276 lines
40 KiB
XML
276 lines
40 KiB
XML
<document id="FF7687A66113F671C44131B758F5248F" ID-CLB-Dataset="4845" ID-DOI="10.11646/zootaxa.502.1.1" ID-GBIF-Dataset="76aa06a8-f646-4c5a-b596-54bfd8875223" ID-ISSN="11755334" ID-Zenodo-Dep="5030148" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624553872186" checkinUser="felipe" docAuthor="Jameson, Mary Liz & Wada, Kaoru" docDate="2004" docId="03A4A725CA44FFEEFEC6F964FD0CFC52" docLanguage="en" docName="zt00502.pdf" docOrigin="Zootaxa 502 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.502.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Peltonotus karubei Muramoto" docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FF9DDF5DCA52FFF6FFCEFF9DFFB5FF8C" masterDocTitle="Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="23" updateTime="1699020591651" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E42E758A112BDCA0D72D125767EC8E50">
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<mods:title id="027B29E207530182F4FD8DE535F52E30">Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia</mods:title>
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<mods:name id="3542F29A997A08A102ED1B33649481DE" type="personal">
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<mods:roleTerm id="FB433DB9BCDF3747D19731BAF4217C73">Author</mods:roleTerm>
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<mods:namePart id="8111D7C294BAC095B5D2BEADE50D0CEA">Jameson, Mary Liz</mods:namePart>
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<mods:affiliation id="10C4C48DE00797A14998DE4E79E4FA32">University of Nebraska State Museum, Division of Entomology, Lincoln, NE U. S. A., 68588 - 0514 mjameson 1 @ unl. edu</mods:affiliation>
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<mods:namePart id="4002A75B7AD2928EF27C430A4C3A164C">Wada, Kaoru</mods:namePart>
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<mods:affiliation id="08068A76B9D754D6E93F37EE46B7DE34">Musashimurayama Daini Junior School, 1460 Midorigaoka, Musashimurayama-shi, Tokyo 208 - 0012 Japan kwada 007 @ hotmail. com</mods:affiliation>
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<mods:typeOfResource id="8D356B2F5DCF75CCD6B415146FF6DB08">text</mods:typeOfResource>
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<mods:title id="9A6C6CED0C917CE31FABB27BCA6AAD79">Zootaxa</mods:title>
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<mods:part id="05EC5DD6FA952CA335BC9E731F78D6C8">
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<mods:date id="626CCB3EFA26E8BEAFB1C8021BD120F3">2004</mods:date>
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<mods:number id="6330F018603812B0362B90F0775DCA3C">2004-05-04</mods:number>
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<mods:number id="CD8C128F629BF1B2DE24A7E93E57415E">502</mods:number>
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<mods:url id="80246251B4AFE15E14BB451B7074C16E">https://biotaxa.org/Zootaxa/article/view/zootaxa.502.1.1</mods:url>
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<mods:classification id="BA77BBC04228A0732FECFED55EAB0791">journal article</mods:classification>
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<mods:identifier id="2D919DCF4095E4A83F939D10FB7F905B" type="CLB-Dataset">4845</mods:identifier>
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<treatment id="03A4A725CA44FFEEFEC6F964FD0CFC52" ID-DOI="http://doi.org/10.5281/zenodo.5248453" ID-GBIF-Taxon="184930989" ID-Zenodo-Dep="5248453" LSID="urn:lsid:plazi:treatment:03A4A725CA44FFEEFEC6F964FD0CFC52" httpUri="http://treatment.plazi.org/id/03A4A725CA44FFEEFEC6F964FD0CFC52" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<subSubSection id="C31745B8CA44FFE0FEC6F964FDCEF89F" box="[264,635,1785,1811]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BB21633CA44FFE0FEC6F964FDCEF89F" blockId="22.[264,635,1785,1846]" box="[264,635,1785,1811]" pageId="22" pageNumber="23">
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<heading id="D0FAA15FCA44FFE0FEC6F964FDCEF89F" bold="true" box="[264,635,1785,1811]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<taxonomicName id="4C0D6DB0CA44FFE0FEC6F964FDCEF89F" authority="Muramoto" authorityName="Muramoto" box="[264,635,1785,1811]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="karubei">
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<emphasis id="B979CA21CA44FFE0FEC6F964FDCEF89F" bold="true" box="[264,635,1785,1811]" pageId="22" pageNumber="23">
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<emphasis id="B979CA21CA44FFE0FEC6F964FE58F89F" bold="true" box="[264,493,1785,1811]" italics="true" pageId="22" pageNumber="23">Peltonotus karubei</emphasis>
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Muramoto
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31745B8CA44FFE0FEC6F881FE4EF8BA" box="[264,507,1820,1846]" pageId="22" pageNumber="23" type="description">
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<paragraph id="8BB21633CA44FFE0FEC6F881FE4EF8BA" blockId="22.[264,635,1785,1846]" box="[264,507,1820,1846]" pageId="22" pageNumber="23">
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(
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||
<figureCitation id="13360AB6CA44FFE0FEDFF881FEC7F8BA" box="[273,370,1820,1846]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="22" pageNumber="23">Figs. 14</figureCitation>
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,
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<figureCitation id="13360AB6CA44FFE0FEB1F881FE72F8BA" box="[383,455,1820,1846]" captionStart="FIGURES 40–42" captionStartId="57.[264,382,1756,1780]" captionTargetBox="[460,1125,328,1727]" captionTargetId="figure-50@57.[444,1144,309,1732]" captionTargetPageId="57" captionText="FIGURES 40–42. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 40) P.fujiokai, 41) P. gracilipodus and P. podocrassus, 42) P.karubei." figureDoi="http://doi.org/10.5281/zenodo.5030180" httpUri="https://zenodo.org/record/5030180/files/figure.png" pageId="22" pageNumber="23">42a–c</figureCitation>
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,
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<figureCitation id="13360AB6CA44FFE0FE1DF881FE44F8BA" box="[467,497,1820,1846]" captionStart="FIGURE 65" captionStartId="64.[264,368,1270,1294]" captionTargetBox="[337,1251,315,1246]" captionTargetId="figure-27@64.[337,1251,315,1246]" captionTargetPageId="64" captionText="FIGURE 65. Distribution of Peltonotus species in Southeast Asia and Sundaland." figureDoi="http://doi.org/10.5281/zenodo.5030204" httpUri="https://zenodo.org/record/5030204/files/figure.png" pageId="22" pageNumber="23">65</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31745B8CA44FFE1FEC6F8C7FD5DFEFF" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="reference_group">
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<paragraph id="8BB21633CA44FFE1FEC6F8C7FD5DFEFF" blockId="22.[264,1322,1881,1905]" lastBlockId="23.[312,1323,284,371]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<treatmentCitationGroup id="AB1D311DCA44FFE1FEC6F8C7FD5DFEFF" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<treatmentCitation id="0AAC3022CA44FFE0FEC6F8C7FD65F8FC" author="Muramoto, R." box="[264,720,1881,1905]" page="11" pageId="22" pageNumber="23" year="2000">
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<taxonomicName id="4C0D6DB0CA44FFE0FEC6F8C7FD65F8FC" authority="Muramoto 2000: 9 - 11" authorityName="Muramoto" authorityPageNumber="9 - 11" authorityYear="2000" box="[264,720,1881,1905]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="karubei">
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<emphasis id="B979CA21CA44FFE0FEC6F8C7FE67F8FD" box="[264,466,1882,1905]" italics="true" pageId="22" pageNumber="23">Peltonotus karubei</emphasis>
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<bibRefCitation id="EF9C6BC2CA44FFE0FE12F8C4FD65F8FC" author="Muramoto, R." box="[476,720,1881,1904]" pageId="22" pageNumber="23" pagination="9 - 11" refId="ref31943" refString="Muramoto, R. (2000) A new spesies [sic] of the ruteline genus Peltonotus (Coleoptera, Scarabaeidae) from southern Vietnam. Kogane, 1, 9 - 11." type="journal article" year="2000">Muramoto 2000: 9–11</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
.
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<typeStatus id="54B6A891CA44FFE0FD2AF8C4FCF2F8FC" box="[740,839,1881,1904]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
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in collection of Muramoto from South Vietnam,
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<collectingRegion id="49C9D8D1CA45FFE1FEB9FE81FD81FEBF" box="[375,564,284,307]" country="Vietnam" name="Lam Dong" pageId="23" pageNumber="24">Lam Dong Prov.</collectingRegion>
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, Dambri Waterfall near Bao Lok.
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<bibRefCitation id="EF9C6BC2CA45FFE1FC0FFE81FB36FEBF" author="Muramoto, R." box="[961,1155,284,307]" pageId="23" pageNumber="24" pagination="9 - 11" refId="ref31943" refString="Muramoto, R. (2000) A new spesies [sic] of the ruteline genus Peltonotus (Coleoptera, Scarabaeidae) from southern Vietnam. Kogane, 1, 9 - 11." type="journal article" year="2000">Muramoto (2000)</bibRefCitation>
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stated that the description was based on the
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<typeStatus id="54B6A891CA45FFE1FDBBFEA1FD67FEDF" box="[629,722,316,339]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
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and
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<specimenCount id="9D0BDDBACA45FFE1FCC9FEA1FCD4FEDF" box="[775,865,316,339]" pageId="23" pageNumber="24" type="male">six male</specimenCount>
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<typeStatus id="54B6A891CA45FFE1FCA7FEA1FC7AFEDF" box="[873,975,316,339]" pageId="23" pageNumber="24" type="paratype">paratypes</typeStatus>
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(five deposited in the collection of Muramoto, one deposited in WADA).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31745B8CA45FFEEFEC6FE39FCFCFDAA" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BB21633CA45FFE1FEC6FE39FCE5F9E2" blockId="23.[264,1324,420,1886]" pageId="23" pageNumber="24">
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Description (
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<specimenCount id="9D0BDDBACA45FFE1FE6CFE39FDB1FE32" box="[418,516,420,446]" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
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). Length
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<quantity id="4CF5BBD6CA45FFE1FDB1FE39FC9BFE32" box="[639,814,420,446]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.45" metricValueMin="1.35" pageId="23" pageNumber="24" unit="mm" value="14.0" valueMax="14.5" valueMin="13.5">13.5–14.5 mm</quantity>
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. Widest width
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<quantity id="4CF5BBD6CA45FFE1FC22FE39FBF2FE32" box="[1004,1095,420,446]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="23" pageNumber="24" unit="mm" value="6.8">6.8 mm</quantity>
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(based on
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<specimenCount id="9D0BDDBACA45FFE1FB03FE39FE8FFE6A" pageId="23" pageNumber="24" type="generic">1 specimen</specimenCount>
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and
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<bibRefCitation id="EF9C6BC2CA45FFE1FE4EFE51FDFCFE6A" author="Muramoto, R." box="[384,585,460,486]" pageId="23" pageNumber="24" pagination="9 - 11" refId="ref31943" refString="Muramoto, R. (2000) A new spesies [sic] of the ruteline genus Peltonotus (Coleoptera, Scarabaeidae) from southern Vietnam. Kogane, 1, 9 - 11." type="journal article" year="2000">Muramoto 2000</bibRefCitation>
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). Color: Male: Head, pronotum, scutellum, propygidium, pygidium, and venter shining, black; elytra reddishorange with iridescent bloom; elytral suture, base, and lateral margins black. Head (
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<figureCitation id="13360AB6CA45FFE1FC94FD81FC0DFDBA" box="[858,952,540,566]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="23" pageNumber="24">Fig. 14</figureCitation>
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): Surface of frons with base sparsely punctate, middle of frons to apex densely punctate; punctures ocellate, multisetigerous; short setae on disc (1–5 per puncture), moderately long setae (0–1 per puncture) laterally. Surface of clypeus densely punctate; punctures ocellate, multisetigerous; short setae on disc (1–5 per puncture), moderately long setae (0–1 per puncture) laterally. Clypeus laterally weakly constricted toward apex, apex truncate, corners square, beaded; bead weakly arcuate posteriorly. Labrum deeply, narrowly emarginate at middle (
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<figureCitation id="13360AB6CA45FFE1FB08FC91FAA9FCAA" box="[1222,1308,780,806]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="23" pageNumber="24">Fig. 14</figureCitation>
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). Mandibles rounded laterally, inner apex with 2 weak teeth.
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<collectingRegion id="49C9D8D1CA45FFE1FC34FCA9FBE8FCC2" box="[1018,1117,820,846]" country="Myanmar" name="Chin" pageId="23" pageNumber="24">Mentum</collectingRegion>
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with apical half rounded, weakly notched at middle; palpomere 2 dorsoventrally flattened, twice width of palpomere 1, setose; setae dense, moderately long, rufous, not curled at apex, not flattened. Maxilla: mala with weak lamellate setal brush; stipes with setae dense, long, not flattened, not curled at apex; palpomere 2 lacking internomedial bump. Antennal club subequal to segments 2–7. Pronotum: Bead lacking anterior to scutellum. Surface moderately densely punctate, punctures simple. Lateral margin lacking long setae. Elytral sutural length: About 5.0 times length of scutellum. Elytron: Surface with 5 poorly developed, punctate, longitudinal striae between suture and humerus; punctures moderate in size, moderately dense. Intervals with similar sculpturing. Propygidium: Surface densely punctate (disc) and confluently punctate (sides); punctures simple and ocellate, not setigerous. Pygidium: Surface coriaceous and punctate; punctures moderately dense, ocellate and simple, not setigerous. Venter: Prosternal keel triangular; apex produced to middle of protrochanter, rounded. Legs: Foretibia of male bidentate; lateral margin lacking short, dense setae. Foretarsomere 5 of male subequal in length to tarsomeres 1–4; foretarsomere 3–4 with apices expanded, dorsal and ventral apices clothed with dense, short setae. Foreclaws of male with inner claw curved, about 4 times thicker than outer claw; outer claw simply arcuate, about 1/2 the length of inner claw; empodium bulbous at base. Meso and metatibial claws of male with 2 setae. Metatibia of male apical spurs weakly curved spurs; ventral spur produced to middle of metatarsomere 1, dorsal spur produced to base of metatarsomere 2. Parameres:
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<figureCitation id="13360AB6CA45FFE1FD02F9C9FCF9F9E2" box="[716,844,1620,1646]" captionStart="FIGURES 40–42" captionStartId="57.[264,382,1756,1780]" captionTargetBox="[460,1125,328,1727]" captionTargetId="figure-50@57.[444,1144,309,1732]" captionTargetPageId="57" captionText="FIGURES 40–42. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 40) P.fujiokai, 41) P. gracilipodus and P. podocrassus, 42) P.karubei." figureDoi="http://doi.org/10.5281/zenodo.5030180" httpUri="https://zenodo.org/record/5030180/files/figure.png" pageId="23" pageNumber="24">Fig. 42a–c</figureCitation>
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.
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</paragraph>
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<paragraph id="8BB21633CA45FFEEFEF6F9E1FB42FE5A" blockId="23.[264,1324,420,1886]" lastBlockId="24.[264,1323,284,990]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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Diagnosis. Only two species of
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FD62F9E1FC9CF919" box="[684,809,1660,1685]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979CA21CA45FFE1FD62F9E1FC9CF919" box="[684,809,1660,1685]" italics="true" pageId="23" pageNumber="24">Peltonotus</emphasis>
|
||
</taxonomicName>
|
||
are known from
|
||
<collectingCountry id="F31A56A3CA45FFE1FC3DF9E1FBEEF91A" box="[1011,1115,1660,1686]" name="Vietnam" pageId="23" pageNumber="24">Vietnam</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FBA8F9E1FB69F919" box="[1126,1244,1660,1685]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA45FFE1FBA8F9E1FB69F919" box="[1126,1244,1660,1685]" italics="true" pageId="23" pageNumber="24">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FADBF9E1FED4F931" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="B979CA21CA45FFE1FADBF9E1FED4F931" italics="true" pageId="23" pageNumber="24">P. nasutus</emphasis>
|
||
</taxonomicName>
|
||
. These species are easily separated based on elytral coloration (reddishorange with iridescent bloom in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FDFAF951FD19F969" box="[564,684,1740,1765]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA45FFE1FDFAF951FD19F969" box="[564,684,1740,1765]" italics="true" pageId="23" pageNumber="24">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
; black to castaneous with dorsum shining in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FB05F951FE9FF881" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="B979CA21CA45FFE1FB05F951FE9FF881" italics="true" pageId="23" pageNumber="24">P. nasutus</emphasis>
|
||
</taxonomicName>
|
||
), form of the mentum (apical half rounded in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FCA5F969FC51F881" box="[875,996,1780,1805]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA45FFE1FCA5F969FC51F881" box="[875,996,1780,1805]" italics="true" pageId="23" pageNumber="24">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
; apical half quadrate in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FADBF969FED4F8B9" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="B979CA21CA45FFE1FADBF969FED4F8B9" italics="true" pageId="23" pageNumber="24">P. nasutus</emphasis>
|
||
</taxonomicName>
|
||
), form of the mandible in males (rounded laterally in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FC1AF881FBFCF8B9" box="[980,1097,1820,1845]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA45FFE1FC1AF881FBFCF8B9" box="[980,1097,1820,1845]" italics="true" pageId="23" pageNumber="24">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA45FFE1FB96F881FB07F8BA" box="[1112,1202,1820,1846]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="23" pageNumber="24">Fig. 14</figureCitation>
|
||
]; quadrate laterally in
|
||
<taxonomicName id="4C0D6DB0CA45FFE1FE5AF8D9FDA5F8D1" box="[404,528,1860,1885]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="B979CA21CA45FFE1FE5AF8D9FDA5F8D1" box="[404,528,1860,1885]" italics="true" pageId="23" pageNumber="24">P. nasutus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA45FFE1FDD0F8D9FDCCF8D2" box="[542,633,1860,1886]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="23" pageNumber="24">Fig. 17</figureCitation>
|
||
]), form of the labrum (deeply, narrowly emarginate in
|
||
<taxonomicName id="4C0D6DB0CA45FFEEFADBF8D9FED4FEB9" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" lastPageId="24" lastPageNumber="25" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA45FFEEFADBF8D9FED4FEB9" italics="true" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA4AFFEEFEBAFE81FE65FEBA" box="[372,464,284,310]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
|
||
]; bilobed in
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFDA2FE81FD53FEB9" box="[620,742,284,309]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="nasutus">
|
||
<emphasis id="B979CA21CA4AFFEEFDA2FE81FD53FEB9" box="[620,742,284,309]" italics="true" pageId="24" pageNumber="25">P. nasutus</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<figureCitation id="13360AB6CA4AFFEEFD36FE81FCE1FEBA" box="[760,852,284,310]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="24" pageNumber="25">Fig. 17</figureCitation>
|
||
]), and form of the male genitalia (
|
||
<figureCitation id="13360AB6CA4AFFEEFB35FE81FEFBFED2" captionStart="FIGURES 40–42" captionStartId="57.[264,382,1756,1780]" captionTargetBox="[460,1125,328,1727]" captionTargetId="figure-50@57.[444,1144,309,1732]" captionTargetPageId="57" captionText="FIGURES 40–42. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 40) P.fujiokai, 41) P. gracilipodus and P. podocrassus, 42) P.karubei." figureDoi="http://doi.org/10.5281/zenodo.5030180" httpUri="https://zenodo.org/record/5030180/files/figure.png" pageId="24" pageNumber="25">Fig. 42a–c</figureCitation>
|
||
versus
|
||
<figureCitation id="13360AB6CA4AFFEEFE69FED9FD92FED2" box="[423,551,324,350]" captionStart="FIGURES 43–45" captionStartId="58.[264,382,1739,1763]" captionTargetBox="[449,1146,330,1677]" captionTargetId="figure-45@58.[438,1148,312,1680]" captionTargetPageId="58" captionText="FIGURES 43–45. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 43) P. malayensis, 44) P.morio, 45) P. nasutus." figureDoi="http://doi.org/10.5281/zenodo.5030184" httpUri="https://zenodo.org/record/5030184/files/figure.png" pageId="24" pageNumber="25">Fig. 45a–c</figureCitation>
|
||
).
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFDF4FED9FCADFED1" box="[570,792,324,349]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA4AFFEEFDF4FED9FCADFED1" box="[570,792,324,349]" italics="true" pageId="24" pageNumber="25">Peltonotus karubei</emphasis>
|
||
</taxonomicName>
|
||
is separated from other species of
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFB63FED9FA9FFED1" box="[1197,1322,324,349]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979CA21CA4AFFEEFB63FED9FA9FFED1" box="[1197,1322,324,349]" italics="true" pageId="24" pageNumber="25">Peltonotus</emphasis>
|
||
</taxonomicName>
|
||
based on the following characters: labrum deeply, narrowly emarginate at middle apex (
|
||
<figureCitation id="13360AB6CA4AFFEEFEDFFE09FEDFFE22" box="[273,362,404,430]" captionStart="FIGURES 14–18" captionStartId="53.[264,382,1425,1449]" captionTargetBox="[340,1245,341,1398]" captionTargetId="figure-118@53.[337,1249,326,1401]" captionTargetPageId="53" captionText="FIGURES 14–18. Head (dorsal view) showing characters of the labrum, mandible, and clypeus. 14) P.karubei (apex of labrum deeply, narrowly emarginated; apex of mandible rounded laterally), 15) P. malayensis (apex of labrum broadly emarginate; apex of mandible rounded laterally), 16) P. morio (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus without tubercle in male), 17) P. nasutus (apex of labrum bilobed; apex of mandible quadrate laterally with broadly truncate apex; apex of clypeus with weak tubercle in male), 18) P. vittatus (apex of labrum broadly emarginate; apex of mandible obliquely quadrate laterally)." figureDoi="http://doi.org/10.5281/zenodo.5030160" httpUri="https://zenodo.org/record/5030160/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
|
||
); maxilla with palpomere 2 enlarged and with lamellate brush on mala; form of the male genitalia (
|
||
<figureCitation id="13360AB6CA4AFFEEFE26FE21FDDFFE5A" box="[488,618,444,470]" captionStart="FIGURES 40–42" captionStartId="57.[264,382,1756,1780]" captionTargetBox="[460,1125,328,1727]" captionTargetId="figure-50@57.[444,1144,309,1732]" captionTargetPageId="57" captionText="FIGURES 40–42. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 40) P.fujiokai, 41) P. gracilipodus and P. podocrassus, 42) P.karubei." figureDoi="http://doi.org/10.5281/zenodo.5030180" httpUri="https://zenodo.org/record/5030180/files/figure.png" pageId="24" pageNumber="25">Fig. 42a–c</figureCitation>
|
||
); male foretibia bidentate; and setae on the head sparse.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FE79FC93FE72" blockId="24.[264,1323,284,990]" box="[312,806,484,510]" pageId="24" pageNumber="25">
|
||
Distribution (
|
||
<figureCitation id="13360AB6CA4AFFEEFE1BFE79FD9BFE72" box="[469,558,484,510]" captionStart="FIGURE 65" captionStartId="64.[264,368,1270,1294]" captionTargetBox="[337,1251,315,1246]" captionTargetId="figure-27@64.[337,1251,315,1246]" captionTargetPageId="64" captionText="FIGURE 65. Distribution of Peltonotus species in Southeast Asia and Sundaland." figureDoi="http://doi.org/10.5281/zenodo.5030204" httpUri="https://zenodo.org/record/5030204/files/figure.png" pageId="24" pageNumber="25">Fig. 65</figureCitation>
|
||
). Southern
|
||
<collectingCountry id="F31A56A3CA4AFFEEFD75FE79FC97FE72" box="[699,802,484,510]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FD91FCFCFDAA" blockId="24.[264,1323,284,990]" box="[312,841,524,550]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B651C6ECA4AFFEEFEF6FD91FCF0FDAA" ID-GBIF-Occurrence="3337342357" box="[312,837,524,550]" collectionCode="WADA" pageId="24" pageNumber="25" specimenCount="2">
|
||
Locality records (
|
||
<specimenCount id="9D0BDDBACA4AFFEEFDC9FD91FD2FFDAA" box="[519,666,524,550]" pageId="24" pageNumber="25" type="generic">2 specimens</specimenCount>
|
||
) from
|
||
<collectionCode id="ED1C8EF6CA4AFFEEFD27FD91FCF0FDAA" box="[745,837,524,550]" country="Australia" httpUri="http://grbio.org/cool/mnrx-zeat" name="Western Australia Department of Agriculture" pageId="24" pageNumber="25">WADA</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31745B8CA4AFFEEFEF6FDA9FD0CFC52" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FDA9FABFFDC2" blockId="24.[264,1323,284,990]" box="[312,1290,564,590]" pageId="24" pageNumber="25">
|
||
<collectingCountry id="F31A56A3CA4AFFEEFEF6FDA9FE0AFDC2" box="[312,447,564,590]" name="Vietnam" pageId="24" pageNumber="25">VIETNAM</collectingCountry>
|
||
(2):
|
||
<collectingRegion id="49C9D8D1CA4AFFEEFE36FDA9FDC3FDC2" box="[504,630,564,590]" country="Vietnam" name="Lam Dong" pageId="24" pageNumber="25">Lam Dong</collectingRegion>
|
||
(Dambri near Baolok, Dambri Waterfall near Bao Lok).
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FDC1FDD7FDFA" blockId="24.[264,1323,284,990]" box="[312,610,604,630]" pageId="24" pageNumber="25">Temporal Data. May (2).</paragraph>
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FD19FE02FD62" blockId="24.[264,1323,284,990]" pageId="24" pageNumber="25">
|
||
Remarks. This species is known only from the type series (
|
||
<typeStatus id="54B6A891CA4AFFEEFBD6FD19FB35FD12" box="[1048,1152,644,670]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D0BDDBACA4AFFEEFB0CFD19FA9CFD12" box="[1218,1321,644,670]" pageId="24" pageNumber="25" type="male">six male</specimenCount>
|
||
<typeStatus id="54B6A891CA4AFFEEFEC6FD31FECFFD4A" box="[264,378,684,710]" pageId="24" pageNumber="25" type="paratype">paratypes</typeStatus>
|
||
) and an additional specimen we report here from near the type locality. Females are not known.
|
||
</paragraph>
|
||
<paragraph id="8BB21633CA4AFFEEFEF6FD61FD0CFC52" blockId="24.[264,1323,284,990]" pageId="24" pageNumber="25">
|
||
In the description,
|
||
<bibRefCitation id="EF9C6BC2CA4AFFEEFDD6FD61FD5EFC9A" author="Muramoto, R." box="[536,747,764,790]" pageId="24" pageNumber="25" pagination="9 - 11" refId="ref31943" refString="Muramoto, R. (2000) A new spesies [sic] of the ruteline genus Peltonotus (Coleoptera, Scarabaeidae) from southern Vietnam. Kogane, 1, 9 - 11." type="journal article" year="2000">Muramoto (2000)</bibRefCitation>
|
||
compared
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFCBEFD61FC52FC99" box="[880,999,764,789]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA4AFFEEFCBEFD61FC52FC99" box="[880,999,764,789]" italics="true" pageId="24" pageNumber="25">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFBE0FD61FB75FC99" box="[1070,1216,764,789]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="B979CA21CA4AFFEEFBE0FD61FB75FC99" box="[1070,1216,764,789]" italics="true" pageId="24" pageNumber="25">P. pruinosus</emphasis>
|
||
</taxonomicName>
|
||
, and she stated that
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFE41FCB9FD96FCB1" box="[399,547,804,829]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="B979CA21CA4AFFEEFE41FCB9FD96FCB1" box="[399,547,804,829]" italics="true" pageId="24" pageNumber="25">P. pruinosus</emphasis>
|
||
</taxonomicName>
|
||
differed based on the pygidium (shining black in
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFB40FCB9FA97FCB1" box="[1166,1314,804,829]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="B979CA21CA4AFFEEFB40FCB9FA97FCB1" box="[1166,1314,804,829]" italics="true" pageId="24" pageNumber="25">P. pruinosus</emphasis>
|
||
</taxonomicName>
|
||
; opaque black in
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFE1BFCD1FDFBFCE9" box="[469,590,844,870]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="54" phylum="Arthropoda" rank="species" species="karubei">
|
||
P.
|
||
<emphasis id="B979CA21CA4AFFEEFE3BFCD1FDFBFCE9" box="[501,590,844,869]" italics="true" pageId="24" pageNumber="25">karubei</emphasis>
|
||
</taxonomicName>
|
||
) and elytral coloration (black in
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFC27FCD1FBC8FCE9" box="[1001,1149,844,869]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="B979CA21CA4AFFEEFC27FCD1FBC8FCE9" box="[1001,1149,844,869]" italics="true" pageId="24" pageNumber="25">P. pruinosus</emphasis>
|
||
</taxonomicName>
|
||
; reddish in
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFADBFCD1FED4FC01" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="karubei">
|
||
<emphasis id="B979CA21CA4AFFEEFADBFCD1FED4FC01" italics="true" pageId="24" pageNumber="25">P. karubei</emphasis>
|
||
</taxonomicName>
|
||
). Based on rarity of specimens in collections, it is unlikely that comparisons were truly based on
|
||
<emphasis id="B979CA21CA4AFFEEFE71FC01FDEFFC39" box="[447,602,924,949]" italics="true" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFE71FC01FDE2FC39" box="[447,599,924,949]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">P. pruinosus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In this research, we found that
|
||
<taxonomicName id="4C0D6DB0CA4AFFEEFC2EFC01FBC1FC39" box="[992,1140,924,949]" class="Insecta" family="Dynastidae" genus="Peltonotus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="pruinosus">
|
||
<emphasis id="B979CA21CA4AFFEEFC2EFC01FBC1FC39" box="[992,1140,924,949]" italics="true" pageId="24" pageNumber="25">P. pruinosus</emphasis>
|
||
</taxonomicName>
|
||
is known only from a single, female
|
||
<typeStatus id="54B6A891CA4AFFEEFDC6FC59FD8FFC52" box="[520,570,964,990]" pageId="24" pageNumber="25">type</typeStatus>
|
||
specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |