1265 lines
162 KiB
XML
1265 lines
162 KiB
XML
<document id="3501F074AD4E089F3545EA34CB3A4785" ID-DOI="10.11646/zootaxa.4048.2.3" ID-GBIF-Dataset="124ae87a-8c30-42b6-9f06-6f49c0a5b7cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="239892" ID-ZooBank="0964C238-A12F-4334-B42A-26726A66996F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461346931879" checkinUser="plazi" docAuthor="Brown, Rafe M., Layola, Louise Abigail De, Ii, Antonio Lorenzo, Diesmos, Mae Lowe L. & Diesmos, Arvin C." docDate="2015" docId="03AC76345E096B463BF4F8E4FB1FFB44" docLanguage="en" docName="zt04048p210.pdf" docOrigin="Zootaxa 4048 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Platymantis quezoni Brown, Layola, Ii, Diesmos & Diesmos, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="204" masterDocId="FF950E4C5E0A6B4B3B63FFE6FFA7FFBE" masterDocTitle="A new species of limestone karst inhabiting forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae: subgenus Lupacolus) from southern Luzon Island, Philippines" masterLastPageNumber="210" masterPageNumber="191" pageNumber="194" updateTime="1698620707286" updateUser="plazi">
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<mods:mods id="441480F551B3D9099E91F28EE369B915" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E5875B85A78236135F84FF18AC722435">
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<mods:title id="6F9327BC66444B09779589C71858D2E2">A new species of limestone karst inhabiting forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae: subgenus Lupacolus) from southern Luzon Island, Philippines</mods:title>
|
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</mods:titleInfo>
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<mods:name id="7E590CC445E1883832387752FE49E8AF" type="personal">
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<mods:role id="444D8F6AD82F645F95AD9B9A987DDE44">
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<mods:roleTerm id="726FBBEE1829D4FBBD569878CBB250FE">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="CD895A77010B9C141D56A20734193E8B">Brown, Rafe M.</mods:namePart>
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</mods:name>
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<mods:name id="40DA2750CCB9EE8CDC91518CD4D62C93" type="personal">
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<mods:role id="21D94A522E42A212494AD6A01061ECBA">
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<mods:roleTerm id="63EF9312FA71063DEBEB844C416BA593">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="AE6551C5359AC868E2CCEE0EE51F8954">Layola, Louise Abigail De</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A1E3FEED6B30C66500F6872DC8D52B5E" type="personal">
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<mods:role id="1E24463C88D6800EA12BFCB4169BB5E7">
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<mods:roleTerm id="03E3734633DCEC4A3DD150F46D1F5962">Author</mods:roleTerm>
|
||
</mods:role>
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||
<mods:namePart id="C208B0F0B573D22BA56E158168E32F63">Ii, Antonio Lorenzo</mods:namePart>
|
||
</mods:name>
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||
<mods:name id="F89B06AB27036B0A12D4FEF125CA7330" type="personal">
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<mods:roleTerm id="672B7CB06F0D608B83EBA6E2176D4ED3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="87A39F31F60FAACAA7FD41B8EBBE70F8">Diesmos, Mae Lowe L.</mods:namePart>
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||
</mods:name>
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<mods:name id="B7A09F395D07656EC6AFA57C3528C670" type="personal">
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<mods:roleTerm id="15CFB322B8FC010BCE2FDA9268EE0A20">Author</mods:roleTerm>
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</mods:role>
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||
<mods:namePart id="112637D17B5484EDE28F2E04C0FE917E">Diesmos, Arvin C.</mods:namePart>
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||
</mods:name>
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<mods:typeOfResource id="73B1A94920E64A66996983E59FA46B1C">text</mods:typeOfResource>
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<mods:relatedItem id="77B3C70C025448D75695DDCB5170309E" type="host">
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<mods:titleInfo id="102BED034333FF32597CA793560D7922">
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<mods:title id="C3A7B34CE70F076A1F9680EBB003079D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8752F0EF01D999D5C48C3F3283948B1F">
|
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<mods:date id="DED819EAD166B48E05430435B4017660">2015</mods:date>
|
||
<mods:detail id="B11ADB2ECB072F6F2A5B9486FFE10C25" type="volume">
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||
<mods:number id="94CEC9744D6FE13865EDEE73D6581E4D">4048</mods:number>
|
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</mods:detail>
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<mods:detail id="D37EF203B4601EE51F9480DFC048259C" type="issue">
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<mods:number id="048A1EDC00DD4A5D41074779C1DD4889">2</mods:number>
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<mods:end id="1D4B758720FF51366E3BC85AF6641146">210</mods:end>
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<mods:classification id="E2E1CC6314D081FE419BA2C096B1C8BA">journal article</mods:classification>
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<mods:identifier id="6E8DBA5A7231CC55F118D729E583B78C" type="DOI">10.11646/zootaxa.4048.2.3</mods:identifier>
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<mods:identifier id="17F4FD7D74A09DE3EF3D79624D8F5C3D" type="GBIF-Dataset">124ae87a-8c30-42b6-9f06-6f49c0a5b7cc</mods:identifier>
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<mods:identifier id="8897880457F5370D9BC1258CBD378FB0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E8ED2F07CCCABD889D194BAF4D7495F1" type="Zenodo-Dep">239892</mods:identifier>
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<mods:identifier id="0EB8F6476E352C57BD87F38810898A2B" type="ZooBank">0964C238-A12F-4334-B42A-26726A66996F</mods:identifier>
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</mods:mods>
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<treatment id="03AC76345E096B463BF4F8E4FB1FFB44" ID-DOI="http://doi.org/10.5281/zenodo.5681237" ID-GBIF-Taxon="119645477" ID-Zenodo-Dep="5681237" LSID="urn:lsid:plazi:treatment:03AC76345E096B463BF4F8E4FB1FFB44" httpUri="http://treatment.plazi.org/id/03AC76345E096B463BF4F8E4FB1FFB44" lastPageId="13" lastPageNumber="204" pageId="3" pageNumber="194">
|
||
<subSubSection id="C31F94A95E096B483BF4F8E4FEA7F8DD" pageId="3" pageNumber="194" type="nomenclature">
|
||
<paragraph id="8BBAC7225E096B483BF4F8E4FE4FF8A5" blockId="3.[151,592,1794,1891]" box="[151,488,1794,1819]" pageId="3" pageNumber="194">
|
||
<heading id="D0F2704E5E096B483BF4F8E4FE4FF8A5" bold="true" box="[151,488,1794,1819]" fontSize="10" level="2" pageId="3" pageNumber="194" reason="2">
|
||
<emphasis id="B9711B305E096B483BF4F8E4FE4FF8A5" bold="true" box="[151,488,1794,1819]" pageId="3" pageNumber="194">
|
||
<taxonomicName id="4C05BCA15E096B483BF4F8E4FE26F8A5" ID-CoL="4JSRM" box="[151,385,1794,1819]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="194" phylum="Chordata" rank="species" species="quezoni" status="sp. nov.">
|
||
<emphasis id="B9711B305E096B483BF4F8E4FE26F8A5" bold="true" box="[151,385,1794,1819]" italics="true" pageId="3" pageNumber="194">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A242A64B5E096B483AECF8E4FE4FF8A5" box="[399,488,1794,1819]" pageId="3" pageNumber="194" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E096B483BF4F8C1FEA7F8DD" blockId="3.[151,592,1794,1891]" pageId="3" pageNumber="194">
|
||
“
|
||
<taxonomicName id="4C05BCA15E096B483BC0F8C0FE8EF881" box="[163,297,1830,1855]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="194" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9711B305E096B483BC0F8C0FE8EF881" box="[163,297,1830,1855]" italics="true" pageId="3" pageNumber="194">Platymantis</emphasis>
|
||
</taxonomicName>
|
||
sp. 27” Brown
|
||
<emphasis id="B9711B305E096B483AB9F8C1FDB5F881" box="[474,530,1830,1855]" italics="true" pageId="3" pageNumber="194">et al.</emphasis>
|
||
2015
|
||
<figureCitation id="133EDBA75E096B483BF4F8ACFEA7F8DD" box="[151,256,1866,1891]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="4.[151,244,1449,1470]" captionStartId-1="5.[151,244,1547,1568]" captionStartId-2="6.[151,244,1685,1706]" captionTargetBox-0="[217,1369,411,1418]" captionTargetBox-1="[249,1334,199,1486]" captionTargetBox-2="[307,1275,204,1636]" captionTargetId-0="figure@4.[217,1369,411,1425]" captionTargetId-1="figure@5.[241,1345,191,1525]" captionTargetId-2="figure@6.[289,1297,191,1663]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." captionText-1="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." captionText-2="FIGURE 4. Oblique view illustration of lateral head of (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817) and (B) P. dorsalis (KU 300356). Scale bars = 2 mm; Illustrations by M. Garfield." httpUri-0="https://zenodo.org/record/239894/files/figure.png" httpUri-1="https://zenodo.org/record/239895/files/figure.png" httpUri-2="https://zenodo.org/record/239896/files/figure.png" pageId="3" pageNumber="194">Figs. 2–4</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F94A95E096B4F3BF4F874FE8FFEDB" lastPageId="4" lastPageNumber="195" pageId="3" pageNumber="194" type="materials_examined">
|
||
<paragraph id="8BBAC7225E096B483BF4F874FA3AF84D" blockId="3.[151,1437,1938,2035]" pageId="3" pageNumber="194">
|
||
<emphasis id="B9711B305E096B483BF4F874FEACF815" bold="true" box="[151,267,1938,1963]" pageId="3" pageNumber="194">
|
||
<typeStatus id="54BE79805E096B483BF4F874FEA1F815" box="[151,262,1938,1963]" pageId="3" pageNumber="194" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="ED145FE75E096B483A70F875FEF7F814" box="[275,336,1939,1962]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="3" pageNumber="194">
|
||
<collectingCountry id="F31287B25E096B483A70F875FEF7F814" box="[275,336,1939,1962]" name="Panama" pageId="3" pageNumber="194">PNM</collectingCountry>
|
||
</collectionCode>
|
||
9817 (RMB 4067; formerly KU 339542), adult male, collected on limestone outcrop in original forest (19:00 hr) on
|
||
<date id="FFBBE1E25E096B483A1FF851FDF1F871" box="[380,598,1974,1999]" pageId="3" pageNumber="194" value="2001-11-27">
|
||
27 November
|
||
<quantity id="4CFD6AC75E096B48397DF851FDF1F871" box="[542,598,1975,1999]" metricMagnitude="1" metricUnit="m" metricValue="5.08254" pageId="3" pageNumber="194" unit="in" value="2001.0">2001</quantity>
|
||
</date>
|
||
in Barangay Malinao Ilaya, Municipality of Atimonan, Quezon Province, Luzon Island,
|
||
<collectingCountry id="F31287B25E096B483A54F83CFE12F84D" box="[311,437,2010,2035]" name="Philippines" pageId="3" pageNumber="194">Philippines</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE31A1E55E096B483AA7F83DFD9CF84D" box="[452,571,2011,2035]" direction="north" orientation="latitude" pageId="3" pageNumber="194" precision="55" value="13.989">N 13.989°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE31A1E55E096B483925F83DFD6FF84D" box="[582,712,2011,2035]" direction="east" orientation="longitude" pageId="3" pageNumber="194" precision="55" value="121.818">E 121.818°</geoCoordinate>
|
||
;
|
||
<quantity id="4CFD6AC75E096B4839B0F83CFCBEF84C" box="[723,793,2010,2035]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="194" unit="m" value="275.0">275 m</quantity>
|
||
elevation; datum: WGS 85), by
|
||
<collectionCode id="ED145FE75E096B483FE7F83DFB74F84C" box="[1156,1235,2011,2034]" httpUri="http://grbio.org/cool/yq8n-xdc0" name="Microbiological Research Laboratory, Soil and Water Section, Department of Land Development" pageId="3" pageNumber="194">MLLD</collectionCode>
|
||
and G. V. A. Gee.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E0E6B4F3BA4FF71FE8FFEDB" blockId="4.[151,1436,151,357]" pageId="4" pageNumber="195">
|
||
<emphasis id="B9711B305E0E6B4F3BA4FF71FDABFF11" bold="true" box="[199,524,151,176]" pageId="4" pageNumber="195">
|
||
<typeStatus id="54BE79805E0E6B4F3BA4FF71FE98FF0E" box="[199,319,151,176]" pageId="4" pageNumber="195" type="paratype">Paratypes</typeStatus>
|
||
(paratopotypes).
|
||
</emphasis>
|
||
Sixteen males:
|
||
<collectionCode id="ED145FE75E0E6B4F39A5FF7EFCB5FF0E" box="[710,786,152,176]" httpUri="http://grbio.org/cool/57yf-fp36" name="Texas Memorial Museum, Texas Natural History Collection" pageId="4" pageNumber="195">TNHC</collectionCode>
|
||
61989–94, bearing the same data as the
|
||
<typeStatus id="54BE79805E0E6B4F3F8FFF71FAF6FF0E" box="[1260,1361,151,176]" pageId="4" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
;
|
||
<collectionCode id="ED145FE75E0E6B4F3E03FF7EFA3BFF11" box="[1376,1436,152,175]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="4" pageNumber="195">
|
||
<collectingCountry id="F31287B25E0E6B4F3E03FF7EFA3BFF11" box="[1376,1436,152,175]" name="Panama" pageId="4" pageNumber="195">PNM</collectingCountry>
|
||
</collectionCode>
|
||
9910–12, collected on
|
||
<date id="FFBBE1E25E0E6B4F3AFBFF5AFDF1FF6A" box="[408,598,188,213]" pageId="4" pageNumber="195" value="2013-10-19">19 October 2013</date>
|
||
by
|
||
<collectionCode id="ED145FE75E0E6B4F39E2FF5AFD68FF6A" box="[641,719,188,212]" pageId="4" pageNumber="195">LADL</collectionCode>
|
||
,
|
||
<collectionCode id="ED145FE75E0E6B4F39B9FF5AFCA5FF6A" box="[730,770,188,212]" pageId="4" pageNumber="195">AL</collectionCode>
|
||
,
|
||
<collectionCode id="ED145FE75E0E6B4F386DFF5BFCC7FF6A" box="[782,864,189,212]" httpUri="http://grbio.org/cool/yq8n-xdc0" name="Microbiological Research Laboratory, Soil and Water Section, Department of Land Development" pageId="4" pageNumber="195">MLLD</collectionCode>
|
||
, and Jason Fernandez;
|
||
<collectionCode id="ED145FE75E0E6B4F3F05FF5BFB04FF6A" box="[1126,1187,189,212]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="4" pageNumber="195">
|
||
<collectingCountry id="F31287B25E0E6B4F3F05FF5BFB04FF6A" box="[1126,1187,189,212]" name="Panama" pageId="4" pageNumber="195">PNM</collectingCountry>
|
||
</collectionCode>
|
||
9913–9916, collected on
|
||
<date id="FFBBE1E25E0E6B4F3BDEFF39FE30FF46" box="[189,407,223,248]" pageId="4" pageNumber="195" value="2014-12-15">15 December 2014</date>
|
||
by
|
||
<collectionCode id="ED145FE75E0E6B4F3AA4FF06FDB5FF46" box="[455,530,224,248]" pageId="4" pageNumber="195">LADL</collectionCode>
|
||
and
|
||
<collectionCode id="ED145FE75E0E6B4F392DFF06FDD0FF46" box="[590,631,224,248]" pageId="4" pageNumber="195">AL</collectionCode>
|
||
; KU 328735 and 328736, bearing the same data as the
|
||
<typeStatus id="54BE79805E0E6B4F3E63FF39FAC1FF46" box="[1280,1382,223,248]" pageId="4" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
; KU 339543, collected
|
||
<date id="FFBBE1E25E0E6B4F3A09FEE2FDBAFEA2" box="[362,541,260,285]" pageId="4" pageNumber="195" value="2011-01-09">9 January 2011</date>
|
||
; three females (
|
||
<collectionCode id="ED145FE75E0E6B4F39B6FEE2FCB3FEA2" box="[725,788,260,284]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="4" pageNumber="195">
|
||
<collectingCountry id="F31287B25E0E6B4F39B6FEE2FCB3FEA2" box="[725,788,260,284]" name="Panama" pageId="4" pageNumber="195">PNM</collectingCountry>
|
||
</collectionCode>
|
||
9907–09, collected on
|
||
<date id="FFBBE1E25E0E6B4F3F47FEE2FB42FEA2" box="[1060,1253,260,285]" pageId="4" pageNumber="195" value="2013-10-19">19 October 2013</date>
|
||
by
|
||
<collectionCode id="ED145FE75E0E6B4F3E77FEE2FAC5FEA2" box="[1300,1378,260,284]" pageId="4" pageNumber="195">LADL</collectionCode>
|
||
,
|
||
<collectionCode id="ED145FE75E0E6B4F3E13FEE2FA3FFEA2" box="[1392,1432,260,284]" pageId="4" pageNumber="195">AL</collectionCode>
|
||
,
|
||
<collectionCode id="ED145FE75E0E6B4F3BF4FECEFF41FE81" box="[151,230,296,319]" httpUri="http://grbio.org/cool/yq8n-xdc0" name="Microbiological Research Laboratory, Soil and Water Section, Department of Land Development" pageId="4" pageNumber="195">MLLD</collectionCode>
|
||
and Jason Fernandez; one juvenile (KU 339544, bearing the same data as the
|
||
<typeStatus id="54BE79805E0E6B4F3F36FEC1FB1BFEFE" box="[1109,1212,295,320]" pageId="4" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
), all collected at the
|
||
<typeStatus id="54BE79805E0E6B4F3BF4FEABFF60FEDB" box="[151,199,333,357]" pageId="4" pageNumber="195">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7A97AA5E0E6B4F3BF4FA4FFDD2FA47" httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="4" pageNumber="195" targetBox="[217,1369,411,1418]" targetPageId="4">
|
||
<paragraph id="8BBAC7225E0E6B4F3BF4FA4FFDD2FA47" blockId="4.[151,1435,1449,1529]" pageId="4" pageNumber="195">
|
||
<emphasis id="B9711B305E0E6B4F3BF4FA4FFEA9FA00" bold="true" box="[151,270,1449,1470]" pageId="4" pageNumber="195">FIGURE 2.</emphasis>
|
||
Ventral views of hand (A) and foot (B) of the adult male holotype of
|
||
<taxonomicName id="4C05BCA15E0E6B4F38DEFA4FFAA7FA00" box="[957,1280,1449,1470]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="quezoni" status="sp. nov." subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E0E6B4F38DEFA4FFB88FA00" box="[957,1071,1449,1470]" italics="true" pageId="4" pageNumber="195">Platymantis</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0E6B4F3F5CFA4FFB04FA00" box="[1087,1187,1449,1470]" italics="true" pageId="4" pageNumber="195">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E0E6B4F3FD0FA4FFAA7FA00" box="[1203,1280,1449,1470]" italics="true" pageId="4" pageNumber="195">quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A242A64B5E0E6B4F3E68FA4FFAFFFA00" box="[1291,1368,1449,1470]" pageId="4" pageNumber="195" rank="species">
|
||
<emphasis id="B9711B305E0E6B4F3E68FA4FFAFFFA00" bold="true" box="[1291,1368,1449,1470]" pageId="4" pageNumber="195">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(PNM 9817), and adult male specimens of (C, D)
|
||
<taxonomicName id="4C05BCA15E0E6B4F3958FA20FCA6FA65" box="[571,769,1478,1499]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E0E6B4F3958FA20FCA6FA65" box="[571,769,1478,1499]" italics="true" pageId="4" pageNumber="195">Platymantis dorsalis</emphasis>
|
||
</taxonomicName>
|
||
(KU 300356), (E, F)
|
||
<emphasis id="B9711B305E0E6B4F38B5FA20FB8AFA65" box="[982,1069,1478,1499]" italics="true" pageId="4" pageNumber="195">P. p a e n g i</emphasis>
|
||
(KU 300206), and (G, H)
|
||
<taxonomicName id="4C05BCA15E0E6B4F3E52FA20FA3CFA65" box="[1329,1435,1478,1499]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="spelaeus">
|
||
<emphasis id="B9711B305E0E6B4F3E52FA20FA3CFA65" box="[1329,1435,1478,1499]" italics="true" pageId="4" pageNumber="195">P. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
(KU 300438) for comparison. Scale bars = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31F94A95E0E6B433BA4F9AFFAC5FAEA" lastPageId="8" lastPageNumber="199" pageId="4" pageNumber="195" type="diagnosis">
|
||
<paragraph id="8BBAC7225E0E6B4F3BA4F9AFFD54F818" blockId="4.[151,1436,1609,1958]" pageId="4" pageNumber="195">
|
||
<emphasis id="B9711B305E0E6B4F3BA4F9AFFEE6F9DC" bold="true" box="[199,321,1609,1634]" pageId="4" pageNumber="195">Diagnosis.</emphasis>
|
||
<emphasis id="B9711B305E0E6B4F3A2CF9AFFDE7F9DC" box="[335,576,1609,1634]" italics="true" pageId="4" pageNumber="195">
|
||
<taxonomicName id="4C05BCA15E0E6B4F3A2CF9AFFD9BF9DC" box="[335,572,1609,1634]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="195" phylum="Chordata" rank="species" species="quezoni" status="sp. nov.">Platymantis quezoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B9711B305E0E6B4F392DF9AFFD11F9DC" bold="true" box="[590,694,1609,1634]" pageId="4" pageNumber="195">
|
||
<taxonomicNameLabel id="A242A64B5E0E6B4F392DF9AFFD17F9DC" box="[590,688,1609,1634]" pageId="4" pageNumber="195" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
can be distinguished from other
|
||
<collectingCountry id="F31287B25E0E6B4F3F2CF9AFFB65F9DC" box="[1103,1218,1609,1634]" name="Philippines" pageId="4" pageNumber="195">Philippine</collectingCountry>
|
||
congeners by the following suite of characters: body size
|
||
<quantity id="4CFD6AC75E0E6B4F3937F988FD51F93B" box="[596,758,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.3899999999999997" metricValueMin="2.21" pageId="4" pageNumber="195" unit="mm" value="28.0" valueMax="33.9" valueMin="22.1">22.1–33.9 mm</quantity>
|
||
SVL for males,
|
||
<quantity id="4CFD6AC75E0E6B4F38CDF988FBF7F93B" box="[942,1104,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="3.675" metricValueMax="3.97" metricValueMin="3.38" pageId="4" pageNumber="195" unit="mm" value="36.75" valueMax="39.7" valueMin="33.8">33.8–39.7 mm</quantity>
|
||
for females; terminal discs of fingers and toes slightly expanded (
|
||
<figureCitation id="133EDBA75E0E6B4F3956F977FDDBF914" box="[565,636,1681,1706]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="4" pageNumber="195">Fig. 2</figureCitation>
|
||
A,B); Finger I longer than Finger II; fingers and toes lacking dermal flanges along sides of digits, subarticular tubercles large and highly protuberant (
|
||
<figureCitation id="133EDBA75E0E6B4F3F42F953FBC2F970" box="[1057,1125,1717,1742]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="4" pageNumber="195">Fig. 2</figureCitation>
|
||
A,B); skin smooth, but with scattered and raised dermal tubercles (
|
||
<figureCitation id="133EDBA75E0E6B4F390BF93FFD19F94C" box="[616,702,1753,1778]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="4" pageNumber="195">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="133EDBA75E0E6B4F39B1F93CFD44F94C" box="[722,739,1754,1778]" captionStart="FIGURE 4" captionStartId="6.[151,244,1685,1706]" captionTargetBox="[307,1275,204,1636]" captionTargetId="figure@6.[289,1297,191,1663]" captionTargetPageId="6" captionText="FIGURE 4. Oblique view illustration of lateral head of (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817) and (B) P. dorsalis (KU 300356). Scale bars = 2 mm; Illustrations by M. Garfield." httpUri="https://zenodo.org/record/239896/files/figure.png" pageId="4" pageNumber="195">4</figureCitation>
|
||
); dorsolateral folds, mid-dorsal ridges, and other cephalic protuberances absent; color pattern medium to dark brown with black spots (
|
||
<figureCitation id="133EDBA75E0E6B4F389AF91BFB99F8A8" box="[1017,1086,1789,1814]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="4" pageNumber="195">Fig. 3</figureCitation>
|
||
); advertisement call a series of invariant low-frequency stridulated one-syllable barks: “Iyohk, iyohk, iyohk…” (
|
||
<figureCitation id="133EDBA75E0E6B4F3F33F8C7FB3FF884" box="[1104,1176,1825,1850]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="4" pageNumber="195">Fig. 6</figureCitation>
|
||
) and its restriction to terrestrial microhabitats in forest covering limestone karst formations (
|
||
<figureCitation id="133EDBA75E0E6B4F3898F8A3FBEEF8E0" box="[1019,1097,1861,1886]" captionStart="FIGURE 5" captionStartId="7.[151,244,1518,1539]" captionTargetBox="[250,1335,449,1488]" captionTargetId="figure@7.[243,1344,443,1496]" captionTargetPageId="7" captionText="FIGURE 5. Limestone karst forest habitat of Platymantis (Lupacolus) quezoni, sp. nov., at the type locality: (A) limestone bolder where the holotype (PNM 9817) was recorded and collected; (B) appearance of habitat in late afternoon (~ 16: 30). Note darkened caves and crevices beneath limestone over-hangs from where P. q u e z o n i initiate vocalizations in the late afternoons (16: 30 – 17: 30 hr) from concealed perches inside crevices and small caves. Photos: RMB." httpUri="https://zenodo.org/record/239897/files/figure.png" pageId="4" pageNumber="195">Fig. 5</figureCitation>
|
||
). Diagnostic character state differences are summarized in
|
||
<tableCitation id="C687F2995E0E6B4F3A94F88FFDEBF83C" box="[503,588,1897,1922]" captionStart="TABLE 1" captionStartId="3.[151,231,259,280]" captionText="TABLE 1. — Summary of body size and selected qualitative diagnostic characters (+ present, – absent) in Platymantis (Lupacolus) quezoni, sp. nov., and other species of Philippine limestone karst forest frogs. Sample size, body size (Snout – vent length, SVL, in mm), former species group (Brown et al. 1997 a; Alcala & Brown, 1999), evolutionary affinities indicated by phylogenetically-defined subgenus (Brown et al. 2015), and general geographical provided for reference. The call of P. (Tahananpuno) bayani is still unknown (Siler et al. 2009)." pageId="4" pageNumber="195">Table 1</tableCitation>
|
||
and we provide detailed comparisons, emphasizing diagnostic differences between the new species and its close relatives below.
|
||
</paragraph>
|
||
<caption id="DF7A97AA5E0F6B4E3BF4F9EDFD24F983" httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="5" pageNumber="196" targetBox="[249,1334,199,1486]" targetPageId="5">
|
||
<paragraph id="8BBAC7225E0F6B4E3BF4F9EDFD24F983" blockId="5.[151,1435,1547,1597]" pageId="5" pageNumber="196">
|
||
<emphasis id="B9711B305E0F6B4E3BF4F9EDFEA9F99E" bold="true" box="[151,270,1547,1568]" pageId="5" pageNumber="196">FIGURE 3.</emphasis>
|
||
Photographs of the (A)
|
||
<taxonomicName id="4C05BCA15E0F6B4E3A9DF9EDFCE6F99E" box="[510,833,1547,1568]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="quezoni" status="sp. nov." subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E0F6B4E3A9DF9EDFDD7F99E" box="[510,624,1547,1568]" italics="true" pageId="5" pageNumber="196">Platymantis</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0F6B4E39E3F9EDFD43F99E" box="[640,740,1547,1568]" italics="true" pageId="5" pageNumber="196">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E0F6B4E3997F9EDFCE6F99E" box="[756,833,1547,1568]" italics="true" pageId="5" pageNumber="196">quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9711B305E0F6B4E382FF9EDFC06F99E" bold="true" box="[844,929,1547,1568]" pageId="5" pageNumber="196">
|
||
<taxonomicNameLabel id="A242A64B5E0F6B4E382FF9EDFC3CF99E" box="[844,923,1547,1568]" pageId="5" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBAC7225E0F6B4E3BA4F98CFE88F855" blockId="5.[151,1437,1642,2027]" pageId="5" pageNumber="196">
|
||
<emphasis id="B9711B305E0F6B4E3BA4F98CFECBF93D" bold="true" box="[199,364,1642,1667]" pageId="5" pageNumber="196">Comparisons.</emphasis>
|
||
We assign the new species to the subgenus
|
||
<emphasis id="B9711B305E0F6B4E3811F98CFC40F93D" box="[882,999,1642,1667]" italics="true" pageId="5" pageNumber="196">Lupacolus</emphasis>
|
||
Brown
|
||
<emphasis id="B9711B305E0F6B4E3F25F98DFB26F93D" box="[1094,1153,1642,1667]" italics="true" pageId="5" pageNumber="196">et al.</emphasis>
|
||
2015 on the basis of its placement (as “
|
||
<taxonomicName id="4C05BCA15E0F6B4E3A27F968FE6DF919" box="[324,458,1678,1703]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9711B305E0F6B4E3A27F968FE6DF919" box="[324,458,1678,1703]" italics="true" pageId="5" pageNumber="196">Platymantis</emphasis>
|
||
</taxonomicName>
|
||
sp. 27”) in Clade R of the genus
|
||
<taxonomicName id="4C05BCA15E0F6B4E385FF968FC65F919" box="[828,962,1678,1703]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9711B305E0F6B4E385FF968FC65F919" box="[828,962,1678,1703]" italics="true" pageId="5" pageNumber="196">Platymantis</emphasis>
|
||
</taxonomicName>
|
||
(Brown
|
||
<emphasis id="B9711B305E0F6B4E3F46F969FBF1F919" box="[1061,1110,1678,1703]" italics="true" pageId="5" pageNumber="196">et al</emphasis>
|
||
. 2015:
|
||
<figureCitation id="133EDBA75E0F6B4E3FC9F968FB49F919" box="[1194,1262,1678,1703]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="5" pageNumber="196">Fig. 2</figureCitation>
|
||
). The subgenus
|
||
<emphasis id="B9711B305E0F6B4E3BF4F954FEABF975" box="[151,268,1714,1739]" italics="true" pageId="5" pageNumber="196">Lupacolus</emphasis>
|
||
is largely equivalent of the former
|
||
<taxonomicName id="4C05BCA15E0F6B4E39D2F955FC8DF975" box="[689,810,1714,1739]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E0F6B4E39D2F955FC8DF975" box="[689,810,1714,1739]" italics="true" pageId="5" pageNumber="196">P. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
Group of
|
||
<bibRefCitation id="EF94BAD35E0F6B4E38CAF955FB3AF975" box="[937,1181,1714,1739]" pageId="5" pageNumber="196" refString="Brown, W. C., Brown, R. M. & Alcala, A. C. (1997 a) Species of the hazelae group of Platymantis from the Philippines, with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405 - 421." type="journal article">
|
||
Brown
|
||
<emphasis id="B9711B305E0F6B4E3F63F955FB9BF975" box="[1024,1084,1714,1739]" italics="true" pageId="5" pageNumber="196">et al.</emphasis>
|
||
(1997a)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF94BAD35E0F6B4E3FBFF954FF44F951" author="Alcala" pageId="5" pageNumber="196" refString="Brown W. C., Alcala, A. C. & Diesmos A. C. (1999) Four new species of the genus Platymantis (Amphibia: Ranidae) from Luzon Island, Philippines. Proceedings of the California Academy of Sciences, 51, 449 - 460." type="journal article" year="1999">Alcala & Brown (1999)</bibRefCitation>
|
||
, and contains the named taxa
|
||
<emphasis id="B9711B305E0F6B4E3953F931FE73F8AD" italics="true" pageId="5" pageNumber="196">
|
||
<taxonomicName id="4C05BCA15E0F6B4E3953F931FD6FF950" box="[560,712,1751,1774]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E39B0F931FCEDF951" box="[723,842,1750,1775]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3836F931FC5EF951" box="[853,1017,1750,1775]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3F67F931FB27F951" box="[1028,1152,1750,1775]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3FE8F931FAAFF950" box="[1163,1288,1751,1774]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3E70F931FADAF950" box="[1299,1405,1751,1774]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="paengi">P. paengi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3EE4F931FEE3F8AD" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="pseudodorsalis">P. pseudodorsalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3A30F91DFE73F8AD" box="[339,468,1786,1811]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="spelaeus">P. spelaeus</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<emphasis id="B9711B305E0F6B4E3978F91DFD25F8AC" box="[539,642,1786,1811]" italics="true" pageId="5" pageNumber="196">P. t a y l o r i</emphasis>
|
||
but does not include
|
||
<taxonomicName id="4C05BCA15E0F6B4E38ECF91DFC47F8AD" box="[911,992,1786,1811]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="biak">
|
||
<emphasis id="B9711B305E0F6B4E38ECF91DFC47F8AD" box="[911,992,1786,1811]" italics="true" pageId="5" pageNumber="196">P. biak</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9711B305E0F6B4E3890F91DFABBF8AC" box="[1011,1308,1786,1811]" italics="true" pageId="5" pageNumber="196">
|
||
<taxonomicName id="4C05BCA15E0F6B4E3890F91DFBDBF8AD" box="[1011,1148,1786,1811]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="levigatus">P. levigatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0F6B4E3FEFF91DFABBF8AC" box="[1164,1308,1787,1810]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="pygmaeus">P. pygmaeus</taxonomicName>
|
||
</emphasis>
|
||
(placed in subgenus
|
||
<emphasis id="B9711B305E0F6B4E3A65F8F8FE07F889" box="[262,416,1822,1847]" italics="true" pageId="5" pageNumber="196">Lahatnanguri</emphasis>
|
||
; Clade O of Brown
|
||
<emphasis id="B9711B305E0F6B4E39E1F8F9FD1DF889" box="[642,698,1822,1847]" italics="true" pageId="5" pageNumber="196">et al.</emphasis>
|
||
2015:
|
||
<figureCitation id="133EDBA75E0F6B4E3864F8F8FCEEF889" box="[775,841,1822,1847]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="5" pageNumber="196">Fig. 2</figureCitation>
|
||
) or
|
||
<taxonomicName id="4C05BCA15E0F6B4E3815F8F9FBAAF888" box="[886,1037,1823,1846]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9711B305E0F6B4E3815F8F9FBAAF888" box="[886,1037,1823,1846]" italics="true" pageId="5" pageNumber="196">P. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
(subgenus
|
||
<taxonomicName id="4C05BCA15E0F6B4E3FEFF8F8FAB4F889" box="[1164,1299,1822,1847]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">Platymantis</taxonomicName>
|
||
, clade name Tagomukhus; Clade N of Brown
|
||
<emphasis id="B9711B305E0F6B4E3973F8A5FDE5F8E5" box="[528,578,1858,1883]" italics="true" pageId="5" pageNumber="196">et al</emphasis>
|
||
. 2015:
|
||
<figureCitation id="133EDBA75E0F6B4E39FBF8A4FD7AF8E5" box="[664,733,1858,1883]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="5" pageNumber="196">Fig. 2</figureCitation>
|
||
). Members of the genus
|
||
<taxonomicName id="4C05BCA15E0F6B4E3894F8A4FBDAF8E5" box="[1015,1149,1858,1883]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="196" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9711B305E0F6B4E3894F8A4FBDAF8E5" box="[1015,1149,1858,1883]" italics="true" pageId="5" pageNumber="196">Platymantis</emphasis>
|
||
</taxonomicName>
|
||
, subgenus
|
||
<emphasis id="B9711B305E0F6B4E3F9FF8A4FAD6F8E5" box="[1276,1393,1858,1883]" italics="true" pageId="5" pageNumber="196">Lupacolus</emphasis>
|
||
, all possess the following combination of characters: body size moderate (~
|
||
<quantity id="4CFD6AC75E0F6B4E38B2F880FBE8F8C0" box="[977,1103,1894,1919]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="5" pageNumber="196" unit="mm" value="35.0" valueMax="45.0" valueMin="25.0">25–45 mm</quantity>
|
||
SVL in males); digital discs non-expanded to minimally expanded; first finger longer than second; fingers and toes lacking dermal flanges along sides of digits; subarticular tubercles large and highly protuberant (
|
||
<bibRefCitation id="EF94BAD35E0F6B4E38BFF849FB1BF878" author="Brown" box="[988,1212,1966,1991]" pageId="5" pageNumber="196" refString="Brown, W. C., Brown, R. M. & Alcala, A. C. (1997 a) Species of the hazelae group of Platymantis from the Philippines, with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405 - 421." type="journal article" year="1997" yearSuffix="a">
|
||
Brown
|
||
<emphasis id="B9711B305E0F6B4E3F50F849FBCAF879" box="[1075,1133,1966,1991]" italics="true" pageId="5" pageNumber="196">et al.</emphasis>
|
||
1997
|
||
<emphasis id="B9711B305E0F6B4E3FCEF849FB1BF878" box="[1197,1212,1967,1990]" italics="true" pageId="5" pageNumber="196">a</emphasis>
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF94BAD35E0F6B4E3FA2F849FB69F878" author="Brown" box="[1217,1230,1967,1990]" pageId="5" pageNumber="196" refString="Brown, W. C., Alcala, A. C., Diesmos, A. C. & Alcala, E. (1997 c) Species of the guentheri group of Platymantis from the Philippines with descriptions of four new species. Proceedings of the California Academy of Sciences, 50, 1 - 20." type="journal article" year="1997" yearSuffix="c">c</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E0F6B4E3FBCF848FF74F855" author="Alcala" pageId="5" pageNumber="196" refString="Brown W. C., Alcala, A. C. & Diesmos A. C. (1999) Four new species of the genus Platymantis (Amphibia: Ranidae) from Luzon Island, Philippines. Proceedings of the California Academy of Sciences, 51, 449 - 460." type="journal article" year="1999">Alcala & Brown 1999</bibRefCitation>
|
||
;
|
||
<figureCitation id="133EDBA75E0F6B4E3BBEF834FE86F855" box="[221,289,2002,2027]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="5" pageNumber="196">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7A97AA5E0C6B4D3BF4F973FCFBF979" httpUri="https://zenodo.org/record/239896/files/figure.png" pageId="6" pageNumber="197" targetBox="[307,1275,204,1636]" targetPageId="6">
|
||
<paragraph id="8BBAC7225E0C6B4D3BF4F973FCFBF979" blockId="6.[151,1436,1685,1735]" pageId="6" pageNumber="197">
|
||
<emphasis id="B9711B305E0C6B4D3BF4F973FEABF914" bold="true" box="[151,268,1685,1706]" pageId="6" pageNumber="197">FIGURE 4.</emphasis>
|
||
Oblique view illustration of lateral head of (A)
|
||
<taxonomicName id="4C05BCA15E0C6B4D39BBF973FBB0F914" box="[728,1047,1685,1706]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="quezoni" status="sp. nov." subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E0C6B4D39BBF973FCECF914" box="[728,843,1685,1706]" italics="true" pageId="6" pageNumber="197">Platymantis</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0C6B4D383AF973FC1AF914" box="[857,957,1685,1706]" italics="true" pageId="6" pageNumber="197">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E0C6B4D38A9F973FBB0F914" box="[970,1047,1685,1706]" italics="true" pageId="6" pageNumber="197">quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9711B305E0C6B4D3F42F973FBD3F914" bold="true" box="[1057,1140,1685,1706]" pageId="6" pageNumber="197">
|
||
<taxonomicNameLabel id="A242A64B5E0C6B4D3F42F973FBC9F914" box="[1057,1134,1685,1706]" pageId="6" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
holotype (PNM 9817) and (B)
|
||
<taxonomicName id="4C05BCA15E0C6B4D3BF4F954FF5CF979" box="[151,251,1714,1735]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E0C6B4D3BF4F954FF5CF979" box="[151,251,1714,1735]" italics="true" pageId="6" pageNumber="197">P. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
(KU 300356). Scale bars = 2 mm; Illustrations by M. Garfield.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBAC7225E0C6B4C3BA4F8FFFF77FE36" blockId="6.[151,1436,1816,2021]" lastBlockId="7.[151,1436,151,392]" lastPageId="7" lastPageNumber="198" pageId="6" pageNumber="197">
|
||
In overall phenotypic similarity
|
||
<taxonomicName id="4C05BCA15E0C6B4D3953F8FEFCB3F88E" box="[560,788,1816,1841]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0C6B4D3953F8FEFCB3F88E" box="[560,788,1816,1841]" italics="true" pageId="6" pageNumber="197">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
most resembles the related species
|
||
<emphasis id="B9711B305E0C6B4D3FC0F8FFFA3BF88F" box="[1187,1436,1816,1841]" italics="true" pageId="6" pageNumber="197">
|
||
<taxonomicName id="4C05BCA15E0C6B4D3FC0F8FFFAA9F88E" box="[1187,1294,1817,1840]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="paengi">P. paengi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0C6B4D3E7AF8FFFA30F88F" box="[1305,1431,1816,1841]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="spelaeus">P. spelaeus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
(all three belong to the subgenus
|
||
<emphasis id="B9711B305E0C6B4D3927F8DAFD1FF8EB" box="[580,696,1852,1877]" italics="true" pageId="6" pageNumber="197">Lupacolus</emphasis>
|
||
), the unrelated
|
||
<emphasis id="B9711B305E0C6B4D38E6F8DBFBDBF8EB" box="[901,1148,1852,1877]" italics="true" pageId="6" pageNumber="197">
|
||
<taxonomicName id="4C05BCA15E0C6B4D38E6F8DBFC79F8EB" box="[901,990,1852,1877]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="biak">P. biak</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0C6B4D3891F8DBFBDBF8EB" box="[1010,1148,1852,1877]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="insulatus">P. insulatus</taxonomicName>
|
||
</emphasis>
|
||
(both of the subgenus
|
||
<emphasis id="B9711B305E0C6B4D3BF4F886FE96F8C7" box="[151,305,1888,1913]" italics="true" pageId="6" pageNumber="197">Lahatnanguri</emphasis>
|
||
), and
|
||
<taxonomicName id="4C05BCA15E0C6B4D3AEBF887FE5FF8C7" box="[392,504,1888,1913]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="bayani">
|
||
<emphasis id="B9711B305E0C6B4D3AEBF887FE5FF8C7" box="[392,504,1888,1913]" italics="true" pageId="6" pageNumber="197">P. bayani</emphasis>
|
||
</taxonomicName>
|
||
(subgenus
|
||
<emphasis id="B9711B305E0C6B4D39EAF886FC85F8C7" box="[649,802,1888,1913]" italics="true" pageId="6" pageNumber="197">Tahananpuno</emphasis>
|
||
). All six taxa share the general mottled limestonecamouflaged color pattern and all exclusively inhabit limestone karst microhabitats.
|
||
<taxonomicName id="4C05BCA15E0C6B4D3F06F862FAECF822" box="[1125,1355,1924,1949]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0C6B4D3F06F862FAECF822" box="[1125,1355,1924,1949]" italics="true" pageId="6" pageNumber="197">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
shares with the other members of the subgenus
|
||
<emphasis id="B9711B305E0C6B4D390FF84EFD47F87F" box="[620,736,1960,1985]" italics="true" pageId="6" pageNumber="197">Lupacolus</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0C6B4D3990F84FFC4CF87F" box="[755,1003,1960,1985]" italics="true" pageId="6" pageNumber="197">
|
||
<taxonomicName id="4C05BCA15E0C6B4D3990F84FFCC7F87E" box="[755,864,1961,1984]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="paengi">P. paengi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0C6B4D380EF84FFC4CF87F" box="[877,1003,1960,1985]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="spelaeus">P. spelaeus</taxonomicName>
|
||
</emphasis>
|
||
) minimally expanded terminal digital discs, whereas these two species of subgenus
|
||
<emphasis id="B9711B305E0C6B4D39F7F82AFC89F85B" box="[660,814,1996,2021]" italics="true" pageId="6" pageNumber="197">Lahatnanguri</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0C6B4D385DF82BFBB0F85B" box="[830,1047,1996,2021]" italics="true" pageId="6" pageNumber="197">
|
||
<taxonomicName id="4C05BCA15E0C6B4D385DF82BFC2BF85B" box="[830,908,1996,2021]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="biak">P. biak</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0C6B4D38F4F82BFBB0F85B" box="[919,1047,1996,2021]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="197" phylum="Chordata" rank="species" species="insulatus">P. insulatus</taxonomicName>
|
||
</emphasis>
|
||
), and all members of the subgenus
|
||
<emphasis id="B9711B305E0D6B4C3BF4FF71FE97FF0E" box="[151,304,151,176]" italics="true" pageId="7" pageNumber="198">Tahananpuno</emphasis>
|
||
(including
|
||
<taxonomicName id="4C05BCA15E0D6B4C3AD1FF7FFDBEFF0E" box="[434,537,151,176]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="bayani">
|
||
<emphasis id="B9711B305E0D6B4C3AD1FF7FFDBEFF0E" box="[434,537,151,176]" italics="true" pageId="7" pageNumber="198">P. bayani</emphasis>
|
||
</taxonomicName>
|
||
) have widely expanded terminal discs of the fingers and toes (
|
||
<bibRefCitation id="EF94BAD35E0D6B4C3FB9FF7EFF74FF6A" author="Brown" pageId="7" pageNumber="198" refString="Brown, W. C. & Alcala, A. C. (1970) A new species of the genus Platymantis (Ranidae) with a list of the amphibians known from South Gigante Island, Philippines. Occasional Papers of the California Academy of Sciences, 84, 1 - 7." type="journal article" year="1970">Brown & Alcala, 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E0D6B4C3BBCFF5BFE18FF6A" author="Brown" box="[223,447,188,213]" pageId="7" pageNumber="198" refString="Brown, W. C., Brown, R. M. & Alcala, A. C. (1997 a) Species of the hazelae group of Platymantis from the Philippines, with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405 - 421." type="journal article" year="1997" yearSuffix="a">
|
||
Brown
|
||
<emphasis id="B9711B305E0D6B4C3A57FF5BFEC1FF6B" box="[308,358,188,213]" italics="true" pageId="7" pageNumber="198">et al</emphasis>
|
||
. 1997a
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF94BAD35E0D6B4C3AA7FF5AFE76FF6A" author="Brown" box="[452,465,188,212]" pageId="7" pageNumber="198" refString="Brown, W. C., Alcala, A. C., Diesmos, A. C. & Alcala, E. (1997 c) Species of the guentheri group of Platymantis from the Philippines with descriptions of four new species. Proceedings of the California Academy of Sciences, 50, 1 - 20." type="journal article" year="1997" yearSuffix="c">c</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E0D6B4C3ABEFF5AFD30FF6A" author="Siler" box="[477,663,188,213]" pageId="7" pageNumber="198" refString="Siler, C. D., Alcala, A. C., Diesmos, A. C. & Brown, R. M. (2009) A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from eastern Samar Island, Philippines. Herpetologica, 65, 92 - 104. http: // dx. doi. org / 10.1655 / 08 - 040 R. 1" type="journal article" year="2009">
|
||
Siler
|
||
<emphasis id="B9711B305E0D6B4C3978FF5BFDF3FF6B" box="[539,596,188,213]" italics="true" pageId="7" pageNumber="198">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
, 2010). Additionally,
|
||
<taxonomicName id="4C05BCA15E0D6B4C38ECFF5BFC5FFF6B" box="[911,1016,188,213]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="bayani">
|
||
<emphasis id="B9711B305E0D6B4C38ECFF5BFC5FFF6B" box="[911,1016,188,213]" italics="true" pageId="7" pageNumber="198">P. bayani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F6BFF5BFB2EFF6B" box="[1032,1161,188,213]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="insulatus">
|
||
<emphasis id="B9711B305E0D6B4C3F6BFF5BFB2EFF6B" box="[1032,1161,188,213]" italics="true" pageId="7" pageNumber="198">P. insulatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C05BCA15E0D6B4C3FA9FF5BFAE3FF6A" box="[1226,1348,188,213]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="subSpecies" species="spel" subSpecies="aeus">
|
||
<emphasis id="B9711B305E0D6B4C3FA9FF5BFAE3FF6A" box="[1226,1348,188,213]" italics="true" pageId="7" pageNumber="198">P. spel aeus</emphasis>
|
||
</taxonomicName>
|
||
further differ by their much larger body sizes, which do not overlap the size range of
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F47FF07FB3FFF46" box="[1060,1176,225,248]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0D6B4C3F47FF07FB3FFF46" box="[1060,1176,225,248]" italics="true" pageId="7" pageNumber="198">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C687F2995E0D6B4C3FC9FF39FAA6FF46" box="[1194,1281,223,248]" captionStart="TABLE 1" captionStartId="3.[151,231,259,280]" captionText="TABLE 1. — Summary of body size and selected qualitative diagnostic characters (+ present, – absent) in Platymantis (Lupacolus) quezoni, sp. nov., and other species of Philippine limestone karst forest frogs. Sample size, body size (Snout – vent length, SVL, in mm), former species group (Brown et al. 1997 a; Alcala & Brown, 1999), evolutionary affinities indicated by phylogenetically-defined subgenus (Brown et al. 2015), and general geographical provided for reference. The call of P. (Tahananpuno) bayani is still unknown (Siler et al. 2009)." pageId="7" pageNumber="198">Table 1</tableCitation>
|
||
). Finally, the advertisement call of the new species (a rapid series of loud, harsh, single-syllable barks) differs from that of
|
||
<taxonomicName id="4C05BCA15E0D6B4C3EE4FEE3FF60FEFE" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="biak">
|
||
<emphasis id="B9711B305E0D6B4C3EE4FEE3FF60FEFE" italics="true" pageId="7" pageNumber="198">P. biak</emphasis>
|
||
</taxonomicName>
|
||
(a much slower, low amplitude series of tonal notes),
|
||
<taxonomicName id="4C05BCA15E0D6B4C3844FECFFC00FEFE" box="[807,935,295,320]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="insulatus">
|
||
<emphasis id="B9711B305E0D6B4C3844FECFFC00FEFE" box="[807,935,295,320]" italics="true" pageId="7" pageNumber="198">P. insulatus</emphasis>
|
||
</taxonomicName>
|
||
(a long clicking train of brief atonal pulses),
|
||
<taxonomicName id="4C05BCA15E0D6B4C3BF4FEABFEB3FEDB" box="[151,276,332,357]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="spelaeus">
|
||
<emphasis id="B9711B305E0D6B4C3BF4FEABFEB3FEDB" box="[151,276,332,357]" italics="true" pageId="7" pageNumber="198">P. spelaeus</emphasis>
|
||
</taxonomicName>
|
||
(a four-syllable, multinote complex call) and
|
||
<taxonomicName id="4C05BCA15E0D6B4C387DFEABFC20FEDA" box="[798,903,333,356]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="paengi">
|
||
<emphasis id="B9711B305E0D6B4C387DFEABFC20FEDA" box="[798,903,333,356]" italics="true" pageId="7" pageNumber="198">P. paengi</emphasis>
|
||
</taxonomicName>
|
||
(tonal, frequency sweep, amplitude modulated call).
|
||
</paragraph>
|
||
<caption id="DF7A97AA5E0D6B4C3BF4FA08FCE5F9E5" httpUri="https://zenodo.org/record/239897/files/figure.png" pageId="7" pageNumber="198" targetBox="[250,1335,449,1488]" targetPageId="7">
|
||
<paragraph id="8BBAC7225E0D6B4C3BF4FA08FCE5F9E5" blockId="7.[151,1436,1518,1627]" pageId="7" pageNumber="198">
|
||
<emphasis id="B9711B305E0D6B4C3BF4FA08FEA9F9BD" bold="true" box="[151,270,1518,1539]" pageId="7" pageNumber="198">FIGURE 5.</emphasis>
|
||
Limestone karst forest habitat of
|
||
<taxonomicName id="4C05BCA15E0D6B4C393DFA08FC06F9BD" box="[606,929,1518,1539]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="quezoni" status="sp. nov." subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E0D6B4C393DFA08FD76F9BD" box="[606,721,1518,1539]" italics="true" pageId="7" pageNumber="198">Platymantis</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0D6B4C3983FA08FCE3F9BD" box="[736,836,1518,1539]" italics="true" pageId="7" pageNumber="198">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E0D6B4C3837FA08FC06F9BD" box="[852,929,1518,1539]" italics="true" pageId="7" pageNumber="198">quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9711B305E0D6B4C38CEFA08FBA5F9BD" bold="true" box="[941,1026,1518,1539]" pageId="7" pageNumber="198">
|
||
<taxonomicNameLabel id="A242A64B5E0D6B4C38CEFA08FC5BF9BD" box="[941,1020,1518,1539]" pageId="7" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
at the type locality: (A) limestone bolder where the holotype (PNM 9817) was recorded and collected; (B) appearance of habitat in late afternoon (~16:30). Note darkened caves and crevices beneath limestone over-hangs from where
|
||
<emphasis id="B9711B305E0D6B4C398EF9CFFCE9F980" box="[749,846,1577,1598]" italics="true" pageId="7" pageNumber="198">P. q u e z o n i</emphasis>
|
||
initiate vocalizations in the late afternoons (16:30–17:30 hr) from concealed perches inside crevices and small caves. Photos: RMB.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBAC7225E0D6B4C3BA4F94DFBC7F822" blockId="7.[151,1437,1707,2020]" pageId="7" pageNumber="198">
|
||
<taxonomicName id="4C05BCA15E0D6B4C3BA4F94DFE0CF97A" box="[199,427,1707,1732]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0D6B4C3BA4F94DFE0CF97A" box="[199,427,1707,1732]" italics="true" pageId="7" pageNumber="198">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from the remaining members of the subgenus
|
||
<emphasis id="B9711B305E0D6B4C3F0DF94DFAA0F97A" box="[1134,1287,1707,1732]" italics="true" pageId="7" pageNumber="198">Tahananpuno</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0D6B4C3E74F94BFD22F956" italics="true" pageId="7" pageNumber="198">
|
||
<taxonomicName id="4C05BCA15E0D6B4C3E74F94BFA3FF97A" box="[1303,1432,1707,1732]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="diesmosi">P. diesmosi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3BF4F937FE89F956" box="[151,302,1743,1768]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="luzonensis">P. luzonensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3A5DF937FE43F956" box="[318,484,1745,1768]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="negrosensis">P. negrosensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3A90F937FD26F956" box="[499,641,1743,1768]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="guentheri">P. guentheri</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05BCA15E0D6B4C39A7F937FC8CF956" box="[708,811,1743,1768]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="rabori">
|
||
<emphasis id="B9711B305E0D6B4C39A7F937FC8CF956" box="[708,811,1743,1768]" italics="true" pageId="7" pageNumber="198">P. rabori</emphasis>
|
||
</taxonomicName>
|
||
) by the presence of minimally expanded (vs. widely expanded) terminal digital discs of the fingers and toes. The terrestrial limestone karst microhabitat of
|
||
<taxonomicName id="4C05BCA15E0D6B4C3E4BF913FA3BF8B2" box="[1320,1436,1781,1804]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0D6B4C3E4BF913FA3BF8B2" box="[1320,1436,1781,1804]" italics="true" pageId="7" pageNumber="198">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
further distinguishes the new species from all of these arboreal tree frog taxa except
|
||
<emphasis id="B9711B305E0D6B4C3FE0F8FFFAA6F88E" box="[1155,1281,1815,1840]" italics="true" pageId="7" pageNumber="198">P. d i e s m o s i</emphasis>
|
||
(a terrestrial species calling exclusively from cliff ledges on Mt. Malino;
|
||
<bibRefCitation id="EF94BAD35E0D6B4C3828F8DAFBCEF8EA" author="Brown" box="[843,1129,1851,1876]" pageId="7" pageNumber="198" refString="Brown, R. M & Gonzales, J. C. (2007) A new forest frog of the genus Platymantis (Amphibia; Anura; Ranidae) from the Bicol Peninsula of Luzon Island, Philippines. Copeia, 2007, 251 - 266. http: // dx. doi. org / 10.1643 / 0045 - 8511 (2007) 7 [251: ANFFOT] 2.0. CO; 2" type="journal article" year="2007">Brown & Gonzales 2007</bibRefCitation>
|
||
) and its advertisement call likewise distinguishes
|
||
<taxonomicName id="4C05BCA15E0D6B4C3AF7F887FDA0F8C6" box="[404,519,1889,1912]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E0D6B4C3AF7F887FDA0F8C6" box="[404,519,1889,1912]" italics="true" pageId="7" pageNumber="198">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
from the tree frogs with amplitude modulated calls (
|
||
<emphasis id="B9711B305E0D6B4C3F3BF887FA3BF8C6" box="[1112,1436,1887,1912]" italics="true" pageId="7" pageNumber="198">
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F3BF887FB4CF8C6" box="[1112,1259,1887,1912]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="luzonensis">P. luzonensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F95F887FA3FF8C6" box="[1270,1432,1889,1912]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="negrosensis">P. negrosensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05BCA15E0D6B4C3BA5F863FE8DF822" box="[198,298,1923,1948]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="rabori">
|
||
<emphasis id="B9711B305E0D6B4C3BA5F863FE8DF822" box="[198,298,1923,1948]" italics="true" pageId="7" pageNumber="198">P. rabori</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C05BCA15E0D6B4C3A0AF863FE48F822" box="[361,495,1923,1948]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="guentheri">
|
||
<emphasis id="B9711B305E0D6B4C3A0AF863FE48F822" box="[361,495,1923,1948]" italics="true" pageId="7" pageNumber="198">P. guentheri</emphasis>
|
||
</taxonomicName>
|
||
, an arboreal species with a tonal, frequency sweep call.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E0D6B4C3BA4F841FA3AF85A" blockId="7.[151,1437,1707,2020]" pageId="7" pageNumber="198">
|
||
The new species is distinguished from all species of
|
||
<collectingCountry id="F31287B25E0D6B4C384EF841FC07F87E" box="[813,928,1959,1984]" name="Philippines" pageId="7" pageNumber="198">Philippine</collectingCountry>
|
||
cloud frogs (formerly the
|
||
<emphasis id="B9711B305E0D6B4C3FB4F84FFAE0F87E" box="[1239,1351,1959,1984]" italics="true" pageId="7" pageNumber="198">P. ha z e l a e</emphasis>
|
||
Group species) of the subgenus
|
||
<emphasis id="B9711B305E0D6B4C3ACEF82DFD9FF85A" box="[429,568,1995,2020]" italics="true" pageId="7" pageNumber="198">Tirahanulap</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E0D6B4C3924F82BFA3AF85A" box="[583,1437,1995,2020]" italics="true" pageId="7" pageNumber="198">
|
||
<taxonomicName id="4C05BCA15E0D6B4C3924F82BFD1DF85A" box="[583,698,1995,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="hazelae">P. hazelae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C39A6F82BFC8EF85A" box="[709,809,1997,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="isarog">P. isarog</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3856F82BFC00F85A" box="[821,935,1995,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="lawtoni">P. lawtoni</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C38D2F82BFB9BF85A" box="[945,1084,1997,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="montanus">P. montanus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F24F82BFB46F85A" box="[1095,1249,1997,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="panayensis">P. panayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E0D6B4C3F8FF82BFADAF85A" box="[1260,1405,1995,2020]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="198" phylum="Chordata" rank="species" species="polillensis">P. polillensis</taxonomicName>
|
||
, P.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B433BF4FF71FDD4FEDA" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
<emphasis id="B9711B305E026B433BF4FF71FEF4FF0E" box="[151,339,151,176]" italics="true" pageId="8" pageNumber="199">
|
||
<taxonomicName id="4C05BCA15E026B433BF4FF71FEE8FF0E" box="[151,335,151,176]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="sierramadrensis">sierramadrensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05BCA15E026B433AEFFF7FFD93FF0E" box="[396,564,151,176]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="subterrestris">
|
||
<emphasis id="B9711B305E026B433AEFFF7FFD93FF0E" box="[396,564,151,176]" italics="true" pageId="8" pageNumber="199">P. subterrestris</emphasis>
|
||
</taxonomicName>
|
||
) by its larger body size (vs. much smaller in these species, typically ~
|
||
<quantity id="4CFD6AC75E026B433E36FF7EFF64FF6A" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.8" metricValueMin="2.0" pageId="8" pageNumber="199" unit="mm" value="24.0" valueMax="28.0" valueMin="20.0">20–28 mm</quantity>
|
||
in males), by unadorned digits (vs. digits with wide dermal fringes), by strongly protuberant (vs. ventrally flattened) subarticular tubercles, by possessing a longer first finger than the second (vs. second longest) and its terrestrial limestone karst microhabitat (vs. shrub-layer vegetation of montane forests). Species of the subgenus
|
||
<emphasis id="B9711B305E026B433BF4FEC1FE85FEFE" box="[151,290,295,320]" italics="true" pageId="8" pageNumber="199">Tirahanulap</emphasis>
|
||
all possess tonal, constant frequency calls with complex harmonic structure, contrasting sharply with the loud, harsh barking calls of
|
||
<emphasis id="B9711B305E026B433A9AFEABFDD4FEDA" box="[505,627,333,356]" italics="true" pageId="8" pageNumber="199">
|
||
<taxonomicName id="4C05BCA15E026B433A9AFEABFDC8FEDA" box="[505,623,333,356]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">P. quezoni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B433BA4FE96FAF0FD19" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
From remaining members of the subgenus
|
||
<emphasis id="B9711B305E026B4339ABFE89FC9AFE36" box="[712,829,367,392]" italics="true" pageId="8" pageNumber="199">Lupacolus</emphasis>
|
||
(previously referred to as the
|
||
<taxonomicName id="4C05BCA15E026B433FCCFE89FA3BFE36" box="[1199,1436,367,392]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E026B433FCCFE89FA3BFE36" box="[1199,1436,367,392]" italics="true" pageId="8" pageNumber="199">Platymantis dorsalis</emphasis>
|
||
</taxonomicName>
|
||
Group) (
|
||
<emphasis id="B9711B305E026B433B99FE73FB6DFE13" box="[250,1226,404,429]" italics="true" pageId="8" pageNumber="199">
|
||
<taxonomicName id="4C05BCA15E026B433B99FE73FE31FE12" box="[250,406,405,428]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433AC0FE73FDB9FE13" box="[419,542,404,429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B43394FFE73FD74FE13" box="[556,723,404,429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433983FE73FCC7FE13" box="[736,864,404,429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B43380EFE73FC48FE12" box="[877,1007,405,428]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B43389FFE73FB60FE13" box="[1020,1223,404,429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="pseudodorsalis">P. pseudodorsalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05BCA15E026B433E66FE73FACDFE12" box="[1285,1386,404,429]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="subSpecies" species="tayl" subSpecies="ori">
|
||
<emphasis id="B9711B305E026B433E66FE73FACDFE12" box="[1285,1386,404,429]" italics="true" pageId="8" pageNumber="199">P. tayl ori</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C05BCA15E026B433EE4FE73FF49FE6E" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E026B433EE4FE73FF49FE6E" italics="true" pageId="8" pageNumber="199">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by from all species except
|
||
<taxonomicName id="4C05BCA15E026B433987FE5FFCC0FE6E" box="[740,871,439,464]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="levigatus">
|
||
<emphasis id="B9711B305E026B433987FE5FFCC0FE6E" box="[740,871,439,464]" italics="true" pageId="8" pageNumber="199">P. levigatus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of dorsal folds and/or ridges, and limitation of dermal tubercles to its eyelids (vs. presence dermal folds/ridges and widely distributed tubercles over most dorsal body surfaces), and from all
|
||
<emphasis id="B9711B305E026B433913FE19FD43FDA6" box="[624,740,511,536]" italics="true" pageId="8" pageNumber="199">Lupacolus</emphasis>
|
||
except
|
||
<taxonomicName id="4C05BCA15E026B433823FDE7FC0EFDA6" box="[832,937,513,536]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="paengi">
|
||
<emphasis id="B9711B305E026B433823FDE7FC0EFDA6" box="[832,937,513,536]" italics="true" pageId="8" pageNumber="199">P. paengi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B9711B305E026B433887FDE7FBC1FDA6" box="[996,1126,511,536]" italics="true" pageId="8" pageNumber="199">P. s p e l a e us,</emphasis>
|
||
by its terrestrial limestone forest microhabitat (vs. much more generalist inhabitant of leaf litter of forest floor and/or herb-layer vegetation in
|
||
<emphasis id="B9711B305E026B433BF4FDAFFBF4FDDE" box="[151,1107,583,608]" italics="true" pageId="8" pageNumber="199">
|
||
<taxonomicName id="4C05BCA15E026B433BF4FDAFFE97FDDE" box="[151,304,585,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="cagayensis">P. cagayensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433A5FFDAFFE12FDDE" box="[316,437,583,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="dorsalis">P. dorsalis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433AA2FDAFFDC1FDDE" box="[449,614,583,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="indeprensus">P. indeprensus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433911FDAFFD57FDDE" box="[626,752,583,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="mimulus">P. mimulus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433998FDAFFCDDFDDE" box="[763,890,585,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="naomiae">P. naomiae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B4338E5FDAFFBE8FDDE" box="[902,1103,583,608]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="pseudodorsalis">P. pseudodorsalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B9711B305E026B433FE9FDAFFB4AFDDE" box="[1162,1261,583,608]" italics="true" pageId="8" pageNumber="199">P. t a yl or i</emphasis>
|
||
). Other than
|
||
<taxonomicName id="4C05BCA15E026B433EE4FDAFFF56FD3B" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E026B433EE4FDAFFF56FD3B" italics="true" pageId="8" pageNumber="199">P. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C05BCA15E026B433A51FD8BFE5AFD3B" box="[306,509,620,645]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="pseudodorsalis">
|
||
<emphasis id="B9711B305E026B433A51FD8BFE5AFD3B" box="[306,509,620,645]" italics="true" pageId="8" pageNumber="199">P. pseudodorsalis</emphasis>
|
||
</taxonomicName>
|
||
(which possess frequency sweep calls), these species possess complex, multisyllable call structure, which contrasts markedly with the simpler, single-syllable barking calls of
|
||
<taxonomicName id="4C05BCA15E026B433FBEFD77FAF7FD16" box="[1245,1360,657,680]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E026B433FBEFD77FAF7FD16" box="[1245,1360,657,680]" italics="true" pageId="8" pageNumber="199">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B433BA4FD52FD28FC1B" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
The new species differs from
|
||
<taxonomicName id="4C05BCA15E026B43397BFD52FC86FD72" box="[536,801,692,717]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9711B305E026B43397BFD52FC86FD72" box="[536,801,692,717]" italics="true" pageId="8" pageNumber="199">Platymantis corrugatus</emphasis>
|
||
</taxonomicName>
|
||
(subgenus
|
||
<taxonomicName id="4C05BCA15E026B4338C2FD52FB8FFD73" box="[929,1064,692,717]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="genus">Platymantis</taxonomicName>
|
||
, clade name Tagomukhus) by its smooth to shagreened skin texture (vs. dorsal skin with elongate dermal folds), by its lateral head coloration similar to other dorsolateral surfaces (vs. presence of a dark “mask” coloration on the lateral side of the head of
|
||
<taxonomicName id="4C05BCA15E026B433EE4FD1BFEB5FC86" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9711B305E026B433EE4FD1BFEB5FC86" italics="true" pageId="8" pageNumber="199">P. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
) and by its exclusive limestone karst microhabitat (vs. a variety of terrestrial forest
|
||
<typeStatus id="54BE79805E026B433E74FCC6FAF0FC86" box="[1303,1367,800,824]" pageId="8" pageNumber="199">types</typeStatus>
|
||
). The advertisement call of
|
||
<taxonomicName id="4C05BCA15E026B433AF4FCA3FDA8FCE2" box="[407,527,837,860]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E026B433AF4FCA3FDA8FCE2" box="[407,527,837,860]" italics="true" pageId="8" pageNumber="199">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
is surprisingly quite similar to that of
|
||
<taxonomicName id="4C05BCA15E026B433887FCA3FBD8FCE2" box="[996,1151,837,860]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9711B305E026B433887FCA3FBD8FCE2" box="[996,1151,837,860]" italics="true" pageId="8" pageNumber="199">P. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
(a loud series of single “quacks”) except that
|
||
<taxonomicName id="4C05BCA15E026B433AF1FC8FFD8EFC3E" box="[402,553,873,896]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9711B305E026B433AF1FC8FFD8EFC3E" box="[402,553,873,896]" italics="true" pageId="8" pageNumber="199">P. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
is generally crepuscular whereas
|
||
<emphasis id="B9711B305E026B4338CBFC8FFBBFFC3E" box="[936,1048,873,896]" italics="true" pageId="8" pageNumber="199">P. qu e z o ni</emphasis>
|
||
calls beyond sunset and well into night-time hours if precipitation is sufficient.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B433BA4FC49FAC5FAEA" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
The final set of comparisons for the recognition of
|
||
<taxonomicName id="4C05BCA15E026B433869FC57FCDAFC76" box="[778,893,945,968]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E026B433869FC57FCDAFC76" box="[778,893,945,968]" italics="true" pageId="8" pageNumber="199">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
is to the group of species comprising a recently discovered clade for which the subgenus
|
||
<emphasis id="B9711B305E026B43391EFC32FCB0FC53" box="[637,791,980,1005]" italics="true" pageId="8" pageNumber="199">Lahatnanguri</emphasis>
|
||
was established (Brown
|
||
<emphasis id="B9711B305E026B433F23FC33FBDAFC53" box="[1088,1149,980,1005]" italics="true" pageId="8" pageNumber="199">et al.</emphasis>
|
||
2015). The new species differs from tree frogs of this clade by its terrestrial limestone karst microhabitat and possession of minimallyexpanded terminal discs of the fingers and toes (vs. widely expanded discs in the arboreal species
|
||
<emphasis id="B9711B305E026B433F9AFBFBFF5EFBE6" italics="true" pageId="8" pageNumber="199">
|
||
<taxonomicName id="4C05BCA15E026B433F9AFBFBFADCFB8B" box="[1273,1403,1052,1077]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="banahao">P. banahao</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E026B433EE4FBFBFF5EFBE6" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="cornutus">P. cornutus</taxonomicName>
|
||
</emphasis>
|
||
), from the forest floor leaf litter-inhabiting, miniature species
|
||
<taxonomicName id="4C05BCA15E026B4338D4FBA7FBE4FBE6" box="[951,1091,1089,1112]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="pygmaeus">
|
||
<emphasis id="B9711B305E026B4338D4FBA7FBE4FBE6" box="[951,1091,1089,1112]" italics="true" pageId="8" pageNumber="199">P. pygmaeus</emphasis>
|
||
</taxonomicName>
|
||
(SVL
|
||
<quantity id="4CFD6AC75E026B433FF3FBD9FAAEFBE9" box="[1168,1289,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="8" pageNumber="199" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 mm</quantity>
|
||
in males) by its much larger body size (
|
||
<quantity id="4CFD6AC75E026B433AB1FB82FDF5FBC2" box="[466,594,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.4" metricValueMin="2.2" pageId="8" pageNumber="199" unit="mm" value="28.0" valueMax="34.0" valueMin="22.0">22–34 mm</quantity>
|
||
) and limestone karst microhabitat, and from the semi-aquatic species
|
||
<taxonomicName id="4C05BCA15E026B433EE4FB83FF5BFB1E" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="levigatus">
|
||
<emphasis id="B9711B305E026B433EE4FB83FF5BFB1E" italics="true" pageId="8" pageNumber="199">P. levigatus</emphasis>
|
||
</taxonomicName>
|
||
by its light brown, mottled dorsal coloration (vs. dark, more uniformly colored dorsum), by its drier, limestone karst microhabitats (vs. moist, riparian environments). Finally,
|
||
<taxonomicName id="4C05BCA15E026B43388FFB4BFBC5FB7A" box="[1004,1122,1197,1220]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E026B43388FFB4BFBC5FB7A" box="[1004,1122,1197,1220]" italics="true" pageId="8" pageNumber="199">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
differs from the limestone karst frogs of the subgenus
|
||
<emphasis id="B9711B305E026B433AB9FB29FDD3FB56" box="[474,628,1231,1256]" italics="true" pageId="8" pageNumber="199">Lahatnanguri</emphasis>
|
||
by its minimally-expanded terminal discs of the fingers and toes (vs. markedly expanded in
|
||
<taxonomicName id="4C05BCA15E026B433AFEFB13FE4BFAB3" box="[413,492,1268,1293]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="biak">
|
||
<emphasis id="B9711B305E026B433AFEFB13FE4BFAB3" box="[413,492,1268,1293]" italics="true" pageId="8" pageNumber="199">P. biak</emphasis>
|
||
</taxonomicName>
|
||
, and widely expanded in
|
||
<taxonomicName id="4C05BCA15E026B433874FB13FC3DFAB3" box="[791,922,1268,1293]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="insulatus">
|
||
<emphasis id="B9711B305E026B433874FB13FC3DFAB3" box="[791,922,1268,1293]" italics="true" pageId="8" pageNumber="199">P. insulatus</emphasis>
|
||
</taxonomicName>
|
||
), the presence of distinct transverse banding patterns of the thigh and tibia (vs. absent or vague in
|
||
<taxonomicName id="4C05BCA15E026B433868FAFFFC28FA8E" box="[779,911,1303,1328]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="insulatus">
|
||
<emphasis id="B9711B305E026B433868FAFFFC28FA8E" box="[779,911,1303,1328]" italics="true" pageId="8" pageNumber="199">P. insulatus</emphasis>
|
||
</taxonomicName>
|
||
), and its simple, single-syllable barking calls advertisement call (vs. atonal clicking pulses in
|
||
<taxonomicName id="4C05BCA15E026B4339D2FADBFC96FAEB" box="[689,817,1340,1365]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="insulatus">
|
||
<emphasis id="B9711B305E026B4339D2FADBFC96FAEB" box="[689,817,1340,1365]" italics="true" pageId="8" pageNumber="199">P. insulatus</emphasis>
|
||
</taxonomicName>
|
||
and the slow series of low, tonal notes in
|
||
<taxonomicName id="4C05BCA15E026B433E65FADBFAF5FAEB" box="[1286,1362,1340,1365]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="199" phylum="Chordata" rank="species" species="biak">
|
||
<emphasis id="B9711B305E026B433E65FADBFAF5FAEB" box="[1286,1362,1340,1365]" italics="true" pageId="8" pageNumber="199">P. biak</emphasis>
|
||
</taxonomicName>
|
||
.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F94A95E026B413BA4FAB9FB19FCAB" lastPageId="10" lastPageNumber="201" pageId="8" pageNumber="199" type="description">
|
||
<paragraph id="8BBAC7225E026B433BA4FAB9FE2EF903" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
<emphasis id="B9711B305E026B433BA4FAB9FE41FAC6" bold="true" box="[199,486,1375,1400]" pageId="8" pageNumber="199">
|
||
Description of
|
||
<typeStatus id="54BE79805E026B433A14FAB9FE47FAC6" box="[375,480,1375,1400]" pageId="8" pageNumber="199" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
A mature male (
|
||
<figureCitation id="133EDBA75E026B4339C0FAB9FD40FAC6" box="[675,743,1375,1400]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="8" pageNumber="199">Fig. 3</figureCitation>
|
||
); habitus moderate; head as wide as body, head length 43.9% snout–vent length; snout protruding moderately beyond lower jaw, snout tip moderately pointed in dorsal and lateral aspects; snout 39.1% head length; eyes moderate and anterolaterally protruding, beyond width of head in dorsal aspect; eye diameter 15.1% snout–vent length, 34.4% head length; lips flared, labial region slightly swollen (
|
||
<figureCitation id="133EDBA75E026B433BFCFA09FF44F9B6" box="[159,227,1519,1544]" captionStart="FIGURE 4" captionStartId="6.[151,244,1685,1706]" captionTargetBox="[307,1275,204,1636]" captionTargetId="figure@6.[289,1297,191,1663]" captionTargetPageId="6" captionText="FIGURE 4. Oblique view illustration of lateral head of (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817) and (B) P. dorsalis (KU 300356). Scale bars = 2 mm; Illustrations by M. Garfield." httpUri="https://zenodo.org/record/239896/files/figure.png" pageId="8" pageNumber="199">Fig. 4</figureCitation>
|
||
A); interorbital region flat; eye diameter 87.9% snout length, eye–naris distance 74.5% eye diameter; pupil horizontally elliptical; canthus rostralis strait, only sightly medially bowed; loreal region moderately concave, sloping ventrolaterally to labia; nostrils oriented laterally; internarial region slightly concave; tympanic annulus distinct, 60.1% eye diameter; supratympanic fold moderately protuberant, swollen and thick, extending from posterior corner of eye across entire dorsal margin of tympanum, along postero-dorsal edge of annulus, and to supra-axillary region.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B433BA4F92EFE11F807" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="199">
|
||
Tongue elongate, wider posteriorly, with shallow posterior notch, a single small anterior lingual papilla, and narrow anterior attachment; choanae situated at anterolateral edge of palate, irregularly ovoid, separated by distance six greater than diameter of single choana, partially concealed in ventral aspect by maxillary shelf; dentigerous process of vomer distinct, with four teeth visible on left process, six on right; dentigerous process angled slightly anterolaterally, approximately at 20–30º incline with closest (posterior) points separated by distance equal to
|
||
<date id="FFBBE1E25E026B433B99F89AFEBAF82A" box="[250,285,1916,1940]" pageId="8" pageNumber="199">2X</date>
|
||
diameter of single choana, their most distant (anterior) ends separated adjacent choana by a distance equivalent to one choana.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E026B423BA4F822FD0EFE4B" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="200" pageId="8" pageNumber="199">
|
||
Hands and feet large; manus length 52.6% pes length; limbs well developed; tibia length 63.4% snout–vent length, 111.9% pes length; fingers slender, long; terminal discs moderately expanded (
|
||
<figureCitation id="133EDBA75E036B423FF5FF71FB7AFF0E" box="[1174,1245,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="9" pageNumber="200">Fig. 2</figureCitation>
|
||
A) with circummarginal grooves evident around distal edge of terminal discs; dorsal surfaces of terminal phalanges with small, cutaneous supra-articular flaps between ultimate and penultimate phalanges; relative lengths of fingers II <I <IV <III; subarticular tubercles prominent, convex, smooth; one subarticular tubercle below Finger I and II, two tubercles under Finger III and IV; supernumerary tubercles present at bases of all digits, distinct, round, prominent for Finger II–IV but elongate and modified into tubercular ridge under Finger I; inner metacarpal tubercle moderately enlarged, irregularly ovoid, with raised nearly spade-like anteromedial edge (
|
||
<figureCitation id="133EDBA75E036B423F93FE89FA91FE36" box="[1264,1334,367,392]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="9" pageNumber="200">Fig. 2</figureCitation>
|
||
A); outer metacarpal tubercle consists of two substructures: an inner (medial) tubercle, slightly smaller and less pronounces than inner metacarpal tubercle; and outter postaxial tubercle, equal in size to a supernumerary tubercle; forearms slender, not hypertrophied; nuptial pads absent.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E036B423BA4FE19FAD1FD73" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="200">
|
||
Skin of dorsal surfaces of head, trunk, and limbs smooth to slightly shagreened; dermal ornamentation minimal, consisting of one enlarged tubercle per eyelid, surrounded by a few much smaller palpebral tubercles, and a cluster of low, swollen post-symphysial tubercles, in a line from the posterior edge of the tympanic annulus to the posterior end of the supratympanic tubercular ridge (
|
||
<figureCitation id="133EDBA75E036B42399EFD8AFCE3FD3A" box="[765,836,620,645]" captionStart="FIGURE 4" captionStartId="6.[151,244,1685,1706]" captionTargetBox="[307,1275,204,1636]" captionTargetId="figure@6.[289,1297,191,1663]" captionTargetPageId="6" captionText="FIGURE 4. Oblique view illustration of lateral head of (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817) and (B) P. dorsalis (KU 300356). Scale bars = 2 mm; Illustrations by M. Garfield." httpUri="https://zenodo.org/record/239896/files/figure.png" pageId="9" pageNumber="200">Fig. 4</figureCitation>
|
||
A); skin of dorsal surfaces of limbs, hands and feet smooth with the exception of a shagreened texture at the knee; skin of chin, throat, and undersurfaces of limbs smooth; skin becoming increasing granular from sternal to inguinal regions; ventral surfaces of thighs granular.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E036B423BA4FD3EFE7EFC1B" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="200">
|
||
Toes long, narrow (
|
||
<figureCitation id="133EDBA75E036B423AD2FD31FE5CFD4E" box="[433,507,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="9" pageNumber="200">Fig. 2</figureCitation>
|
||
B); terminal discs of toes moderately expanded, with circum-marginal grooves evident along distal margin of terminal pad; supra-articular cutaneous flaps present above ultimate-penultimate phalangeal articulations; plantar surface of foot smooth, with well-developed, prominently protuberant subarticular tubercles (
|
||
<figureCitation id="133EDBA75E036B423A6FFCA2FEF5FCE2" box="[268,338,836,861]" captionStart="FIGURE 2" captionStartId="4.[151,244,1449,1470]" captionTargetBox="[217,1369,411,1418]" captionTargetId="figure@4.[217,1369,411,1425]" captionTargetPageId="4" captionText="FIGURE 2. Ventral views of hand (A) and foot (B) of the adult male holotype of Platymantis (Lupacolus) quezoni, sp. nov. (PNM 9817), and adult male specimens of (C, D) Platymantis dorsalis (KU 300356), (E, F) P. p a e n g i (KU 300206), and (G, H) P. spelaeus (KU 300438) for comparison. Scale bars = 2 mm." httpUri="https://zenodo.org/record/239894/files/figure.png" pageId="9" pageNumber="200">Fig. 2</figureCitation>
|
||
B); supernumerary tubercles absent; relative lengths of toes I <II <V <III <IV; outer metatarsal tubercle round, small, but moderately prominent; inner metatarsal tubercle elongate, prominent, length 2´diameter of outter metatarsal tubercle.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E036B423BA4FC49FDF3FBC2" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="200">
|
||
<emphasis id="B9711B305E036B423BA4FC49FDC8FC76" bold="true" box="[199,623,943,968]" pageId="9" pageNumber="200">
|
||
Measurements of
|
||
<typeStatus id="54BE79805E036B423AF8FC49FDA5FC76" box="[411,514,943,968]" pageId="9" pageNumber="200" type="holotype">holotype</typeStatus>
|
||
(in mm).
|
||
</emphasis>
|
||
Snout–vent length 33.7; head length 14.8; head width 13.9; snout length 5.8; upper arm length 5.9; forearm length 7.5; femur length 18.1; tibia length 21.7; tarsus length 10.9; pes length 19.4; manus length 10.2; Finger I length 5.9; Finger III length 7.7; Toe IV length 12.6; eye–narial distance 3.8; interorbital distance 3.2; internarial distance 3.0; eye diameter 5.1; horizontal tympanic annulus diameter 3.1; Finger I disc width 1.0; Finger III disc width 1.2; Toe IV disc width 1.3; Finger III penultimate phalange width 0.8; Toe IV penultimate phalange width 0.7.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E036B423BA4FB61FE62F95E" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="200">
|
||
<emphasis id="B9711B305E036B423BA4FB61FD05FB1E" bold="true" box="[199,674,1159,1184]" pageId="9" pageNumber="200">
|
||
Coloration of
|
||
<typeStatus id="54BE79805E036B423A17FB61FE7CFB1E" box="[372,475,1159,1184]" pageId="9" pageNumber="200" type="holotype">holotype</typeStatus>
|
||
in preservative.
|
||
</emphasis>
|
||
Dorsal surfaces of head, body, and limbs dark brown with black blotches; anterior portions of head darker greenish brown; limbs banded with black bars; interorbital bar present but faint: a thin, irregular, black line; lateral surfaces of head darker than dorsum, nearly greenish brown with black markings; black bar extends across dorsal half of canthus rostralis to anterior edge of eye, around margin of the palpebrum, and across suptratympanic fold; tympanum medium brownish-green with black covering dorsal-most half; lips with alternating dark green and black vertical labial bars, bordered anteriorly by black lower edge of canthal and subocular regions; three bold black bars radiate out of orbit dorsally, and extend ventrally to labial region to form the most conspicuous labial bars; transverse limb bars of irregular widths, with an overall darker coloration (compared to dorsum), due to a greater proportion of surface covered with black markings; upper arm almost devoid of black blotches, forearms each with two transverse black bars, thighs with five, tibia with three, tarsus with two; anterior flank coloration similar to dorsum on left, but predominantly black on right due to aggregation of black blotches, both sides progressively fading to less strongly patterned ventral color: dark brown blotches on throat, sternal region tan, inguinal region tan with steel bluish markings; ventral surfaces of limbs solid dark reddish brown; ventral surfaces of hands and feed dark reddish brown with light cream to tan subarticular tubercles; dorsal surfaces of hands are light brown, with alternating bands black bars on Fingers II–IV, and light tan on Finger I; dorsal surfaces of feet with solid redish brown above Toes I–III, boldly alternating light brown and black on Fingers IV and V.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E036B413BA4F90AFD37FEFE" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,1977]" lastPageId="10" lastPageNumber="201" pageId="9" pageNumber="200">
|
||
<emphasis id="B9711B305E036B423BA4F90AFE04F8BB" bold="true" box="[199,419,1772,1797]" pageId="9" pageNumber="200">Coloration in life.</emphasis>
|
||
Dorsal surfaces of head, body, and limbs variably mottled shades of dark brown and greenish-gray, a few scattered bright gold spots, and with bold black markings scattered throughout but congregating irregularly into the following characteristics: distinct black suprascapular bar, interorbital bar, transverse limb bars (on thigh, tibia, shank, and upper arm and forearm), a dark bar from the nares, along the canthus, aligned with the enlarged pupil at night (
|
||
<figureCitation id="133EDBA75E036B42398EF89AFC91F82A" box="[749,822,1916,1941]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="9" pageNumber="200">Fig. 3</figureCitation>
|
||
), and posteriorly across the dorsal-most half of the tympanum; upper lips with distinct black labial bars; pupil bordered by bright yellow ciliary ring, iris silver; flanks with enlarged black blotches and interrupted by reduced dark brown of dorsal surfaces, grading ventrally across flank to cream ground color with black blotches on anterior portion of flanks; dorsal surfaces of digits mostly black, with small brown spots. Ventrum very dark gray with black blotches on throat and sternum; liver nearly bluish gray through translucent ventral skin of chest and groin; ventral surfaces of legs nearly solid black, fading to dark purplish gray on ventral tibia and shank; ventral surfaces of hands and feet dark purplish gray with lighter gray subarticular tubercles and distal digital discs.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E006B413BA4FEAAFD11FD16" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
<emphasis id="B9711B305E006B413BA4FEAAFE9CFEDB" bold="true" box="[199,315,332,357]" pageId="10" pageNumber="201">Variation.</emphasis>
|
||
Overall color pattern consistently mottled across the
|
||
<typeStatus id="54BE79805E006B4138B7FEABFBA3FEDB" box="[980,1028,333,357]" pageId="10" pageNumber="201">type</typeStatus>
|
||
series, typical of limestone karst inhabiting
|
||
<taxonomicName id="4C05BCA15E006B413A71FE89FE3FFE36" box="[274,408,367,392]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9711B305E006B413A71FE89FE3FFE36" box="[274,408,367,392]" italics="true" pageId="10" pageNumber="201">Platymantis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF94BAD35E006B413AC9FE96FD10FE36" author="Brown" box="[426,695,367,392]" pageId="10" pageNumber="201" refString="Brown, W. C. & Alcala, A. C. (1970) A new species of the genus Platymantis (Ranidae) with a list of the amphibians known from South Gigante Island, Philippines. Occasional Papers of the California Academy of Sciences, 84, 1 - 7." type="journal article" year="1970">Brown & Alcala, 1970</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF94BAD35E006B4139A6FE96FCAAFE36" author="Alcala" box="[709,781,368,392]" pageId="10" pageNumber="201" refString="Brown, W. C. & Alcala, A. C. (1982 a) A new cave Platymantis (Amphibia: Ranidae) from the Philippine Islands. Proceedings of the Biological Society of Washington, 95, 386 - 391." type="journal article" year="1982" yearSuffix="a">1982a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E006B413878FE89FC46FE36" author="Siler" box="[795,993,367,392]" pageId="10" pageNumber="201" refString="Siler, C. D., Alcala, A. C., Diesmos, A. C. & Brown, R. M. (2009) A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from eastern Samar Island, Philippines. Herpetologica, 65, 92 - 104. http: // dx. doi. org / 10.1655 / 08 - 040 R. 1" type="journal article" year="2009">
|
||
Siler
|
||
<emphasis id="B9711B305E006B413839FE97FC29FE36" box="[858,910,367,392]" italics="true" pageId="10" pageNumber="201">et al</emphasis>
|
||
., 2009
|
||
</bibRefCitation>
|
||
, 2010), with dorsal ground color light tannish yellow or mustard (e.g., KU 328735;
|
||
<figureCitation id="133EDBA75E006B4139F0FE72FD72FE12" box="[659,725,404,429]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="10" pageNumber="201">Fig. 3</figureCitation>
|
||
) to dark greenish-gray with black blotches (the
|
||
<typeStatus id="54BE79805E006B413F8CFE72FAF1FE13" box="[1263,1366,404,429]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F31287B25E006B413E03FE73FA3AFE12" box="[1376,1437,405,428]" name="Panama" pageId="10" pageNumber="201">PNM</collectingCountry>
|
||
9817;
|
||
<figureCitation id="133EDBA75E006B413B81FE51FE8EFE6E" box="[226,297,439,464]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="10" pageNumber="201">Fig. 3</figureCitation>
|
||
). The degree and concentration of dorsal black markings ranges, from specimens nearly covered dorsally in black markings (KU 339543, TNHC 61993), to individuals predominantly light gray to tan, with few black markings (KU 61989, 61991, 61992;
|
||
<figureCitation id="133EDBA75E006B41391EFE19FD1AFDA6" box="[637,701,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,244,1547,1568]" captionTargetBox="[249,1334,199,1486]" captionTargetId="figure@5.[241,1345,191,1525]" captionTargetPageId="5" captionText="FIGURE 3. Photographs of the (A) Platymantis (Lupacolus) quezoni, sp. nov., holotype (PNM 9817; SVL = 33.7) and (B) a male paratype (KU 328735; SVL = 31.3). Photos: RMB." httpUri="https://zenodo.org/record/239895/files/figure.png" pageId="10" pageNumber="201">Fig. 3</figureCitation>
|
||
B). The suprascapular bar ranges from a distinct w-shaped pattern (the
|
||
<typeStatus id="54BE79805E006B413BA9FDC2FE97FD83" box="[202,304,548,573]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F31287B25E006B413A58FDC3FEDFFD82" box="[315,376,549,572]" name="Panama" pageId="10" pageNumber="201">PNM</collectingCountry>
|
||
9817; TNHC 61990; KU 328736) to a few irregular, disconnected blotches (TNHC 61991) to an inverted v-shaped formation (KU 339543). Ventral surfaces less variable, predominantly as described for the
|
||
<typeStatus id="54BE79805E006B413BF4FD8AFF5BFD3B" box="[151,252,620,645]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
|
||
, but a few
|
||
<typeStatus id="54BE79805E006B413AEFFD8BFE50FD3B" box="[396,503,621,645]" pageId="10" pageNumber="201" type="paratype">paratypes</typeStatus>
|
||
(KU 328735, 328736, TNHC 61990, 61994) possess darker throats, owing to aggregation of black blotches as in the
|
||
<typeStatus id="54BE79805E006B41392EFD69FD14FD16" box="[589,691,655,680]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E006B413BA4FD52FB19FCAB" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
Summaries of univariate morphological variation in the series are presented in
|
||
<tableCitation id="C687F2995E006B413F22FD52FB3FFD72" box="[1089,1176,692,717]" captionStart="TABLE 2" captionStartId="11.[151,231,151,172]" captionTargetBox="[165,1267,231,2011]" captionTargetPageId="11" captionText="TABLE 2. — Summary of univariate morphological variation (Range, above; mean ± 1 S. D., below) among mensural characters in the type series of Platymantis (Lupacolus) quezoni, sp. nov." httpUri="http://table.plazi.org/id/DF7A97AA5E016B403BF4FF71FD15FF74" pageId="10" pageNumber="201" tableUuid="DF7A97AA5E016B403BF4FF71FD15FF74">Table 2</tableCitation>
|
||
. Females (
|
||
<collectingCountry id="F31287B25E006B413E73FD52FAE8FD72" box="[1296,1359,692,716]" name="Panama" pageId="10" pageNumber="201">PNM</collectingCountry>
|
||
9907– 0 9, 9914) are considerably larger and more robust (
|
||
<tableCitation id="C687F2995E006B4139BEFD31FC92FD4E" box="[733,821,727,752]" captionStart="TABLE 2" captionStartId="11.[151,231,151,172]" captionTargetBox="[165,1267,231,2011]" captionTargetPageId="11" captionText="TABLE 2. — Summary of univariate morphological variation (Range, above; mean ± 1 S. D., below) among mensural characters in the type series of Platymantis (Lupacolus) quezoni, sp. nov." httpUri="http://table.plazi.org/id/DF7A97AA5E016B403BF4FF71FD15FF74" pageId="10" pageNumber="201" tableUuid="DF7A97AA5E016B403BF4FF71FD15FF74">Table 2</tableCitation>
|
||
), and the single juvenile (KU 339544; SVL
|
||
<quantity id="4CFD6AC75E006B413E4DFD3EFA33FD4E" box="[1326,1428,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="10" pageNumber="201" unit="mm" value="24.3">24.3 mm</quantity>
|
||
) was more brightly patterned in life, with bolder and contrasting light tan versus black blotches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F94A95E006B463BA4FCF9FA31FBD4" lastPageId="13" lastPageNumber="204" pageId="10" pageNumber="201" type="distribution">
|
||
<paragraph id="8BBAC7225E006B413BA4FCF9FC15FC3E" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
<emphasis id="B9711B305E006B413BA4FCF9FEC7FC86" bold="true" box="[199,352,799,824]" pageId="10" pageNumber="201">Distribution.</emphasis>
|
||
<taxonomicName id="4C05BCA15E006B413A08FCF9FDF4FC86" box="[363,595,799,824]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E006B413A08FCF9FDF4FC86" box="[363,595,799,824]" italics="true" pageId="10" pageNumber="201">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54BE79805E006B4138E3FCC6FC17FC86" box="[896,944,800,824]" pageId="10" pageNumber="201">type</typeStatus>
|
||
locality but may be more wide spread in scattered karst outcrops of northern portions of the Bicol Peninsula (
|
||
<bibRefCitation id="EF94BAD35E006B4138FBFCA2FB2EFCE3" author="Restificar" box="[920,1161,836,861]" pageId="10" pageNumber="201" refString="Restificar, S. D. F., Day, M. J. & Urich, P. B. (2006) Protection of karst in the Philippines. Acta Carsologica, 35, 121 - 130. http: // dx. doi. org / 10.3986 / ac. v 35 i 1.248" type="journal article" year="2006">
|
||
Restificar
|
||
<emphasis id="B9711B305E006B413F6DFCA3FBE1FCE3" box="[1038,1094,836,861]" italics="true" pageId="10" pageNumber="201">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E006B413FF6FCA2FABFFCE2" author="MGB" box="[1173,1304,836,860]" pageId="10" pageNumber="201" refString="Mines and Geosciences Bureau (MGB) (2010) Geology of the Philippines. Mines and Geosciences Bureau, & Department of the Environment and Natural Resources, Quezon City, 311 pp." type="book" year="2010">MGB 2010</bibRefCitation>
|
||
), especially on limestone karst formations in the nearby Bondoc Peninsula (
|
||
<figureCitation id="133EDBA75E006B41383CFC81FC07FC3E" box="[863,928,871,896]" captionStart="FIGURE 1" captionStartId="1.[151,244,1917,1938]" captionTargetBox="[244,1343,453,1886]" captionTargetId="figure@1.[231,1356,443,1896]" captionTargetPageId="1" captionText="FIGURE 1. Known occurrence of Platymantis (Lupacolus) quezoni, sp. nov., in the Quezon Protected Landscape, southern Luzon Island, Philippines (black dot). The inset shows the location of Luzon Island (darkly shaded) within the Philippines. Note the adjacent position of the Bondoc Peninsula (a geological formation characterized by extensive limestone karst formations; MGB 2010), where we suspect additional P. q u e z o n i populations may occur (see text)." httpUri="https://zenodo.org/record/239893/files/figure.png" pageId="10" pageNumber="201">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E006B413BA4FC6AFADEFBC2" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
<emphasis id="B9711B305E006B413BA4FC6AFDB9FC1B" bold="true" box="[199,542,908,933]" pageId="10" pageNumber="201">Ecology and natural history.</emphasis>
|
||
<taxonomicName id="4C05BCA15E006B413945FC6AFCACFC1A" box="[550,779,908,933]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E006B413945FC6AFCACFC1A" box="[550,779,908,933]" italics="true" pageId="10" pageNumber="201">Platymantis quezoni</emphasis>
|
||
</taxonomicName>
|
||
occurs in mature secondary-growth and mixed secondary growth combined with original forest, on the limestone karst substrates of Quezon Protected Landscape and most likely in other surrounding areas (
|
||
<figureCitation id="133EDBA75E006B413979FC32FDFAFC52" box="[538,605,980,1005]" captionStart="FIGURE 1" captionStartId="1.[151,244,1917,1938]" captionTargetBox="[244,1343,453,1886]" captionTargetId="figure@1.[231,1356,443,1896]" captionTargetPageId="1" captionText="FIGURE 1. Known occurrence of Platymantis (Lupacolus) quezoni, sp. nov., in the Quezon Protected Landscape, southern Luzon Island, Philippines (black dot). The inset shows the location of Luzon Island (darkly shaded) within the Philippines. Note the adjacent position of the Bondoc Peninsula (a geological formation characterized by extensive limestone karst formations; MGB 2010), where we suspect additional P. q u e z o n i populations may occur (see text)." httpUri="https://zenodo.org/record/239893/files/figure.png" pageId="10" pageNumber="201">Fig. 1</figureCitation>
|
||
). Both males and females were frequently encountered climbing vertical limestone rock faces, on narrow shelves of limestone at the base of karst cliffs, perched on top of large limestone boulders, in cracks and crevices, and around the entrances to small caves (
|
||
<figureCitation id="133EDBA75E006B413880FBFAFB8FFB8B" box="[995,1064,1052,1077]" captionStart="FIGURE 5" captionStartId="7.[151,244,1518,1539]" captionTargetBox="[250,1335,449,1488]" captionTargetId="figure@7.[243,1344,443,1496]" captionTargetPageId="7" captionText="FIGURE 5. Limestone karst forest habitat of Platymantis (Lupacolus) quezoni, sp. nov., at the type locality: (A) limestone bolder where the holotype (PNM 9817) was recorded and collected; (B) appearance of habitat in late afternoon (~ 16: 30). Note darkened caves and crevices beneath limestone over-hangs from where P. q u e z o n i initiate vocalizations in the late afternoons (16: 30 – 17: 30 hr) from concealed perches inside crevices and small caves. Photos: RMB." httpUri="https://zenodo.org/record/239897/files/figure.png" pageId="10" pageNumber="201">Fig. 5</figureCitation>
|
||
B). No specimens were observed climbing on vegetation growing from limestone, in contrast to the common scansorial habits of species of the other members of the subgenus
|
||
<emphasis id="B9711B305E006B413AD8FB82FD88FBC3" box="[443,559,1124,1149]" italics="true" pageId="10" pageNumber="201">Lupacolus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF94BAD35E006B41395DFB82FCBEFBC3" author="Brown" box="[574,793,1124,1149]" pageId="10" pageNumber="201" refString="Brown, W. C., Brown, R. M. & Alcala, A. C. (1997 a) Species of the hazelae group of Platymantis from the Philippines, with descriptions of two new species. Proceedings of the California Academy of Sciences, 49, 405 - 421." type="journal article" year="1997" yearSuffix="a">
|
||
Brown
|
||
<emphasis id="B9711B305E006B4139F0FB83FD6BFBC3" box="[659,716,1124,1149]" italics="true" pageId="10" pageNumber="201">et al.</emphasis>
|
||
1997
|
||
<emphasis id="B9711B305E006B413869FB82FCBEFBC3" box="[778,793,1124,1149]" italics="true" pageId="10" pageNumber="201">a</emphasis>
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF94BAD35E006B41387DFB82FC8AFBC3" author="Brown" box="[798,813,1124,1149]" pageId="10" pageNumber="201" refString="Brown, W. C., Alcala, A. C. & Diesmos, A. C. (1997 b) A new species of the genus Platymantis (Amphibia: Ranidae) from Luzon Island, Philippines. Proceedings of the Biological Society of Washington, 110, 18 - 23." type="journal article" year="1997" yearSuffix="b">b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E006B41385FFB82FB99FBC2" author="Alcala" box="[828,1086,1124,1149]" pageId="10" pageNumber="201" refString="Brown W. C., Alcala, A. C. & Diesmos A. C. (1999) Four new species of the genus Platymantis (Amphibia: Ranidae) from Luzon Island, Philippines. Proceedings of the California Academy of Sciences, 51, 449 - 460." type="journal article" year="1999">Alcala & Brown, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF94BAD35E006B413F29FB83FACAFBC2" author="Brown" box="[1098,1389,1124,1149]" pageId="10" pageNumber="201" refString="Brown, R. M & Gonzales, J. C. (2007) A new forest frog of the genus Platymantis (Amphibia; Anura; Ranidae) from the Bicol Peninsula of Luzon Island, Philippines. Copeia, 2007, 251 - 266. http: // dx. doi. org / 10.1643 / 0045 - 8511 (2007) 7 [251: ANFFOT] 2.0. CO; 2" type="journal article" year="2007">Brown & Gonzales, 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E006B413BA4FB61FC74FA22" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
Sympatric but non-syntopic anuran species include
|
||
<taxonomicName id="4C05BCA15E006B41386FFB61FC64FB1E" box="[780,963,1159,1184]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="marina">
|
||
<emphasis id="B9711B305E006B41386FFB61FC64FB1E" box="[780,963,1159,1184]" italics="true" pageId="10" pageNumber="201">Rhinella marina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B4138B1FB61FBECFB1E" box="[978,1099,1159,1184]" class="Amphibia" family="Bufonidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Bufonidae</taxonomicName>
|
||
, introduced),
|
||
<taxonomicName id="4C05BCA15E006B413F86FB61FEA2FB7B" class="Amphibia" family="Dicroglossidae" genus="Hoplobatrachus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="rugulosus">
|
||
<emphasis id="B9711B305E006B413F86FB61FEA2FB7B" italics="true" pageId="10" pageNumber="201">Hoplobatrachus rugulosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B413A74FB4AFE63FB7B" box="[279,452,1196,1221]" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Dicroglossidae</taxonomicName>
|
||
, introduced),
|
||
<taxonomicName id="4C05BCA15E006B413900FB4AFCB9FB7B" box="[611,798,1196,1221]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis id="B9711B305E006B413900FB4AFCB9FB7B" box="[611,798,1196,1221]" italics="true" pageId="10" pageNumber="201">Kaloula pulchra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B413853FB4AFC6CFB7B" box="[816,971,1196,1221]" class="Amphibia" family="Microhylidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Microhylidae</taxonomicName>
|
||
, introduced),
|
||
<emphasis id="B9711B305E006B413F08FB4BFB69FB7A" box="[1131,1230,1197,1220]" italics="true" pageId="10" pageNumber="201">
|
||
<taxonomicName id="4C05BCA15E006B413F08FB4BFB6DFB7A" box="[1131,1226,1197,1220]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="picta">K. picta</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05BCA15E006B413E69FB4BFA3CFB7A" box="[1290,1435,1197,1220]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="conjuncta">
|
||
<emphasis id="B9711B305E006B413E69FB4BFA3CFB7A" box="[1290,1435,1197,1220]" italics="true" pageId="10" pageNumber="201">K. conjuncta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B413BFDFB29FE9DFB56" box="[158,314,1231,1256]" class="Amphibia" family="Microhylidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Microhylidae</taxonomicName>
|
||
, native & endemic to the
|
||
<collectingCountry id="F31287B25E006B413912FB29FD53FB56" box="[625,756,1231,1256]" name="Philippines" pageId="10" pageNumber="201">Philippines</collectingCountry>
|
||
),
|
||
<emphasis id="B9711B305E006B413869FB37FF7FFAB3" italics="true" pageId="10" pageNumber="201">
|
||
<taxonomicName id="4C05BCA15E006B413869FB37FBF7FB56" box="[778,1104,1231,1256]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="macrocephalus">Limnonectes macrocephalus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E006B413F3CFB37FAAEFB56" box="[1119,1289,1231,1256]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="woodworthi">L. woodworthi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E006B413E7BFB29FF7FFAB3" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="laevis">Occidozyga laevis</taxonomicName>
|
||
</emphasis>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B413B88FB12FE3FFAB3" box="[235,408,1268,1293]" class="Amphibia" family="Dicroglossidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Dicroglossidae</taxonomicName>
|
||
, native & endemic),
|
||
<taxonomicName id="4C05BCA15E006B4139F7FB13FC30FAB3" box="[660,919,1268,1293]" class="Amphibia" family="Ranidae" genus="Sanguirana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="luzonensis">
|
||
<emphasis id="B9711B305E006B4139F7FB13FC30FAB3" box="[660,919,1268,1293]" italics="true" pageId="10" pageNumber="201">Sanguirana luzonensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B4138C9FB12FBADFAB3" box="[938,1034,1268,1293]" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Ranidae</taxonomicName>
|
||
, native & endemic),
|
||
<taxonomicName id="4C05BCA15E006B413E65FB12FF54FA8E" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="201" phylum="Arthropoda" rank="species" species="pardalis">
|
||
<emphasis id="B9711B305E006B413E65FB12FF54FA8E" italics="true" pageId="10" pageNumber="201">Rhacophorus pardalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C05BCA15E006B413A52FAF1FDE8FA8E" box="[305,591,1303,1328]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="leucomystax">
|
||
<emphasis id="B9711B305E006B413A52FAF1FDE8FA8E" box="[305,591,1303,1328]" italics="true" pageId="10" pageNumber="201">Polypedates leucomystax</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C05BCA15E006B413901FAF1FCB7FA8E" box="[610,784,1303,1328]" class="Amphibia" family="Rhacophoridae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Rhacophoridae</taxonomicName>
|
||
, native, non-endemic). Syntopic native endemic species (collected in the immediate vicinity or adjacent microhabitats) include
|
||
<taxonomicName id="4C05BCA15E006B4138A8FADBFB70FAEB" box="[971,1239,1340,1365]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="dorsalis" subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E006B4138A8FADBFC7AFAEA" box="[971,989,1341,1364]" italics="true" pageId="10" pageNumber="201">P.</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E006B413891FADAFBC1FAEB" box="[1010,1126,1340,1365]" italics="true" pageId="10" pageNumber="201">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E006B413F19FADAFB70FAEB" box="[1146,1239,1340,1365]" italics="true" pageId="10" pageNumber="201">dorsalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E006B413F86FADBFEB2FAC6" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="corrugatus" subGenus="Platymantis">
|
||
P. (
|
||
<emphasis id="B9711B305E006B413E6EFADAFA34FAEB" box="[1293,1427,1340,1365]" italics="true" pageId="10" pageNumber="201">Platymantis</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E006B413BF4FA87FEB2FAC6" box="[151,277,1377,1400]" italics="true" pageId="10" pageNumber="201">corrugatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E006B413A52FA87FD3DFAC6" box="[305,666,1375,1400]" class="Amphibia" family="Ranidae" genus="Sanguirana" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="luzonensis" subGenus="Tahananpuno">
|
||
P. (
|
||
<emphasis id="B9711B305E006B413A04FAB9FDA7FAC6" box="[359,512,1375,1400]" italics="true" pageId="10" pageNumber="201">Tahananpuno</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E006B413941FAB9FD3DFAC6" box="[546,666,1375,1400]" italics="true" pageId="10" pageNumber="201">luzonensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C05BCA15E006B4139D5FAB9FC70FAC6" authority="Lupalolus" authorityName="Lupalolus" box="[694,983,1375,1400]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="genus">
|
||
Platymantis (
|
||
<emphasis id="B9711B305E006B41383DFAB9FC69FAC6" box="[862,974,1375,1400]" italics="true" pageId="10" pageNumber="201">Lupalolus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
n. sp. 14 (Brown
|
||
<emphasis id="B9711B305E006B413F9BFA87FA9DFAC6" box="[1272,1338,1375,1400]" italics="true" pageId="10" pageNumber="201">et al</emphasis>
|
||
. 2015) (
|
||
<taxonomicName id="4C05BCA15E006B413BFDFA62FEC9FA23" box="[158,366,1412,1437]" class="Amphibia" family="Ceratobatrachidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="family">Ceratobatrachidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C05BCA15E006B413AE0FA62FDBAFA22" box="[387,541,1412,1437]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="picta">
|
||
<emphasis id="B9711B305E006B413AE0FA62FDBAFA22" box="[387,541,1412,1437]" italics="true" pageId="10" pageNumber="201">Kaloula picta</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C05BCA15E006B413939FA63FC6BFA23" box="[602,972,1412,1437]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="kalingensis">
|
||
<emphasis id="B9711B305E006B413939FA63FDD3FA22" box="[602,628,1413,1436]" italics="true" pageId="10" pageNumber="201">K.</emphasis>
|
||
cf.
|
||
<emphasis id="B9711B305E006B4139FDFA62FCBBFA23" box="[670,796,1412,1437]" italics="true" pageId="10" pageNumber="201">kalingensis</emphasis>
|
||
(Microhylidae)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E006B413BA4FA41FB33F807" blockId="10.[151,1437,151,1977]" pageId="10" pageNumber="201">
|
||
<emphasis id="B9711B305E006B413BA4FA41FE08FA01" bold="true" box="[199,431,1447,1472]" pageId="10" pageNumber="201">Advertisement call.</emphasis>
|
||
At the
|
||
<typeStatus id="54BE79805E006B413967FA4FFD93FA7F" box="[516,564,1449,1473]" pageId="10" pageNumber="201">type</typeStatus>
|
||
locality, when visited in dry-season months (December and January), calling activity was heard during and following rainfall; in the rainy months of July through September, the new species called continuously in the afternoon and early evenings. Advertisement calls were recorded on two visits to the
|
||
<typeStatus id="54BE79805E006B413BF4F9F3FF60F993" box="[151,199,1557,1581]" pageId="10" pageNumber="201">type</typeStatus>
|
||
locality, at ambient temperatures of 22.5–22.6°C on the first occasion (
|
||
<date id="FFBBE1E25E006B413881F9F2FB4EF992" box="[994,1257,1556,1581]" pageId="10" pageNumber="201" value="2001-11-27" valueMax="2001-11-28" valueMin="2001-11-27">27–28 November 2001</date>
|
||
), and 23.1°C on our second visit. (
|
||
<date id="FFBBE1E25E006B413A03F9DEFDB7F9EE" box="[352,528,1592,1616]" pageId="10" pageNumber="201" value="2011-01-09">9 January 2011</date>
|
||
). The following description is based on vouchered calls of seven collected males (
|
||
<collectingCountry id="F31287B25E006B413BFDF9BAFF7AF9CA" box="[158,221,1628,1652]" name="Panama" pageId="10" pageNumber="201">PNM</collectingCountry>
|
||
9817,
|
||
<typeStatus id="54BE79805E006B413A4FF9BAFE35F9CB" box="[300,402,1628,1653]" pageId="10" pageNumber="201" type="holotype">holotype</typeStatus>
|
||
; TNHC 61989–92, KU 328735, 328736,
|
||
<typeStatus id="54BE79805E006B413809F9BAFC7DF9CA" box="[874,986,1628,1652]" pageId="10" pageNumber="201" type="paratype">paratypes</typeStatus>
|
||
). Neither ambient noises of wind in the canopy of mature forest in the area, nor advertisement calls of congeners (
|
||
<taxonomicName id="4C05BCA15E006B413F6CF967FB2EF926" box="[1039,1161,1663,1688]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E006B413F6CF967FB2EF926" box="[1039,1161,1663,1688]" italics="true" pageId="10" pageNumber="201">P. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
calls frequently at this locality) disrupted the activity of calling male
|
||
<taxonomicName id="4C05BCA15E006B4139F8F943FCAAF902" box="[667,781,1701,1724]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="201" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E006B4139F8F943FCAAF902" box="[667,781,1701,1724]" italics="true" pageId="10" pageNumber="201">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
. Calling activity begins in the late afternoon (16:30–17:30 hr) and typically lasts 3–5 hr. In the late afternoon, frogs call intermittently from relatively deep inside limestone crevices, or from within small caves. As the forest becomes darker, additional males begin calling and eventually can be seen on the surface of limestone, or calling from just inside crevices. As more individuals move to exposed perches, calling around limestone aggregations reaches maximal intensity. Periods of intense calling energy are separated by 30–120 s of silence, before males call again. Because all our recordings were made at the same locality, at very similar temperatures, and in almost identical atmospheric conditions (immediately following a heavy late afternoon rain), no temperature corrections were undertaken for this description.
|
||
</paragraph>
|
||
<caption id="DF7A97AA5E016B403BF4FF71FD15FF74" ID-Table-UUID="DF7A97AA5E016B403BF4FF71FD15FF74" httpUri="http://table.plazi.org/id/DF7A97AA5E016B403BF4FF71FD15FF74" pageId="11" pageNumber="202" targetBox="[165,1267,231,2011]" targetIsTable="true" targetPageId="11">
|
||
<paragraph id="8BBAC7225E016B403BF4FF71FD15FF74" blockId="11.[151,1435,151,202]" pageId="11" pageNumber="202">
|
||
<emphasis id="B9711B305E016B403BF4FF71FF5FFF12" bold="true" box="[151,248,151,172]" pageId="11" pageNumber="202">TABLE 2</emphasis>
|
||
.—Summary of univariate morphological variation (Range, above; mean ± 1 S.D., below) among mensural characters in the type series of
|
||
<taxonomicName id="4C05BCA15E016B403A7FFF53FDFBFF74" box="[284,604,181,202]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="202" phylum="Chordata" rank="species" species="quezoni" status="sp. nov." subGenus="Lupacolus">
|
||
<emphasis id="B9711B305E016B403A7FFF53FE28FF74" box="[284,399,181,202]" italics="true" pageId="11" pageNumber="202">Platymantis</emphasis>
|
||
(
|
||
<emphasis id="B9711B305E016B403AFEFF53FDA6FF74" box="[413,513,181,202]" italics="true" pageId="11" pageNumber="202">Lupacolus</emphasis>
|
||
)
|
||
<emphasis id="B9711B305E016B40396CFF53FDFBFF74" box="[527,604,181,202]" italics="true" pageId="11" pageNumber="202">quezoni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A242A64B5E016B403906FF53FD15FF74" box="[613,690,181,202]" pageId="11" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBAC7225E016B403839FF01FB40F865" pageId="11" pageNumber="202">
|
||
<table id="F90535825E0194B43BC6FF01FB54F865" box="[165,1267,231,2011]" gridcols="3" gridrows="26" pageId="11" pageNumber="202">
|
||
<tr id="3535C5605E0194B43BC6FF01FB54FF42" box="[165,1267,231,252]" gridrow="0" pageId="11" pageNumber="202" rowspan-0="1">
|
||
<th id="76E4AC1C5E0194B43839FF01FC6BFF42" box="[858,972,231,252]" gridcol="1" gridrow="0" pageId="11" pageNumber="202">Female</th>
|
||
<th id="76E4AC1C5E0194B43FE1FF01FB54FF42" box="[1154,1267,231,252]" gridcol="2" gridrow="0" pageId="11" pageNumber="202">Male</th>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FEEAFB54FE9F" box="[165,1267,268,289]" gridrow="1" pageId="11" pageNumber="202" rowspan-0="1">
|
||
<td id="76E4AC1C5E0194B43839FEEAFC6BFE9F" box="[858,972,268,289]" gridcol="1" gridrow="1" pageId="11" pageNumber="202">
|
||
<emphasis id="B9711B305E016B403839FEEAFCC1FE9F" box="[858,870,268,289]" italics="true" pageId="11" pageNumber="202">n</emphasis>
|
||
= 4
|
||
</td>
|
||
<td id="76E4AC1C5E0194B43FE1FEEAFB54FE9F" box="[1154,1267,268,289]" gridcol="2" gridrow="1" pageId="11" pageNumber="202">
|
||
<emphasis id="B9711B305E016B403FE1FEEAFB29FE9F" box="[1154,1166,268,289]" italics="true" pageId="11" pageNumber="202">n</emphasis>
|
||
= 16
|
||
</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FED4FB54FED7" box="[165,1267,306,361]" gridrow="2" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FED4FE1BFED7" box="[165,444,306,361]" gridcol="0" gridrow="2" pageId="11" pageNumber="202">Snout–Vent Length</th>
|
||
<td id="76E4AC1C5E0194B43839FED4FC6BFED7" box="[858,972,306,361]" gridcol="1" gridrow="2" pageId="11" pageNumber="202">32.4–39.7 (37.8 ± 2.7)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FED4FB54FED7" box="[1154,1267,306,361]" gridcol="2" gridrow="2" pageId="11" pageNumber="202">22.1–33.9 (29.9 ± 4.9)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FE9CFB54FE0F" box="[165,1267,378,433]" gridrow="3" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FE9CFE1BFE0F" box="[165,444,378,433]" gridcol="0" gridrow="3" pageId="11" pageNumber="202">Head Length</th>
|
||
<td id="76E4AC1C5E0194B43839FE9CFC6BFE0F" box="[858,972,378,433]" gridcol="1" gridrow="3" pageId="11" pageNumber="202">14.3–14.3 (15.8 ± 1.0)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FE9CFB54FE0F" box="[1154,1267,378,433]" gridcol="2" gridrow="3" pageId="11" pageNumber="202">9.4–15.1 (12.4 ± 2.6)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FE27FB54FE47" box="[165,1267,449,505]" gridrow="4" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FE27FE1BFE47" box="[165,444,449,505]" gridcol="0" gridrow="4" pageId="11" pageNumber="202">Snout Length</th>
|
||
<td id="76E4AC1C5E0194B43839FE27FC6BFE47" box="[858,972,449,505]" gridcol="1" gridrow="4" pageId="11" pageNumber="202">5.3–7.3 (6.5 ± 0.9)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FE27FB54FE47" box="[1154,1267,449,505]" gridcol="2" gridrow="4" pageId="11" pageNumber="202">4.7–6.6 (5.7 ± 0.7)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FDEFFB54FDFF" box="[165,1267,521,577]" gridrow="5" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FDEFFE1BFDFF" box="[165,444,521,577]" gridcol="0" gridrow="5" pageId="11" pageNumber="202">Interorbital Distance</th>
|
||
<td id="76E4AC1C5E0194B43839FDEFFC6BFDFF" box="[858,972,521,577]" gridcol="1" gridrow="5" pageId="11" pageNumber="202">3.0–3.5 (3.4 ± 0.2)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FDEFFB54FDFF" box="[1154,1267,521,577]" gridcol="2" gridrow="5" pageId="11" pageNumber="202">1.9–3.0 (2.6 ± 0.4)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FDB7FB54FD36" box="[165,1267,593,648]" gridrow="6" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FDB7FE1BFD36" box="[165,444,593,648]" gridcol="0" gridrow="6" pageId="11" pageNumber="202">Internarial Distance</th>
|
||
<td id="76E4AC1C5E0194B43839FDB7FC6BFD36" box="[858,972,593,648]" gridcol="1" gridrow="6" pageId="11" pageNumber="202">2.4–3.5 (3.0 ± 0.5)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FDB7FB54FD36" box="[1154,1267,593,648]" gridcol="2" gridrow="6" pageId="11" pageNumber="202">1.9–3.7 (2.3 ± 0.4)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FD7EFB54FD6E" box="[165,1267,664,720]" gridrow="7" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FD7EFE1BFD6E" box="[165,444,664,720]" gridcol="0" gridrow="7" pageId="11" pageNumber="202">Eye Diameter</th>
|
||
<td id="76E4AC1C5E0194B43839FD7EFC6BFD6E" box="[858,972,664,720]" gridcol="1" gridrow="7" pageId="11" pageNumber="202">5.0–6.0 (5.5 ± 0.4)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FD7EFB54FD6E" box="[1154,1267,664,720]" gridcol="2" gridrow="7" pageId="11" pageNumber="202">3.5–5.2 (4.4 ± 0.7)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FD06FB54FCA6" box="[165,1267,736,792]" gridrow="8" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FD06FE1BFCA6" box="[165,444,736,792]" gridcol="0" gridrow="8" pageId="11" pageNumber="202">Tympanic Annulus Diameter</th>
|
||
<td id="76E4AC1C5E0194B43839FD06FC6BFCA6" box="[858,972,736,792]" gridcol="1" gridrow="8" pageId="11" pageNumber="202">3.1–3.6 (3.3 ± 0.2)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FD06FB54FCA6" box="[1154,1267,736,792]" gridcol="2" gridrow="8" pageId="11" pageNumber="202">1.9–3.1 (2.6 ± 0.5)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FCCEFB54FCDE" box="[165,1267,808,864]" gridrow="9" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FCCEFE1BFCDE" box="[165,444,808,864]" gridcol="0" gridrow="9" pageId="11" pageNumber="202">Eye–Tympanum Distance</th>
|
||
<td id="76E4AC1C5E0194B43839FCCEFC6BFCDE" box="[858,972,808,864]" gridcol="1" gridrow="9" pageId="11" pageNumber="202">0.5–1.5 (0.8 ± 0.3)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FCCEFB54FCDE" box="[1154,1267,808,864]" gridcol="2" gridrow="9" pageId="11" pageNumber="202">0.3–1.4 (0.7 ± 0.2)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FC96FB54FC19" box="[165,1267,880,935]" gridrow="10" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FC96FE1BFC19" box="[165,444,880,935]" gridcol="0" gridrow="10" pageId="11" pageNumber="202">Head Width</th>
|
||
<td id="76E4AC1C5E0194B43839FC96FC6BFC19" box="[858,972,880,935]" gridcol="1" gridrow="10" pageId="11" pageNumber="202">14.8–17.2 (16.4 ± 1.1)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FC96FB54FC19" box="[1154,1267,880,935]" gridcol="2" gridrow="10" pageId="11" pageNumber="202">9.2–14.3 (11.9 ± 2.4)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FC51FB54FC51" box="[165,1267,951,1007]" gridrow="11" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FC51FE1BFC51" box="[165,444,951,1007]" gridcol="0" gridrow="11" pageId="11" pageNumber="202">Upper Arm Length</th>
|
||
<td id="76E4AC1C5E0194B43839FC51FC6BFC51" box="[858,972,951,1007]" gridcol="1" gridrow="11" pageId="11" pageNumber="202">6.2–8.4 (7.6 ± 0.9)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FC51FB54FC51" box="[1154,1267,951,1007]" gridcol="2" gridrow="11" pageId="11" pageNumber="202">4.3–6.8 (5.2 ± 0.9)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FC19FB54FB89" box="[165,1267,1023,1079]" gridrow="12" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FC19FE1BFB89" box="[165,444,1023,1079]" gridcol="0" gridrow="12" pageId="11" pageNumber="202">Forearm Length</th>
|
||
<td id="76E4AC1C5E0194B43839FC19FC6BFB89" box="[858,972,1023,1079]" gridcol="1" gridrow="12" pageId="11" pageNumber="202">8.9–10.9 (10.2 ± 0.9)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FC19FB54FB89" box="[1154,1267,1023,1079]" gridcol="2" gridrow="12" pageId="11" pageNumber="202">5.5–9.4 (7.7 ± 1.4)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FBA1FB54FBC0" box="[165,1267,1095,1150]" gridrow="13" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FBA1FE1BFBC0" box="[165,444,1095,1150]" gridcol="0" gridrow="13" pageId="11" pageNumber="202">Femur Length</th>
|
||
<td id="76E4AC1C5E0194B43839FBA1FC6BFBC0" box="[858,972,1095,1150]" gridcol="1" gridrow="13" pageId="11" pageNumber="202">15.4–21.7 (19.4 ± 2.7)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FBA1FB54FBC0" box="[1154,1267,1095,1150]" gridcol="2" gridrow="13" pageId="11" pageNumber="202">10.5–17.4 (14.7 ± 2.9)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FB68FB54FB78" box="[165,1267,1166,1222]" gridrow="14" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FB68FE1BFB78" box="[165,444,1166,1222]" gridcol="0" gridrow="14" pageId="11" pageNumber="202">Tibia Length</th>
|
||
<td id="76E4AC1C5E0194B43839FB68FC6BFB78" box="[858,972,1166,1222]" gridcol="1" gridrow="14" pageId="11" pageNumber="202">18.3–23.4 (21.3 ± 2.4)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FB68FB54FB78" box="[1154,1267,1166,1222]" gridcol="2" gridrow="14" pageId="11" pageNumber="202">13.0–19.1 (16.8 ± 2.6)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FB30FB54FAB0" box="[165,1267,1238,1294]" gridrow="15" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FB30FE1BFAB0" box="[165,444,1238,1294]" gridcol="0" gridrow="15" pageId="11" pageNumber="202">Tarsus Length</th>
|
||
<td id="76E4AC1C5E0194B43839FB30FC6BFAB0" box="[858,972,1238,1294]" gridcol="1" gridrow="15" pageId="11" pageNumber="202">10.2–13.0 (12.0 ± 1.2)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FB30FB54FAB0" box="[1154,1267,1238,1294]" gridcol="2" gridrow="15" pageId="11" pageNumber="202">7.4–11.4 (9.6 ± 1.7)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FAF8FB54FAE8" box="[165,1267,1310,1366]" gridrow="16" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FAF8FE1BFAE8" box="[165,444,1310,1366]" gridcol="0" gridrow="16" pageId="11" pageNumber="202">Foot Length</th>
|
||
<td id="76E4AC1C5E0194B43839FAF8FC6BFAE8" box="[858,972,1310,1366]" gridcol="1" gridrow="16" pageId="11" pageNumber="202">18.2–23.9 (22.0 ± 2.6)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FAF8FB54FAE8" box="[1154,1267,1310,1366]" gridcol="2" gridrow="16" pageId="11" pageNumber="202">12.8–19.4 (16.6 ± 2.8)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FA80FB54FA23" box="[165,1267,1382,1437]" gridrow="17" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FA80FE1BFA23" box="[165,444,1382,1437]" gridcol="0" gridrow="17" pageId="11" pageNumber="202">Hand Length</th>
|
||
<td id="76E4AC1C5E0194B43839FA80FC6BFA23" box="[858,972,1382,1437]" gridcol="1" gridrow="17" pageId="11" pageNumber="202">9.8–12.8 (11.5 ± 1.2)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FA80FB54FA23" box="[1154,1267,1382,1437]" gridcol="2" gridrow="17" pageId="11" pageNumber="202">6.7–10.0 (8.6 ± 1.4)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FA4BFB54FA5B" box="[165,1267,1453,1509]" gridrow="18" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FA4BFE1BFA5B" box="[165,444,1453,1509]" gridcol="0" gridrow="18" pageId="11" pageNumber="202">Toe 4 Length</th>
|
||
<td id="76E4AC1C5E0194B43839FA4BFC6BFA5B" box="[858,972,1453,1509]" gridcol="1" gridrow="18" pageId="11" pageNumber="202">11.8–15.0 (14.2 ± 1.6)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FA4BFB54FA5B" box="[1154,1267,1453,1509]" gridcol="2" gridrow="18" pageId="11" pageNumber="202">8.8–12.7 (11.3 ± 1.5)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6FA13FB54F993" box="[165,1267,1525,1581]" gridrow="19" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6FA13FE1BF993" box="[165,444,1525,1581]" gridcol="0" gridrow="19" pageId="11" pageNumber="202">Finger 1 Length</th>
|
||
<td id="76E4AC1C5E0194B43839FA13FC6BF993" box="[858,972,1525,1581]" gridcol="1" gridrow="19" pageId="11" pageNumber="202">4.4–6.3 (5.6 ± 0.8)</td>
|
||
<td id="76E4AC1C5E0194B43FE1FA13FB54F993" box="[1154,1267,1525,1581]" gridcol="2" gridrow="19" pageId="11" pageNumber="202">2.8–5.2 (4.0 ± 1.1)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F9DBFB54F9CA" box="[165,1267,1597,1652]" gridrow="20" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F9DBFE1BF9CA" box="[165,444,1597,1652]" gridcol="0" gridrow="20" pageId="11" pageNumber="202">Finger 3 Length</th>
|
||
<td id="76E4AC1C5E0194B43839F9DBFC6BF9CA" box="[858,972,1597,1652]" gridcol="1" gridrow="20" pageId="11" pageNumber="202">6.4–8.4 (7.7 ± 0.9)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F9DBFB54F9CA" box="[1154,1267,1597,1652]" gridcol="2" gridrow="20" pageId="11" pageNumber="202">4.1–6.6 (5.5 ± 1.1)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F963FB54F902" box="[165,1267,1669,1724]" gridrow="21" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F963FE1BF902" box="[165,444,1669,1724]" gridcol="0" gridrow="21" pageId="11" pageNumber="202">Finger 1 Disc Width</th>
|
||
<td id="76E4AC1C5E0194B43839F963FC6BF902" box="[858,972,1669,1724]" gridcol="1" gridrow="21" pageId="11" pageNumber="202">0.6–1.2 (0.9 ± 0.3)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F963FB54F902" box="[1154,1267,1669,1724]" gridcol="2" gridrow="21" pageId="11" pageNumber="202">0.5–1.2 (0.8 ± 0.3)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F92BFB54F8BA" box="[165,1267,1741,1796]" gridrow="22" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F92BFE1BF8BA" box="[165,444,1741,1796]" gridcol="0" gridrow="22" pageId="11" pageNumber="202">Finger 3 Disk Width</th>
|
||
<td id="76E4AC1C5E0194B43839F92BFC6BF8BA" box="[858,972,1741,1796]" gridcol="1" gridrow="22" pageId="11" pageNumber="202">0.8–1.6 (1.2 ± 0.3)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F92BFB54F8BA" box="[1154,1267,1741,1796]" gridcol="2" gridrow="22" pageId="11" pageNumber="202">0.1–1.2 (1.3 ± 0.2)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F8F2FB54F8F2" box="[165,1267,1812,1868]" gridrow="23" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F8F2FE1BF8F2" box="[165,444,1812,1868]" gridcol="0" gridrow="23" pageId="11" pageNumber="202">Toe 4 Disc Width</th>
|
||
<td id="76E4AC1C5E0194B43839F8F2FC6BF8F2" box="[858,972,1812,1868]" gridcol="1" gridrow="23" pageId="11" pageNumber="202">1.0–1.6 (1.3 ± 0.2)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F8F2FB54F8F2" box="[1154,1267,1812,1868]" gridcol="2" gridrow="23" pageId="11" pageNumber="202">0.6–1.3 (0.9 ± 0.2)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F8BAFB54F82D" box="[165,1267,1884,1939]" gridrow="24" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F8BAFE1BF82D" box="[165,444,1884,1939]" gridcol="0" gridrow="24" pageId="11" pageNumber="202">Penultimate Phalanx Width, Finger 3</th>
|
||
<td id="76E4AC1C5E0194B43839F8BAFC6BF82D" box="[858,972,1884,1939]" gridcol="1" gridrow="24" pageId="11" pageNumber="202">0.4–0.6 (0.5 ± 0.1)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F8BAFB54F82D" box="[1154,1267,1884,1939]" gridcol="2" gridrow="24" pageId="11" pageNumber="202">0.3–0.5 (0.4 ± 0.1)</td>
|
||
</tr>
|
||
<tr id="3535C5605E0194B43BC6F842FB54F865" box="[165,1267,1956,2011]" gridrow="25" pageId="11" pageNumber="202">
|
||
<th id="76E4AC1C5E0194B43BC6F842FE1BF865" box="[165,444,1956,2011]" gridcol="0" gridrow="25" pageId="11" pageNumber="202">Penultimate Phalanx Width, Toe 4</th>
|
||
<td id="76E4AC1C5E0194B43839F842FC6BF865" box="[858,972,1956,2011]" gridcol="1" gridrow="25" pageId="11" pageNumber="202">0.5–0.8 (0.6 ± 0.1)</td>
|
||
<td id="76E4AC1C5E0194B43FE1F842FB54F865" box="[1154,1267,1956,2011]" gridcol="2" gridrow="25" pageId="11" pageNumber="202">0.4–0.6 (0.5 ± 0.1)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF7A97AA5E066B473BF4F9B2FEFCF960" httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="12" pageNumber="203" targetBox="[233,1366,219,1587]" targetPageId="12">
|
||
<paragraph id="8BBAC7225E066B473BF4F9B2FEFCF960" blockId="12.[151,1436,1620,1758]" pageId="12" pageNumber="203">
|
||
<emphasis id="B9711B305E066B473BF4F9B2FEAAF9D7" bold="true" box="[151,269,1620,1641]" pageId="12" pageNumber="203">FIGURE 6.</emphasis>
|
||
Advertisement call of
|
||
<emphasis id="B9711B305E066B473A8EF9B2FD10F9D7" box="[493,695,1620,1641]" italics="true" pageId="12" pageNumber="203">
|
||
<taxonomicName id="4C05BCA15E066B473A8EF9B2FD14F9D7" box="[493,691,1620,1641]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="quezoni" status="sp. nov.">Platymantis quezoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A242A64B5E066B4739DDF9B2FCACF9D7" box="[702,779,1620,1641]" pageId="12" pageNumber="203" rank="species">
|
||
<emphasis id="B9711B305E066B4739DDF9B2FCACF9D7" bold="true" box="[702,779,1620,1641]" pageId="12" pageNumber="203">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3-call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of
|
||
<taxonomicName id="4C05BCA15E066B473A97F94AFDFFF97F" box="[500,600,1708,1729]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="dorsalis">
|
||
<emphasis id="B9711B305E066B473A97F94AFDFFF97F" box="[500,600,1708,1729]" italics="true" pageId="12" pageNumber="203">P. dorsalis</emphasis>
|
||
</taxonomicName>
|
||
between those of
|
||
<taxonomicName id="4C05BCA15E066B473867F94AFCC2F97F" box="[772,869,1708,1729]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="12" pageNumber="203" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E066B473867F94AFCC2F97F" box="[772,869,1708,1729]" italics="true" pageId="12" pageNumber="203">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
, and constant ambient background vocalizations of cicadas (4–5 kHz) in Fig. D.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBAC7225E066B463BA4F8C9FBEEFE30" blockId="12.[151,1437,1839,2008]" lastBlockId="13.[151,1437,151,1274]" lastPageId="13" lastPageNumber="204" pageId="12" pageNumber="203">
|
||
Males produce a largely invariant and unique advertisement call, sounding to the human ear like a long continuous series of low-frequency one-syllable barking notes ranging from: “yox, yox, yox…” to “iyohk, iyohk, iyohk …” Male calling behavior continues for up to several minutes of uninterrupted, steadily paced vocalizations. Calling begins when a previously resting male takes a more erect posture, raises the head, and takes several inhalations, buccal pumping while filling the lungs. The male then calls with a single note (described below) on expiration by contraction of body wall musculature, which fills the vocal sac. The elastic vocal sac then recoils, forcing air back into the lungs, with the process repeats over a series of up to a few minutes, with air being shuttled back and forth between the vocal sac and the lungs, and the multi-note call lasting a minute or more. Calls were separated by intervals of 12–78 (
|
||
<emphasis id="B9711B305E076B463971FEECFD87FE98" bold="true" box="[530,544,266,294]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 31.3 ± 22.7 SD; n =
|
||
<date id="FFBBE1E25E076B463842FEE2FCF7FE98" box="[801,848,260,294]" pageId="13" pageNumber="204" value="2013-10">
|
||
13
|
||
<emphasis id="B9711B305E076B463821FEECFCF7FE98" bold="true" box="[834,848,266,294]" pageId="13" pageNumber="204">x</emphasis>
|
||
</date>
|
||
) s. Note repetition rate ([total number of notes – 1]/ time from beginning of first note to beginning of last) ranged from 0.37–1.17 (
|
||
<emphasis id="B9711B305E076B463F76FED7FB84FEF3" bold="true" box="[1045,1059,305,333]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 0.80 ± 0.28 SD; n = 7) notes/s. Apart from a slightly slower initial “warm-up” of a few seconds, no modulation of note repetition rate was detected across the call, and inter-note intervals were slightly variable, but not predictably so.
|
||
</paragraph>
|
||
<paragraph id="8BBAC7225E076B463BA4FE7FFA31FBD4" blockId="13.[151,1437,151,1274]" pageId="13" pageNumber="204">
|
||
Individual notes have a complex structure of three main elements (
|
||
<figureCitation id="133EDBA75E076B4638D6FE7FFC5EFE0C" box="[949,1017,409,434]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
A,D), some of which are so brief that the call’s complex structure is nearly indiscernible to the human ear. Each note consist of a brief prefix, a second, croaking, main syllable, and a brief suffix differing from the prefix (
|
||
<figureCitation id="133EDBA75E076B4638C5FE07FC4BFE44" box="[934,1004,481,506]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
A). In the majority of calls, the prefix takes the form of a tonal frequency arc, which rapidly climbs from an initial frequency of approximately 2.2–2.6 (
|
||
<emphasis id="B9711B305E076B463BC7FDD6FF15FDF2" bold="true" box="[164,178,560,588]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 2.4 ± 0.08 SD; n = 7) kHz, reaches a maximum at 2.8–3.2 (
|
||
<emphasis id="B9711B305E076B463805FDD6FCD3FDF2" bold="true" box="[870,884,560,588]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 2.9 ± 0.11 SD; n = 7) kHz, and then declines in a mirror-image fashion over a period of 30–41 (
|
||
<emphasis id="B9711B305E076B4639A3FDB1FD69FDCD" bold="true" box="[704,718,599,627]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 34 ± 4.3 SD; n = 7) ms. The final frequency of this variable tonal prefix (in most instances a full tonal arc, in others more of a decreasing frequency sweep) is identical in apparent range and peak frequency to the upper frequency of the call’s second syllable (
|
||
<figureCitation id="133EDBA75E076B463FC0FD7DFB4DFD0A" box="[1187,1258,667,692]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
D). The second element, approximately 3–3.5 x duration of the first, is a harsh croaking syllable composed of two nearly equal intensity non-modulated frequency bands (
|
||
<figureCitation id="133EDBA75E076B46391BFD05FD1BFD42" box="[632,700,739,764]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
A,D). The two constant-frequency components consist of a lower fundamental frequency band spanning 0.9–1.7 kHz and an upper (dominant) frequency ranging across 2.2–2.8 kHz. Peak frequencies of these components varied from 2.25 to 1.4 (
|
||
<emphasis id="B9711B305E076B4638D0FCD4FC66FCF0" bold="true" box="[947,961,818,846]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 1.3 ± 0.05 SD; n = 7) kHz for the first band and 2.5 to 2.8 (
|
||
<emphasis id="B9711B305E076B463AE7FCBFFE35FCCB" bold="true" box="[388,402,857,885]" pageId="13" pageNumber="204">x</emphasis>
|
||
= 2.6 ± 0.09 SD; n = 7) kHz in the second. The third note element is a highly variable tonal suffix, again typically of the same frequency as the upper frequency band of the main, second croaking syllable, and is a variable tonal ascending sweep which, like the call’s prefix, typically sweeps from ~2.4 to 2.9 kHz (
|
||
<figureCitation id="133EDBA75E076B463E0CFC7BFF01FC64" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
D). The spectral structure of notes (
|
||
<figureCitation id="133EDBA75E076B46395DFC27FD25FC64" box="[574,642,961,986]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
A,D) consists of a pair of nearly equally emphasized frequencies (the lower, fundamental, and the upper, which is the first multiple of the fundamental). The FFT (power spectrum) demonstrates that the majority of energy in each call is found in the second frequency component (the dominant frequency, in contrast to the lower, fundamental frequency). Finally, the calls of
|
||
<taxonomicName id="4C05BCA15E076B463F47FBC9FB30FBFB" box="[1060,1175,1070,1094]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="204" phylum="Chordata" rank="species" species="quezoni">
|
||
<emphasis id="B9711B305E076B463F47FBC9FB30FBFB" box="[1060,1175,1070,1094]" italics="true" pageId="13" pageNumber="204">P. quezoni</emphasis>
|
||
</taxonomicName>
|
||
have complex spectral structure (
|
||
<figureCitation id="133EDBA75E076B463A6BFBB7FEECFBD4" box="[264,331,1105,1130]" captionStart="FIGURE 6" captionStartId="12.[151,244,1620,1641]" captionTargetBox="[233,1366,219,1587]" captionTargetId="figure@12.[217,1369,191,1598]" captionTargetPageId="12" captionText="FIGURE 6. Advertisement call of Platymantis quezoni, sp. nov. An expanded audiospectogram and waveform (A: frequency [kHz] vs. time [ms]); the Fast-Fourier Transformation (B: Power spectrum of A: relative amplitude vs. frequency); a 3 - call non-expanded oscillogram (C: relative amplitude vs. time) and the corresponding sonogram (D: audiospectrogram, frequency vs. time). Note alternating frequency sweep calls of P. dorsalis between those of P. quezoni, and constant ambient background vocalizations of cicadas (4 – 5 kHz) in Fig. D." httpUri="https://zenodo.org/record/239898/files/figure.png" pageId="13" pageNumber="204">Fig. 6</figureCitation>
|
||
D), with two detectable harmonics in most recordings and three in at least one call (TNHC 61990).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F94A95E076B463BA4FB93FB1FFB44" pageId="13" pageNumber="204" type="etymology">
|
||
<paragraph id="8BBAC7225E076B463BA4FB93FB1FFB44" blockId="13.[151,1437,151,1274]" pageId="13" pageNumber="204">
|
||
<emphasis id="B9711B305E076B463BA4FB93FEE9FB33" bold="true" box="[199,334,1141,1166]" pageId="13" pageNumber="204">Etymology.</emphasis>
|
||
The specific epithet is a patronym in the genitive singular, honoring Manuel Luis Molina Quezon. Quezon served as president of the Commonwealth of the
|
||
<collectingCountry id="F31287B25E076B463851FB7FFC17FB0C" box="[818,944,1177,1202]" name="Philippines" pageId="13" pageNumber="204">Philippines</collectingCountry>
|
||
during the American colonial period from 1935 through the conclusion of the Second World War. An exemplary statesman, he led the struggle for
|
||
<collectingCountry id="F31287B25E076B463E4AFB5BFA3BFB68" box="[1321,1436,1213,1238]" name="Philippines" pageId="13" pageNumber="204">Philippine</collectingCountry>
|
||
independence from American rule. Suggested common name: Quezon Limestone Forest Frog.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |