C044D3D3807FFBB1B1A32E4FC03FA20" box="[796,943,1449,1473]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB1B1A32E4FC03FA20" box="[796,943,1449,1473]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
) (
<figureCitation id="133F2A3B3807FFBB1BCA32E5FB9DFA20" box="[972,1073,1448,1473]" captionStart="FIGURE 1" captionStartId="3.[151,250,1513,1535]" captionTargetBox="[159,1426,301,1491]" captionTargetId="figure@3.[159,1427,301,1491]" captionTargetPageId="3" captionText="FIGURE 1. Anolis limon sp. nov. Top: male holotype (MHUA-R 11760); Bottom: female paratype (MHUA-R 10939)." httpUri="https://zenodo.org/record/251957/files/figure.png" pageId="5" pageNumber="206">Figure 1</figureCitation>
and 
<figureCitation id="133F2A3B3807FFBB1C7632E4FB2DFA20" box="[1136,1153,1449,1473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1957,1979]" captionTargetBox="[394,1185,193,1932]" captionTargetId="figure@4.[394,1185,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dewlap of Anolis limon sp. nov. and closest putative relatives. Top: Anolis limon sp. nov. (Photo: JPHG). Middle: Anolis chocorum (Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: Anolis ibanezi (Photo: Steven Poe)." httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="5" pageNumber="206">2</figureCitation>
). Furthermore, 
<taxonomicName id="4C044D3D3807FFBB1D3E32E4FA30FA20" box="[1336,1436,1449,1473]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3807FFBB1D3E32E4FA30FA20" box="[1336,1436,1449,1473]" italics="true" pageId="5" pageNumber="206">A. limon</emphasis>
</taxonomicName>
exhibits fewer scales across the snout than 
<taxonomicName id="4C044D3D3807FFBB1A703280FCA9FA04" box="[630,773,1485,1509]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB1A703280FCA9FA04" box="[630,773,1485,1509]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
(mean = 9.7, range = 
<quantity id="4CFC9B5B3807FFBB1BFA3280FBFCFA07" box="[1020,1104,1485,1510]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="5" pageNumber="206" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">8–11 in</quantity>
<taxonomicName id="4C044D3D3807FFBB1C503280FB19FA04" box="[1110,1205,1485,1509]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3807FFBB1C503280FB19FA04" box="[1110,1205,1485,1509]" italics="true" pageId="5" pageNumber="206">A. limon</emphasis>
</taxonomicName>
; mean = 12, range = 
<quantity id="4CFC9B5B3807FFBB189132BDFF56F9E8" box="[151,250,1520,1545]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="5" pageNumber="206" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
<taxonomicName id="4C044D3D3807FFBB190732BCFE3CF9E8" box="[257,400,1521,1545]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3807FFBB190732BCFE3CF9E8" box="[257,400,1521,1545]" italics="true" pageId="5" pageNumber="206">A. chocorum</emphasis>
</taxonomicName>
) (
<figureCitation id="133F2A3B3807FFBB19A132BDFDA6F9E8" box="[423,522,1520,1545]" captionStart="FIGURE 4" captionStartId="7.[151,250,1931,1953]" captionTargetBox="[152,1436,625,1908]" captionTargetId="figure@7.[151,1436,625,1909]" captionTargetPageId="7" captionText="FIGURE 4. Comparisons of head scales between Anolis limon sp. nov. and Anolis chocorum A) lateral, B) ventral, and C) dorsal view of head of A. limon sp. nov. (paratype, MHUA-R 11270); D) lateral, E) ventral, and F) dorsal view of head of A. chocorum (MHUA-R 10075; Colombia, Chocó, Nuqui, Estación Biológica El Amargal)." httpUri="https://zenodo.org/record/251960/files/figure.png" pageId="5" pageNumber="206">Figure 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7B66363806FFBA189130E8FB76F838" httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="4" pageNumber="205" targetBox="[394,1185,193,1932]" targetPageId="4">
<paragraph id="8BBB36BE3806FFBA189130E8FB76F838" blockId="4.[151,1437,1956,2009]" pageId="4" pageNumber="205">
<emphasis id="B970EAAC3806FFBA189130E8FEBEF85B" bold="true" box="[151,274,1957,1979]" pageId="4" pageNumber="205">FIGURE 2.</emphasis>
Dewlap of 
<taxonomicName id="4C044D3D3806FFBA198F30E9FDA6F85B" box="[393,522,1956,1978]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3806FFBA198F30E9FDA6F85B" box="[393,522,1956,1978]" italics="true" pageId="4" pageNumber="205">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73806FFBA1A1630E8FDCCF85B" box="[528,608,1957,1978]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B970EAAC3806FFBA1A1630E8FDCCF85B" bold="true" box="[528,608,1957,1978]" pageId="4" pageNumber="205">sp. nov.</emphasis>
</taxonomicNameLabel>
and closest putative relatives. Top: 
<taxonomicName id="4C044D3D3806FFBA1BCA30E9FBE1F85B" box="[972,1101,1956,1978]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3806FFBA1BCA30E9FBE1F85B" box="[972,1101,1956,1978]" italics="true" pageId="4" pageNumber="205">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73806FFBA1C5430E8FB0FF85B" box="[1106,1187,1957,1978]" pageId="4" pageNumber="205" rank="species">
<emphasis id="B970EAAC3806FFBA1C5430E8FB0FF85B" bold="true" box="[1106,1187,1957,1978]" pageId="4" pageNumber="205">sp. nov.</emphasis>
</taxonomicNameLabel>
(Photo: JPHG). Middle: 
<taxonomicName id="4C044D3D3806FFBA1891308EFEEFF838" box="[151,323,1987,2009]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3806FFBA1891308EFEEFF838" box="[151,323,1987,2009]" italics="true" pageId="4" pageNumber="205">Anolis chocorum</emphasis>
</taxonomicName>
(Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: 
<taxonomicName id="4C044D3D3806FFBA1B74308EFBAEF838" box="[882,1026,1987,2009]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="205" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC3806FFBA1B74308EFBAEF838" box="[882,1026,1987,2009]" italics="true" pageId="4" pageNumber="205">Anolis ibanezi</emphasis>
</taxonomicName>
(Photo: Steven Poe).
</paragraph>
</caption>
<caption id="DF7B66363807FFBB1891337FFE85FB45" httpUri="https://zenodo.org/record/251959/files/figure.png" pageId="5" pageNumber="206" targetBox="[151,1435,193,1051]" targetPageId="5">
<paragraph id="8BBB36BE3807FFBB1891337FFE85FB45" blockId="5.[151,1436,1074,1188]" pageId="5" pageNumber="206">
<emphasis id="B970EAAC3807FFBB1891337FFEB9FBA6" bold="true" box="[151,277,1074,1096]" pageId="5" pageNumber="206">FIGURE 3.</emphasis>
Holotype of 
<taxonomicName id="4C044D3D3807FFBB19A7337FFD89FBA9" box="[417,549,1074,1096]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="206" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3807FFBB19A7337FFD89FBA9" box="[417,549,1074,1096]" italics="true" pageId="5" pageNumber="206">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73807FFBB1A2A337FFDD3FBA6" box="[556,639,1074,1095]" pageId="5" pageNumber="206" rank="species">
<emphasis id="B970EAAC3807FFBB1A2A337FFDD3FBA6" bold="true" box="[556,639,1074,1095]" pageId="5" pageNumber="206">sp. nov.</emphasis>
</taxonomicNameLabel>
(MHUA-R 11760, adult male). A) lateral, B) dorsal, and C) ventral view of head; D) detail of dorsal scales at mid-dorsum of body, arrow indicate the middorsal scales; E) detail of ventral scales at middorsum of body; F) detail of flank scales at mid-dorsum of body; G) ventral view of cloacal region showing enlarged postcloacal scales.
</paragraph>
</caption>
<subSubSection id="C31E65353807FFB918C13158FCC3FDDD" lastPageId="7" lastPageNumber="208" pageId="5" pageNumber="206" type="description">
<paragraph id="8BBB36BE3807FFB818C13158FC62FEDE" blockId="5.[151,1437,1232,2014]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206">
<emphasis id="B970EAAC3807FFBB18C13158FC94F9CF" bold="true" box="[199,824,1557,1582]" pageId="5" pageNumber="206">
Description of external morphology of the 
<typeStatus id="54BF881C3807FFBB1AD73158FC94F9CF" box="[721,824,1557,1582]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
</emphasis>
(
<typeStatus id="54BF881C3807FFBB1B4F3158FC07F9CC" box="[841,939,1557,1581]" pageId="5" pageNumber="206" type="paratype">paratype</typeStatus>
variation in parentheses) (
<figureCitation id="133F2A3B3807FFBB1CD83158FAEDF9CC" box="[1246,1345,1557,1582]" captionStart="FIGURE 3" captionStartId="5.[151,250,1074,1096]" captionTargetBox="[151,1435,193,1051]" captionTargetId="figure@5.[151,1436,193,1053]" captionTargetPageId="5" captionText="FIGURE 3. Holotype of Anolis limon sp. nov. (MHUA-R 11760, adult male). A) lateral, B) dorsal, and C) ventral view of head; D) detail of dorsal scales at mid-dorsum of body, arrow indicate the middorsal scales; E) detail of ventral scales at middorsum of body; F) detail of flank scales at mid-dorsum of body; G) ventral view of cloacal region showing enlarged postcloacal scales." httpUri="https://zenodo.org/record/251959/files/figure.png" pageId="5" pageNumber="206">Figure 3</figureCitation>
and 
<figureCitation id="133F2A3B3807FFBB1D7B3158FA22F9CC" box="[1405,1422,1557,1581]" captionStart="FIGURE 4" captionStartId="7.[151,250,1931,1953]" captionTargetBox="[152,1436,625,1908]" captionTargetId="figure@7.[151,1436,625,1909]" captionTargetPageId="7" captionText="FIGURE 4. Comparisons of head scales between Anolis limon sp. nov. and Anolis chocorum A) lateral, B) ventral, and C) dorsal view of head of A. limon sp. nov. (paratype, MHUA-R 11270); D) lateral, E) ventral, and F) dorsal view of head of A. chocorum (MHUA-R 10075; Colombia, Chocó, Nuqui, Estación Biológica El Amargal)." httpUri="https://zenodo.org/record/251960/files/figure.png" pageId="5" pageNumber="206">4</figureCitation>
): Morphological dimensions are shown in 
<tableCitation id="C68603053807FFBB1A7E3175FD7CF9B0" box="[632,720,1592,1617]" captionStart="TABLE 1" captionStartId="6.[151,239,368,391]" captionTargetBox="[159,1357,450,1134]" captionTargetPageId="6" captionText="TABLE 1. Ranges of morphological measurements of the holotype, males and females of Anolis limon sp. nov. and Anolis chocorum. For abbreviations see materials and methods. Dimensions are given in millimeters." httpUri="http://table.plazi.org/id/DF7B66363804FFB81891363DFB61FE46" pageId="5" pageNumber="206" tableUuid="DF7B66363804FFB81891363DFB61FE46">Table 1</tableCitation>
. Dorsal head scales smooth; weak frontal depression; rostral canthus distinct; seven postrostrals (5–7); two sublabial scales slightly enlarged, (one enlarged sublabial scales on each side); seven postmentals (6–7); interparietal separated from supraorbital semicircles by four scales (2–4); three scales between supraorbital semicircles (2–3); nine supralabials to center of eye (8–9); eight infralabials to center of eye (7–9); one superciliary scale enlarged followed by granular scales; none scales rows between suboculars and supralabial (1); ten scales across snout between second canthals (8–11); six loreal rows (5–7); circumnasal in contact with supralabial, separated from rostral by one scale (not in contact with supralabial in MHUA-R 11267); rostral slightly overlapping mental; mental broader than rostral, extending posterolaterally beyond rostral; posterior border of mental concave (convex in all 
<typeStatus id="54BF881C3807FFBB1BAE3014FBB5F890" box="[936,1049,1881,1905]" pageId="5" pageNumber="206" type="paratype">paratypes</typeStatus>
); mental partially divided; scales surrounding the interparietal are smooth; some enlarged scales in supraocular disc, the rest granular. Dorsal scales keeled; two rows of slightly enlarged middorsal scales (0–2); nine dorsal scales counted longitudinally in 5% of SVL (8–11); nine ventral scales counted longitudinally in 5% of SVL (8–12) (see 
<bibRefCitation id="EF954B4F3807FFBB1C2F308BFB3EF83C" author="Poe" box="[1065,1170,1989,2013]" pageId="5" pageNumber="206" refString="Poe, S. (2004) Phylogeny of Anoles. Herpetological Monographs, 18, 37. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
for a description of this character); ventrals smooth, each scale bordered posteriorly by two scales (i.e., in transverse rows). Anterior part of the arm with unicarinate scales, elbow with multicarinate scales; thigh with unicarinate scales, knee with multicarinate scales; postcloacal greatly scales enlarged (absent in 
<typeStatus id="54BF881C3804FFB81B5637ADFC6DFF19" box="[848,961,224,248]" pageId="6" pageNumber="207" type="paratype">paratypes</typeStatus>
); scales on dewlap in rows of three scales (3–4); 22 expanded lamellae under third and four phalanges of fourth toe (20–22) (second and third phalanges of 
<bibRefCitation id="EF954B4F3804FFB81891366AFED2FEA1" author="Williams" box="[151,382,295,320]" pageId="6" pageNumber="207" refString="Williams, E. E., Rand, H., Rand, A. &amp; O'Hara, J. (1995) A computer approach to the comparison and identification of species in difficult taxonomic groups. Breviora, 502, 1 - 47." type="journal article" year="1995">
Williams 
<emphasis id="B970EAAC3804FFB819053664FE90FEA1" box="[259,316,296,320]" italics="true" pageId="6" pageNumber="207">et al.</emphasis>
1995
</bibRefCitation>
); tail with keeled scales and a double middorsal row.
</paragraph>
<caption id="DF7B66363804FFB81891363DFB61FE46" ID-Table-UUID="DF7B66363804FFB81891363DFB61FE46" httpUri="http://table.plazi.org/id/DF7B66363804FFB81891363DFB61FE46" pageId="6" pageNumber="207" targetBox="[159,1357,450,1134]" targetIsTable="true" targetPageId="6">
<paragraph id="8BBB36BE3804FFB81891363DFB61FE46" blockId="6.[151,1435,367,423]" pageId="6" pageNumber="207">
<emphasis id="B970EAAC3804FFB81891363DFEA9FE66" bold="true" box="[151,261,368,391]" pageId="6" pageNumber="207">TABLE 1</emphasis>
. Ranges of morphological measurements of the holotype, males and females of 
<taxonomicName id="4C044D3D3804FFB81C863622FAA7FE66" box="[1152,1291,367,391]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3804FFB81C863622FAA7FE66" box="[1152,1291,367,391]" italics="true" pageId="6" pageNumber="207">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73804FFB81D12363DFAC0FE66" box="[1300,1388,368,391]" pageId="6" pageNumber="207" rank="species">
<emphasis id="B970EAAC3804FFB81D12363DFAC0FE66" bold="true" box="[1300,1388,368,391]" pageId="6" pageNumber="207">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C044D3D3804FFB8189136C2FEE1FE46" box="[151,333,399,423]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3804FFB8189136C2FEE1FE46" box="[151,333,399,423]" italics="true" pageId="6" pageNumber="207">Anolis chocorum</emphasis>
</taxonomicName>
. For abbreviations see materials and methods. Dimensions are given in millimeters.
</paragraph>
</caption>
<paragraph id="8BBB36BE3804FFB81921368FFB59FB8F" pageId="6" pageNumber="207">
<table id="F904C41E380400411899368FFAE1FB8F" box="[159,1357,450,1134]" gridcols="6" gridrows="17" pageId="6" pageNumber="207">
<tr id="353434FC380400411899368FFAE1FE39" box="[159,1357,450,472]" gridrow="0" pageId="6" pageNumber="207" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<th id="76E55D80380400411921368FFE04FE39" box="[295,424,450,472]" gridcol="1" gridrow="0" pageId="6" pageNumber="207">
<taxonomicName id="4C044D3D3804FFB81921368FFE04FE39" box="[295,424,450,472]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3804FFB81921368FFE04FE39" box="[295,424,450,472]" italics="true" pageId="6" pageNumber="207">Anolis limon</emphasis>
</taxonomicName>
</th>
<th id="76E55D80380400411B98368FFBE6FE39" box="[926,1098,450,472]" gridcol="4" gridrow="0" pageId="6" pageNumber="207">
<taxonomicName id="4C044D3D3804FFB81B98368FFBE6FE39" box="[926,1098,450,472]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3804FFB81B98368FFBE6FE39" box="[926,1098,450,472]" italics="true" pageId="6" pageNumber="207">Anolis chocorum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="353434FC38040041189936A1FAE1FDE3" box="[159,1357,492,514]" gridrow="1" pageId="6" pageNumber="207">
<th id="76E55D8038040041189936A1FEADFDE3" box="[159,257,492,514]" gridcol="0" gridrow="1" pageId="6" pageNumber="207">Character</th>
<td id="76E55D8038040041192136A1FE04FDE3" box="[295,424,492,514]" gridcol="1" gridrow="1" pageId="6" pageNumber="207">Holotype</td>
<td id="76E55D803804004119D036A1FD97FDE3" box="[470,571,492,514]" gridcol="2" gridrow="1" pageId="6" pageNumber="207">Paratype*</td>
<td id="76E55D80380400411A8736A1FC8DFDE3" box="[641,801,492,514]" gridcol="3" gridrow="1" pageId="6" pageNumber="207">Females (n = 4)</td>
<td id="76E55D80380400411B9836A1FBE6FDE3" box="[926,1098,492,514]" gridcol="4" gridrow="1" pageId="6" pageNumber="207">Males (n = 7)</td>
<td id="76E55D80380400411C7036A1FAE1FDE3" box="[1142,1357,492,514]" gridcol="5" gridrow="1" pageId="6" pageNumber="207">Females (n = 6)</td>
</tr>
<tr id="353434FC3804004118993558FAE1FDCA" box="[159,1357,533,555]" gridrow="2" pageId="6" pageNumber="207">
<th id="76E55D803804004118993558FEADFDCA" box="[159,257,533,555]" gridcol="0" gridrow="2" pageId="6" pageNumber="207">SVL</th>
<td id="76E55D803804004119213558FE04FDCA" box="[295,424,533,555]" gridcol="1" gridrow="2" pageId="6" pageNumber="207">78.60</td>
<td id="76E55D803804004119D03558FD97FDCA" box="[470,571,533,555]" gridcol="2" gridrow="2" pageId="6" pageNumber="207">74.50</td>
<td id="76E55D80380400411A873558FC8DFDCA" box="[641,801,533,555]" gridcol="3" gridrow="2" pageId="6" pageNumber="207">71.21–81.78</td>
<td id="76E55D80380400411B983558FBE6FDCA" box="[926,1098,533,555]" gridcol="4" gridrow="2" pageId="6" pageNumber="207">71.92–80.64</td>
<td id="76E55D80380400411C703558FAE1FDCA" box="[1142,1357,533,555]" gridcol="5" gridrow="2" pageId="6" pageNumber="207">58.61–74.27(n=8)</td>
</tr>
<tr id="353434FC3804004118993573FAE1FDB5" box="[159,1357,574,596]" gridrow="3" pageId="6" pageNumber="207">
<th id="76E55D803804004118993573FEADFDB5" box="[159,257,574,596]" gridcol="0" gridrow="3" pageId="6" pageNumber="207">TL</th>
<td id="76E55D803804004119213573FE04FDB5" box="[295,424,574,596]" gridcol="1" gridrow="3" pageId="6" pageNumber="207">193.00</td>
<td id="76E55D803804004119D03573FD97FDB5" box="[470,571,574,596]" gridcol="2" gridrow="3" pageId="6" pageNumber="207">182.00</td>
<td id="76E55D80380400411A873573FC8DFDB5" box="[641,801,574,596]" gridcol="3" gridrow="3" pageId="6" pageNumber="207">180.97–201.92</td>
<td id="76E55D80380400411B983573FBE6FDB5" box="[926,1098,574,596]" gridcol="4" gridrow="3" pageId="6" pageNumber="207">179.17–203.55</td>
<td id="76E55D80380400411C703573FAE1FDB5" box="[1142,1357,574,596]" gridcol="5" gridrow="3" pageId="6" pageNumber="207">124.82–202.87(n=7)</td>
</tr>
<tr id="353434FC3804004118993525FAE1FD9F" box="[159,1357,616,638]" gridrow="4" pageId="6" pageNumber="207">
<th id="76E55D803804004118993525FEADFD9F" box="[159,257,616,638]" gridcol="0" gridrow="4" pageId="6" pageNumber="207">HL</th>
<td id="76E55D803804004119213525FE04FD9F" box="[295,424,616,638]" gridcol="1" gridrow="4" pageId="6" pageNumber="207">20.10</td>
<td id="76E55D803804004119D03525FD97FD9F" box="[470,571,616,638]" gridcol="2" gridrow="4" pageId="6" pageNumber="207">18.70</td>
<td id="76E55D80380400411A873525FC8DFD9F" box="[641,801,616,638]" gridcol="3" gridrow="4" pageId="6" pageNumber="207">19.37–21.5</td>
<td id="76E55D80380400411B983525FBE6FD9F" box="[926,1098,616,638]" gridcol="4" gridrow="4" pageId="6" pageNumber="207">17.56–21.34</td>
<td id="76E55D80380400411C703525FAE1FD9F" box="[1142,1357,616,638]" gridcol="5" gridrow="4" pageId="6" pageNumber="207">17.22–18.89</td>
</tr>
<tr id="353434FC38040041189935DCFAE1FD46" box="[159,1357,657,679]" gridrow="5" pageId="6" pageNumber="207">
<th id="76E55D8038040041189935DCFEADFD46" box="[159,257,657,679]" gridcol="0" gridrow="5" pageId="6" pageNumber="207">HW</th>
<td id="76E55D8038040041192135DCFE04FD46" box="[295,424,657,679]" gridcol="1" gridrow="5" pageId="6" pageNumber="207">10.70</td>
<td id="76E55D803804004119D035DCFD97FD46" box="[470,571,657,679]" gridcol="2" gridrow="5" pageId="6" pageNumber="207">10.00</td>
<td id="76E55D80380400411A8735DCFC8DFD46" box="[641,801,657,679]" gridcol="3" gridrow="5" pageId="6" pageNumber="207">9.37–11.28</td>
<td id="76E55D80380400411B9835DCFBE6FD46" box="[926,1098,657,679]" gridcol="4" gridrow="5" pageId="6" pageNumber="207">8.54–11.63</td>
<td id="76E55D80380400411C7035DCFAE1FD46" box="[1142,1357,657,679]" gridcol="5" gridrow="5" pageId="6" pageNumber="207">8.42–9.48</td>
</tr>
<tr id="353434FC38040041189935F7FAE1FD31" box="[159,1357,698,720]" gridrow="6" pageId="6" pageNumber="207">
<th id="76E55D8038040041189935F7FEADFD31" box="[159,257,698,720]" gridcol="0" gridrow="6" pageId="6" pageNumber="207">JL</th>
<td id="76E55D8038040041192135F7FE04FD31" box="[295,424,698,720]" gridcol="1" gridrow="6" pageId="6" pageNumber="207">18.20</td>
<td id="76E55D803804004119D035F7FD97FD31" box="[470,571,698,720]" gridcol="2" gridrow="6" pageId="6" pageNumber="207">15.60</td>
<td id="76E55D80380400411A8735F7FC8DFD31" box="[641,801,698,720]" gridcol="3" gridrow="6" pageId="6" pageNumber="207">16.96–18.53</td>
<td id="76E55D80380400411B9835F7FBE6FD31" box="[926,1098,698,720]" gridcol="4" gridrow="6" pageId="6" pageNumber="207">15.89–18.3</td>
<td id="76E55D80380400411C7035F7FAE1FD31" box="[1142,1357,698,720]" gridcol="5" gridrow="6" pageId="6" pageNumber="207">15.06–16.70</td>
</tr>
<tr id="353434FC38040041189935A9FAE1FD1B" box="[159,1357,740,762]" gridrow="7" pageId="6" pageNumber="207">
<th id="76E55D8038040041189935A9FEADFD1B" box="[159,257,740,762]" gridcol="0" gridrow="7" pageId="6" pageNumber="207">IoD</th>
<td id="76E55D8038040041192135A9FE04FD1B" box="[295,424,740,762]" gridcol="1" gridrow="7" pageId="6" pageNumber="207">3.40</td>
<td id="76E55D803804004119D035A9FD97FD1B" box="[470,571,740,762]" gridcol="2" gridrow="7" pageId="6" pageNumber="207">3.70</td>
<td id="76E55D80380400411A8735A9FC8DFD1B" box="[641,801,740,762]" gridcol="3" gridrow="7" pageId="6" pageNumber="207">3.05–3.39</td>
<td id="76E55D80380400411B9835A9FBE6FD1B" box="[926,1098,740,762]" gridcol="4" gridrow="7" pageId="6" pageNumber="207">2.41–2.94</td>
<td id="76E55D80380400411C7035A9FAE1FD1B" box="[1142,1357,740,762]" gridcol="5" gridrow="7" pageId="6" pageNumber="207">2.23–2.74</td>
</tr>
<tr id="353434FC3804004118993440FAE1FCC2" box="[159,1357,781,803]" gridrow="8" pageId="6" pageNumber="207">
<th id="76E55D803804004118993440FEADFCC2" box="[159,257,781,803]" gridcol="0" gridrow="8" pageId="6" pageNumber="207">AGL</th>
<td id="76E55D803804004119213440FE04FCC2" box="[295,424,781,803]" gridcol="1" gridrow="8" pageId="6" pageNumber="207">35.90</td>
<td id="76E55D803804004119D03440FD97FCC2" box="[470,571,781,803]" gridcol="2" gridrow="8" pageId="6" pageNumber="207">33.70</td>
<td id="76E55D80380400411A873440FC8DFCC2" box="[641,801,781,803]" gridcol="3" gridrow="8" pageId="6" pageNumber="207">34.16–39.56</td>
<td id="76E55D80380400411B983440FBE6FCC2" box="[926,1098,781,803]" gridcol="4" gridrow="8" pageId="6" pageNumber="207">32.42–38.15</td>
<td id="76E55D80380400411C703440FAE1FCC2" box="[1142,1357,781,803]" gridcol="5" gridrow="8" pageId="6" pageNumber="207">30.01–34.14</td>
</tr>
<tr id="353434FC380400411899347BFAE1FCAD" box="[159,1357,822,844]" gridrow="9" pageId="6" pageNumber="207">
<th id="76E55D80380400411899347BFEADFCAD" box="[159,257,822,844]" gridcol="0" gridrow="9" pageId="6" pageNumber="207">InD</th>
<td id="76E55D80380400411921347BFE04FCAD" box="[295,424,822,844]" gridcol="1" gridrow="9" pageId="6" pageNumber="207">2.50</td>
<td id="76E55D803804004119D0347BFD97FCAD" box="[470,571,822,844]" gridcol="2" gridrow="9" pageId="6" pageNumber="207">2.40</td>
<td id="76E55D80380400411A87347BFC8DFCAD" box="[641,801,822,844]" gridcol="3" gridrow="9" pageId="6" pageNumber="207">2.33–2.61</td>
<td id="76E55D80380400411B98347BFBE6FCAD" box="[926,1098,822,844]" gridcol="4" gridrow="9" pageId="6" pageNumber="207">2.11–2.59</td>
<td id="76E55D80380400411C70347BFAE1FCAD" box="[1142,1357,822,844]" gridcol="5" gridrow="9" pageId="6" pageNumber="207">2.15–2.60</td>
</tr>
<tr id="353434FC380400411899342DFAE1FC97" box="[159,1357,864,886]" gridrow="10" pageId="6" pageNumber="207">
<th id="76E55D80380400411899342DFEADFC97" box="[159,257,864,886]" gridcol="0" gridrow="10" pageId="6" pageNumber="207">SL</th>
<td id="76E55D80380400411921342DFE04FC97" box="[295,424,864,886]" gridcol="1" gridrow="10" pageId="6" pageNumber="207">9.70</td>
<td id="76E55D803804004119D0342DFD97FC97" box="[470,571,864,886]" gridcol="2" gridrow="10" pageId="6" pageNumber="207">9.10</td>
<td id="76E55D80380400411A87342DFC8DFC97" box="[641,801,864,886]" gridcol="3" gridrow="10" pageId="6" pageNumber="207">9.26–10.51</td>
<td id="76E55D80380400411B98342DFBE6FC97" box="[926,1098,864,886]" gridcol="4" gridrow="10" pageId="6" pageNumber="207">8.33–10.65</td>
<td id="76E55D80380400411C70342DFAE1FC97" box="[1142,1357,864,886]" gridcol="5" gridrow="10" pageId="6" pageNumber="207">8.15–9.29</td>
</tr>
<tr id="353434FC38040041189934C4FAE1FC7E" box="[159,1357,905,927]" gridrow="11" pageId="6" pageNumber="207">
<th id="76E55D8038040041189934C4FEADFC7E" box="[159,257,905,927]" gridcol="0" gridrow="11" pageId="6" pageNumber="207">HuL</th>
<td id="76E55D8038040041192134C4FE04FC7E" box="[295,424,905,927]" gridcol="1" gridrow="11" pageId="6" pageNumber="207">11.00</td>
<td id="76E55D803804004119D034C4FD97FC7E" box="[470,571,905,927]" gridcol="2" gridrow="11" pageId="6" pageNumber="207">10.50</td>
<td id="76E55D80380400411A8734C4FC8DFC7E" box="[641,801,905,927]" gridcol="3" gridrow="11" pageId="6" pageNumber="207">10.00–12.09</td>
<td id="76E55D80380400411B9834C4FBE6FC7E" box="[926,1098,905,927]" gridcol="4" gridrow="11" pageId="6" pageNumber="207">10.13–13.27</td>
<td id="76E55D80380400411C7034C4FAE1FC7E" box="[1142,1357,905,927]" gridcol="5" gridrow="11" pageId="6" pageNumber="207">9.34–12.10</td>
</tr>
<tr id="353434FC38040041189934FFFAE1FC29" box="[159,1357,946,968]" gridrow="12" pageId="6" pageNumber="207">
<th id="76E55D8038040041189934FFFEADFC29" box="[159,257,946,968]" gridcol="0" gridrow="12" pageId="6" pageNumber="207">UlL</th>
<td id="76E55D8038040041192134FFFE04FC29" box="[295,424,946,968]" gridcol="1" gridrow="12" pageId="6" pageNumber="207">10.10</td>
<td id="76E55D803804004119D034FFFD97FC29" box="[470,571,946,968]" gridcol="2" gridrow="12" pageId="6" pageNumber="207">9.80</td>
<td id="76E55D80380400411A8734FFFC8DFC29" box="[641,801,946,968]" gridcol="3" gridrow="12" pageId="6" pageNumber="207">9.22–11.21</td>
<td id="76E55D80380400411B9834FFFBE6FC29" box="[926,1098,946,968]" gridcol="4" gridrow="12" pageId="6" pageNumber="207">9.31–12.16</td>
<td id="76E55D80380400411C7034FFFAE1FC29" box="[1142,1357,946,968]" gridcol="5" gridrow="12" pageId="6" pageNumber="207">8.67–11.10</td>
</tr>
<tr id="353434FC3804004118993491FAE1FC13" box="[159,1357,988,1010]" gridrow="13" pageId="6" pageNumber="207">
<th id="76E55D803804004118993491FEADFC13" box="[159,257,988,1010]" gridcol="0" gridrow="13" pageId="6" pageNumber="207">FL</th>
<td id="76E55D803804004119213491FE04FC13" box="[295,424,988,1010]" gridcol="1" gridrow="13" pageId="6" pageNumber="207">18.60</td>
<td id="76E55D803804004119D03491FD97FC13" box="[470,571,988,1010]" gridcol="2" gridrow="13" pageId="6" pageNumber="207">17.10</td>
<td id="76E55D80380400411A873491FC8DFC13" box="[641,801,988,1010]" gridcol="3" gridrow="13" pageId="6" pageNumber="207">17.21–19.76</td>
<td id="76E55D80380400411B983491FBE6FC13" box="[926,1098,988,1010]" gridcol="4" gridrow="13" pageId="6" pageNumber="207">17.52–20.31</td>
<td id="76E55D80380400411C703491FAE1FC13" box="[1142,1357,988,1010]" gridcol="5" gridrow="13" pageId="6" pageNumber="207">14.76–20.12</td>
</tr>
<tr id="353434FC3804004118993348FAE1FBFA" box="[159,1357,1029,1051]" gridrow="14" pageId="6" pageNumber="207">
<th id="76E55D803804004118993348FEADFBFA" box="[159,257,1029,1051]" gridcol="0" gridrow="14" pageId="6" pageNumber="207">TiL</th>
<td id="76E55D803804004119213348FE04FBFA" box="[295,424,1029,1051]" gridcol="1" gridrow="14" pageId="6" pageNumber="207">18.20</td>
<td id="76E55D803804004119D03348FD97FBFA" box="[470,571,1029,1051]" gridcol="2" gridrow="14" pageId="6" pageNumber="207">16.40</td>
<td id="76E55D80380400411A873348FC8DFBFA" box="[641,801,1029,1051]" gridcol="3" gridrow="14" pageId="6" pageNumber="207">17.01–18.48</td>
<td id="76E55D80380400411B983348FBE6FBFA" box="[926,1098,1029,1051]" gridcol="4" gridrow="14" pageId="6" pageNumber="207">17.00–19.40</td>
<td id="76E55D80380400411C703348FAE1FBFA" box="[1142,1357,1029,1051]" gridcol="5" gridrow="14" pageId="6" pageNumber="207">14.96–19.64</td>
</tr>
<tr id="353434FC3804004118993363FAE1FBA5" box="[159,1357,1070,1092]" gridrow="15" pageId="6" pageNumber="207">
<th id="76E55D803804004118993363FEADFBA5" box="[159,257,1070,1092]" gridcol="0" gridrow="15" pageId="6" pageNumber="207">FoL</th>
<td id="76E55D803804004119213363FE04FBA5" box="[295,424,1070,1092]" gridcol="1" gridrow="15" pageId="6" pageNumber="207">25.80</td>
<td id="76E55D803804004119D03363FD97FBA5" box="[470,571,1070,1092]" gridcol="2" gridrow="15" pageId="6" pageNumber="207">23.90</td>
<td id="76E55D80380400411A873363FC8DFBA5" box="[641,801,1070,1092]" gridcol="3" gridrow="15" pageId="6" pageNumber="207">23.85–25.44</td>
<td id="76E55D80380400411B983363FBE6FBA5" box="[926,1098,1070,1092]" gridcol="4" gridrow="15" pageId="6" pageNumber="207">23.59–25.93</td>
<td id="76E55D80380400411C703363FAE1FBA5" box="[1142,1357,1070,1092]" gridcol="5" gridrow="15" pageId="6" pageNumber="207">19.8–25.12</td>
</tr>
<tr id="353434FC3804004118993315FAE1FB8F" box="[159,1357,1112,1134]" gridrow="16" pageId="6" pageNumber="207">
<th id="76E55D803804004118993315FEADFB8F" box="[159,257,1112,1134]" gridcol="0" gridrow="16" pageId="6" pageNumber="207">4tL</th>
<td id="76E55D803804004119213315FE04FB8F" box="[295,424,1112,1134]" gridcol="1" gridrow="16" pageId="6" pageNumber="207">14.90</td>
<td id="76E55D803804004119D03315FD97FB8F" box="[470,571,1112,1134]" gridcol="2" gridrow="16" pageId="6" pageNumber="207">12.90</td>
<td id="76E55D80380400411A873315FC8DFB8F" box="[641,801,1112,1134]" gridcol="3" gridrow="16" pageId="6" pageNumber="207">13.42–13.90</td>
<td id="76E55D80380400411B983315FBE6FB8F" box="[926,1098,1112,1134]" gridcol="4" gridrow="16" pageId="6" pageNumber="207">12.85–15.29</td>
<td id="76E55D80380400411C703315FAE1FB8F" box="[1142,1357,1112,1134]" gridcol="5" gridrow="16" pageId="6" pageNumber="207">10.78–13.68</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBB36BE3804FFB8189133C3FC2AFB44" blockId="6.[151,902,1166,1189]" box="[151,902,1166,1189]" pageId="6" pageNumber="207">
*There was only one male additional to the 
<typeStatus id="54BF881C3804FFB81A6D33C3FD64FB44" box="[619,712,1166,1189]" pageId="6" pageNumber="207" type="holotype">holotype</typeStatus>
during this work.
</paragraph>
<paragraph id="8BBB36BE3804FFB818C13383FF64FA96" blockId="6.[151,1436,1230,2011]" pageId="6" pageNumber="207">
<emphasis id="B970EAAC3804FFB818C13383FE42FB06" bold="true" box="[199,494,1230,1255]" pageId="6" pageNumber="207">Osteological description</emphasis>
(based in MHUA-R 11020). Postcranial skeleton: Interclavicles T-shaped; 24 presacral vertebrae; four lumbar vertebrae; eight caudal vertebrae with transverse processes anterior, gradually lost posteriorly (&quot;alpha&quot; condition in 
<emphasis id="B970EAAC3804FFB81A1C325BFDC7FACF" box="[538,619,1302,1326]" italics="true" pageId="6" pageNumber="207">
<taxonomicName id="4C044D3D3804FFB81A1C325BFDC8FACF" box="[538,612,1302,1326]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="genus">Anolis</taxonomicName>
;
</emphasis>
Etheridge 1959); Caudal autotomy septa absent; five postxiphisternal inscriptional ribs, four pairs attached to dorsal ribs and one pair free (5:1); Two sternal ribs and two xiphisternal ribs.
</paragraph>
<paragraph id="8BBB36BE3804FFB818C132CFFDE8F8AA" blockId="6.[151,1436,1230,2011]" pageId="6" pageNumber="207">Skull: Supraoccipital cresting continuous across supraoccipital; dorsal surface of skull smooth; parietal crests Y-shaped; anterolateral corners of parietal crests reach posterolateral corners of frontal; parietal casque absent; position of pineal foramen at parietal/frontal suture; supratemporal processes leave supraoccipital exposed above; postfrontal present; prefrontal contacts nasal; frontal sutures contacts only with nasals anteriorly; dorsal processes of jugal terminates on lateral aspect of postorbital; contact between jugal and squamosal absent; posteroventral corner of jugal is anterior to posterior edge of jugal; epipterygoid does not contact parietal; pterygoid teeth absent; lateral edges of vomer smooth; maxilla extends posteriorly to ectopterygoid; quadrate lateral shelf absent; premaxilla overlaps nasals laterally; posterior of skull slopes superiorly; crenulation along lateral edges of parietal absent; parietal roof flat; posteriormost tooth is posterior to anterior inferior alveolar foramen; angular processes of articular large; posterior suture of dentary pronged; posterior border of dentary extending within mandibular fossa; splenial large; ventral aspect of anteromedial process of coronoid projects posteriorly; surangular foramen completely in surangular; coronoid labial process present; posterolateral aspect of coronoid terminates anterior to supraangular foramen; angular absent.</paragraph>
<paragraph id="8BBB36BE3804FFB918C1301BFBD0FE31" blockId="6.[151,1436,1230,2011]" lastBlockId="7.[151,1436,151,573]" lastPageId="7" lastPageNumber="208" pageId="6" pageNumber="207">
Color in life: 
<taxonomicName id="4C044D3D3804FFB81966301BFE43F88F" box="[352,495,1878,1902]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="207" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3804FFB81966301BFE43F88F" box="[352,495,1878,1902]" italics="true" pageId="6" pageNumber="207">Anolis limon</emphasis>
</taxonomicName>
is sexually dimorphic in color pattern. The dorsal surface of the body in both sexes is light green. Males have transverse dark green to black bands (five from the neck to the tail base, see 
<figureCitation id="133F2A3B3804FFB81D2A3037FA26F872" box="[1324,1418,1914,1939]" captionStart="FIGURE 1" captionStartId="3.[151,250,1513,1535]" captionTargetBox="[159,1426,301,1491]" captionTargetId="figure@3.[159,1427,301,1491]" captionTargetPageId="3" captionText="FIGURE 1. Anolis limon sp. nov. Top: male holotype (MHUA-R 11760); Bottom: female paratype (MHUA-R 10939)." httpUri="https://zenodo.org/record/251957/files/figure.png" pageId="6" pageNumber="207">Figure 1</figureCitation>
), whereas females have numerous dispersed dark green to bluish small blotches with a few additional small white blotches. The banded/blotched (male/female) pattern continues onto the dorsal surface of limbs and tail. Two aspects of the coloration, observed in the 
<typeStatus id="54BF881C3805FFB91A7A37DAFD4DFF51" box="[636,737,151,176]" pageId="7" pageNumber="208" type="holotype">holotype</typeStatus>
, seem to be related to stress conditions, since they were not present when just caught. The first stress pattern is the brown and yellowish coloration in different parts of the head, and the second is some reticulated brown lines in trunk. The ventral surface of head, body, limbs, and tail is whitish with a diffuse light pattern of scattered bluish and greenish irregular small blotches in both sexes. Eyelids are bright yellow in their anterior and posterior margins, sometimes white at the very margin to the eye. The superior and inferior borders of eye are green like the rest of the body. The iris is light brown to reddish, the pupil is black. In males, the dewlap skin is uniform light tan or yellow near the throat and tan in the distal portion; the border scales are white, and has five transverse lines of light green scales. Females have white scales on the border of the dewlap, while the skin is green fading into yellow with rows of small white scales.
</paragraph>
<paragraph id="8BBB36BE3805FFB918C13691FCC3FDDD" blockId="7.[151,1436,151,573]" pageId="7" pageNumber="208">
Color in preservative. The color of recently preserved specimens is darker than that of living specimens, but the banded/blotched pattern is conserved. Older specimens also conserve the pattern, but green surfaces turn into purple, and the ventral surfaces become grayish (
<figureCitation id="133F2A3B3805FFB91ABD3569FCB7FDDD" box="[699,795,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1957,1979]" captionTargetBox="[394,1185,193,1932]" captionTargetId="figure@4.[394,1185,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Dewlap of Anolis limon sp. nov. and closest putative relatives. Top: Anolis limon sp. nov. (Photo: JPHG). Middle: Anolis chocorum (Photo: Omar Torres-Carvajal; QCAZ 4734). Bottom: Anolis ibanezi (Photo: Steven Poe)." httpUri="https://zenodo.org/record/251958/files/figure.png" pageId="7" pageNumber="208">Figure 2</figureCitation>
and 
<figureCitation id="133F2A3B3805FFB91B573569FCCDFDDD" box="[849,865,548,572]" captionStart="FIGURE 3" captionStartId="5.[151,250,1074,1096]" captionTargetBox="[151,1435,193,1051]" captionTargetId="figure@5.[151,1436,193,1053]" captionTargetPageId="5" captionText="FIGURE 3. Holotype of Anolis limon sp. nov. (MHUA-R 11760, adult male). A) lateral, B) dorsal, and C) ventral view of head; D) detail of dorsal scales at mid-dorsum of body, arrow indicate the middorsal scales; E) detail of ventral scales at middorsum of body; F) detail of flank scales at mid-dorsum of body; G) ventral view of cloacal region showing enlarged postcloacal scales." httpUri="https://zenodo.org/record/251959/files/figure.png" pageId="7" pageNumber="208">3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF7B66363805FFB9189130C6FBB6F83F" httpUri="https://zenodo.org/record/251960/files/figure.png" pageId="7" pageNumber="208" targetBox="[152,1436,625,1908]" targetPageId="7">
<paragraph id="8BBB36BE3805FFB9189130C6FBB6F83F" blockId="7.[151,1437,1931,2014]" pageId="7" pageNumber="208">
<emphasis id="B970EAAC3805FFB9189130C6FEB9F841" bold="true" box="[151,277,1931,1953]" pageId="7" pageNumber="208">FIGURE 4.</emphasis>
Comparisons of head scales between 
<taxonomicName id="4C044D3D3805FFB91AA530C6FC89F840" box="[675,805,1931,1953]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3805FFB91AA530C6FC89F840" box="[675,805,1931,1953]" italics="true" pageId="7" pageNumber="208">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73805FFB91B2830C6FC2DF841" box="[814,897,1931,1952]" pageId="7" pageNumber="208" rank="species">
<emphasis id="B970EAAC3805FFB91B2830C6FC2DF841" bold="true" box="[814,897,1931,1952]" pageId="7" pageNumber="208">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C044D3D3805FFB91BB030C6FBC8F840" box="[950,1124,1931,1953]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3805FFB91BB030C6FBC8F840" box="[950,1124,1931,1953]" italics="true" pageId="7" pageNumber="208">Anolis chocorum</emphasis>
</taxonomicName>
A) lateral, B) ventral, and C) dorsal view of head of 
<taxonomicName id="4C044D3D3805FFB9198230E7FE70F821" box="[388,476,1962,1984]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3805FFB9198230E7FE70F821" box="[388,476,1962,1984]" italics="true" pageId="7" pageNumber="208">A. limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73805FFB919E530E7FD98F85E" box="[483,564,1962,1983]" pageId="7" pageNumber="208" rank="species">
<emphasis id="B970EAAC3805FFB919E530E7FD98F85E" bold="true" box="[483,564,1962,1983]" pageId="7" pageNumber="208">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype, MHUA-R 11270); D) lateral, E) ventral, and F) dorsal view of head of 
<taxonomicName id="4C044D3D3805FFB91D8030E7FF50F83F" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="208" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3805FFB91D8030E7FF50F83F" italics="true" pageId="7" pageNumber="208">A. chocorum</emphasis>
</taxonomicName>
(MHUA-R 10075; Colombia, Chocó, Nuqui, Estación Biológica El Amargal).
</paragraph>
</caption>
<caption id="DF7B6636380AFFB6189133DBFF75FAE9" httpUri="https://zenodo.org/record/251961/files/figure.png" pageId="8" pageNumber="209" targetBox="[158,1424,199,1148]" targetPageId="8">
<paragraph id="8BBB36BE380AFFB6189133DBFF75FAE9" blockId="8.[151,1437,1174,1288]" pageId="8" pageNumber="209">
<emphasis id="B970EAAC380AFFB6189133DBFEBBFB4D" bold="true" box="[151,279,1174,1196]" pageId="8" pageNumber="209">FIGURE 5.</emphasis>
Distribution map for 
<taxonomicName id="4C044D3D380AFFB61A0033DBFD20FB4D" box="[518,652,1174,1196]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC380AFFB61A0033DBFD20FB4D" box="[518,652,1174,1196]" italics="true" pageId="8" pageNumber="209">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D7380AFFB61A9033DAFD47FB4D" box="[662,747,1175,1196]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B970EAAC380AFFB61A9033DAFD47FB4D" bold="true" box="[662,747,1175,1196]" pageId="8" pageNumber="209">sp. nov.</emphasis>
</taxonomicNameLabel>
and closest putative relatives. Circles in white for 
<taxonomicName id="4C044D3D380AFFB61D1033DAFA30FB4D" box="[1302,1436,1174,1196]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380AFFB61D1033DAFA30FB4D" box="[1302,1436,1174,1196]" italics="true" pageId="8" pageNumber="209">A. chocorum</emphasis>
</taxonomicName>
correspond to specimens and observations checked by one of us (JAV). The locality information for the rest of occurrence records were taken from original descriptions (Williams &amp; Duellman 1967, Poe 
<emphasis id="B970EAAC380AFFB61BC53398FC5AFB0B" box="[963,1014,1236,1258]" italics="true" pageId="8" pageNumber="209">et al.</emphasis>
2009) and recent literature (Lotzkat 
<emphasis id="B970EAAC380AFFB61D6F3398FA31FB0B" box="[1385,1437,1236,1258]" italics="true" pageId="8" pageNumber="209">et al.</emphasis>
2013).
</paragraph>
</caption>
<caption id="DF7B6636380AFFB61891307DFC90F884" httpUri="https://zenodo.org/record/251962/files/figure.png" pageId="8" pageNumber="209" targetBox="[389,1219,1338,1792]" targetPageId="8">
<paragraph id="8BBB36BE380AFFB61891307DFC90F884" blockId="8.[151,1436,1840,1893]" pageId="8" pageNumber="209">
<emphasis id="B970EAAC380AFFB61891307DFEBFF8A4" bold="true" box="[151,275,1840,1862]" pageId="8" pageNumber="209">FIGURE 6.</emphasis>
Dispersion of individuals (females) of 
<taxonomicName id="4C044D3D380AFFB61AA5307DFC88F8A7" box="[675,804,1840,1862]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC380AFFB61AA5307DFC88F8A7" box="[675,804,1840,1862]" italics="true" pageId="8" pageNumber="209">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D7380AFFB61B2C307CFCD7F8A7" box="[810,891,1841,1862]" pageId="8" pageNumber="209" rank="species">
<emphasis id="B970EAAC380AFFB61B2C307CFCD7F8A7" bold="true" box="[810,891,1841,1862]" pageId="8" pageNumber="209">sp. nov.</emphasis>
</taxonomicNameLabel>
(green) and 
<taxonomicName id="4C044D3D380AFFB61BFB307DFB05F8A7" box="[1021,1193,1840,1862]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380AFFB61BFB307DFB05F8A7" box="[1021,1193,1840,1862]" italics="true" pageId="8" pageNumber="209">Anolis chocorum</emphasis>
</taxonomicName>
(red) of the first (PC1) and second (PC2) components of PCA for 15 morphological traits.
</paragraph>
</caption>
<subSubSection id="C31E6535380AFFB718C130C1FBF7FE4D" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209" type="distribution">
<paragraph id="8BBB36BE380AFFB618C130C1FB95F80C" blockId="8.[151,1436,1932,2029]" pageId="8" pageNumber="209">
<emphasis id="B970EAAC380AFFB618C130C1FE5BF844" bold="true" box="[199,503,1932,1957]" pageId="8" pageNumber="209">Distribution and ecology.</emphasis>
<taxonomicName id="4C044D3D380AFFB61A0630C0FD3CF844" box="[512,656,1933,1957]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380AFFB61A0630C0FD3CF844" box="[512,656,1933,1957]" italics="true" pageId="8" pageNumber="209">Anolis limon</emphasis>
</taxonomicName>
is only known from some localities in the northeastern slope of the Central Andes in 
<collectingCountry id="F313762E380AFFB6195D30FDFE65F828" box="[347,457,1968,1993]" name="Colombia" pageId="8" pageNumber="209">Colombia</collectingCountry>
between 
<date id="FFBA107E380AFFB61A3230FCFD75F828" box="[564,729,1968,1993]" pageId="8" pageNumber="209" value="1000" valueMax="1800">
1000 and 
<quantity id="4CFC9B5B380AFFB61AA730FCFD75F828" box="[673,729,1969,1993]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="209" unit="m" value="1800.0">1800</quantity>
</date>
m of elevation (
<figureCitation id="133F2A3B380AFFB61B9630FDFC5DF828" box="[912,1009,1968,1993]" captionStart="FIGURE 5" captionStartId="8.[151,250,1174,1196]" captionTargetBox="[158,1424,199,1148]" captionTargetId="figure@8.[151,1436,193,1153]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for Anolis limon sp. nov. and closest putative relatives. Circles in white for A. chocorum correspond to specimens and observations checked by one of us (JAV). The locality information for the rest of occurrence records were taken from original descriptions (Williams &amp; Duellman 1967, Poe et al. 2009) and recent literature (Lotzkat et al. 2013)." httpUri="https://zenodo.org/record/251961/files/figure.png" pageId="8" pageNumber="209">Figure 5</figureCitation>
). We found 
<taxonomicName id="4C044D3D380AFFB61C7F30FDFAABF829" box="[1145,1287,1968,1992]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="209" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380AFFB61C7F30FDFAABF829" box="[1145,1287,1968,1992]" italics="true" pageId="8" pageNumber="209">Anolis limon</emphasis>
</taxonomicName>
in secondary forest and forest edges, near streams and ponds, mostly sleeping on leaves at night.
</paragraph>
<paragraph id="8BBB36BE380BFFB718C137DAFC2AFEA1" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="210">
<emphasis id="B970EAAC380BFFB718C137DAFE4FFF51" bold="true" box="[199,483,151,176]" pageId="9" pageNumber="210">Morphometric analysis.</emphasis>
The principal component analysis shows a clear separation between females for each species (
<figureCitation id="133F2A3B380BFFB718FC37F1FEF3FF34" box="[250,351,188,213]" captionStart="FIGURE 6" captionStartId="8.[151,250,1840,1862]" captionTargetBox="[389,1219,1338,1792]" captionTargetId="figure@8.[367,1219,1338,1819]" captionTargetPageId="8" captionText="FIGURE 6. Dispersion of individuals (females) of Anolis limon sp. nov. (green) and Anolis chocorum (red) of the first (PC 1) and second (PC 2) components of PCA for 15 morphological traits." httpUri="https://zenodo.org/record/251962/files/figure.png" pageId="9" pageNumber="210">Figure 6</figureCitation>
). The first two components explain 79.63% of variance in females (
<tableCitation id="C6860305380BFFB71C7837F1FB75FF35" box="[1150,1241,188,213]" captionStart="TABLE 2" captionStartId="9.[151,239,548,571]" captionTargetBox="[151,1208,632,1485]" captionTargetPageId="9" captionText="TABLE 2. Results of principal component analysis (the first two PC axes) for 15 morphological variables foradult females of Anolis limon sp. nov. and Anolis chocorum." httpUri="http://table.plazi.org/id/DF7B6636380BFFB718913569FD4DFDBD" pageId="9" pageNumber="210" tableUuid="DF7B6636380BFFB718913569FD4DFDBD">Table 2</tableCitation>
). Head variables exhibited the highest loadings in the first component (IoD, HW, SL) and limb dimensions (HuL, UL, FL) exhibited the highest loadings in the second component (see 
<tableCitation id="C6860305380BFFB71AD73649FC8AFEFD" box="[721,806,260,285]" captionStart="TABLE 2" captionStartId="9.[151,239,548,571]" captionTargetBox="[151,1208,632,1485]" captionTargetPageId="9" captionText="TABLE 2. Results of principal component analysis (the first two PC axes) for 15 morphological variables foradult females of Anolis limon sp. nov. and Anolis chocorum." httpUri="http://table.plazi.org/id/DF7B6636380BFFB718913569FD4DFDBD" pageId="9" pageNumber="210" tableUuid="DF7B6636380BFFB718913569FD4DFDBD">Table 2</tableCitation>
). Females of both species show a clear separation in the first component with very few differences in the second (
<figureCitation id="133F2A3B380BFFB71B10366AFCD4FEA1" box="[790,888,295,320]" captionStart="FIGURE 5" captionStartId="8.[151,250,1174,1196]" captionTargetBox="[158,1424,199,1148]" captionTargetId="figure@8.[151,1436,193,1153]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for Anolis limon sp. nov. and closest putative relatives. Circles in white for A. chocorum correspond to specimens and observations checked by one of us (JAV). The locality information for the rest of occurrence records were taken from original descriptions (Williams &amp; Duellman 1967, Poe et al. 2009) and recent literature (Lotzkat et al. 2013)." httpUri="https://zenodo.org/record/251961/files/figure.png" pageId="9" pageNumber="210">Figure 5</figureCitation>
).
</paragraph>
<paragraph id="8BBB36BE380BFFB718C13601FBF7FE4D" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="210">
<emphasis id="B970EAAC380BFFB718C13601FDA7FE84" bold="true" box="[199,523,332,357]" pageId="9" pageNumber="210">Phylogenetic relationships.</emphasis>
Our search strategy resulted in 576 equally-parsimonious trees with length of 31889 steps and a consistency index of 0.12 and a retention index of 0.49. The 50% majority rule consensus of these trees is shown in 
<figureCitation id="133F2A3B380BFFB7199C36D9FE54FE4D" box="[410,504,404,429]" captionStart="FIGURE 7. A 50" captionStartId="10.[151,250,1981,2003]" captionTargetBox="[449,1144,207,1917]" captionTargetId="figure@10.[440,1152,193,1922]" captionTargetPageId="10" captionText="FIGURE 7. A 50 % majority rule consensus showing phylogenetic relationships for dactyloid anoles including Anolis limon sp. nov. Values in the nodes correspond to support values above 50 % from 500 bootstrap replicates." httpUri="https://zenodo.org/record/251963/files/figure.png" pageId="9" pageNumber="210">Figure 7</figureCitation>
(all non-dactyloid species were collapsed in this tree).
</paragraph>
</subSubSection>
<subSubSection id="C31E6535380BFFB518C136FAFC66FC61" lastPageId="11" lastPageNumber="212" pageId="9" pageNumber="210" type="etymology">
<paragraph id="8BBB36BE380BFFB718C136FAFCFFFE15" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="210">
<emphasis id="B970EAAC380BFFB718C136FAFEE2FE31" bold="true" box="[199,334,439,464]" pageId="9" pageNumber="210">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C044D3D380BFFB71A2B36F5FDC1FE31" box="[557,621,440,464]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380BFFB71A2B36F5FDC1FE31" box="[557,621,440,464]" italics="true" pageId="9" pageNumber="210">limon</emphasis>
</taxonomicName>
makes reference to the resemblance of the body color of the females to the green color of the fruits of the lemon tree (
<emphasis id="B970EAAC380BFFB71A983691FCEFFE15" box="[670,835,476,500]" italics="true" pageId="9" pageNumber="210">
<taxonomicName id="4C044D3D380BFFB71A983691FD48FE12" box="[670,740,476,499]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="210" phylum="Tracheophyta" rank="genus">Citrus</taxonomicName>
x lemon
</emphasis>
).
</paragraph>
<caption id="DF7B6636380BFFB718913569FD4DFDBD" ID-Table-UUID="DF7B6636380BFFB718913569FD4DFDBD" httpUri="http://table.plazi.org/id/DF7B6636380BFFB718913569FD4DFDBD" pageId="9" pageNumber="210" targetBox="[151,1208,632,1485]" targetIsTable="true" targetPageId="9">
<paragraph id="8BBB36BE380BFFB718913569FD4DFDBD" blockId="9.[151,1435,548,604]" pageId="9" pageNumber="210">
<emphasis id="B970EAAC380BFFB718913569FEA1FDDA" bold="true" box="[151,269,548,571]" pageId="9" pageNumber="210">TABLE 2.</emphasis>
Results of principal component analysis (the first two PC axes) for 15 morphological variables foradult females of 
<taxonomicName id="4C044D3D380BFFB7190B3509FE39FDBD" box="[269,405,580,604]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC380BFFB7190B3509FE39FDBD" box="[269,405,580,604]" italics="true" pageId="9" pageNumber="210">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D7380BFFB7199D3508FE5DFDBD" box="[411,497,581,604]" pageId="9" pageNumber="210" rank="species">
<emphasis id="B970EAAC380BFFB7199D3508FE5DFDBD" bold="true" box="[411,497,581,604]" pageId="9" pageNumber="210">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C044D3D380BFFB71A223509FD76FDBD" box="[548,730,580,604]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380BFFB71A223509FD76FDBD" box="[548,730,580,604]" italics="true" pageId="9" pageNumber="210">Anolis chocorum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BBB36BE380BFFB71B363535FEB7FA2C" pageId="9" pageNumber="210">
<table id="F904C41E380B004118913535FB14FA2C" box="[151,1208,632,1485]" gridcols="3" gridrows="19" pageId="9" pageNumber="210">
<tr id="353434FC380B004118913535FB14FD6F" box="[151,1208,632,654]" gridrow="0" pageId="9" pageNumber="210" rowspan-0="1">
<th id="76E55D80380B00411B363535FCD2FD6F" box="[816,894,632,654]" gridcol="1" gridrow="0" pageId="9" pageNumber="210">PC 1</th>
<th id="76E55D80380B00411C6C3535FB14FD6F" box="[1130,1208,632,654]" gridcol="2" gridrow="0" pageId="9" pageNumber="210">PC 2</th>
</tr>
<tr id="353434FC380B0041189135ECFB14FD56" box="[151,1208,673,695]" gridrow="1" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189135ECFDE9FD56" box="[151,581,673,695]" gridcol="0" gridrow="1" pageId="9" pageNumber="210">% Variation</th>
<td id="76E55D80380B00411B3635ECFCD2FD56" box="[816,894,673,695]" gridcol="1" gridrow="1" pageId="9" pageNumber="210">60.61%</td>
<td id="76E55D80380B00411C6C35ECFB14FD56" box="[1130,1208,673,695]" gridcol="2" gridrow="1" pageId="9" pageNumber="210">19.00%</td>
</tr>
<tr id="353434FC380B004118913587FB14FD01" box="[151,1208,714,736]" gridrow="2" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913587FDE9FD01" box="[151,581,714,736]" gridcol="0" gridrow="2" pageId="9" pageNumber="210">Eigenvalues</th>
<td id="76E55D80380B00411B363587FCD2FD01" box="[816,894,714,736]" gridcol="1" gridrow="2" pageId="9" pageNumber="210">0.096</td>
<td id="76E55D80380B00411C6C3587FB14FD01" box="[1130,1208,714,736]" gridcol="2" gridrow="2" pageId="9" pageNumber="210">0.003</td>
</tr>
<tr id="353434FC380B0041189135B9FB14FCEB" box="[151,1208,756,778]" gridrow="3" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189135B9FDE9FCEB" box="[151,581,756,778]" gridcol="0" gridrow="3" pageId="9" pageNumber="210">Loadings Head Length</th>
<td id="76E55D80380B00411B3635B9FCD2FCEB" box="[816,894,756,778]" gridcol="1" gridrow="3" pageId="9" pageNumber="210">0.322</td>
<td id="76E55D80380B00411C6C35B9FB14FCEB" box="[1130,1208,756,778]" gridcol="2" gridrow="3" pageId="9" pageNumber="210">0.006</td>
</tr>
<tr id="353434FC380B004118913451FB14FCD3" box="[151,1208,796,818]" gridrow="4" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913451FDE9FCD3" box="[151,581,796,818]" gridcol="0" gridrow="4" pageId="9" pageNumber="210">Head Width</th>
<td id="76E55D80380B00411B363451FCD2FCD3" box="[816,894,796,818]" gridcol="1" gridrow="4" pageId="9" pageNumber="210">0.363</td>
<td id="76E55D80380B00411C6C3451FB14FCD3" box="[1130,1208,796,818]" gridcol="2" gridrow="4" pageId="9" pageNumber="210">0.075</td>
</tr>
<tr id="353434FC380B004118913408FB14FCBA" box="[151,1208,837,859]" gridrow="5" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913408FDE9FCBA" box="[151,581,837,859]" gridcol="0" gridrow="5" pageId="9" pageNumber="210">Jaw Length</th>
<td id="76E55D80380B00411B363408FCD2FCBA" box="[816,894,837,859]" gridcol="1" gridrow="5" pageId="9" pageNumber="210">0.281</td>
<td id="76E55D80380B00411C6C3408FB14FCBA" box="[1130,1208,837,859]" gridcol="2" gridrow="5" pageId="9" pageNumber="210">-0.007</td>
</tr>
<tr id="353434FC380B004118913423FB14FC65" box="[151,1208,878,900]" gridrow="6" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913423FDE9FC65" box="[151,581,878,900]" gridcol="0" gridrow="6" pageId="9" pageNumber="210">Internarinal Distance</th>
<td id="76E55D80380B00411B363423FCD2FC65" box="[816,894,878,900]" gridcol="1" gridrow="6" pageId="9" pageNumber="210">0.097</td>
<td id="76E55D80380B00411C6C3423FB14FC65" box="[1130,1208,878,900]" gridcol="2" gridrow="6" pageId="9" pageNumber="210">-0.009</td>
</tr>
<tr id="353434FC380B0041189134D5FB14FC4F" box="[151,1208,920,942]" gridrow="7" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189134D5FDE9FC4F" box="[151,581,920,942]" gridcol="0" gridrow="7" pageId="9" pageNumber="210">Snout Length</th>
<td id="76E55D80380B00411B3634D5FCD2FC4F" box="[816,894,920,942]" gridcol="1" gridrow="7" pageId="9" pageNumber="210">0.332</td>
<td id="76E55D80380B00411C6C34D5FB14FC4F" box="[1130,1208,920,942]" gridcol="2" gridrow="7" pageId="9" pageNumber="210">-0.109</td>
</tr>
<tr id="353434FC380B00411891348CFB14FC36" box="[151,1208,961,983]" gridrow="8" pageId="9" pageNumber="210">
<th id="76E55D80380B00411891348CFDE9FC36" box="[151,581,961,983]" gridcol="0" gridrow="8" pageId="9" pageNumber="210">Interorbital Distance</th>
<td id="76E55D80380B00411B36348CFCD2FC36" box="[816,894,961,983]" gridcol="1" gridrow="8" pageId="9" pageNumber="210">0.618</td>
<td id="76E55D80380B00411C6C348CFB14FC36" box="[1130,1208,961,983]" gridcol="2" gridrow="8" pageId="9" pageNumber="210">0.049</td>
</tr>
<tr id="353434FC380B0041189134A7FB14FBE1" box="[151,1208,1002,1024]" gridrow="9" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189134A7FDE9FBE1" box="[151,581,1002,1024]" gridcol="0" gridrow="9" pageId="9" pageNumber="210">Snout Vent Length</th>
<td id="76E55D80380B00411B3634A7FCD2FBE1" box="[816,894,1002,1024]" gridcol="1" gridrow="9" pageId="9" pageNumber="210">0.237</td>
<td id="76E55D80380B00411C6C34A7FB14FBE1" box="[1130,1208,1002,1024]" gridcol="2" gridrow="9" pageId="9" pageNumber="210">0.130</td>
</tr>
<tr id="353434FC380B004118913359FB14FBCB" box="[151,1208,1044,1066]" gridrow="10" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913359FDE9FBCB" box="[151,581,1044,1066]" gridcol="0" gridrow="10" pageId="9" pageNumber="210">Tail Length</th>
<td id="76E55D80380B00411B363359FCD2FBCB" box="[816,894,1044,1066]" gridcol="1" gridrow="10" pageId="9" pageNumber="210">0.135</td>
<td id="76E55D80380B00411C6C3359FB14FBCB" box="[1130,1208,1044,1066]" gridcol="2" gridrow="10" pageId="9" pageNumber="210">-0.179</td>
</tr>
<tr id="353434FC380B004118913370FB14FBB2" box="[151,1208,1085,1107]" gridrow="11" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913370FDE9FBB2" box="[151,581,1085,1107]" gridcol="0" gridrow="11" pageId="9" pageNumber="210">Axila Groin Length</th>
<td id="76E55D80380B00411B363370FCD2FBB2" box="[816,894,1085,1107]" gridcol="1" gridrow="11" pageId="9" pageNumber="210">0.249</td>
<td id="76E55D80380B00411C6C3370FB14FBB2" box="[1130,1208,1085,1107]" gridcol="2" gridrow="11" pageId="9" pageNumber="210">0.259</td>
</tr>
<tr id="353434FC380B00411891332BFB14FB9D" box="[151,1208,1126,1148]" gridrow="12" pageId="9" pageNumber="210">
<th id="76E55D80380B00411891332BFDE9FB9D" box="[151,581,1126,1148]" gridcol="0" gridrow="12" pageId="9" pageNumber="210">Humerus Length</th>
<td id="76E55D80380B00411B36332BFCD2FB9D" box="[816,894,1126,1148]" gridcol="1" gridrow="12" pageId="9" pageNumber="210">-0.116</td>
<td id="76E55D80380B00411C6C332BFB14FB9D" box="[1130,1208,1126,1148]" gridcol="2" gridrow="12" pageId="9" pageNumber="210">0.521</td>
</tr>
<tr id="353434FC380B0041189133DDFB14FB47" box="[151,1208,1168,1190]" gridrow="13" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189133DDFDE9FB47" box="[151,581,1168,1190]" gridcol="0" gridrow="13" pageId="9" pageNumber="210">Ulna Length</th>
<td id="76E55D80380B00411B3633DDFCD2FB47" box="[816,894,1168,1190]" gridcol="1" gridrow="13" pageId="9" pageNumber="210">-0.040</td>
<td id="76E55D80380B00411C6C33DDFB14FB47" box="[1130,1208,1168,1190]" gridcol="2" gridrow="13" pageId="9" pageNumber="210">0.470</td>
</tr>
<tr id="353434FC380B0041189133F4FB14FB2E" box="[151,1208,1209,1231]" gridrow="14" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189133F4FDE9FB2E" box="[151,581,1209,1231]" gridcol="0" gridrow="14" pageId="9" pageNumber="210">Femur Length</th>
<td id="76E55D80380B00411B3633F4FCD2FB2E" box="[816,894,1209,1231]" gridcol="1" gridrow="14" pageId="9" pageNumber="210">-0.048</td>
<td id="76E55D80380B00411C6C33F4FB14FB2E" box="[1130,1208,1209,1231]" gridcol="2" gridrow="14" pageId="9" pageNumber="210">0.470</td>
</tr>
<tr id="353434FC380B0041189133AFFB14FB19" box="[151,1208,1250,1272]" gridrow="15" pageId="9" pageNumber="210">
<th id="76E55D80380B0041189133AFFDE9FB19" box="[151,581,1250,1272]" gridcol="0" gridrow="15" pageId="9" pageNumber="210">Tibia Length</th>
<td id="76E55D80380B00411B3633AFFCD2FB19" box="[816,894,1250,1272]" gridcol="1" gridrow="15" pageId="9" pageNumber="210">-0.017</td>
<td id="76E55D80380B00411C6C33AFFB14FB19" box="[1130,1208,1250,1272]" gridcol="2" gridrow="15" pageId="9" pageNumber="210">0.374</td>
</tr>
<tr id="353434FC380B004118913241FB14FAC3" box="[151,1208,1292,1314]" gridrow="16" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913241FDE9FAC3" box="[151,581,1292,1314]" gridcol="0" gridrow="16" pageId="9" pageNumber="210">Foot Length</th>
<td id="76E55D80380B00411B363241FCD2FAC3" box="[816,894,1292,1314]" gridcol="1" gridrow="16" pageId="9" pageNumber="210">0.145</td>
<td id="76E55D80380B00411C6C3241FB14FAC3" box="[1130,1208,1292,1314]" gridcol="2" gridrow="16" pageId="9" pageNumber="210">0.094</td>
</tr>
<tr id="353434FC380B004118913278FB14FAAA" box="[151,1208,1333,1355]" gridrow="17" pageId="9" pageNumber="210">
<th id="76E55D80380B004118913278FDE9FAAA" box="[151,581,1333,1355]" gridcol="0" gridrow="17" pageId="9" pageNumber="210">4th Toe Length</th>
<td id="76E55D80380B00411B363278FCD2FAAA" box="[816,894,1333,1355]" gridcol="1" gridrow="17" pageId="9" pageNumber="210">0.089</td>
<td id="76E55D80380B00411C6C3278FB14FAAA" box="[1130,1208,1333,1355]" gridcol="2" gridrow="17" pageId="9" pageNumber="210">0.026</td>
</tr>
<tr id="353434FC380B0041189132FEFB14FA2C" box="[151,1208,1459,1485]" gridrow="18" pageId="9" pageNumber="210" rowspan-1="1" rowspan-2="1">
<th id="76E55D80380B0041189132FEFDE9FA2C" box="[151,581,1459,1485]" gridcol="0" gridrow="18" pageId="9" pageNumber="210">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="8BBB36BE380BFFB7189132B7FA21F823" blockId="9.[151,1436,1529,1986]" pageId="9" pageNumber="210">
Our phylogenetic results show that 
<taxonomicName id="4C044D3D380BFFB71A2932B4FD6CF9F0" box="[559,704,1529,1553]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380BFFB71A2932B4FD6CF9F0" box="[559,704,1529,1553]" italics="true" pageId="9" pageNumber="210">Anolis limon</emphasis>
</taxonomicName>
is nested within the &quot; 
<taxonomicName id="4C044D3D380BFFB71BBD32B4FB89F9F3" box="[955,1061,1529,1554]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="214" phylum="Chordata" rank="genus">Dactyloa</taxonomicName>
&quot; clade of 
<taxonomicName id="4C044D3D380BFFB71C9A32B4FB48F9F0" box="[1180,1252,1529,1553]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B970EAAC380BFFB71C9A32B4FB48F9F0" box="[1180,1252,1529,1553]" italics="true" pageId="9" pageNumber="210">Anolis</emphasis>
</taxonomicName>
(
<figureCitation id="133F2A3B380BFFB71CF332B4FAF5F9F3" box="[1269,1369,1529,1554]" captionStart="FIGURE 7. A 50" captionStartId="10.[151,250,1981,2003]" captionTargetBox="[449,1144,207,1917]" captionTargetId="figure@10.[440,1152,193,1922]" captionTargetPageId="10" captionText="FIGURE 7. A 50 % majority rule consensus showing phylogenetic relationships for dactyloid anoles including Anolis limon sp. nov. Values in the nodes correspond to support values above 50 % from 500 bootstrap replicates." httpUri="https://zenodo.org/record/251963/files/figure.png" pageId="9" pageNumber="210">Figure 7</figureCitation>
). The inclusion of species scored only for morphological characters probably reduces the support values for many nodes (
<bibRefCitation id="EF954B4F380BFFB71898310CFED9F9BB" author="Huelsenbeck" box="[158,373,1601,1626]" pageId="9" pageNumber="210" refString="Huelsenbeck, J. (1991) When are fossils better than extant taxa in phylogenetic analysis? Systematic Zoology, 40, 458 - 469. http: // dx. doi. org / 10.2307 / 2992240" type="journal article" year="1991">Huelsenbeck 1991</bibRefCitation>
). However, the inclusion of these taxa with a lot of missing data may increase the accuracy of recovered phylogenetic relationships (
<bibRefCitation id="EF954B4F380BFFB71A4B3128FC83F99F" author="Wiens" box="[589,815,1637,1662]" pageId="9" pageNumber="210" refString="Wiens, J. &amp; Tiu, J. (2012) Highly incomplete taxa can rescue phylogenetic analyses from the negative impacts of limited taxon sampling. PLoS One, 7, 1 - 8. http: // dx. doi. org / 10.1371 / journal. pone. 0042925" type="journal article" year="2012">Wiens &amp; Tiu 2012</bibRefCitation>
). The results of our phylogenetic analysis reveal that although 
<taxonomicName id="4C044D3D380BFFB7190731C7FECDF940" box="[257,353,1673,1697]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380BFFB7190731C7FECDF940" box="[257,353,1673,1697]" italics="true" pageId="9" pageNumber="210">A. limon</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C044D3D380BFFB7199F31C7FD85F940" box="[409,553,1673,1697]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380BFFB7199F31C7FD85F940" box="[409,553,1673,1697]" italics="true" pageId="9" pageNumber="210">A. chocorum</emphasis>
</taxonomicName>
are very similar morphologically they are nested in different subclades inside the “ 
<taxonomicName id="4C044D3D380BFFB718CD31E0FE99F927" box="[203,309,1709,1734]" class="Squamata" family="Dactyloidae" genus="Dactyloa" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="214" phylum="Chordata" rank="genus">Dactyloa</taxonomicName>
” clade. Instead, we found that 
<taxonomicName id="4C044D3D380BFFB71A9531E3FD5EF924" box="[659,754,1709,1733]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380BFFB71A9531E3FD5EF924" box="[659,754,1709,1733]" italics="true" pageId="9" pageNumber="210">A. limon</emphasis>
</taxonomicName>
is the sister taxa to 
<taxonomicName id="4C044D3D380BFFB71BD331E3FBE9F924" box="[981,1093,1709,1733]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC380BFFB71BD331E3FBE9F924" box="[981,1093,1709,1733]" italics="true" pageId="9" pageNumber="210">A. ibanezi</emphasis>
</taxonomicName>
, although the nodal support is too low (below 50% of bootstrap replicates). However, we only included molecular data for 
<taxonomicName id="4C044D3D380BFFB71CA1319CFACAF908" box="[1191,1382,1745,1769]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380BFFB71CA1319CFACAF908" box="[1191,1382,1745,1769]" italics="true" pageId="9" pageNumber="210">Anolis chocorum</emphasis>
</taxonomicName>
and a further study (including molecular data) it is necessary to highlights the phylogenetic relationships of mainland green anoles (
<taxonomicName id="4C044D3D380BFFB719463054FE79F8D0" box="[320,469,1817,1841]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC380BFFB719463054FE79F8D0" box="[320,469,1817,1841]" italics="true" pageId="9" pageNumber="210">Anolis limon</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C044D3D380BFFB719EF3057FDF3F8D0" box="[489,607,1817,1841]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="ibanezi">
<emphasis id="B970EAAC380BFFB719EF3057FDF3F8D0" box="[489,607,1817,1841]" italics="true" pageId="9" pageNumber="210">A. ibanezi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C044D3D380BFFB71AA43057FC9BF8D0" box="[674,823,1817,1841]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="210" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC380BFFB71AA43057FC9BF8D0" box="[674,823,1817,1841]" italics="true" pageId="9" pageNumber="210">A. chocorum</emphasis>
</taxonomicName>
). The recent phylogenies of dactyloid anoles (see Castañeda &amp; de 
<bibRefCitation id="EF954B4F380BFFB719513070FE50F8B7" author="Queiroz" box="[343,508,1853,1878]" pageId="9" pageNumber="210" refString="Castaneda, M. R. &amp; de Queiroz, K. (2011) Phylogenetic relationships of the Dactyloa clade of Anolis lizards based on nuclear and mitochondrial DNA sequence data. Molecular phylogenetics and evolution, 61, 784 - 800. http: // dx. doi. org / 10.1016 / j. ympev. 2011.07.004" type="journal article" year="2011">Queiroz, 2011</bibRefCitation>
, 
<bibRefCitation id="EF954B4F380BFFB71A0C3073FDE8F8B7" author="Queiroz" box="[522,580,1854,1878]" pageId="9" pageNumber="210" refString="Castaneda, M. R. &amp; de Queiroz, K. (2013) Phylogeny of the Dactyloa Clade of Anolis Lizards: New Insights from Combining Morphological and Molecular Data. Bulletin of the Museum of Comparative Zoology, 160, 345 - 398. http: // dx. doi. org / 10.3099 / 0027 - 4100 - 160.7.345" type="journal article" year="2013">2013</bibRefCitation>
) only have included the half of dactyloid taxa described (Castañeda &amp; de 
<bibRefCitation id="EF954B4F380BFFB71891302CFE91F89B" author="Queiroz" box="[151,317,1889,1914]" pageId="9" pageNumber="210" refString="Castaneda, M. R. &amp; de Queiroz, K. (2011) Phylogenetic relationships of the Dactyloa clade of Anolis lizards based on nuclear and mitochondrial DNA sequence data. Molecular phylogenetics and evolution, 61, 784 - 800. http: // dx. doi. org / 10.1016 / j. ympev. 2011.07.004" type="journal article" year="2011">Queiroz, 2011</bibRefCitation>
) and in some cases, taxa with many missing data, mostly for molecular data (Castañeda &amp; de Quieroz, 2013). We considered that a better taxa sampling combined with a most complete character sampling (especially for molecular data) it is necessary to improve our understanding of the phylogeny of dactyloid anoles.
</paragraph>
<caption id="DF7B66363808FFB4189130F0FBC0F810" httpUri="https://zenodo.org/record/251963/files/figure.png" pageId="10" pageNumber="211" targetBox="[449,1144,207,1917]" targetPageId="10">
<paragraph id="8BBB36BE3808FFB4189130F0FBC0F810" blockId="10.[151,1436,1980,2033]" pageId="10" pageNumber="211">
<emphasis id="B970EAAC3808FFB4189130F0FEBEF833" bold="true" box="[151,274,1981,2003]" pageId="10" pageNumber="211">FIGURE 7.</emphasis>
A 50% majority rule consensus showing phylogenetic relationships for dactyloid anoles including 
<taxonomicName id="4C044D3D3808FFB41CFE30F1FAD4F833" box="[1272,1400,1980,2002]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="211" phylum="Chordata" rank="species" species="limon" status="sp. nov.">
<emphasis id="B970EAAC3808FFB41CFE30F1FAD4F833" box="[1272,1400,1980,2002]" italics="true" pageId="10" pageNumber="211">Anolis limon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24357D73808FFB41D7B30F0FF6FF810" pageId="10" pageNumber="211" rank="species">
<emphasis id="B970EAAC3808FFB41D7B30F0FF6FF810" bold="true" pageId="10" pageNumber="211">sp. nov.</emphasis>
</taxonomicNameLabel>
Values in the nodes correspond to support values above 50% from 500 bootstrap replicates.
</paragraph>
</caption>
<paragraph id="8BBB36BE3809FFB518C137D5FAC4FF19" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="212">
Our morphometric analyses suggested that the females of 
<taxonomicName id="4C044D3D3809FFB51B5237D5FC4FFF51" box="[852,995,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3809FFB51B5237D5FC4FFF51" box="[852,995,152,176]" italics="true" pageId="11" pageNumber="212">Anolis limon</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C044D3D3809FFB51C1D37D5FB76FF51" box="[1051,1242,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="chocorum">
<emphasis id="B970EAAC3809FFB51C1D37D5FB76FF51" box="[1051,1242,152,176]" italics="true" pageId="11" pageNumber="212">Anolis chocorum</emphasis>
</taxonomicName>
differs mainly in head variables (see 
<tableCitation id="C68603053809FFB5197037F1FE66FF35" box="[374,458,188,213]" captionStart="TABLE 2" captionStartId="9.[151,239,548,571]" captionTargetBox="[151,1208,632,1485]" captionTargetPageId="9" captionText="TABLE 2. Results of principal component analysis (the first two PC axes) for 15 morphological variables foradult females of Anolis limon sp. nov. and Anolis chocorum." httpUri="http://table.plazi.org/id/DF7B6636380BFFB718913569FD4DFDBD" pageId="11" pageNumber="212" tableUuid="DF7B6636380BFFB718913569FD4DFDBD">Table 2</tableCitation>
and 
<figureCitation id="133F2A3B3809FFB51A0537F1FDC9FF34" box="[515,613,188,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1174,1196]" captionTargetBox="[158,1424,199,1148]" captionTargetId="figure@8.[151,1436,193,1153]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for Anolis limon sp. nov. and closest putative relatives. Circles in white for A. chocorum correspond to specimens and observations checked by one of us (JAV). The locality information for the rest of occurrence records were taken from original descriptions (Williams &amp; Duellman 1967, Poe et al. 2009) and recent literature (Lotzkat et al. 2013)." httpUri="https://zenodo.org/record/251961/files/figure.png" pageId="11" pageNumber="212">Figure 5</figureCitation>
). These morphological differences likely are associated to differences in diet and bite force (Herrel 
<emphasis id="B970EAAC3809FFB519C637ACFE54FF19" box="[448,504,224,248]" italics="true" pageId="11" pageNumber="212">et al.</emphasis>
2001a; b), reflecting niche partitioning between species (
<bibRefCitation id="EF954B4F3809FFB51C7D37ADFAF0FF19" author="Harmon" box="[1147,1372,223,248]" pageId="11" pageNumber="212" refString="Harmon, L. J., Kolbe, J. J., Cheverud, J. M. &amp; Losos, J. B. (2005) Convergence and the multidimensional niche. Evolution; international journal of organic evolution, 59, 409 - 421. http: // dx. doi. org / 10.1554 / 04 - 038" type="journal article" year="2005">
Harmon 
<emphasis id="B970EAAC3809FFB51CE637ACFAB4FF19" box="[1248,1304,224,248]" italics="true" pageId="11" pageNumber="212">et al.</emphasis>
2005
</bibRefCitation>
).
</paragraph>
<paragraph id="8BBB36BE3809FFB518C13649FE12FE14" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="212">
The distribution of 
<taxonomicName id="4C044D3D3809FFB519AE3649FD95FEFD" box="[424,569,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3809FFB519AE3649FD95FEFD" box="[424,569,260,284]" italics="true" pageId="11" pageNumber="212">Anolis limon</emphasis>
</taxonomicName>
is restricted to the middle Magdalena region of 
<collectingCountry id="F313762E3809FFB51C6B3649FB77FEFC" box="[1133,1243,260,285]" name="Colombia" pageId="11" pageNumber="212">Colombia</collectingCountry>
(
<figureCitation id="133F2A3B3809FFB51CEB3649FAFEFEFC" box="[1261,1362,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1174,1196]" captionTargetBox="[158,1424,199,1148]" captionTargetId="figure@8.[151,1436,193,1153]" captionTargetPageId="8" captionText="FIGURE 5. Distribution map for Anolis limon sp. nov. and closest putative relatives. Circles in white for A. chocorum correspond to specimens and observations checked by one of us (JAV). The locality information for the rest of occurrence records were taken from original descriptions (Williams &amp; Duellman 1967, Poe et al. 2009) and recent literature (Lotzkat et al. 2013)." httpUri="https://zenodo.org/record/251961/files/figure.png" pageId="11" pageNumber="212">Figure 5</figureCitation>
). This restricted distribution provide additional support for the hypothesis that Magdalena region is an endemism area (Hernández-Camacho 
<emphasis id="B970EAAC3809FFB519933600FE7CFE85" box="[405,464,332,356]" italics="true" pageId="11" pageNumber="212">et al.</emphasis>
1992; Morrone 2001). Several species are known to be restricted to this region, like monkeys (e.g, 
<taxonomicName id="4C044D3D3809FFB5194A363DFD76FE69" authority="Gunther 1877" authorityName="Gunther" authorityYear="1877" box="[332,730,367,392]" class="Mammalia" family="Callitrichidae" genus="Saguinus" kingdom="Animalia" order="Primates" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="leucopus">
<emphasis id="B970EAAC3809FFB5194A363DFD89FE69" box="[332,549,368,392]" italics="true" pageId="11" pageNumber="212">Saguinus leucopus</emphasis>
Günther 1877
</taxonomicName>
), birds (e.g., 
<taxonomicName id="4C044D3D3809FFB51B82363DFB53FE69" authority="Salvin 1897" authorityName="Salvin" authorityYear="1897" box="[900,1279,367,392]" class="Aves" family="Ramphastidae" genus="Capito" kingdom="Animalia" order="Piciformes" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="hypoleucus">
<emphasis id="B970EAAC3809FFB51B82363DFBF2FE69" box="[900,1118,368,392]" italics="true" pageId="11" pageNumber="212">Capito hypoleucus</emphasis>
<bibRefCitation id="EF954B4F3809FFB51C6B3622FB53FE69" author="Salvin" box="[1133,1279,367,392]" pageId="11" pageNumber="212" refString="Salvin, O. (1897) Descriptions of five species of southamerican birds. Bulletin of the British Ornithologists' Club, 7, 15 - 17." type="journal article" year="1897">Salvin 1897</bibRefCitation>
</taxonomicName>
), frogs (e.g., 
<taxonomicName id="4C044D3D3809FFB5189136D9FCE0FE4D" authority="Rivera, Duarte, Rueda &amp; Daza, 2013" authorityName="Rivera, Duarte, Rueda &amp; Daza" authorityYear="2013" box="[151,844,404,429]" class="Amphibia" family="Hylidae" genus="Agalychnis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="terranova">
<emphasis id="B970EAAC3809FFB5189136D9FE20FE4D" box="[151,396,404,428]" italics="true" pageId="11" pageNumber="212">Agalychnis terranova</emphasis>
Rivera, Duarte, Rueda &amp; 
<bibRefCitation id="EF954B4F3809FFB51AC036D8FCE0FE4D" author="Daza" box="[710,844,404,428]" pageId="11" pageNumber="212" refString="Rivera-Correa, M., Duarte-Cubides, F., Rueda-Almonacid, J. V. &amp; Daza, J. M. (2013) A new red-eyed treefrog of Agalychnis (Anura: Hylidae: Phyllomedusinae) from middle Magdalena River valley of Colombia with comments on its phylogenetic position. Zootaxa, 3636 (1), 85 - 100. http: // dx. doi. org / 10.11646 / zootaxa. 3636.1.4" type="journal article" year="2013">Daza, 2013</bibRefCitation>
</taxonomicName>
;, 
<taxonomicName id="4C044D3D3809FFB51B6536D9FA3BFE4D" authority="Grant, Acosta &amp; Rada 2007" authorityName="Grant, Acosta &amp; Rada" authorityYear="2007" box="[867,1431,404,429]" class="Amphibia" family="Aromobatidae" genus="Allobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="niputidea">
<emphasis id="B970EAAC3809FFB51B6536D9FBEEFE4D" box="[867,1090,404,428]" italics="true" pageId="11" pageNumber="212">Allobates niputidea</emphasis>
Grant, Acosta &amp; 
<bibRefCitation id="EF954B4F3809FFB51D1E36D9FA3BFE4D" author="Rada" box="[1304,1431,404,429]" pageId="11" pageNumber="212" refString="Grant, T., Acosta-Galvis, A. R. &amp; Rada, M. (2007) A name for the species of Allobates (Anura: Dendrobatoidea: Aromobatidae) from the Magdalena Valley of Colombia. Copeia, 2007, 844 - 854. http: // dx. doi. org / 10.1643 / 0045 - 8511 (2007) 7 [844: anftso] 2.0. co; 2" type="journal article" year="2007">Rada 2007</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C044D3D3809FFB5189136F5FDB6FE31" authority="Cope 1861" authorityName="Cope" authorityYear="1861" box="[151,538,439,464]" class="Amphibia" family="Dendrobatidae" genus="Dendrobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="truncatus">
<emphasis id="B970EAAC3809FFB5189136F5FE3BFE31" box="[151,407,440,464]" italics="true" pageId="11" pageNumber="212">Dendrobates truncatus</emphasis>
Cope 1861
</taxonomicName>
), turtles (e.g., 
<taxonomicName id="4C044D3D3809FFB51ABC36F5FBE6FE31" authority="Dumeril 1852" authorityName="Dumeril" authorityYear="1852" box="[698,1098,439,464]" class="Reptilia" family="Podocnemididae" genus="Podocnemis" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="lewyana">
<emphasis id="B970EAAC3809FFB51ABC36F5FC0BFE31" box="[698,935,440,464]" italics="true" pageId="11" pageNumber="212">Podocnemis lewyana</emphasis>
Duméril 1852
</taxonomicName>
) and other anole species (e.g., 
<taxonomicName id="4C044D3D3809FFB518913691FE1DFE14" authority="Williams 1975" authorityName="Williams" authorityYear="1975" box="[151,433,476,501]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="ibague">
<emphasis id="B970EAAC3809FFB518913691FEAEFE15" box="[151,258,476,500]" italics="true" pageId="11" pageNumber="212">A. ibague</emphasis>
<bibRefCitation id="EF954B4F3809FFB5190F3691FE1DFE14" author="Williams" box="[265,433,476,501]" pageId="11" pageNumber="212" refString="Williams, E. E. (1975) South American Anolis: Anolis ibague, New Species of the pentaprion group from Colombia. Breviora, 433, 1 - 10." type="journal article" year="1975">Williams 1975</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BBB36BE3809FFB518C136B2FC66FC61" blockId="11.[151,1436,151,896]" pageId="11" pageNumber="212">
The discovery of this new species highlights the need for extensive work with Colombian anoles. Additional research in Colombian herpetological collections supplemented by extensive fieldwork at several sites across the country likely will allow the discovery of new anole taxa. With the description of 
<taxonomicName id="4C044D3D3809FFB51C523505FB4AFD81" box="[1108,1254,584,608]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="limon">
<emphasis id="B970EAAC3809FFB51C523505FB4AFD81" box="[1108,1254,584,608]" italics="true" pageId="11" pageNumber="212">Anolis limon</emphasis>
</taxonomicName>
we have added another species to the Colombian anole fauna for a total of 75 species (
<bibRefCitation id="EF954B4F3809FFB51BE23521FB7AFD65" author="Uetz" box="[996,1238,620,645]" pageId="11" pageNumber="212" refString="Uetz, P. &amp; Hosek, J. (2013) The Reptile Database. Available from: www. reptile-database. org (accessed 20 July 2013)" type="journal article" year="2013">Uetz &amp; Hosek 2013</bibRefCitation>
). Despite limited sampling effort dedicated to anoles in 
<collectingCountry id="F313762E3809FFB51A6935C2FD4CFD49" box="[623,736,655,680]" name="Colombia" pageId="11" pageNumber="212">Colombia</collectingCountry>
, in comparison with other regions (e.g., Greater Antilles) 
<collectingCountry id="F313762E3809FFB5189135F9FEA9FD2C" box="[151,261,692,717]" name="Colombia" pageId="11" pageNumber="212">Colombia</collectingCountry>
has the highest anole diversity of any country (
<bibRefCitation id="EF954B4F3809FFB51B1335F9FC56FD2D" author="Uetz" box="[789,1018,692,717]" pageId="11" pageNumber="212" refString="Uetz, P. &amp; Hosek, J. (2013) The Reptile Database. Available from: www. reptile-database. org (accessed 20 July 2013)" type="journal article" year="2013">Uetz &amp; Hosek 2013</bibRefCitation>
). The high diversity of 
<taxonomicName id="4C044D3D3809FFB51CFB35F9FAE9FD2D" box="[1277,1349,692,716]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="genus">
<emphasis id="B970EAAC3809FFB51CFB35F9FAE9FD2D" box="[1277,1349,692,716]" italics="true" pageId="11" pageNumber="212">Anolis</emphasis>
</taxonomicName>
species in 
<collectingCountry id="F313762E3809FFB518B1359AFE89FD11" box="[183,293,727,752]" name="Colombia" pageId="11" pageNumber="212">Colombia</collectingCountry>
offers an excellent opportunity for addressing evolutionary and ecological questions (see Molina- 
<bibRefCitation id="EF954B4F3809FFB5189135B1FDE6FCF5" author="Zuluaga" box="[151,586,764,789]" pageId="11" pageNumber="212" refString="Molina-Zuluaga, C. &amp; Gutierrez-Cardenas, P. D. A. (2007) Uso nocturno de perchas en dos especies de Anolis (Squamata: Polychrotidae) en un bosque Andino de Colombia. Papeis Avulsos de Zoologia, 47, 273 - 281." type="journal article" year="2007">Zuluaga &amp; Gutiérrez-Cárdenas 2007</bibRefCitation>
; 
<bibRefCitation id="EF954B4F3809FFB51A5B35B1FCD5FCF5" author="Velasco" box="[605,889,764,789]" pageId="11" pageNumber="212" refString="Velasco, J. &amp; Herrel, A. (2007) Ecomorphology of Anolis lizards of the Choc region in Colombia and comparisons with Greater Antillean ecomorphs. Biological Journal of the Linnean Society, 92, 29 - 39. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2007.00885. x" type="journal article" year="2007">Velasco &amp; Herrel 2007</bibRefCitation>
; 
<bibRefCitation id="EF954B4F3809FFB51B8A35B1FA3BFCF5" author="Calderon-Espinosa" box="[908,1431,764,789]" pageId="11" pageNumber="212" refString="Calderon-Espinosa, M. L. &amp; Barragan-Forero, A. (2011) Morphological Diversification in Solitary Endemic Anoles: Anolis concolor and Anolis pinchoti from San Andres and Providence Islands, Colombia. South American Journal of Herpetology, 6, 205 - 210. http: // dx. doi. org / 10.2994 / 057.006.0306" type="journal article" year="2011">Calderón-Espinosa &amp; Barragán-Forero 2011</bibRefCitation>
; Calderón-Espinosa 
<emphasis id="B970EAAC3809FFB51972346CFE03FCD9" box="[372,431,800,824]" italics="true" pageId="11" pageNumber="212">et al.</emphasis>
2013; Moreno-Arias &amp; Urbina-Cardona 2013). We encourage herpetologists to conduct biological studies focused on Colombian anole species. Such studies will contribute to a greater understanding of the ecological and evolutionary causes of the high diversity in 
<collectingCountry id="F313762E3809FFB51B52342AFC6AFC61" box="[852,966,871,896]" name="Colombia" pageId="11" pageNumber="212">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>