treatments-xml/data/03/AD/87/03AD879AF86DFFF3FF63FAB453E8FE26.xml

<document ID-DOI="10.11646/zootaxa.5256.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7755108" ID-ZooBank="DF67CEDA-3845-45FC-8E82-687F14CE2ACE" checkinTime="1679383099089" checkinUser="plazi" docAuthor="Li, Zihe, Jin, Zezhong, Polhemus, Dan A. &amp; Ye, Zhen" docDate="2023" docId="03AD879AF86DFFF3FF63FAB453E8FE26" docLanguage="en" docName="zootaxa.5256.4.2.pdf" docOrigin="Zootaxa 5256 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Valleriola asiatica Li, Polhemus &amp; Ye 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="342" masterDocId="FF94FFE2F866FFFEFFF4FFE2505FFFF7" masterDocTitle="Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia" masterLastPageNumber="344" masterPageNumber="329" pageNumber="340" updateTime="1679383513048" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Two new species of Valleriola (Hemiptera: Heteroptera: Leptopodidae) and taxonomic notes on the tribe Leptopodini Brullé, 1836 from East and Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Zihe</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-0332-4356</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China. &amp; https: // orcid. org / 0000 - 0002 - 0332 - 4356</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jin, Zezhong</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7223-770X</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China. &amp; https: // orcid. org / 0000 - 0001 - 7223 - 770 X</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Polhemus, Dan A.</mods:namePart>
<mods:affiliation>Department of Natural Sciences, Bishop Museum, 1525 Bernice Street, Honolulu, HI 96817, USA.</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ye, Zhen</mods:namePart>
<mods:affiliation>Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-21</mods:number>
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<mods:number>4</mods:number>
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<taxonomicName authority="Li, Polhemus &amp; Ye" authorityName="Li, Polhemus &amp; Ye" authorityYear="2023" box="[151,612,1366,1393]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="340" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis bold="true" box="[151,369,1366,1393]" italics="true" pageId="11" pageNumber="340">Valleriola asiatica</emphasis>
Li, Polhemus &amp; Ye
</taxonomicName>
, 
<taxonomicNameLabel box="[624,718,1367,1393]" pageId="11" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph blockId="11.[151,718,1366,1430]" box="[151,610,1403,1430]" pageId="11" pageNumber="340">
(
<figureCitation box="[159,239,1403,1429]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="11" pageNumber="340">Figs. 2</figureCitation>
, 
<figureCitation box="[251,267,1403,1429]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="11" pageNumber="340">7</figureCitation>
, 
<figureCitation box="[279,352,1403,1429]" captionStart="FIGURES 13–27" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-17@5.[284,1303,181,1919]" captionTargetPageId="5" captionText="FIGURES 13–27. Heads of Leptopus riparius Hsiao, 1964 and Valleriola spp. in frontal view, lateral view and ventral view. (13–15) L. riparius Hsiao, 1964; (16–18) V. asiatica sp. nov.; (19–21) V. buenoi (Usinger, 1942); (22–24) V. bui sp. nov.; (25–27) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7755120" httpUri="https://zenodo.org/record/7755120/files/figure.png" pageId="11" pageNumber="340">16–18</figureCitation>
, 
<figureCitation box="[363,393,1403,1429]" captionStart="FIGURES 28–33" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[397,1190,193,1870]" captionTargetId="figure-23@6.[384,1203,181,1884]" captionTargetPageId="6" captionText="FIGURES 28–33. Hemelytra of Leptopus riparius Hsiao, 1964 and Valleriola spp. (28) right hemelytron of Valleriola; (29) L. riparius Hsiao, 1964; (30) V. asiatica sp. nov.; (31) V. buenoi (Usinger, 1942); (32) V. bui sp. nov.; (33) V. javanica Drake &amp; Hottes, 1951. Sc, subcosta; R, radius; M, media; Cu, cubitus; Cs, claval suture; r, cell R; m, cell M; cu, cell Cu; sc, cell Sc; I, 1st membrane cell; II, 2nd membrane cell; III, 3rd membrane cell; IV, 4th membrane cell. Scale 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755124" httpUri="https://zenodo.org/record/7755124/files/figure.png" pageId="11" pageNumber="340">30</figureCitation>
, 
<figureCitation box="[405,478,1403,1429]" captionStart="FIGURES 34–48" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[170,1413,194,1457]" captionTargetId="figure-17@7.[151,1435,181,1468]" captionTargetPageId="7" captionText="FIGURES 34–48. Left fore femora of Leptopus riparius Hsiao, 1964 and Valleriola spp. in interior view, ventral view and exterior view. (34–36) L. riparius Hsiao, 1964; (40–42) V. buenoi (Usinger, 1942); (37–39) V. asiatica sp. nov.; (43–45) V. bui sp. nov.; (46–48) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm. Red arrows show the positions of large spines." figureDoi="http://doi.org/10.5281/zenodo.7755126" httpUri="https://zenodo.org/record/7755126/files/figure.png" pageId="11" pageNumber="340">37–39</figureCitation>
, 
<figureCitation box="[489,561,1403,1430]" captionStart="FIGURES 51–60" captionStartId="9.[151,264,1879,1903]" captionTargetBox="[187,1398,538,1830]" captionTargetId="figure-152@9.[163,1425,525,1856]" captionTargetPageId="9" captionText="FIGURES 51–60. Right parameres of Leptopus riparius and Valleriola spp. in different views. (51–52) L. riparius Hsiao, 1964, dorsal and interior views; (53–54) V. asiatica sp. nov., dorsal and interior views; (55–56) V. buenoi (Usinger, 1942), dorsal and interior views; (57–59) V. bui sp. nov., dorsal, exterior, and interior views; (60) V. javanica Drake &amp; Hottes, 1951, dorsal view. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7755132" httpUri="https://zenodo.org/record/7755132/files/figure.png" pageId="11" pageNumber="340">53, 54</figureCitation>
, 
<figureCitation box="[573,602,1403,1430]" captionStart="FIGURES 62–65" captionStartId="11.[151,264,1224,1248]" captionTargetBox="[151,1435,181,1191]" captionTargetId="figure-17@11.[151,1435,181,1201]" captionTargetPageId="11" captionText="FIGURES 62–65. In situ photographs of species of Leptopodini. (62) L. riparius Hsiao, 1964; (63) V. bui sp. nov.; (64) V. javanica Drake &amp; Hottes, 1951; (65) V. asiatica sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7755138" httpUri="https://zenodo.org/record/7755138/files/figure.png" pageId="11" pageNumber="340">65</figureCitation>
)
</paragraph>
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<emphasis bold="true" box="[151,385,1474,1500]" pageId="11" pageNumber="340">Material examined.</emphasis>
<materialsCitation collectingDate="2017-07-28" collectionCode="NKUM" collectorName="Zhen Ye" country="China" location="Bawangling Nature Reserve" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus box="[392,501,1475,1501]" pageId="11" pageNumber="340">Holotype</typeStatus>
: male, 
<collectingCountry box="[579,671,1475,1501]" name="China" pageId="11" pageNumber="340">CHINA</collectingCountry>
, 
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<collectingRegion box="[682,840,1474,1501]" country="China" name="Hainan" pageId="11" pageNumber="340">Hainan Prov.</collectingRegion>
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, 
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, 
<date box="[1184,1332,1475,1501]" pageId="11" pageNumber="340" value="2017-07-28">
<collectingDate box="[1184,1332,1475,1501]" pageId="11" pageNumber="340" value="2017-07-28">28 July 2017</collectingDate>
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, 
<collectorName box="[1343,1436,1475,1501]" pageId="11" pageNumber="340">Zhen Ye</collectorName>
leg. (
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)
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; 
<materialsCitation box="[322,1328,1510,1537]" collectingDate="2017-07-28" collectionCode="NKUM" collectorName="Zhen Ye" country="China" location="Bawangling Nature Reserve" pageId="11" pageNumber="340" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hainan" typeStatus="paratype">
<typeStatus box="[322,435,1511,1537]" pageId="11" pageNumber="340">Paratypes</typeStatus>
: 
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, 
<specimenCount box="[552,665,1511,1537]" pageId="11" pageNumber="340" type="female">3 females</specimenCount>
, 
<collectingCountry box="[678,770,1511,1537]" name="China" pageId="11" pageNumber="340">CHINA</collectingCountry>
, 
<emphasis bold="true" box="[783,943,1510,1537]" pageId="11" pageNumber="340">
<collectingRegion box="[783,943,1510,1537]" country="China" name="Hainan" pageId="11" pageNumber="340">Hainan Prov.</collectingRegion>
</emphasis>
, same data as holotype (
<collectionCode box="[1228,1320,1511,1537]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="11" pageNumber="340">NKUM</collectionCode>
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</materialsCitation>
; 
<materialsCitation collectingDate="2019-08-18" collectionCode="NKUM" collectorName="Yanfei Li" country="Laos" location="Sae Pra Waterfall" pageId="11" pageNumber="340" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Attapu" typeStatus="paratype">
<specimenCount box="[1341,1432,1511,1537]" pageId="11" pageNumber="340" type="male">3 males</specimenCount>
, 
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, 
<collectingCountry box="[275,351,1547,1573]" name="Laos" pageId="11" pageNumber="340">LAOS</collectingCountry>
, 
<emphasis bold="true" box="[363,447,1546,1572]" pageId="11" pageNumber="340">
<collectingRegion box="[363,447,1546,1572]" country="Laos" name="Attapu" pageId="11" pageNumber="340">Attapu</collectingRegion>
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, 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FE39F9E952CAF9D2" box="[461,661,1547,1573]" country="Laos" name="Sae Pra Waterfall" pageId="11" pageNumber="340" stateProvince="Attapu">Sae Pra Waterfall</location>
, 
<date box="[672,854,1547,1573]" pageId="11" pageNumber="340" value="2019-08-18">
<collectingDate box="[672,854,1547,1573]" pageId="11" pageNumber="340" value="2019-08-18">18 August 2019</collectingDate>
</date>
, 
<collectorName box="[865,971,1547,1573]" pageId="11" pageNumber="340">Yanfei Li</collectorName>
leg. (
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</materialsCitation>
. 
<materialsCitation collectingDate="1998-05-09" collectionCode="T" country="Vietnam" county="Suoi Vang" elevation="1430" latitude="11.990556" location="Dalat" longLatPrecision="21" longitude="108.36833" municipality="Gold Stream" pageId="11" pageNumber="340" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
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, 
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, 
<collectingRegion country="Vietnam" name="Lam Dong" pageId="11" pageNumber="340">
<emphasis bold="true" pageId="11" pageNumber="340">Lam Dong Prov</emphasis>
.
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, stream below 
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hydro project (= 
<collectingMunicipality box="[787,936,1583,1609]" pageId="11" pageNumber="340">Gold Stream</collectingMunicipality>
), 
<quantity box="[956,1032,1583,1609]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="340" unit="km" value="16.0">16 km</quantity>
. N. of 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FBADF9CD54C7F9BE" box="[1113,1176,1583,1609]" country="Vietnam" county="Suoi Vang" latitude="11.990556" longLatPrecision="21" longitude="108.36833" municipality="Gold Stream" name="Dalat" pageId="11" pageNumber="340" stateProvince="Lam Dong">Dalat</location>
, 
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<elevation box="[1189,1281,1583,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="340" unit="m" value="1430.0">1430 m</elevation>
</quantity>
., 
<geoCoordinate box="[1298,1431,1583,1610]" degrees="11" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="340" precision="15" seconds="26" value="11.990556">11°59'26&quot;N</geoCoordinate>
, 
<geoCoordinate box="[151,296,1619,1645]" degrees="108" direction="east" minutes="22" orientation="longitude" pageId="11" pageNumber="340" precision="15" seconds="06" value="108.36833">108°22'06&quot;E</geoCoordinate>
, 
<date box="[309,448,1619,1645]" pageId="11" pageNumber="340" value="1998-05-09">
<collectingDate box="[309,448,1619,1645]" pageId="11" pageNumber="340" value="1998-05-09">9 May 1998</collectingDate>
</date>
, CL 3078, J. 
<collectionCode box="[599,617,1619,1645]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[628,850,1619,1645]" collectionCode="USNM" collectorName="Polhemus" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<collectorName box="[628,738,1619,1645]" pageId="11" pageNumber="340">Polhemus</collectorName>
(
<collectionCode box="[754,841,1619,1645]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Vietnam" elevation="825" location="Pongour Falls" pageId="11" pageNumber="340" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCount box="[861,951,1619,1645]" pageId="11" pageNumber="340" type="male">7 males</specimenCount>
, 
<specimenCount box="[963,1075,1619,1645]" pageId="11" pageNumber="340" type="female">6 females</specimenCount>
, 
<collectingCountry box="[1086,1218,1619,1645]" name="Vietnam" pageId="11" pageNumber="340">VIETNAM</collectingCountry>
, 
<collectingRegion box="[1230,1428,1619,1645]" country="Vietnam" name="Lam Dong" pageId="11" pageNumber="340">
<emphasis bold="true" box="[1230,1422,1619,1645]" pageId="11" pageNumber="340">Lam Dong Prov</emphasis>
.
</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FF63F9955169F966" box="[151,310,1655,1681]" country="Vietnam" name="Pongour Falls" pageId="11" pageNumber="340" stateProvince="Lam Dong">Pongour Falls</location>
, 
<quantity box="[322,396,1655,1681]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="11" pageNumber="340" unit="km" value="62.0">62 km</quantity>
. 
<locationDeviation box="[407,549,1655,1681]" location="Dalat" pageId="11" pageNumber="340">SW of Dalat</locationDeviation>
, 
<quantity box="[561,637,1655,1682]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="11" pageNumber="340" unit="m" value="825.0">
<specimenCount box="[561,637,1655,1682]" pageId="11" pageNumber="340" type="generic">
<elevation box="[561,637,1655,1682]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="11" pageNumber="340" unit="m" value="825.0">825 m</elevation>
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</materialsCitation>
<materialsCitation box="[637,1315,1655,1682]" collectingDate="1998-05-13" collectingDateMax="1998-05-16" collectingDateMin="1998-05-13" collectionCode="T" country="Vietnam" latitude="11.688611" longLatPrecision="21" longitude="108.265274" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
., 
<geoCoordinate box="[653,785,1655,1681]" degrees="11" direction="north" minutes="41" orientation="latitude" pageId="11" pageNumber="340" precision="15" seconds="19" value="11.688611">11°41'19&quot;N</geoCoordinate>
, 
<geoCoordinate box="[798,943,1655,1682]" degrees="108" direction="east" minutes="15" orientation="longitude" pageId="11" pageNumber="340" precision="15" seconds="55" value="108.265274">108°15'55&quot;E</geoCoordinate>
, 
<date box="[954,1149,1655,1681]" pageId="11" pageNumber="340" value="1998-05-13" valueMax="1998-05-16" valueMin="1998-05-13">
<collectingDate box="[954,1149,1655,1681]" pageId="11" pageNumber="340" value="1998-05-13" valueMax="1998-05-16" valueMin="1998-05-13">13–16 May 1998</collectingDate>
</date>
, CL 3091, J. 
<collectionCode box="[1297,1315,1655,1681]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation collectionCode="BPBM, USNM" collectorName="Polhemus" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<collectorName box="[1326,1436,1655,1681]" pageId="11" pageNumber="340">Polhemus</collectorName>
(
<collectionCode box="[159,241,1691,1717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="340" type="Museum">BPBM</collectionCode>
, 
<collectionCode box="[252,338,1691,1717]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Vietnam" elevation="110" location="River" municipality="Cat Tien National Park" pageId="11" pageNumber="340" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Dong Nai" typeStatus="paratype">
<specimenCount box="[356,433,1691,1717]" pageId="11" pageNumber="340" type="male">1 male</specimenCount>
, 
<specimenCount box="[444,543,1691,1717]" pageId="11" pageNumber="340" type="female">1 female</specimenCount>
, 
<collectingCountry box="[553,685,1691,1717]" name="Vietnam" pageId="11" pageNumber="340">VIETNAM</collectingCountry>
, 
<collectingRegion box="[695,877,1690,1717]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="340">
<emphasis bold="true" box="[695,871,1690,1717]" pageId="11" pageNumber="340">Dong Nai Prov</emphasis>
.
</collectingRegion>
, 
<collectingMunicipality box="[891,1149,1691,1717]" pageId="11" pageNumber="340">Cat Tien National Park</collectingMunicipality>
, 
<collectingRegion box="[1160,1268,1691,1717]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="340">Dong Nai</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FB0FF9795564F942" box="[1275,1339,1691,1717]" country="Vietnam" municipality="Cat Tien National Park" name="River" pageId="11" pageNumber="340" stateProvince="Dong Nai">River</location>
, cascade upstream of park HQ, 
<quantity box="[413,490,1727,1753]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="340" unit="m" value="110.0">
<specimenCount box="[413,490,1727,1753]" pageId="11" pageNumber="340" type="generic">
<elevation box="[413,490,1727,1753]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="340" unit="m" value="110.0">110 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation box="[490,828,1727,1753]" collectingDate="1998-05-07" collectionCode="T" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
., 
<date box="[509,652,1727,1753]" pageId="11" pageNumber="340" value="1998-05-07">
<collectingDate box="[509,652,1727,1753]" pageId="11" pageNumber="340" value="1998-05-07">7 May 1998</collectingDate>
</date>
, CL 3073, J. 
<collectionCode box="[810,828,1727,1753]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[842,1065,1727,1753]" collectionCode="USNM" collectorName="Polhemus" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
<collectorName box="[842,952,1727,1753]" pageId="11" pageNumber="340">Polhemus</collectorName>
(
<collectionCode box="[970,1057,1727,1753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Vietnam" elevation="120" location="River" municipality="Cat Tien National Park" pageId="11" pageNumber="340" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Dong Nai" typeStatus="paratype">
<specimenCount box="[1079,1159,1727,1753]" pageId="11" pageNumber="340" type="male">1 male</specimenCount>
, 
<specimenCount box="[1173,1287,1727,1753]" pageId="11" pageNumber="340" type="female">3 females</specimenCount>
, 
<collectingCountry box="[1300,1432,1727,1753]" name="Vietnam" pageId="11" pageNumber="340">VIETNAM</collectingCountry>
, 
<collectingRegion box="[151,335,1762,1789]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="340">
<emphasis bold="true" box="[151,329,1762,1789]" pageId="11" pageNumber="340">Dong Nai Prov</emphasis>
.
</collectingRegion>
, 
<collectingMunicipality box="[351,612,1763,1789]" pageId="11" pageNumber="340">Cat Tien National Park</collectingMunicipality>
, 
<collectingRegion box="[624,734,1763,1789]" country="Vietnam" name="Dong Nai" pageId="11" pageNumber="340">Dong Nai</collectingRegion>
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FD12F9015379F90A" box="[742,806,1763,1789]" country="Vietnam" municipality="Cat Tien National Park" name="River" pageId="11" pageNumber="340" stateProvince="Dong Nai">River</location>
, cascade area behind first ranger station, 
<quantity box="[1271,1347,1763,1789]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="340" unit="m" value="120.0">
<specimenCount box="[1271,1347,1763,1789]" pageId="11" pageNumber="340" type="generic">
<elevation box="[1271,1347,1763,1789]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="340" unit="m" value="120.0">120 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation collectingDate="1998-05-08" collectionCode="T" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
., 
<date pageId="11" pageNumber="340" value="1998-05-08">
<collectingDate pageId="11" pageNumber="340" value="1998-05-08">8 May 1998</collectingDate>
</date>
, CL 3077, J. 
<collectionCode box="[357,375,1799,1825]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[386,699,1799,1825]" collectionCode="BPBM, USNM" collectorName="Polhemus" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Dong Nai" typeStatus="paratype">
<collectorName box="[386,496,1799,1825]" pageId="11" pageNumber="340">Polhemus</collectorName>
(
<collectionCode box="[512,594,1799,1825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="340" type="Museum">BPBM</collectionCode>
, 
<collectionCode box="[605,691,1799,1825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Vietnam" elevation="200" location="Nam Ceung" municipality="Nam Na River" pageId="11" pageNumber="340" specimenCount="10" specimenCount-female="7" specimenCount-male="3" stateProvince="Lai Chau" typeStatus="paratype">
<specimenCount box="[710,799,1799,1825]" pageId="11" pageNumber="340" type="male">3 males</specimenCount>
, 
<specimenCount box="[810,921,1799,1825]" pageId="11" pageNumber="340" type="female">7 females</specimenCount>
, 
<collectingCountry box="[932,1064,1799,1825]" name="Vietnam" pageId="11" pageNumber="340">VIETNAM</collectingCountry>
, 
<collectingRegion box="[1075,1258,1798,1825]" country="Vietnam" name="Lai Chau" pageId="11" pageNumber="340">
<emphasis bold="true" box="[1075,1252,1798,1825]" pageId="11" pageNumber="340">Lai Châu Prov</emphasis>
.
</collectingRegion>
, 
<collectingMunicipality box="[1273,1436,1799,1825]" pageId="11" pageNumber="340">Nam Na River</collectingMunicipality>
and 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FF3CF8C9510EF8B2" box="[200,337,1835,1861]" country="Vietnam" municipality="Nam Na River" name="Nam Ceung" pageId="11" pageNumber="340" stateProvince="Lai Chau">Nam Ceung</location>
stream trib. 
<quantity box="[484,581,1835,1862]" metricMagnitude="4" metricUnit="m" metricValue="1.55" pageId="11" pageNumber="340" unit="km" value="15.5">15.5 km</quantity>
. N. of 
<collectingRegion box="[662,770,1835,1861]" country="Vietnam" name="Lai Chau" pageId="11" pageNumber="340">Lai Châu</collectingRegion>
, 
<quantity box="[783,861,1835,1861]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="340" unit="m" value="200.0">
<specimenCount box="[783,861,1835,1861]" pageId="11" pageNumber="340" type="generic">
<elevation box="[783,861,1835,1861]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="340" unit="m" value="200.0">200 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation collectingDate="2000-04-11" collectionCode="T" collectorName="Polhemus" country="Vietnam" latitude="22.147778" longLatPrecision="21" longitude="103.1925" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Lai Chau" typeStatus="paratype">
., 
<geoCoordinate box="[879,1014,1835,1862]" degrees="22" direction="north" minutes="08" orientation="latitude" pageId="11" pageNumber="340" precision="15" seconds="52" value="22.147778">22°08'52&quot;N</geoCoordinate>
, 
<geoCoordinate box="[1027,1171,1835,1861]" degrees="103" direction="east" minutes="11" orientation="longitude" pageId="11" pageNumber="340" precision="15" seconds="33" value="103.1925">103°11'33&quot;E</geoCoordinate>
, 
<date box="[1184,1345,1835,1861]" pageId="11" pageNumber="340" value="2000-04-11">
<collectingDate box="[1184,1345,1835,1861]" pageId="11" pageNumber="340" value="2000-04-11">11 April 2000</collectingDate>
</date>
, 10:00– 13:00 hrs., CL 4409, D. A. 
<collectorName box="[470,583,1871,1897]" pageId="11" pageNumber="340">Polhemus</collectorName>
, J. 
<collectionCode box="[624,642,1871,1897]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[656,1150,1871,1897]" collectionCode="BPBM, USNM" collectorName="Polhemus &amp; P. Nguyen" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Lai Chau" typeStatus="paratype">
<collectorName box="[656,766,1871,1897]" pageId="11" pageNumber="340">Polhemus</collectorName>
and 
<collectorName box="[825,943,1871,1897]" pageId="11" pageNumber="340">P. Nguyen</collectorName>
(
<collectionCode box="[960,1042,1871,1897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="340" type="Museum">BPBM</collectionCode>
, 
<collectionCode box="[1056,1142,1871,1897]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Vietnam" county="Ya Rung" elevation="345" location="An Khe" municipality="Ngoe Ba Riv." pageId="11" pageNumber="340" specimenCount="22" specimenCount-female="11" specimenCount-immature="1" specimenCount-male="10" stateProvince="Gia Lai" typeStatus="paratype">
<specimenCount box="[1163,1269,1871,1897]" pageId="11" pageNumber="340" type="male">10 males</specimenCount>
, 
<specimenCount box="[1282,1409,1871,1897]" pageId="11" pageNumber="340" type="female">11 females</specimenCount>
, 
<specimenCount pageId="11" pageNumber="340" type="immature">1 immature</specimenCount>
, 
<collectingCountry box="[271,403,1907,1933]" name="Vietnam" pageId="11" pageNumber="340">VIETNAM</collectingCountry>
, 
<collectingRegion box="[414,573,1906,1933]" country="Vietnam" name="Gia Lai" pageId="11" pageNumber="340">
<emphasis bold="true" box="[414,567,1906,1933]" pageId="11" pageNumber="340">Gia Lai Prov</emphasis>
.
</collectingRegion>
, 
<collectingCounty box="[587,684,1907,1933]" pageId="11" pageNumber="340">Ya Rung</collectingCounty>
waterfall, 
<collectingMunicipality box="[803,956,1907,1933]" pageId="11" pageNumber="340">Ngoe Ba Riv.</collectingMunicipality>
, 
<quantity box="[966,1039,1907,1933]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="340" unit="km" value="30.0">30 km</quantity>
. S. of 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FBA6F89154F3F87A" box="[1106,1196,1907,1933]" country="Vietnam" county="Ya Rung" municipality="Ngoe Ba Riv." name="An Khe" pageId="11" pageNumber="340" stateProvince="Gia Lai">An Khe</location>
, 
<quantity box="[1207,1282,1907,1934]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="11" pageNumber="340" unit="m" value="345.0">
<specimenCount box="[1207,1282,1907,1934]" pageId="11" pageNumber="340" type="generic">
<elevation box="[1207,1282,1907,1934]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="11" pageNumber="340" unit="m" value="345.0">345 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation collectingDate="2001-03-10" collectionCode="T" country="Vietnam" latitude="13.767222" longLatPrecision="21" longitude="108.52473" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Gia Lai" typeStatus="paratype">
., 
<geoCoordinate box="[1297,1432,1907,1933]" degrees="13" direction="north" minutes="46" orientation="latitude" pageId="11" pageNumber="340" precision="15" seconds="02" value="13.767222">13°46'02&quot;N</geoCoordinate>
, 
<geoCoordinate box="[151,297,1943,1969]" degrees="108" direction="east" minutes="31" orientation="longitude" pageId="11" pageNumber="340" precision="15" seconds="29" value="108.52473">108°31'29&quot;E</geoCoordinate>
, 
<date box="[312,493,1943,1969]" pageId="11" pageNumber="340" value="2001-03-10">
<collectingDate box="[312,493,1943,1969]" pageId="11" pageNumber="340" value="2001-03-10">10 March 2001</collectingDate>
</date>
, CL 4287, J. 
<collectionCode box="[655,673,1943,1969]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="340">T</collectionCode>
</materialsCitation>
. 
<materialsCitation box="[688,1170,1943,1969]" collectionCode="BPBM, USNM" collectorName="Polhemus &amp; P. Nguyen" country="Vietnam" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Gia Lai" typeStatus="paratype">
<collectorName box="[688,798,1943,1969]" pageId="11" pageNumber="340">Polhemus</collectorName>
&amp; 
<collectorName box="[842,961,1943,1969]" pageId="11" pageNumber="340">P. Nguyen</collectorName>
(
<collectionCode box="[979,1061,1943,1969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="340" type="Museum">BPBM</collectionCode>
, 
<collectionCode box="[1076,1162,1943,1969]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="340" type="Museum">USNM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation country="Cambodia" elevation="95" location="An Long Rot river" pageId="11" pageNumber="340" specimenCount="7" specimenCount-female="4" specimenCount-immature="1" specimenCount-male="2" stateProvince="Kampong Speu" typeStatus="paratype">
<specimenCount box="[1185,1278,1943,1969]" pageId="11" pageNumber="340" type="male">2 males</specimenCount>
, 
<specimenCount box="[1293,1407,1943,1969]" pageId="11" pageNumber="340" type="female">4 females</specimenCount>
, 
<specimenCount pageId="11" pageNumber="340" type="immature">1 immature</specimenCount>
, 
<collectingCountry box="[274,429,1979,2005]" name="Cambodia" pageId="11" pageNumber="340">CAMBODIA</collectingCountry>
, 
<collectingRegion box="[442,697,1978,2005]" country="Cambodia" name="Kampong Speu" pageId="11" pageNumber="340">
<emphasis bold="true" box="[442,691,1978,2005]" pageId="11" pageNumber="340">Kampong Spoe Prov</emphasis>
.
</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:03AD879AF86DFFF3FF63FAB453E8FE26:8EDB6057F86DFFF5FD3CF85953C3F822" box="[712,924,1979,2005]" country="Cambodia" name="An Long Rot river" pageId="11" pageNumber="340" stateProvince="Kampong Speu">An Long Rot river</location>
at granite rapids, ~ 
<quantity box="[1145,1229,1979,2006]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="340" unit="km" value="4.5">4.5 km</quantity>
. 
<locationDeviation location="Chambok village" pageId="11" pageNumber="340">NW of Chambok village</locationDeviation>
, 
<quantity box="[241,303,2015,2042]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="340" unit="m" value="95.0">
<specimenCount box="[241,303,2015,2042]" pageId="11" pageNumber="340" type="generic">
<elevation box="[241,303,2015,2042]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="11" pageNumber="340" unit="m" value="95.0">95 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation box="[303,1398,2015,2042]" collectingDate="2016-05-21" collectionCode="BPBM" collectorName="Polhemus" country="Cambodia" latitude="12.412499" longLatPrecision="21" longitude="104.09028" pageId="11" pageNumber="340" specimenCount="1" stateProvince="Kampong Speu" typeStatus="paratype">
., 
<geoCoordinate box="[319,453,2015,2042]" degrees="12" direction="north" minutes="24" orientation="latitude" pageId="11" pageNumber="340" precision="15" seconds="45" value="12.412499">12°24'45&quot;N</geoCoordinate>
, 
<geoCoordinate box="[465,624,2015,2042]" degrees="104" direction="east" minutes="05" orientation="longitude" pageId="11" pageNumber="340" precision="15" seconds="025" value="104.09028">104°05'025&quot;E</geoCoordinate>
, 
<date box="[636,788,2015,2041]" pageId="11" pageNumber="340" value="2016-05-21">
<collectingDate box="[636,788,2015,2041]" pageId="11" pageNumber="340" value="2016-05-21">21 May 2016</collectingDate>
</date>
, 11:00–12:15 hrs., CL 6034, D. A. 
<collectorName box="[1181,1291,2015,2041]" pageId="11" pageNumber="340">Polhemus</collectorName>
(
<collectionCode box="[1306,1390,2015,2041]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="11" pageNumber="340" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="341" type="diagnosis">
<paragraph blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="341">
<emphasis bold="true" box="[199,322,150,176]" pageId="12" pageNumber="341">Diagnosis.</emphasis>
Body length: 6.04–6.50, ground color black with yellowish spots and stripes dorsally. Venter of head and labiomere II with dense, long, white hairs. Femora yellowish with black spots and stripes. Parameres slightly twisted, apical part spoon-like, with long, straight hairs.
</paragraph>
<paragraph blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="341">
<emphasis bold="true" box="[199,427,258,285]" pageId="12" pageNumber="341">Comparative notes</emphasis>
. 
<taxonomicName authority="Li &amp; Jin &amp; Polhemus &amp; Ye, 2023" authorityName="Li &amp; Jin &amp; Polhemus &amp; Ye" authorityYear="2023" box="[443,651,259,285]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[443,651,259,285]" italics="true" pageId="12" pageNumber="341">Valleriola asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" box="[659,751,259,285]" pageId="12" pageNumber="341">
<taxonomicNameLabel box="[659,751,259,285]" pageId="12" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the largest species of 
<taxonomicName authorityName="Brulle" authorityYear="1836" box="[1034,1171,259,285]" class="Insecta" family="Leptopodidae" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="339" phylum="Arthropoda" rank="tribe" tribe="Leptopodini">Leptopodini</taxonomicName>
known from Southeast Asia, with the body length of both sexes exceeding 6.0 mm. In comparison to the similar 
<taxonomicName authorityName="Distant" authorityYear="1909" box="[1194,1372,295,321]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="cicindeloides">
<emphasis box="[1194,1372,295,321]" italics="true" pageId="12" pageNumber="341">V. cicindeloides</emphasis>
</taxonomicName>
from 
<collectingCountry box="[151,211,331,357]" name="India" pageId="12" pageNumber="341">India</collectingCountry>
, 
<taxonomicName authority="Li &amp; Jin &amp; Polhemus &amp; Ye, 2023" authorityName="Li &amp; Jin &amp; Polhemus &amp; Ye" authorityYear="2023" box="[223,340,331,356]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[223,340,331,356]" italics="true" pageId="12" pageNumber="341">V. asiatica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[347,438,331,357]" pageId="12" pageNumber="341" rank="species">
<emphasis bold="true" box="[347,373,331,357]" pageId="12" pageNumber="341">sp</emphasis>
. 
<emphasis bold="true" box="[387,431,331,357]" pageId="12" pageNumber="341">nov</emphasis>
.
</taxonomicNameLabel>
may be distinguished by its darker grey ground color, and a shorter antennal segment III in relation to segment II, the ratio of segment II/segment III being 1.00/2.00– 
<quantity box="[1039,1119,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="5.588" pageId="12" pageNumber="341" unit="in" value="2.2">2.20 in</quantity>
<taxonomicName authority="Li &amp; Jin &amp; Polhemus &amp; Ye, 2023" authorityName="Li &amp; Jin &amp; Polhemus &amp; Ye" authorityYear="2023" box="[1128,1247,367,392]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[1128,1247,367,392]" italics="true" pageId="12" pageNumber="341">V. asiatica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1256,1348,367,393]" pageId="12" pageNumber="341" rank="species">
<emphasis bold="true" box="[1256,1282,367,393]" pageId="12" pageNumber="341">sp</emphasis>
. 
<emphasis bold="true" box="[1298,1342,367,393]" pageId="12" pageNumber="341">nov</emphasis>
.
</taxonomicNameLabel>
, versus 1.00/3.00 in 
<taxonomicName authorityName="Distant" authorityYear="1909" box="[294,469,403,429]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="cicindeloides">
<emphasis box="[294,469,403,429]" italics="true" pageId="12" pageNumber="341">V. cicindeloides</emphasis>
</taxonomicName>
. Based on figures made by the late J. T. Polhemus of the 
<typeStatus box="[1117,1215,403,429]" pageId="12" pageNumber="341">holotype</typeStatus>
of 
<taxonomicName authorityName="Distant" authorityYear="1909" box="[1254,1430,403,429]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="cicindeloides">
<emphasis box="[1254,1430,403,429]" italics="true" pageId="12" pageNumber="341">V. cicindeloides</emphasis>
</taxonomicName>
, housed in the Natural History Museum, London, the male parameres of the two species are rather similar in form, including the presence of a small, angular projection on the basal third of the paramere when seen in inner ventral view. The third author (DP) has in his collection additional species of similar size and appearance to 
<taxonomicName authority="Li &amp; Jin &amp; Polhemus &amp; Ye, 2023" authorityName="Li &amp; Jin &amp; Polhemus &amp; Ye" authorityYear="2023" box="[1280,1396,511,536]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="341" phylum="Arthropoda" rank="species" species="asiatica" status="sp. nov.">
<emphasis box="[1280,1396,511,536]" italics="true" pageId="12" pageNumber="341">V. asiatica</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="12" pageNumber="341">
<taxonomicNameLabel pageId="12" pageNumber="341" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the Indonesian islands of Timor, Sumba, Celebes and Sumatra, as well as 
<collectingCountry box="[1109,1180,547,573]" name="Nepal" pageId="12" pageNumber="341">Nepal</collectingCountry>
. All of these examples have relatively large body sizes (over 
<quantity box="[581,672,582,609]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="341" unit="mm" value="5.5">5.5 mm</quantity>
); dark-brown to dark-blackish-grey dorsal ground coloration; a pair of pale spots arranged horizontally across the posterior portion of the corium on each hemelytron; and the legs and lateral pronotum set with numerous erect, pale setae that are much longer than the diameters of the leg segments on which they arise. Within these populations, however, there are geographically correlated differences in paramere shape in comparison to the Indochinese material, and they are therefore not presently considered conspecific. Instead, they will be dealt with in a separate work already partially completed.
</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="342" pageId="12" pageNumber="341" type="description">
<paragraph blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="341">
<emphasis bold="true" box="[199,421,798,825]" pageId="12" pageNumber="341">Description. Male.</emphasis>
Color: ground color black (
<figureCitation box="[741,822,799,825]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[836,852,799,825]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
). Head mainly black, clypeus yellowish with black marks; bucculae yellowish; labiomere I completely hidden beneath bucculae, labiomere II yellowish-white with black apex, labiomeres III–IV black; compound eyes and ocelli black to dark-red; antennomere I yellowish with black stripes, antennomeres II–IV dark-brown (
<figureCitation box="[698,780,907,933]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[795,811,907,933]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
, 
<figureCitation box="[826,900,907,933]" captionStart="FIGURES 13–27" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-17@5.[284,1303,181,1919]" captionTargetPageId="5" captionText="FIGURES 13–27. Heads of Leptopus riparius Hsiao, 1964 and Valleriola spp. in frontal view, lateral view and ventral view. (13–15) L. riparius Hsiao, 1964; (16–18) V. asiatica sp. nov.; (19–21) V. buenoi (Usinger, 1942); (22–24) V. bui sp. nov.; (25–27) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7755120" httpUri="https://zenodo.org/record/7755120/files/figure.png" pageId="12" pageNumber="341">16–18</figureCitation>
). Pronotum mainly blackish-brown, collar and callus black, median part of collar with a yellow spot, posterior lobe of pronotum blackish-brown, posterior margin yellowish, median part with a yellowish stripe, posterior angles of pronotum shinning black; scutellum blackish, with a yellowish apex (
<figureCitation box="[412,492,1015,1041]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[504,520,1015,1041]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
). Hemelytra mainly blackish-brown, clavus with a yellowish stripe in middle and a yellowish spot on apex, lateral margin of hemelytra pale, spots and stripes on corium and membrane yellowish, middle part of corium with two spots on both sides of R+M vein, sometimes the outer one absent; apical part of sc cell and membrane cell IV with yellowish spots, sometimes with a small spot in r cell (
<figureCitation box="[1150,1236,1123,1149]" captionStart="FIGURES 28–33" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[397,1190,193,1870]" captionTargetId="figure-23@6.[384,1203,181,1884]" captionTargetPageId="6" captionText="FIGURES 28–33. Hemelytra of Leptopus riparius Hsiao, 1964 and Valleriola spp. (28) right hemelytron of Valleriola; (29) L. riparius Hsiao, 1964; (30) V. asiatica sp. nov.; (31) V. buenoi (Usinger, 1942); (32) V. bui sp. nov.; (33) V. javanica Drake &amp; Hottes, 1951. Sc, subcosta; R, radius; M, media; Cu, cubitus; Cs, claval suture; r, cell R; m, cell M; cu, cell Cu; sc, cell Sc; I, 1st membrane cell; II, 2nd membrane cell; III, 3rd membrane cell; IV, 4th membrane cell. Scale 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755124" httpUri="https://zenodo.org/record/7755124/files/figure.png" pageId="12" pageNumber="341">Fig. 30</figureCitation>
). Legs yellowish, femora with black stripe and spots; tibiae and tarsi blackish-brown, except fore tibia and tarsus yellowish-brown (
<figureCitation box="[159,238,1195,1221]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[250,324,1195,1221]" captionStart="FIGURES 34–48" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[170,1413,194,1457]" captionTargetId="figure-17@7.[151,1435,181,1468]" captionTargetPageId="7" captionText="FIGURES 34–48. Left fore femora of Leptopus riparius Hsiao, 1964 and Valleriola spp. in interior view, ventral view and exterior view. (34–36) L. riparius Hsiao, 1964; (40–42) V. buenoi (Usinger, 1942); (37–39) V. asiatica sp. nov.; (43–45) V. bui sp. nov.; (46–48) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm. Red arrows show the positions of large spines." figureDoi="http://doi.org/10.5281/zenodo.7755126" httpUri="https://zenodo.org/record/7755126/files/figure.png" pageId="12" pageNumber="341">37–39</figureCitation>
). Abdominal segments black, with white sutures. Paramere yellowish, infuscated apically (
<figureCitation captionStart="FIGURES 51–60" captionStartId="9.[151,264,1879,1903]" captionTargetBox="[187,1398,538,1830]" captionTargetId="figure-152@9.[163,1425,525,1856]" captionTargetPageId="9" captionText="FIGURES 51–60. Right parameres of Leptopus riparius and Valleriola spp. in different views. (51–52) L. riparius Hsiao, 1964, dorsal and interior views; (53–54) V. asiatica sp. nov., dorsal and interior views; (55–56) V. buenoi (Usinger, 1942), dorsal and interior views; (57–59) V. bui sp. nov., dorsal, exterior, and interior views; (60) V. javanica Drake &amp; Hottes, 1951, dorsal view. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7755132" httpUri="https://zenodo.org/record/7755132/files/figure.png" pageId="12" pageNumber="341">Figs. 53, 54</figureCitation>
). Structural characteristics: body elongated, bearing sparse, short, pale setae dorsally (
<figureCitation box="[1141,1220,1231,1257]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[1231,1247,1231,1257]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
). Head short and wide, dorsally bearing dense, short pubescence except two circular areas, venter of head bearing dense, long, white hairs, with three spines beneath compound eyes on each side; compound eyes distinctly large, without sparse, short, spine-like setae; bucculae clearly visible; labium short, labiomere I completely hidden beneath bucculae, labiomere II bearing dense, long, white hairs, with two spines on each side; antennomere I short and stout, antennomeres II–IV extremely long and slender (
<figureCitation box="[479,560,1411,1437]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[574,590,1411,1437]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
, 
<figureCitation box="[604,678,1411,1437]" captionStart="FIGURES 13–27" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[284,1303,181,1919]" captionTargetId="figure-17@5.[284,1303,181,1919]" captionTargetPageId="5" captionText="FIGURES 13–27. Heads of Leptopus riparius Hsiao, 1964 and Valleriola spp. in frontal view, lateral view and ventral view. (13–15) L. riparius Hsiao, 1964; (16–18) V. asiatica sp. nov.; (19–21) V. buenoi (Usinger, 1942); (22–24) V. bui sp. nov.; (25–27) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7755120" httpUri="https://zenodo.org/record/7755120/files/figure.png" pageId="12" pageNumber="341">16–18</figureCitation>
). Pronotum sub-triangular, densely punctured except callus; callus relatively rough, distinctly swollen, with a deep impression on median line; posterior angles of pronotum glossy, without punctures; scutellum triangular, medially with a sub-circular depression, laterally serrated, with a distinct process on each side (
<figureCitation box="[411,494,1519,1545]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[218,1371,207,1895]" captionTargetId="figure-23@2.[200,1386,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Habitus. (1) Leptopus riparius Hsiao, 1964; 2–5. Valleriola spp.: (2) V. asiatica sp. nov.; (3) V. buenoi (Usinger, 1942); (4) V. bui sp. nov.; (5) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755112" httpUri="https://zenodo.org/record/7755112/files/figure.png" pageId="12" pageNumber="341">Figs. 2</figureCitation>
, 
<figureCitation box="[510,526,1519,1545]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1173,1197]" captionTargetBox="[170,1419,194,1132]" captionTargetId="figure-17@3.[151,1435,181,1149]" captionTargetPageId="3" captionText="FIGURES 6–10. Habitus of Leptopus riparius Hsiao, 1964 and Valleriola spp. (6) L. riparius Hsiao, 1964; (7) V. asiatica sp. nov.; (8) V. buenoi (Usinger, 1942); (9) V. bui sp. nov.; (10) V. javanica Drake &amp; Hottes, 1951. Scale 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755114" httpUri="https://zenodo.org/record/7755114/files/figure.png" pageId="12" pageNumber="341">7</figureCitation>
). Hemelytra elongated, clavus and outer cell of corium with dense punctures; membrane opaque, with four closed cells (
<figureCitation box="[622,706,1555,1581]" captionStart="FIGURES 28–33" captionStartId="6.[151,264,1909,1933]" captionTargetBox="[397,1190,193,1870]" captionTargetId="figure-23@6.[384,1203,181,1884]" captionTargetPageId="6" captionText="FIGURES 28–33. Hemelytra of Leptopus riparius Hsiao, 1964 and Valleriola spp. (28) right hemelytron of Valleriola; (29) L. riparius Hsiao, 1964; (30) V. asiatica sp. nov.; (31) V. buenoi (Usinger, 1942); (32) V. bui sp. nov.; (33) V. javanica Drake &amp; Hottes, 1951. Sc, subcosta; R, radius; M, media; Cu, cubitus; Cs, claval suture; r, cell R; m, cell M; cu, cell Cu; sc, cell Sc; I, 1st membrane cell; II, 2nd membrane cell; III, 3rd membrane cell; IV, 4th membrane cell. Scale 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7755124" httpUri="https://zenodo.org/record/7755124/files/figure.png" pageId="12" pageNumber="341">Fig. 30</figureCitation>
). Legs long and slender, fore coxa strong, slightly elongated, with two long spines anteriorly directed; fore femur incrassate, ventrally with seven large spines (three spines on the outer side and four on the inner side) and two rows of dense, small, black spines; fore tibia ventrally with a row of dense, brown spines (
<figureCitation box="[319,459,1663,1689]" captionStart="FIGURES 34–48" captionStartId="7.[151,264,1492,1516]" captionTargetBox="[170,1413,194,1457]" captionTargetId="figure-17@7.[151,1435,181,1468]" captionTargetPageId="7" captionText="FIGURES 34–48. Left fore femora of Leptopus riparius Hsiao, 1964 and Valleriola spp. in interior view, ventral view and exterior view. (34–36) L. riparius Hsiao, 1964; (40–42) V. buenoi (Usinger, 1942); (37–39) V. asiatica sp. nov.; (43–45) V. bui sp. nov.; (46–48) V. javanica Drake &amp; Hottes, 1951. Scale 0.5 mm. Red arrows show the positions of large spines." figureDoi="http://doi.org/10.5281/zenodo.7755126" httpUri="https://zenodo.org/record/7755126/files/figure.png" pageId="12" pageNumber="341">Figs. 37–39</figureCitation>
). Middle and hind legs distinctly elongated. Abdomen slender, tubular-like. Paramere slightly twisted, with a spoon-like apex, covered by dense punctures and long, straight hairs (
<figureCitation box="[1195,1332,1698,1725]" captionStart="FIGURES 51–60" captionStartId="9.[151,264,1879,1903]" captionTargetBox="[187,1398,538,1830]" captionTargetId="figure-152@9.[163,1425,525,1856]" captionTargetPageId="9" captionText="FIGURES 51–60. Right parameres of Leptopus riparius and Valleriola spp. in different views. (51–52) L. riparius Hsiao, 1964, dorsal and interior views; (53–54) V. asiatica sp. nov., dorsal and interior views; (55–56) V. buenoi (Usinger, 1942), dorsal and interior views; (57–59) V. bui sp. nov., dorsal, exterior, and interior views; (60) V. javanica Drake &amp; Hottes, 1951, dorsal view. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7755132" httpUri="https://zenodo.org/record/7755132/files/figure.png" pageId="12" pageNumber="341">Figs. 53, 54</figureCitation>
).
</paragraph>
<paragraph blockId="12.[151,1437,150,2013]" pageId="12" pageNumber="341">
<emphasis bold="true" box="[199,293,1735,1761]" pageId="12" pageNumber="341">Female.</emphasis>
Color and structural characteristics similar to male. Genital segments of female hidden in abdominal segment VII, not visible in vitro.
</paragraph>
<paragraph blockId="12.[151,1437,150,2013]" lastBlockId="13.[151,1436,150,465]" lastPageId="13" lastPageNumber="342" pageId="12" pageNumber="341">
<emphasis bold="true" box="[199,455,1807,1833]" pageId="12" pageNumber="341">Measurements. Male.</emphasis>
Body length: 6.26–6.50 (
<typeStatus box="[736,840,1807,1833]" pageId="12" pageNumber="341">holotype</typeStatus>
: 6.50), body width across hemelytra: 1.70–1.84. Head width: 1.31–1.40, head length: 0.78–0.83, head width about 1.69 times head length; antenna about 1.05 times as long as body, length of antennomeres I–IV: 0.41, 1.36, 2.69, 2.38. Pronotum width across collar: 0.57–0.59, pronotum width across posterior angles: 1.63–1.64, pronotum length: 1.24–1.29; scutellum width: 0.74–0.77, scutellum length: 0.68–0.72; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 2.28, 1.69 and 0.60 (0.16+0.24+0.20), middle leg: 2.44, 2.26 and 0.62 (0.16+0.21+0.25), hind leg: 2.57, 3.52 and 0.82 (0.14+0.31+0.37). 
<emphasis bold="true" box="[1273,1368,1987,2013]" pageId="12" pageNumber="341">Female.</emphasis>
Body length: 6.04–6.35, body width across hemelytra: 1.67–1.79. Head width: 1.32–1.35, head length: 0.77–0.82, head width about 1.06 times head length; antenna about 1.18 times as long as body, length of antennomeres I–IV: 0.37, 1.21, 2.74, 2.42. Pronotum width across collar: 0.59–0.62, pronotum width across posterior angles: 1.61–1.68; pronotum length: 1.20–1.26; scutellum width: 0.78–0.81, scutellum length: 0.75–0.79; length of femur, tibia and tarsus (tarsomeres I+II+III): fore leg: 2.21, 1.66 and 0.57 (0.13+0.26+0.18), middle leg: 2.50, 2.43 and 0.69 (0.14+0.33+0.22), hind leg: 2.66, 3.66 and 0.77 (0.17+0.36+0.24).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="342" type="etymology">
<paragraph blockId="13.[151,1436,150,465]" pageId="13" pageNumber="342">
<emphasis bold="true" box="[199,334,367,393]" pageId="13" pageNumber="342">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName authorityName="Li &amp; Jin &amp; Polhemus &amp; Ye" authorityYear="2023" box="[583,672,367,392]" class="Insecta" family="Leptopodidae" genus="Valleriola" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="342" phylum="Arthropoda" rank="species" species="asiatica">
<emphasis box="[583,672,367,392]" italics="true" pageId="13" pageNumber="342">asiatica</emphasis>
</taxonomicName>
” refers to the wide distribution of this species in Southeast Asia; an adjective.
</paragraph>
</subSubSection>
<subSubSection box="[199,951,438,465]" pageId="13" pageNumber="342" type="distribution">
<paragraph blockId="13.[151,1436,150,465]" box="[199,951,438,465]" pageId="13" pageNumber="342">
<emphasis bold="true" box="[199,353,438,464]" pageId="13" pageNumber="342">Distribution.</emphasis>
<collectingCountry box="[359,426,439,465]" name="China" pageId="13" pageNumber="342">China</collectingCountry>
(
<collectingRegion box="[441,528,439,465]" country="China" name="Hainan" pageId="13" pageNumber="342">Hainan</collectingRegion>
), 
<collectingCountry box="[547,604,439,465]" name="Laos" pageId="13" pageNumber="342">Laos</collectingCountry>
, 
<collectingCountry box="[615,713,439,465]" name="Vietnam" pageId="13" pageNumber="342">Vietnam</collectingCountry>
, 
<collectingCountry box="[723,838,439,465]" name="Cambodia" pageId="13" pageNumber="342">Cambodia</collectingCountry>
(
<figureCitation box="[854,937,439,465]" captionStart="FIGURE 61" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[191,1397,181,1920]" captionTargetId="figure-23@10.[191,1397,181,1920]" captionTargetPageId="10" captionText="FIGURE 61. Distribution records of the tribe Leptopodini Brullé, 1836 in Southeast Asia and China." figureDoi="http://doi.org/10.5281/zenodo.7755136" httpUri="https://zenodo.org/record/7755136/files/figure.png" pageId="13" pageNumber="342">Fig. 61</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>