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<document id="BE8CEB3237E209D1639AEFA61CDECCFA" ID-DOI="10.1649/0010-065X-72.1.97" ID-ISSN="1938-4394" ID-Zenodo-Dep="4836701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622211867469" checkinUser="carolina" docAuthor="Miller, Kelly B. &amp; Wolfe, G. William" docDate="2018" docId="03A1932F092DFF9A839BCD37FCB5FAFD" docLanguage="en" docName="TheColeopBull.72.1.113-117.pdf" docOrigin="The Coleopterists Bulletin 72 (1)" docSource="http://dx.doi.org/10.1649/0010-065x-72.1.97" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Desmopachria singhae Miller and Wolfe 2018, new species" docType="treatment" docVersion="4" lastPageNumber="106" masterDocId="FF98EB570924FF938321CE69FFB6FFCE" masterDocTitle="Nine New Species in the Desmopachria nitida Species-Group of the Neotropical Genus Desmopachria Babington, 1841 (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Hyphydrini)" masterLastPageNumber="112" masterPageNumber="97" pageNumber="106" updateTime="1698949393783" updateUser="plazi">
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<mods:title id="44315013187AE72DCD1475948AF30ABE">Nine New Species in the Desmopachria nitida Species-Group of the Neotropical Genus Desmopachria Babington, 1841 (Coleoptera: Adephaga: Dytiscidae: Hydroporinae: Hyphydrini)</mods:title>
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<mods:namePart id="E27C99BB63E96FE7EE7C87190E2CA4D8">Miller, Kelly B.</mods:namePart>
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<mods:namePart id="F61D4A063D91BEE48465E0CF1AB78EC7">Wolfe, G. William</mods:namePart>
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<mods:title id="61E4785218AFD5DAF576D5387A38B3A9">The Coleopterists Bulletin</mods:title>
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<mods:date id="2ED7217E3C642B4DEC33CBAE571E8986">2018</mods:date>
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<paragraph id="8BB72239092DFF9A839BCD37FE73FC40" blockId="9.[171,618,862,990]" pageId="9" pageNumber="106">
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<taxonomicName id="4C0859BA092DFF9A839BCD37FDE1FCBA" ID-CoL="8M232" authority="Miller and Wolfe" authorityName="Miller and Wolfe" authorityYear="2018" box="[186,599,862,884]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="singhae" status="sp. nov.">
<emphasis id="B97CFE2B092DFF9A839BCD37FE28FCBD" bold="true" box="[186,414,862,883]" italics="true" pageId="9" pageNumber="106">Desmopachria singhae</emphasis>
Miller and Wolfe
</taxonomicName>
,
<taxonomicNameLabel id="A24F4350092DFF9A826ECD10FE73FC40" box="[335,453,889,910]" pageId="9" pageNumber="106" rank="species">new species</taxonomicNameLabel>
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Zoobank.org/
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urn:lsid:zoobank.org:act:
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<figureCitation id="13333EBC092DFF9A821FCDA0FE05FC10" box="[318,435,968,990]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Figs. 2628</figureCitation>
,
<figureCitation id="13333EBC092DFF9A829FCDA0FE60FC10" box="[446,470,969,990]" captionStart="Figs" captionStartId="13.[175,214,1279,1298]" captionTargetBox="[151,1173,194,1263]" captionTargetId="figure-175@13.[151,1173,194,1263]" captionTargetPageId="13" captionText="Figs. 7172. Distribution of Desmopachria species in the D. nitida species-group. 71) D. curseenae, D. singhae, and D. vohrae; 72) D. lloydi." figureDoi="http://doi.org/10.5281/zenodo.4836725" httpUri="https://zenodo.org/record/4836725/files/figure.png" pageId="9" pageNumber="106">71</figureCitation>
)
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<subSubSection id="C31271B2092DFF9A838ECD93FD9DFBE4" pageId="9" pageNumber="106" type="materials_examined">
<paragraph id="8BB72239092DFF9A838ECD93FD9DFBE4" blockId="9.[151,638,1018,1730]" pageId="9" pageNumber="106">
<materialsCitation id="3B602864092DFF9A838ECD93FD91FBE4" country="Venezuela" latitude="10.52365" location="El Pilar" longLatPrecision="1" longitude="-63.117832" pageId="9" pageNumber="106" specimenCount="1" stateProvince="Sucre" typeStatus="holotype">
<emphasis id="B97CFE2B092DFF9A838ECD93FEF7FBC1" bold="true" box="[175,321,1018,1039]" pageId="9" pageNumber="106">
<typeStatus id="54B39C9B092DFF9A838ECD93FF57FBC1" box="[175,225,1018,1039]" pageId="9" pageNumber="106">Type</typeStatus>
Locality.
</emphasis>
<collectingCountry id="F31F62A9092DFF9A8268CD93FE1BFBC1" box="[329,429,1018,1039]" name="Venezuela" pageId="9" pageNumber="106">Venezuela</collectingCountry>
,
<collectingRegion id="49CCECDB092DFF9A8299CD93FE46FBC1" box="[440,496,1018,1039]" country="Venezuela" name="Sucre" pageId="9" pageNumber="106">Sucre</collectingRegion>
, approx.
<quantity id="4CF08FDC092DFF9A816ACD93FDCBFBC1" box="[587,637,1018,1039]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="106" unit="km" value="5.0">5 km</quantity>
SE
<location id="8ED774E2092DFF9A839ACA7CFEB3FBE4" LSID="urn:lsid:plazi:treatment:03A1932F092DFF9A839BCD37FCB5FAFD:8ED774E2092DFF9A839ACA7CFEB3FBE4" box="[187,261,1045,1066]" country="Venezuela" latitude="10.52365" longLatPrecision="1" longitude="-63.117832" name="El Pilar" pageId="9" pageNumber="106" stateProvince="Sucre">El Pilar</location>
,
<geoCoordinate id="EE3C44FE092DFF9A8231CA7CFE25FBE4" box="[272,403,1044,1066]" degrees="10" direction="north" minutes="31.419" orientation="latitude" pageId="9" pageNumber="106" precision="1" value="10.52365">10°31.419ʹ N</geoCoordinate>
,
<geoCoordinate id="EE3C44FE092DFF9A82BECA7CFD91FBE4" box="[415,551,1044,1066]" degrees="63" direction="west" minutes="07.070" orientation="longitude" pageId="9" pageNumber="106" precision="1" value="-63.117832">63°07.070ʹ W</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A838ECA46FDBEFA76" pageId="9" pageNumber="106" type="diagnosis">
<paragraph id="8BB72239092DFF9A838ECA46FDBEFA76" blockId="9.[151,638,1018,1730]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECA46FEA1FB8A" bold="true" box="[175,279,1071,1092]" pageId="9" pageNumber="106">Diagnosis.</emphasis>
This species differs from all other species in the group in the male genitalia. In particular, the male lateral lobe is apically only shallowly bifid (
<figureCitation id="13333EBC092DFF9A8238CA16FED0FB5A" box="[281,358,1151,1172]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 28</figureCitation>
). The male median lobe is deeply bifid in ventral aspect with the lateral rami apically elongate and slender, apically pointed with each ramus apically directed laterally to a very narrow point (
<figureCitation id="13333EBC092DFF9A823FCA80FED3FB30" box="[286,357,1257,1278]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 27</figureCitation>
). In lateral aspect, the median lobe is elongate and apically strongly sinuate to the narrowly pointed and curved apex (
<figureCitation id="13333EBC092DFF9A8100CB77FDC5FAFD" box="[545,627,1310,1331]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 26</figureCitation>
). Specimens are relatively pale with the head and pronotum yellow-orange and the posterior margin of the pronotum narrowly dark brown. The elytra are brown with darker brown along the anterior margin and medially along the suture.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A838ECBD7FCA7FDFE" pageId="9" pageNumber="106" type="description">
<paragraph id="8BB72239092DFF9A838ECBD7FB22FE08" blockId="9.[151,638,1018,1730]" lastBlockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A838ECBD7FE64FA1D" bold="true" box="[175,466,1470,1491]" pageId="9" pageNumber="106">Description. Measurements:</emphasis>
TL =
<quantity id="4CF08FDC092DFF9A8109CBD7FDCCFA1D" box="[552,634,1470,1491]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="106" unit="mm" value="1.6">1.6 mm</quantity>
, GW =
<quantity id="4CF08FDC092DFF9A83FDCBB1FE90FA23" box="[220,294,1496,1517]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="106" unit="mm" value="1.1">1.1 mm</quantity>
, HW =
<quantity id="4CF08FDC092DFF9A8255CBB1FE0BFA23" box="[372,445,1496,1517]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="106" unit="mm" value="0.6">0.6 mm</quantity>
, EW =
<quantity id="4CF08FDC092DFF9A8129CBB1FDE4FA23" box="[520,594,1496,1517]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="106" unit="mm" value="0.3">0.3 mm</quantity>
, TL/ GW = 1.5, HW/EW = 2.12.3.
<emphasis id="B97CFE2B092DFF9A82F1CB9AFDF3F9C6" bold="true" box="[464,581,1523,1544]" pageId="9" pageNumber="106">Coloration:</emphasis>
Head and pronotum yellow-orange, posterior margin of pronotum medially narrowly darker brown. Elytron brown, darker brown along anterior margin and medially along suture. Ventral surfaces and appendages yellow-orange except metacoxal plates brown.
<emphasis id="B97CFE2B092DFF9A83C1C8FBFEC0F969" bold="true" box="[224,374,1682,1703]" pageId="9" pageNumber="106">Male genitalia:</emphasis>
Male median lobe in lateral aspect with elongate, broadly rounded lobate base, apical portion elongate, tapered and curved, apex slender, sharply recurved, apically sharply pointed (
<figureCitation id="13333EBC092DFF9A8195CE9EFCB4FEC2" box="[692,770,247,268]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 26</figureCitation>
). Median lobe in ventral aspect broad, lateral margins broadly convex to apically convergent to elongate slender, apically sharply pointed rami, medial emargination deep (
<figureCitation id="13333EBC092DFF9A80D2CF2FFB88FE95" box="[1011,1086,326,347]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 27</figureCitation>
). Lateral lobe in lateral aspect broad, apically abruptly curved, apex oblique, apically shallowly and broadly bifid, each ramus sharply pointed (
<figureCitation id="13333EBC092DFF9A8763CFFFFB3FFE65" box="[1090,1161,406,427]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 28</figureCitation>
); in ventral aspect narrow, sinuate, apically pointed.
</paragraph>
<paragraph id="8BB72239092DFF9A81E4CFA2FCA7FDFE" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CFA2FC9DFE2E" bold="true" box="[709,811,459,480]" pageId="9" pageNumber="106">Variation.</emphasis>
The darkness of the coloration of the posterior margin of the pronotum and the elytral suture varies somewhat among the few specimens examined.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A81E4CC5CFC57FDAB" pageId="9" pageNumber="106" type="etymology">
<paragraph id="8BB72239092DFF9A81E4CC5CFC57FDAB" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CC5CFC8EFD84" bold="true" box="[709,824,565,586]" pageId="9" pageNumber="106">Etymology.</emphasis>
This species is named
<taxonomicName id="4C0859BA092DFF9A870DCC5CFBC0FD85" authorityName="Miller and Wolfe" authorityYear="2018" box="[1068,1142,565,587]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="singhae">
<emphasis id="B97CFE2B092DFF9A870DCC5CFBC0FD85" box="[1068,1142,565,587]" italics="true" pageId="9" pageNumber="106">singhae</emphasis>
</taxonomicName>
in honor of Dr. Vinita Singh, MD.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A81E4CC02FBE2FD24" pageId="9" pageNumber="106" type="distribution">
<paragraph id="8BB72239092DFF9A81E4CC02FCBFFD7B" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CC02FCF0FD4E" bold="true" box="[709,838,619,640]" pageId="9" pageNumber="106">Distribution.</emphasis>
This species is known only from one locality in
<collectingRegion id="49CCECDB092DFF9A8040CCECFC21FD54" box="[865,919,645,666]" country="Venezuela" name="Sucre" pageId="9" pageNumber="106">Sucre</collectingRegion>
in northern
<collectingCountry id="F31F62A9092DFF9A8713CCECFB25FD54" box="[1074,1171,645,666]" name="Venezuela" pageId="9" pageNumber="106">Venezuela</collectingCountry>
(
<figureCitation id="13333EBC092DFF9A8195CCC9FD4AFD7B" box="[692,764,672,693]" captionStart="Figs" captionStartId="13.[175,214,1279,1298]" captionTargetBox="[151,1173,194,1263]" captionTargetId="figure-175@13.[151,1173,194,1263]" captionTargetPageId="13" captionText="Figs. 7172. Distribution of Desmopachria species in the D. nitida species-group. 71) D. curseenae, D. singhae, and D. vohrae; 72) D. lloydi." figureDoi="http://doi.org/10.5281/zenodo.4836725" httpUri="https://zenodo.org/record/4836725/files/figure.png" pageId="9" pageNumber="106">Fig. 71</figureCitation>
).
</paragraph>
<paragraph id="8BB72239092DFF9A81E4CCD3FBE2FD24" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CCD3FCAFFD01" bold="true" box="[709,793,698,719]" pageId="9" pageNumber="106">Habitat.</emphasis>
According to label data, specimens were collected from a “forested swamp by road.”
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A81E4CC99FCA0FC17" pageId="9" pageNumber="106" type="discussion">
<paragraph id="8BB72239092DFF9A81E4CC99FCA0FC17" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CC99FC8EFCCB" bold="true" box="[709,824,752,773]" pageId="9" pageNumber="106">Comments.</emphasis>
The main character uniting the
<taxonomicName id="4C0859BA092DFF9A818CCD63FCB3FCD1" authorityName="Babington" authorityYear="1841" box="[685,773,777,799]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="nitida">
<emphasis id="B97CFE2B092DFF9A818CCD63FCB3FCD1" box="[685,773,777,799]" italics="true" pageId="9" pageNumber="106">D. nitida</emphasis>
</taxonomicName>
species-group is the deeply bifid lateral lobe of the male aedeagus.
<taxonomicName id="4C0859BA092DFF9A8097CD4DFB22FCF4" authorityName="Miller and Wolfe" authorityYear="2018" box="[950,1172,804,826]" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="singhae">
<emphasis id="B97CFE2B092DFF9A8097CD4DFB22FCF4" box="[950,1172,804,826]" italics="true" pageId="9" pageNumber="106">Desmopachria singhae</emphasis>
</taxonomicName>
species has the lateral lobe apically bifid, but not deeply so (
<figureCitation id="13333EBC092DFF9A803ACD33FCD3FCA1" box="[795,869,858,879]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 32</figureCitation>
). However, the median lobe is somewhat similar to other species in the genus, including its deep medial emargination in ventral aspect (
<figureCitation id="13333EBC092DFF9A81D6CDC3FCF6FC71" box="[759,832,938,959]" captionStart="Figs" captionStartId="2.[197,236,1466,1485]" captionTargetBox="[325,1043,194,1444]" captionTargetId="figure-0@2.[322,1044,194,1451]" captionTargetPageId="2" captionText="Figs. 132. Male genitalia of Desmopachria species in the D. nitida species-group. D. curseenae: Median lobe 1) Right lateral aspect; 2) Ventral aspect; Right lateral lobe 3) Right lateral aspect; 4) Ventral aspect. D. delongi: Median lobe 5) Right lateral aspect; 6) Ventral aspect; Right lateral lobe 7) Right lateral aspect; 8) Ventral aspect. D. gingerae: Median lobe 9) Right lateral aspect; 10) Ventral aspect; 11) Right lateral lobe, right lateral aspect. D. gyrationi: Median lobe 12) Right lateral aspect; 13) Ventral aspect; 14) Right lateral lobe, right lateral aspect. D. hardyae: Median lobe 15) Right lateral aspect; 16) Ventral aspect; 17) Right lateral lobe, right lateral aspect. D. kemptonae: Median lobe 18) Right lateral aspect; 19) Ventral aspect; Right lateral lobe 20) Right lateral aspect; 21) Ventral aspect. D. lloydi: 22) Median lobe, ventral aspect; 23) Apices of median lobe, anterior aspect; Right lateral lobe 24) Right lateral aspect; 25) Ventral aspect. D. singhae: Median lobe 26) Right lateral aspect; 27) Ventral aspect; 28) Right lateral lobe, right lateral aspect. D. vohrae: Median lobe 29) Right lateral aspect; 30) Ventral aspect; Right lateral lobe 31) Right lateral aspect; 32) Ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.4836705" httpUri="https://zenodo.org/record/4836705/files/figure.png" pageId="9" pageNumber="106">Fig. 31</figureCitation>
), suggesting it should be placed in this group.
</paragraph>
</subSubSection>
<subSubSection id="C31271B2092DFF9A81E4CDB6FCB5FAFD" pageId="9" pageNumber="106" type="materials_examined">
<paragraph id="8BB72239092DFF9A81E4CDB6FCB5FAFD" blockId="9.[685,1172,194,1331]" pageId="9" pageNumber="106">
<emphasis id="B97CFE2B092DFF9A81E4CDB6FCEFFC3A" bold="true" box="[709,857,991,1012]" pageId="9" pageNumber="106">Type Material.</emphasis>
<materialsCitation id="3B602864092DFF9A807CCDB6FC98FB60" collectingDate="2010-01-29" collectionCode="MIZA, KUNHM-ENT" collectorName="Short" country="Venezuela" elevation="2" latitude="10.52365" location="Sucre State" longLatPrecision="1" longitude="-63.117832" pageId="9" pageNumber="106" specimenCode="SEMC0891088" specimenCount="2" stateProvince="Sucre" typeStatus="holotype">
<typeStatus id="54B39C9B092DFF9A807CCDB6FC03FC3A" box="[861,949,991,1012]" pageId="9" pageNumber="106" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED19BAFC092DFF9A80F0CDB6FBA6FC3A" box="[977,1040,991,1012]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="106" type="Museum">MIZA</collectionCode>
, male labeled, “
<collectingCountry id="F31F62A9092DFF9A8199CD90FCFFFBC0" box="[696,841,1017,1038]" name="Venezuela" pageId="9" pageNumber="106">VENEZUELA</collectingCountry>
:
<collectingRegion id="49CCECDB092DFF9A804CCD90FC45FBC0" box="[877,1011,1017,1038]" country="Venezuela" name="Sucre" pageId="9" pageNumber="106">Sucre State</collectingRegion>
<geoCoordinate id="EE3C44FE092DFF9A8735CD90FB39FBC0" box="[1044,1167,1017,1039]" degrees="10" direction="north" minutes="31.419" orientation="latitude" pageId="9" pageNumber="106" precision="1" value="10.52365">10°31.419ʹN</geoCoordinate>
,
<geoCoordinate id="EE3C44FE092DFF9A818CCA7DFC9BFBE7" box="[685,813,1043,1065]" degrees="63" direction="west" minutes="07.070" orientation="longitude" pageId="9" pageNumber="106" precision="1" value="-63.117832">63°07.070ʹW</geoCoordinate>
,
<quantity id="4CF08FDC092DFF9A8016CA7DFCEDFBE7" box="[823,859,1044,1065]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="106" unit="m" value="2.0">
<elevation id="0025C50A092DFF9A8016CA7DFCEDFBE7" box="[823,859,1044,1065]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="106" unit="m" value="2.0">2 m</elevation>
</quantity>
ca.
<quantity id="4CF08FDC092DFF9A80A2CA7DFC02FBE7" box="[899,948,1044,1065]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="106" unit="km" value="5.0">5 km</quantity>
SE EL Pilar;
<date id="FFB604F9092DFF9A8718CA7DFB22FBE7" box="[1081,1172,1044,1065]" pageId="9" pageNumber="106" value="2010-01-29">
<collectingDate id="EFF2FD11092DFF9A8718CA7DFB22FBE7" box="[1081,1172,1044,1065]" pageId="9" pageNumber="106" value="2010-01-29">29.i.2010</collectingDate>
</date>
leg.
<collectorName id="26FD47EF092DFF9A81F5CA47FCB1FB8D" box="[724,775,1070,1091]" pageId="9" pageNumber="106">Short</collectorName>
&amp; Garcia; forested swamp by road; VZ10- 01129-04A/
<specimenCode id="DBAE8A42092DFF9A8008CA20FC0BFB90" box="[809,957,1097,1118]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="106" type="Museum">SEMC0891088</specimenCode>
<collectionCode id="ED19BAFC092DFF9A80E7CA20FBE0FB90" box="[966,1110,1097,1118]" pageId="9" pageNumber="106">KUNHM-ENT</collectionCode>
[label with barcode]/
<typeStatus id="54B39C9B092DFF9A806DCA0AFC7FFBB6" box="[844,969,1123,1144]" pageId="9" pageNumber="106" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C0859BA092DFF9A80F6CA0AFC73FB5D" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="singhae">
<emphasis id="B97CFE2B092DFF9A80F6CA0AFD6DFB5D" italics="true" pageId="9" pageNumber="106">Desmopachria singhae</emphasis>
Miller and Wolfe, 2017
</taxonomicName>
[red label with black line border].”
</materialsCitation>
<materialsCitation id="3B602864092DFF9A8016CAF0FCB5FAFD" collectionCode="SEMC" country="Venezuela" pageId="9" pageNumber="106" specimenCode="SEMC0891096, SEMC0891080" specimenCount="3" stateProvince="Sucre" typeStatus="paratype">
<typeStatus id="54B39C9B092DFF9A8016CAF0FC20FB60" box="[823,918,1177,1198]" pageId="9" pageNumber="106" type="paratype">Paratypes</typeStatus>
: Two labeled same as
<typeStatus id="54B39C9B092DFF9A8755CAF0FD5DFB06" pageId="9" pageNumber="106" type="holotype">holotype</typeStatus>
, but with different specimen barcode labels [
<specimenCode id="DBAE8A42092DFF9A8192CAA7FCFCFB2D" box="[691,842,1230,1251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="106" type="Museum">SEMC0891096</specimenCode>
and
<specimenCode id="DBAE8A42092DFF9A80AACAA7FB94FB2D" box="[907,1058,1230,1251]" collectionCode="SEMC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="106" type="Museum">SEMC0891080</specimenCode>
] and each with “…
<typeStatus id="54B39C9B092DFF9A8035CA81FC3CFB33" box="[788,906,1256,1277]" pageId="9" pageNumber="106" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C0859BA092DFF9A8085CA8EFC26FAD6" authority="Miller and Wolfe, 2017" authorityName="Miller and Wolfe" authorityYear="2017" class="Insecta" family="Dytiscidae" genus="Desmopachria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="106" phylum="Arthropoda" rank="species" species="singhae">
<emphasis id="B97CFE2B092DFF9A8085CA8EFB22FB33" box="[932,1172,1255,1277]" italics="true" pageId="9" pageNumber="106">Desmopachria singhae</emphasis>
Miller and Wolfe, 2017
</taxonomicName>
[blue label with black line border].”
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>