615 lines
77 KiB
XML
615 lines
77 KiB
XML
<document id="379ABFF1EC938AFB164546124D8C9F21" ID-CLB-Dataset="45472" ID-DOI="10.5281/zenodo.215278" ID-GBIF-Dataset="5c0fc0e6-8fa2-48ca-af05-b44898f6b636" ID-ISSN="1175-5326" ID-Zenodo-Dep="215278" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460437642047" checkinUser="plazi" docAuthor="Bauer, Aaron M. & Doughty, Paul" docDate="2012" docId="039ED94ABC232D30FF18FEC22C1CFAA7" docLanguage="en" docName="zt03187p042.pdf" docOrigin="Zootaxa 3187" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyrtodactylus kimberleyensis Bauer & Doughty, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="38" masterDocId="FFA7A132BC212D37FF8FFF8E2914FFAB" masterDocTitle="A new bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from the Kimberley region, Western Australia" masterLastPageNumber="42" masterPageNumber="32" pageNumber="34" updateTime="1698332440166" updateUser="ExternalLinkService">
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<mods:mods id="DD199504A8A44375DA5EA6D957005C52" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="60B6EE1760785234A2CC28A0D1244FB1">
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<mods:title id="1B3B57562C89048529D858F9881F38DB">A new bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from the Kimberley region, Western Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="4750EA5A682628060BCECAC604F8EABC" type="personal">
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<mods:namePart id="953290E582A090E7325D6A25EE3F3259">Bauer, Aaron M.</mods:namePart>
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<mods:roleTerm id="88B1485091E2B707380107141262D703">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C1629970B4D555FAC708043BA0269A41">Doughty, Paul</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="76CE53778FF04026DEA83F37CD4E59D4">text</mods:typeOfResource>
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<mods:title id="E39F43D236A17B45A7C915E38ACFA64C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="5FD477B511B1A4D052DB230E618F5DBB">
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<mods:date id="B018F46ADA43681E44BD1939CDCF4537">2012</mods:date>
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<treatment id="039ED94ABC232D30FF18FEC22C1CFAA7" ID-DOI="http://doi.org/10.5281/zenodo.6168522" ID-GBIF-Taxon="119507003" ID-Zenodo-Dep="6168522" LSID="urn:lsid:plazi:treatment:039ED94ABC232D30FF18FEC22C1CFAA7" httpUri="http://treatment.plazi.org/id/039ED94ABC232D30FF18FEC22C1CFAA7" lastPageId="7" lastPageNumber="38" pageId="2" pageNumber="34">
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<subSubSection id="C32D3BD7BC232D35FF18FEC229EBFE0C" pageId="2" pageNumber="34" type="nomenclature">
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<paragraph id="8B88685CBC232D35FF18FEC22B48FECE" blockId="2.[151,604,331,423]" box="[151,604,331,358]" pageId="2" pageNumber="34">
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<heading id="D0C0DF30BC232D35FF18FEC22B48FECE" bold="true" box="[151,604,331,358]" fontSize="11" level="1" pageId="2" pageNumber="34" reason="1">
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<emphasis id="B943B44EBC232D35FF18FEC22B48FECE" bold="true" box="[151,604,331,358]" pageId="2" pageNumber="34">
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<taxonomicName id="4C3713DFBC232D35FF18FEC228E3FECD" box="[151,503,332,358]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
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<emphasis id="B943B44EBC232D35FF18FEC228E3FECD" bold="true" box="[151,503,332,358]" italics="true" pageId="2" pageNumber="34">Cyrtodactylus kimberleyensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2700935BC232D35FE71FEC52B48FECE" box="[510,604,331,357]" pageId="2" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B88685CBC232D35FF18FEE029EBFE0C" blockId="2.[151,604,331,423]" pageId="2" pageNumber="34">
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Kimberley Bent-toed Gecko
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<figureCitation id="130C74D9BC232D35FF18FE0129EBFE0C" box="[151,255,399,423]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="2.[151,250,1470,1493]" captionStartId-1="3.[151,250,1340,1363]" captionStartId-2="4.[151,250,1306,1329]" captionStartId-3="5.[151,250,1437,1460]" captionTargetBox-0="[151,1436,621,1449]" captionTargetBox-1="[151,1436,692,1319]" captionTargetBox-2="[369,1154,517,1267]" captionTargetBox-3="[151,1436,447,1411]" captionTargetId-0="figure@2.[151,1436,621,1449]" captionTargetId-1="figure@3.[151,1436,692,1320]" captionTargetId-2="figure@4.[354,1233,517,1285]" captionTargetId-3="figure@5.[151,1436,447,1411]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 2. Adult female holotype of Cyrtodactylus kimberleyensis sp. nov. (WAM R 164144). Scale bar = 10 mm." captionText-1="FIGURE 3. Head of Cyrtodactylus kimberleyensis sp. nov. A) oblique view of right side of face showing circumnarial and labial scales; B) ventral view of chin and throat showing mental and post-mental scales." captionText-2="FIGURE 4. Ventral view of right pes of Cyrtodactylus kimberleyensis sp. nov. showing disposition of the subdigital lamellae. Roman numerals indicate digit numbers. Scale bar = 2 mm." captionText-3="FIGURE 5. Photo in life of the holotype of Cyrtodactylus kimberleyensis sp. nov. (WAM R 164144). Photo—R. Browne-Cooper." httpUri-0="https://zenodo.org/record/215280/files/figure.png" httpUri-1="https://zenodo.org/record/215281/files/figure.png" httpUri-2="https://zenodo.org/record/215282/files/figure.png" httpUri-3="https://zenodo.org/record/215283/files/figure.png" pageId="2" pageNumber="34">Figs. 2–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D3BD7BC232D35FF18FE5629C2FD9C" pageId="2" pageNumber="34" type="materials_examined">
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<paragraph id="8B88685CBC232D35FF18FE5629C2FD9C" blockId="2.[151,1436,471,567]" pageId="2" pageNumber="34">
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<emphasis id="B943B44EBC232D35FF18FE56281FFE5B" bold="true" box="[151,267,472,496]" pageId="2" pageNumber="34">
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<typeStatus id="548CD6FEBC232D35FF18FE562812FE5B" box="[151,262,472,496]" pageId="2" pageNumber="34" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED26F099BC232D35FE99FE56284EFE5B" LSID="urn:lsid:biocol.org:col:34981" box="[278,346,472,496]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="34">WAM</collectionCode>
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R164144, gravid adult female (
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<figureCitation id="130C74D9BC232D35FD5BFE592A09FE44" box="[724,797,471,495]" captionStart="FIGURE 2" captionStartId="2.[151,250,1470,1493]" captionTargetBox="[151,1436,621,1449]" captionTargetId="figure@2.[151,1436,621,1449]" captionTargetPageId="2" captionText="FIGURE 2. Adult female holotype of Cyrtodactylus kimberleyensis sp. nov. (WAM R 164144). Scale bar = 10 mm." httpUri="https://zenodo.org/record/215280/files/figure.png" pageId="2" pageNumber="34">Fig. 2</figureCitation>
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);
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<collectingCountry id="F32028CCBC232D35FCB8FE592AB5FE44" box="[823,929,471,495]" name="Australia" pageId="2" pageNumber="34">Australia</collectingCountry>
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, Western
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<collectingCountry id="F32028CCBC232D35FB99FE592D94FE44" box="[1046,1152,471,495]" name="Australia" pageId="2" pageNumber="34">Australia</collectingCountry>
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, Kimberley region, East Montalivet Island (
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<geoCoordinate id="EE030E9BBC232D35FEE4FE7228D0FDBF" box="[363,452,507,532]" direction="south" orientation="latitude" pageId="2" pageNumber="34" precision="925" value="-14.266666">14º16’S</geoCoordinate>
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,
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<geoCoordinate id="EE030E9BBC232D35FE5FFE722B2CFDBF" box="[464,568,507,532]" direction="east" orientation="longitude" pageId="2" pageNumber="34" precision="925" value="125.3">125º18’E</geoCoordinate>
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), collected with a funnel trap in a vine thicket by R. Browne-Cooper,
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<date id="FF894E9CBC232D35FAB1FE7229C7FD9C" pageId="2" pageNumber="34" value="2007-04-26">26 April 2007</date>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF4838D4BC232D35FF18FA302C38FA7F" box="[151,1324,1470,1493]" httpUri="https://zenodo.org/record/215280/files/figure.png" pageId="2" pageNumber="34" targetBox="[151,1436,621,1449]" targetPageId="2">
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<paragraph id="8B88685CBC232D35FF18FA302C38FA7F" blockId="2.[151,1324,1470,1493]" box="[151,1324,1470,1493]" pageId="2" pageNumber="34">
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<emphasis id="B943B44EBC232D35FF18FA302807FA7F" bold="true" box="[151,275,1470,1493]" pageId="2" pageNumber="34">FIGURE 2.</emphasis>
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Adult female holotype of
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<taxonomicName id="4C3713DFBC232D35FD91FA312A5DFA7E" box="[542,841,1471,1493]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
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<emphasis id="B943B44EBC232D35FD91FA312A5DFA7E" box="[542,841,1471,1493]" italics="true" pageId="2" pageNumber="34">Cyrtodactylus kimberleyensis</emphasis>
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</taxonomicName>
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<emphasis id="B943B44EBC232D35FCC0FA312AB5FA7F" bold="true" box="[847,929,1471,1492]" pageId="2" pageNumber="34">
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<taxonomicNameLabel id="A2700935BC232D35FCC0FA312AB5FA7F" box="[847,929,1471,1492]" pageId="2" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(WAM R164144). Scale bar = 10 mm.
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</paragraph>
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</caption>
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<subSubSection id="C32D3BD7BC232D35FF48F9AB2C89F8BE" pageId="2" pageNumber="34" type="diagnosis">
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<paragraph id="8B88685CBC232D35FF48F9AB2C89F8BE" blockId="2.[151,1437,1573,2030]" pageId="2" pageNumber="34">
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<emphasis id="B943B44EBC232D35FF48F9AB282EF995" bold="true" box="[199,314,1573,1598]" pageId="2" pageNumber="34">Diagnosis</emphasis>
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. A small sized
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<taxonomicName id="4C3713DFBC232D35FE68F9AB2B90F996" box="[487,644,1573,1597]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="34" phylum="Chordata" rank="genus">
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<emphasis id="B943B44EBC232D35FE68F9AB2B90F996" box="[487,644,1573,1597]" italics="true" pageId="2" pageNumber="34">Cyrtodactylus</emphasis>
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</taxonomicName>
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(female gravid at SVL of
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<quantity id="4CCFC5B9BC232D35FC25F9AB2AE9F996" box="[938,1021,1573,1597]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="34" unit="mm" value="45.0">45 mm</quantity>
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); body slender; limbs and digits relatively long, slender; partly regenerated tail longer than SVL; one pair of greatly enlarged postmental scales in contact with one another behind mental, a smaller pair of enlarged chin shields (second postmentals) lateral to these; small, mostly keeled tubercles in 16–18 longitudinal rows on dorsum; 36 scales across mid-venter between lowest rows of flank tubercles; ventrolateral folds weakly developed and atuberculate; six broad basal lamellae and nine narrow distal lamellae beneath digit IV of pes; dorsal and basal portion of tail only with flattened and keeled tubercles. Color pattern of small diffuse light and dark spots or blotches, no dark transverse bands or large dark blotches.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D3BD7BC232D32FF48F8AF2C34FE2C" lastPageId="5" lastPageNumber="37" pageId="2" pageNumber="34" type="description">
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<paragraph id="8B88685CBC232D34FF48F8AF2B48FDCB" blockId="2.[151,1437,1573,2030]" lastBlockId="3.[151,1437,151,608]" lastPageId="3" lastPageNumber="35" pageId="2" pageNumber="34">
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<emphasis id="B943B44EBC232D35FF48F8AF28F0F891" bold="true" box="[199,484,1825,1850]" pageId="2" pageNumber="34">
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Description of
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<typeStatus id="548CD6FEBC232D35FEF9F8AC28CBF891" box="[374,479,1826,1850]" pageId="2" pageNumber="34" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Adult female. SVL
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<quantity id="4CCFC5B9BC232D35FD49F8AF2A38F892" box="[710,812,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="4.51" pageId="2" pageNumber="34" unit="mm" value="45.1">45.1 mm</quantity>
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; TailL
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<quantity id="4CCFC5B9BC232D35FCF5F8AF2ACAF892" box="[890,990,1825,1849]" metricMagnitude="-2" metricUnit="m" metricValue="5.33" pageId="2" pageNumber="34" unit="mm" value="53.3">53.3 mm</quantity>
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(terminal
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<quantity id="4CCFC5B9BC232D35FBDEF8AF2DB3F892" box="[1105,1191,1825,1849]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="2" pageNumber="34" unit="mm" value="5.3">5.3 mm</quantity>
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regenerated); mass in life
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<quantity id="4CCFC5B9BC232D35FF4AF8CB2817F8F5" box="[197,259,1861,1886]" metricMagnitude="-3" metricUnit="kg" metricValue="2.5" pageId="2" pageNumber="34" unit="g" value="2.5">2.5 g</quantity>
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. Head moderately long (HeadL/SVL ratio 0.29), narrow (HeadW/HeadL = 0.58), somewhat depressed (HeadH/HeadL = 0.34), distinct from neck. Loreal region weakly inflated, canthus rostralis not prominent. Snout elongate (SnEye/HeadL = 0.42), somewhat accuminate; longer than eye diameter (OrbD/SnEye = 0.58); scales on snout small, oval, conical with apex directed posteriorly, mostly homogeneous, distinctly larger than those on crown, interorbital and occipital regions. Eye moderately large (OrbD/HeadL = 0.24); pupil vertical with crenelated margins; supraciliaries short, lacking spines or projections. Ear opening horizontally elliptical (right ear opening partly occluded by fold of skin), large (EarL/HeadL = 0.10); eye to ear distance much greater than diameter of eyes (EyeEar/OrbD = 1.42). Rostral much wider (
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<quantity id="4CCFC5B9BC222D34FD41FF512A33FF5C" box="[718,807,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="35" unit="mm" value="1.9">1.9 mm</quantity>
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) than deep (1.0 mm), rostral crease about 2/3 height of rostral. Two slightly enlarged, rhomboidal supranasals separated by a single, smaller, pentagonal internasal. Rostral in contact with first supralabials, nostrils, internasal and supranasals. Nostrils oval, slightly posterolaterallydirected, each surrounded by supranasal, rostral, first supralabial and two postnasals (
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<figureCitation id="130C74D9BC222D34FBD0FEC22DB6FECF" box="[1119,1186,332,356]" captionStart="FIGURE 3" captionStartId="3.[151,250,1340,1363]" captionTargetBox="[151,1436,692,1319]" captionTargetId="figure@3.[151,1436,692,1320]" captionTargetPageId="3" captionText="FIGURE 3. Head of Cyrtodactylus kimberleyensis sp. nov. A) oblique view of right side of face showing circumnarial and labial scales; B) ventral view of chin and throat showing mental and post-mental scales." httpUri="https://zenodo.org/record/215281/files/figure.png" pageId="3" pageNumber="35">Fig. 3</figureCitation>
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A). Two to three rows of small scales separate orbit from supralabials. Mental triangular with very acute posteriorly-oriented apex separating anterior 2/3 of postmentals, narrower (
|
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<quantity id="4CCFC5B9BC222D34FD02FE1A2BF1FE07" box="[653,741,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="35" unit="mm" value="1.8">1.8 mm</quantity>
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) than deep (
|
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<quantity id="4CCFC5B9BC222D34FCE0FE1A2ADEFE07" box="[879,970,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="35" unit="mm" value="2.2">2.2 mm</quantity>
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). A single pair of enlarged postmentals in contact behind mental, each bordered anteromedially by mental, anterolaterally by first infralabial, posterolaterally by an enlarged lateral chinshield (or second postmental), and posteriorly by one or two enlarged granules (
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||
<figureCitation id="130C74D9BC222D34FAB8FE522C6DFE5F" box="[1335,1401,476,500]" captionStart="FIGURE 3" captionStartId="3.[151,250,1340,1363]" captionTargetBox="[151,1436,692,1319]" captionTargetId="figure@3.[151,1436,692,1320]" captionTargetPageId="3" captionText="FIGURE 3. Head of Cyrtodactylus kimberleyensis sp. nov. A) oblique view of right side of face showing circumnarial and labial scales; B) ventral view of chin and throat showing mental and post-mental scales." httpUri="https://zenodo.org/record/215281/files/figure.png" pageId="3" pageNumber="35">Fig. 3</figureCitation>
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B). Supralabials to midorbital position 7 (right) to 8 (left); enlarged supralabials to angle of jaws 9 (left) to 10 (right). Infralabials 9 (right) to 10 (left). Interorbital scale rows across narrowest point of frontal bone 11; 33 scales between left and right supraciliary rows.
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</paragraph>
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<caption id="DF4838D4BC222D34FF18FAB22D04FADA" httpUri="https://zenodo.org/record/215281/files/figure.png" pageId="3" pageNumber="35" targetBox="[151,1436,692,1319]" targetPageId="3">
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<paragraph id="8B88685CBC222D34FF18FAB22D04FADA" blockId="3.[151,1435,1340,1394]" pageId="3" pageNumber="35">
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<emphasis id="B943B44EBC222D34FF18FAB22800FAF9" bold="true" box="[151,276,1340,1363]" pageId="3" pageNumber="35">FIGURE 3.</emphasis>
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Head of
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<taxonomicName id="4C3713DFBC222D34FEF7FAB32BB0FAF8" box="[376,676,1341,1363]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="35" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
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<emphasis id="B943B44EBC222D34FEF7FAB32BB0FAF8" box="[376,676,1341,1363]" italics="true" pageId="3" pageNumber="35">Cyrtodactylus kimberleyensis</emphasis>
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</taxonomicName>
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<emphasis id="B943B44EBC222D34FD22FAB32A14FAF8" bold="true" box="[685,768,1341,1363]" pageId="3" pageNumber="35">
|
||
<taxonomicNameLabel id="A2700935BC222D34FD22FAB32A14FAF8" box="[685,768,1341,1363]" pageId="3" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) oblique view of right side of face showing circumnarial and labial scales; B) ventral view of chin and throat showing mental and post-mental scales.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B88685CBC222D34FF48FA392C5FF944" blockId="3.[151,1436,1463,2028]" pageId="3" pageNumber="35">
|
||
Body slender, elongate (TrunkL/SVL = 0.42) lacking well defined ventrolateral folds, although small discrete folds clearly present in life. Dorsal scales largely homogeneous, conical with slightly posteriorly-oriented apices; somewhat irregularly distributed tubercles (2–6 times size of adjacent scales), each wider than long, extending from shoulder region on to tail base, smallest on flanks, largest over sacrum, smaller tubercles on postocular region, crown, occiput, and nape; most dorsal tubercles bearing a keel, those on flanks conical, often lacking a distinct keel, those on posterior trunk and sacral region most prominent; tubercles in 16–18 rows at midbody, typically separated from one another by 1–3 dorsal granules (
|
||
<figureCitation id="130C74D9BC222D34FD51F9012A30F90C" box="[734,804,1679,1703]" captionStart="FIGURE 2" captionStartId="2.[151,250,1470,1493]" captionTargetBox="[151,1436,621,1449]" captionTargetId="figure@2.[151,1436,621,1449]" captionTargetPageId="2" captionText="FIGURE 2. Adult female holotype of Cyrtodactylus kimberleyensis sp. nov. (WAM R 164144). Scale bar = 10 mm." httpUri="https://zenodo.org/record/215280/files/figure.png" pageId="3" pageNumber="35">Fig. 2</figureCitation>
|
||
). Ventral scales larger than dorsals, smooth, oval to subtriangular and subimbricate, largest on posterior abdomen and in precloacal region. Midbody scale rows across belly to lowest rows of tubercles ~36. Gular region with homogeneous, smooth, juxtaposed granular scales.
|
||
</paragraph>
|
||
<paragraph id="8B88685CBC222D34FF48F9722BB8F8F0" blockId="3.[151,1436,1463,2028]" pageId="3" pageNumber="35">No precloacal or femoral pores. No precloacal groove. Anteroventral scales of thigh much larger than posteroventral and posterior, but lacking a distinct single row of enlarged femoral scales. Two slightly enlarged, smooth postcloacal spurs, anterior larger than posterior.</paragraph>
|
||
<paragraph id="8B88685CBC222D33FF48F8E92BD5FE07" blockId="3.[151,1436,1463,2028]" lastBlockId="4.[151,1437,151,428]" lastPageId="4" lastPageNumber="36" pageId="3" pageNumber="35">
|
||
Scales on palm and sole smooth, rounded to oval, flattened to slightly domed. Scalation on dorsal surfaces of limbs similar to body dorsum with enlarged, conical, usually keeled tubercles interspersed among smaller scales (except for proximal portion of forearm, which lacks tubercles); tubercles separated from one another by 1–2 small scales, or in direct contact with one another. Fore and hindlimbs moderately short, slender (ForeaL/SVL = 0.12; CrusL/SVL = 0.14). Digits long, slender, inflected at interphalangeal joints, especially antepenultimate joint of longer digits, all bearing robust, slightly recurved claws. Basal subdigital lamellae broad, squarish to rectangular with rounded corners, distalmost generally largest, without scansorial surfaces (1-3-4-5-3 left manus, 1-4-4-5-3 right manus; 2-4-5-6-5 left and right pes); narrow lamellae distal to digital inflection and not including ventral claw sheath: 6-5(tip missing)-8-7-7 (left manus), 6-7-8-8-7 (right manus), 6-7-9-9-9 (left pes), 7-7-9-9-7 (right pes) (
|
||
<figureCitation id="130C74D9BC252D33FAE1FEA929BCFECF" captionStart="FIGURE 4" captionStartId="4.[151,250,1306,1329]" captionTargetBox="[369,1154,517,1267]" captionTargetId="figure@4.[354,1233,517,1285]" captionTargetPageId="4" captionText="FIGURE 4. Ventral view of right pes of Cyrtodactylus kimberleyensis sp. nov. showing disposition of the subdigital lamellae. Roman numerals indicate digit numbers. Scale bar = 2 mm." httpUri="https://zenodo.org/record/215282/files/figure.png" pageId="4" pageNumber="36">Fig. 4</figureCitation>
|
||
); very weakly developed interdigital webbing between digits (except IV and V). Relative length of digits: IV>III>II>V>I (manus); V~IV~III>II>I (pes). Mostly original tail, long, slightly depressed, gently tapering to pointed tip; longer than SVL (TailL/SVL = 1.18).
|
||
</paragraph>
|
||
<caption id="DF4838D4BC252D33FF18FA942BE7FAE4" httpUri="https://zenodo.org/record/215282/files/figure.png" pageId="4" pageNumber="36" targetBox="[369,1154,517,1267]" targetPageId="4">
|
||
<paragraph id="8B88685CBC252D33FF18FA942BE7FAE4" blockId="4.[151,1437,1306,1359]" pageId="4" pageNumber="36">
|
||
<emphasis id="B943B44EBC252D33FF18FA942807FA9B" bold="true" box="[151,275,1306,1329]" pageId="4" pageNumber="36">FIGURE 4.</emphasis>
|
||
Ventral view of right pes of
|
||
<taxonomicName id="4C3713DFBC252D33FDB8FA942A77FA9B" box="[567,867,1306,1328]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="36" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC252D33FDB8FA942A77FA9B" box="[567,867,1306,1328]" italics="true" pageId="4" pageNumber="36">Cyrtodactylus kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC252D33FCE6FA952AAFFA9B" bold="true" box="[873,955,1307,1328]" pageId="4" pageNumber="36">
|
||
<taxonomicNameLabel id="A2700935BC252D33FCE6FA952AAFFA9B" box="[873,955,1307,1328]" pageId="4" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
showing disposition of the subdigital lamellae. Roman numerals indicate digit numbers. Scale bar = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B88685CBC252D33FF48FA112C51F9C7" blockId="4.[151,1437,1439,2004]" pageId="4" pageNumber="36">Scales of tail dorsum squarish, flat, juxtaposed to weakly imbricate, arranged in distinct segments of 8–9 scale rows basally, decreasing to 7, then 6 scale rows distally. Pygal segments bearing 6 strongly keeled tubercles, each 5–6 times the size of adjacent scales and separated from each other by 1–2 scales, in a transverse row; next three tail segments with four tubercles, followed by a single segment with two enlarged, flattened, keeled scales in position of tubercles, but on left side only; no tubercles more distally. Subcaudals approximately 1.5 times size of dorsal caudals, squarish proximally, becoming rounded to oval distally. No enlarged median subcaudal plates.</paragraph>
|
||
<paragraph id="8B88685CBC252D33FF48F9F62C5CF91F" blockId="4.[151,1437,1439,2004]" pageId="4" pageNumber="36">Measurements (in mm): SVL 45.1, ForeaL 5.6, CrusL 6.1, TailL 53.3, TailW 3.5, TrunkL 19.1, HeadL 13.0, HeadW 7.5, HeadH 4.4, OrbD 3.2, EyeEar 4.5, SnEye 5.5, NarEye 3.5, Interorb 4.5, EarL 1.3, Internar 1.2.</paragraph>
|
||
<paragraph id="8B88685CBC252D33FF48F9312A26F8CC" blockId="4.[151,1437,1439,2004]" pageId="4" pageNumber="36">Coloration in ethanol: Dorsum light brown mottled with small darker blotches and tubercles. A series of small rounded white blotches on lower flanks, neck and lower part of side of head. Two dark blotches on occiput and a broken, heart-shaped marking on the crown. A broad bold dark brown streak from posterior midpoint of orbit to above ear, confluent with a narrower incomplete stripe continuing on to shoulder. A pair of dark brown canthal streaks and several dark stray marks on other areas of head.</paragraph>
|
||
<paragraph id="8B88685CBC252D32FF48F8FA2AE2FF7F" blockId="4.[151,1437,1439,2004]" lastBlockId="5.[151,1436,151,391]" lastPageId="5" lastPageNumber="37" pageId="4" pageNumber="36">Rostral mostly medium brown. Labials mostly brown with some scales bearing white patches. Limbs similar to dorsum, light brown with darker mottling and distinct white blotches. Digits with alternating light and dark markings. Soles and palms pale grayish brown. Tail light brown with a pair of paravertebral dark brown stripes proximally breaking into paired dashes at approximately midpoint of tail. Distal 70% of tail with alternating white to cream and mid-brown markings. Regenerated tail tip without regular pattern.</paragraph>
|
||
<paragraph id="8B88685CBC242D32FF48FF6E2BBBFEB7" blockId="5.[151,1436,151,391]" pageId="5" pageNumber="37">Venter beige with cream to whitish spots under throat and posterior border of thighs. Underside of tail uniform brown with white speckling along entire length.</paragraph>
|
||
<paragraph id="8B88685CBC242D32FF48FEA92C34FE2C" blockId="5.[151,1436,151,391]" pageId="5" pageNumber="37">
|
||
Color in life (based on live photograph of holotytpe,
|
||
<figureCitation id="130C74D9BC242D32FC96FEA92A49FE94" box="[793,861,295,319]" captionStart="FIGURE 5" captionStartId="5.[151,250,1437,1460]" captionTargetBox="[151,1436,447,1411]" captionTargetId="figure@5.[151,1436,447,1411]" captionTargetPageId="5" captionText="FIGURE 5. Photo in life of the holotype of Cyrtodactylus kimberleyensis sp. nov. (WAM R 164144). Photo—R. Browne-Cooper." httpUri="https://zenodo.org/record/215283/files/figure.png" pageId="5" pageNumber="37">Fig. 5</figureCitation>
|
||
): Similar to that in preservative, pale areas of body with a pale grayish to grayish-pink suffusion, and pale spots on flanks and limbs more evident than in preserved specimen. Pale coloration of tail base pale yellowish-cream to straw. Iris silvery with dark reticulations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4838D4BC242D32FF18FA1329A9FA78" httpUri="https://zenodo.org/record/215283/files/figure.png" pageId="5" pageNumber="37" targetBox="[151,1436,447,1411]" targetPageId="5">
|
||
<paragraph id="8B88685CBC242D32FF18FA1329A9FA78" blockId="5.[151,1436,1437,1491]" pageId="5" pageNumber="37">
|
||
<emphasis id="B943B44EBC242D32FF18FA132806FA18" bold="true" box="[151,274,1437,1460]" pageId="5" pageNumber="37">FIGURE 5.</emphasis>
|
||
Photo in life of the holotype of
|
||
<taxonomicName id="4C3713DFBC242D32FDD8FA132A96FA18" box="[599,898,1437,1459]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC242D32FDD8FA132A96FA18" box="[599,898,1437,1459]" italics="true" pageId="5" pageNumber="37">Cyrtodactylus kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC242D32FC06FA102ACEFA18" bold="true" box="[905,986,1438,1459]" pageId="5" pageNumber="37">
|
||
<taxonomicNameLabel id="A2700935BC242D32FC06FA102ACEFA18" box="[905,986,1438,1459]" pageId="5" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(WAM R164144). Photo—R. Browne-Cooper.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32D3BD7BC242D32FF48F9912DF3F99D" box="[199,1255,1566,1591]" pageId="5" pageNumber="37" type="etymology">
|
||
<paragraph id="8B88685CBC242D32FF48F9912DF3F99D" blockId="5.[151,1436,1566,2023]" box="[199,1255,1566,1591]" pageId="5" pageNumber="37">
|
||
<emphasis id="B943B44EBC242D32FF48F991285AF99C" bold="true" box="[199,334,1567,1591]" pageId="5" pageNumber="37">Etymology.</emphasis>
|
||
The specific epithet refers to the Kimberley region of northern Western
|
||
<collectingCountry id="F32028CCBC242D32FBF7F9902DF7F99D" box="[1144,1251,1566,1590]" name="Australia" pageId="5" pageNumber="37">Australia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D3BD7BC242D30FF48F9CC2C1CFAA7" lastPageId="7" lastPageNumber="39" pageId="5" pageNumber="37" type="distribution">
|
||
<paragraph id="8B88685CBC242D32FF48F9CC28FEF84D" blockId="5.[151,1436,1566,2023]" pageId="5" pageNumber="37">
|
||
<emphasis id="B943B44EBC242D32FF48F9CC2B8DF9F0" bold="true" box="[199,665,1602,1627]" pageId="5" pageNumber="37">Distribution, reproduction and habitat.</emphasis>
|
||
The only known specimen of
|
||
<taxonomicName id="4C3713DFBC242D32FC65F9CD2DA4F9F1" box="[1002,1200,1602,1626]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC242D32FC65F9CD2DA4F9F1" box="[1002,1200,1602,1626]" italics="true" pageId="5" pageNumber="37">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC242D32FB39F9CD2C1BF9F0" bold="true" box="[1206,1295,1603,1627]" pageId="5" pageNumber="37">
|
||
<taxonomicNameLabel id="A2700935BC242D32FB39F9CD2C1BF9F0" box="[1206,1295,1603,1627]" pageId="5" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is from East Montalivet Island (
|
||
<figureCitation id="130C74D9BC242D32FEE0F9E828A5F9D5" box="[367,433,1638,1662]" captionStart="FIGURE 1" captionStartId="1.[151,250,1267,1290]" captionTargetBox="[159,1426,510,1248]" captionTargetId="figure@1.[151,1436,505,1254]" captionTargetPageId="1" captionText="FIGURE 1. Map of the Kimberley region, Western Australia, showing East Montalivet Island (arrow), the type locality of Cyrtodactylus kimberleyensis sp. nov. was collected." httpUri="https://zenodo.org/record/215279/files/figure.png" pageId="5" pageNumber="37">Fig. 1</figureCitation>
|
||
), although it is possible the taxon occurs on the Kimberley mainland as well. The
|
||
<typeStatus id="548CD6FEBC242D32FAEEF9E829D3F908" pageId="5" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
is a gravid female containing a single egg measuring
|
||
<quantity id="4CCFC5B9BC242D32FCA0F9042A93F909" box="[815,903,1674,1698]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="5" pageNumber="37" unit="mm" value="9.4">9.4 mm</quantity>
|
||
x
|
||
<quantity id="4CCFC5B9BC242D32FC2AF9042AEBF909" box="[933,1023,1674,1698]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="37" unit="mm" value="5.3">5.3 mm</quantity>
|
||
. Like most gekkotans
|
||
<taxonomicName id="4C3713DFBC242D32FB70F9042C88F909" box="[1279,1436,1674,1698]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC242D32FB70F9042C88F909" box="[1279,1436,1674,1698]" italics="true" pageId="5" pageNumber="37">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
spp. typically produce two eggs per clutch. Single egg clutches are uncommon in the
|
||
<taxonomicName id="4C3713DFBC242D32FBEBF92029CEF941" authority="Marquet et al. 1990" authorityName="Marquet et al." authorityYear="1990" class="Reptilia" family="Gekkonidae" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="family">
|
||
Gekkonidae (
|
||
<bibRefCitation id="EFA615ADBC242D32FB74F92029C7F941" author="Marquet" pageId="5" pageNumber="37" refString="Marquet, P. A., Bozinovic, F., Medel, R. G., Werner, Y. L. & Jaksic, F. M. (1990) Ecology of Garthia gaudichaudi, a gecko endemic to the semiarid region of Chile. Journal of Herpetology, 24, 431 - 434." type="journal article" year="1990">Marquet et al. 1990</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but do occur, chiefly in miniaturized lineages. If a single egg clutch is normal for
|
||
<taxonomicName id="4C3713DFBC242D32FBFAF95D2C2FF941" box="[1141,1339,1746,1770]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC242D32FBFAF95D2C2FF941" box="[1141,1339,1746,1770]" italics="true" pageId="5" pageNumber="37">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC242D32FACCF95D2C88F940" bold="true" box="[1347,1436,1747,1771]" pageId="5" pageNumber="37">
|
||
<taxonomicNameLabel id="A2700935BC242D32FACCF95D2C88F940" box="[1347,1436,1747,1771]" pageId="5" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it would be the first member of its genus known to do so. The specimen was collected in late April, very late in the tropical wet season, and it is possible the female had produced other clutches earlier in the season, as occurs in many other tropical geckos (
|
||
<bibRefCitation id="EFA615ADBC242D32FE5BF8B02BF8F8FD" author="Inger" box="[468,748,1854,1878]" pageId="5" pageNumber="37" refString="Inger, R. F. & Greenberg, B. (1966) Annual reproductive patterns of lizards from a Bornean rainforest. Ecology, 47, 1007 - 1021." type="journal article" year="1966">Inger & Greenberg 1966</bibRefCitation>
|
||
). The
|
||
<typeStatus id="548CD6FEBC242D32FCBEF8B02A87F8FD" box="[817,915,1854,1878]" pageId="5" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
was collected from a vine thicket on a lateritic slope (R. Browne-Cooper, pers. comm.;
|
||
<figureCitation id="130C74D9BC242D32FDD2F8EC2BB4F8D1" box="[605,672,1890,1914]" captionStart="FIGURE 6" captionStartId="6.[151,250,1867,1890]" captionTargetBox="[151,1436,879,1843]" captionTargetId="figure@6.[151,1436,879,1843]" captionTargetPageId="6" captionText="FIGURE 6. Vine thicket habitat of Cyrtodactylus kimberleyensis sp. nov. on East Montalivet Island, Western Australia. Photo — M. Bamford." httpUri="https://zenodo.org/record/215284/files/figure.png" pageId="5" pageNumber="37">Fig. 6</figureCitation>
|
||
). Other reptiles also occurring on the island are the gekkotan lizards
|
||
<taxonomicName id="4C3713DFBC242D32FF18F80928D5F835" authority="Gray" authorityName="Gray" box="[151,449,1926,1950]" class="Reptilia" family="Gekkonidae" genus="Heteronotia" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="binoei">
|
||
<emphasis id="B943B44EBC242D32FF18F8092878F835" box="[151,364,1926,1950]" italics="true" pageId="5" pageNumber="37">Heteronotia binoei</emphasis>
|
||
(Gray)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3713DFBC242D32FE73F8082BF4F835" authority="Kluge" authorityName="Kluge" box="[508,736,1926,1950]" class="Reptilia" family="Pygopodidae" genus="Delma" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="borea">
|
||
<emphasis id="B943B44EBC242D32FE73F8082B9BF835" box="[508,655,1926,1950]" italics="true" pageId="5" pageNumber="37">Delma borea</emphasis>
|
||
Kluge
|
||
</taxonomicName>
|
||
, the skinks
|
||
<taxonomicName id="4C3713DFBC242D32FCE4F8082D61F835" authority="Storr" authorityName="Storr" box="[875,1141,1926,1950]" class="Reptilia" family="Scincidae" genus="Carlia" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="johnstonei">
|
||
<emphasis id="B943B44EBC242D32FCE4F8082D26F835" box="[875,1074,1926,1950]" italics="true" pageId="5" pageNumber="37">Carlia johnstonei</emphasis>
|
||
Storr
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC242D32FB0DF8092C8CF835" authority="Mitchell" authorityName="Mitchell" box="[1154,1432,1926,1950]" class="Reptilia" family="Scincidae" genus="Carlia" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="triacantha">
|
||
<emphasis id="B943B44EBC242D32FB0DF8092C0EF835" box="[1154,1306,1926,1950]" italics="true" pageId="5" pageNumber="37">C. triacantha</emphasis>
|
||
(Mitchell)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC242D32FF18F82528D6F869" authority="Gray" authorityName="Gray" box="[151,450,1962,1986]" class="Reptilia" family="Scincidae" genus="Ctenotus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="inornatus">
|
||
<emphasis id="B943B44EBC242D32FF18F825287AF869" box="[151,366,1963,1986]" italics="true" pageId="5" pageNumber="37">Ctenotus inornatus</emphasis>
|
||
(Gray)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC242D32FE41F8252A75F869" authority="Boulenger" authorityName="Boulenger" box="[462,865,1962,1986]" class="Reptilia" family="Scincidae" genus="Eremiascincus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="isolepis">
|
||
<emphasis id="B943B44EBC242D32FE41F8252BDBF869" box="[462,719,1962,1986]" italics="true" pageId="5" pageNumber="37">Eremiascincus isolepis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC242D32FCE2F8252DBEF869" authority="Boulenger" authorityName="Boulenger" box="[877,1194,1962,1986]" class="Reptilia" family="Scincidae" genus="Lerista" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="walkeri">
|
||
<emphasis id="B943B44EBC242D32FCE2F8252D0CF869" box="[877,1048,1962,1986]" italics="true" pageId="5" pageNumber="37">Lerista walkeri</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
, and the elapid snake
|
||
<taxonomicName id="4C3713DFBC242D32FF18F85E28F2F84D" authority="Gunther" authorityName="Gunther" box="[151,486,1998,2023]" class="Reptilia" family="Elapidae" genus="Parasuta" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="37" phylum="Chordata" rank="species" species="nigriceps">
|
||
<emphasis id="B943B44EBC242D32FF18F85E2878F84D" box="[151,364,1999,2023]" italics="true" pageId="5" pageNumber="37">Parasuta nigriceps</emphasis>
|
||
(Günther)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B88685CBC272D31FF48FF192BA7FC9C" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="38">
|
||
<emphasis id="B943B44EBC272D31FF48FF192B5EFF1B" bold="true" box="[199,586,151,176]" pageId="6" pageNumber="38">Comparisons with other species.</emphasis>
|
||
The diagnosis of
|
||
<taxonomicName id="4C3713DFBC272D31FC9BFF162AA5FF1B" box="[788,945,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC272D31FC9BFF162AA5FF1B" box="[788,945,152,176]" italics="true" pageId="6" pageNumber="38">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
spp. known only from female specimens is generally quite difficult because the presence/absence, number, and distribution of precloacal and femoral pores in males are useful and convenient characters that often distinguish species from one another (
|
||
<bibRefCitation id="EFA615ADBC272D31FB1AFF512C09FF5C" author="Bauer" box="[1173,1309,223,247]" pageId="6" pageNumber="38" refString="Bauer, A. M. (2003) Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma). Proceedings of the California Academy of Sciences, 54, 463 - 498." type="journal article" year="2003">Bauer 2003</bibRefCitation>
|
||
). However, the very small size
|
||
<taxonomicName id="4C3713DFBC272D31FEFCFE8B2B2EFEB7" box="[371,570,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC272D31FEFCFE8B2B2EFEB7" box="[371,570,260,284]" italics="true" pageId="6" pageNumber="38">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC272D31FDCCFE8A2B89FEB7" bold="true" box="[579,669,260,284]" pageId="6" pageNumber="38">
|
||
<taxonomicNameLabel id="A2700935BC272D31FDCCFE8A2B89FEB7" box="[579,669,260,284]" pageId="6" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(confirmed as an adult because the sole known specimen is gravid) easily separates it from the majority of its congeners. Most gekkotans show little sexual dimorphism in overall body length, with slightly larger females in approximately two-thirds of species investigated (
|
||
<bibRefCitation id="EFA615ADBC272D31FB37FEC22C23FECF" author="Fitch" box="[1208,1335,332,356]" pageId="6" pageNumber="38" refString="Fitch, H. S. (1981) Sexual size differences in reptiles. The University of Kansas Museum of Natural History Miscellaneous Publication, (70), [4], 1 - 72." type="journal article" year="1981">Fitch 1981</bibRefCitation>
|
||
). In only two species of
|
||
<taxonomicName id="4C3713DFBC272D31FEB0FEFE28C8FE23" box="[319,476,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC272D31FEB0FEFE28C8FE23" box="[319,476,368,392]" italics="true" pageId="6" pageNumber="38">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
are size and maturity data known for large samples (
|
||
<taxonomicName id="4C3713DFBC272D31FBBEFEFE2DDFFE23" box="[1073,1227,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B943B44EBC272D31FBBEFEFE2DDFFE23" box="[1073,1227,368,392]" italics="true" pageId="6" pageNumber="38">C. malayanus</emphasis>
|
||
</taxonomicName>
|
||
(de Rooij), n=
|
||
<specimenCount id="9D31A3D5BC272D31FAFCFEE129CFFE07" pageId="6" pageNumber="38" type="male">131 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D31A3D5BC272D31FF64FE1A2868FE07" box="[235,380,404,428]" pageId="6" pageNumber="38" type="female">162 females</specimenCount>
|
||
;
|
||
<taxonomicName id="4C3713DFBC272D31FE04FE1B2B65FE06" authority="Inger" authorityName="Inger" box="[395,625,404,429]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="pubisulcus">
|
||
<emphasis id="B943B44EBC272D31FE04FE1B2B3DFE07" box="[395,553,404,428]" italics="true" pageId="6" pageNumber="38">C. pubisulcus</emphasis>
|
||
Inger
|
||
</taxonomicName>
|
||
, n=
|
||
<specimenCount id="9D31A3D5BC272D31FD11FE1A2A1EFE07" box="[670,778,404,428]" pageId="6" pageNumber="38" type="male">56 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D31A3D5BC272D31FC95FE1A2A89FE07" box="[794,925,404,428]" pageId="6" pageNumber="38" type="female">62 females</specimenCount>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC272D31FC22FE1B2DC7FE07" author="Inger" box="[941,1235,404,429]" pageId="6" pageNumber="38" refString="Inger, R. F. & Greenberg, B. (1966) Annual reproductive patterns of lizards from a Bornean rainforest. Ecology, 47, 1007 - 1021." type="journal article" year="1966">Inger & Greenberg 1966</bibRefCitation>
|
||
). In both species females were slightly larger on average and attained somewhat greater maximum sizes than males. A review of recent descriptions of new
|
||
<taxonomicName id="4C3713DFBC272D31FE4BFE522B75FE5F" box="[452,609,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC272D31FE4BFE522B75FE5F" box="[452,609,476,500]" italics="true" pageId="6" pageNumber="38">Cyrtodactylus</emphasis>
|
||
</taxonomicName>
|
||
based on samples containing both males and females suggests no intrageneric directionality of sexual dimorphism in SVL, and no clear examples of strong size asymmetry between genders. In
|
||
<taxonomicName id="4C3713DFBC272D31FF78FDAB2886FD97" box="[247,402,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="pubisulcus">
|
||
<emphasis id="B943B44EBC272D31FF78FDAB2886FD97" box="[247,402,548,572]" italics="true" pageId="6" pageNumber="38">C. pubisulcus</emphasis>
|
||
</taxonomicName>
|
||
the minimum size at which females were gravid was 89.6% of maximum SVL and in
|
||
<taxonomicName id="4C3713DFBC272D31FA0DFDAB2804FDCB" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="malayanus">
|
||
<emphasis id="B943B44EBC272D31FA0DFDAB2804FDCB" italics="true" pageId="6" pageNumber="38">C. malayanus</emphasis>
|
||
</taxonomicName>
|
||
it was 83.1%. (
|
||
<bibRefCitation id="EFA615ADBC272D31FE33FDC92BC7FDF4" author="Inger" box="[444,723,583,607]" pageId="6" pageNumber="38" refString="Inger, R. F. & Greenberg, B. (1966) Annual reproductive patterns of lizards from a Bornean rainforest. Ecology, 47, 1007 - 1021." type="journal article" year="1966">Inger & Greenberg 1966</bibRefCitation>
|
||
). Most studies of size at maturity in geckos suggest that females reach maturity at 73–82% of maximum size (
|
||
<bibRefCitation id="EFA615ADBC272D31FD14FDE22A3DFD2F" author="Parker" box="[667,809,620,644]" pageId="6" pageNumber="38" refString="Parker, W. S. (1972) Aspects of the ecology of a Sonoran Desert population of the western banded gecko, Coleonyx variegatus (Sauria, Eublepharinae). American Midland Naturalist, 88, 209 - 224." type="journal article" year="1972">Parker 1972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC272D31FCB8FDE22AB0FD2F" author="Vitt" box="[823,932,620,644]" pageId="6" pageNumber="38" refString="Vitt, L. J. (1986) Reproductive tactics of sympatric gekkonid lizards with a comment on the evolutionary and ecological consequences of invariant clutch size. Copeia, 1986, 773 - 786." type="journal article" year="1986">Vitt 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC272D31FC3EFDE32D79FD2F" author="How" box="[945,1133,620,644]" pageId="6" pageNumber="38" refString="How, R. A., Dell, J. & Wellington, B. D. (1986) Comparative biology of eight species of Diplodactylus gecko in western Australia. Herpetologica, 42, 471 - 482." type="journal article" year="1986">
|
||
How
|
||
<emphasis id="B943B44EBC272D31FC7FFDE32D37FD2F" box="[1008,1059,620,644]" italics="true" pageId="6" pageNumber="38">et al</emphasis>
|
||
. 1986
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC272D31FBF5FDE22C10FD2F" author="Selcer" box="[1146,1284,620,644]" pageId="6" pageNumber="38" refString="Selcer, K. W. (1986) Life history of a successful colonizer: the Mediterranean gecko, Hemidactylus turcicus, in southern Texas. Copeia, 1986, 956 - 962." type="journal article" year="1986">Selcer 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC272D31FA9EFDE229C5FD0C" author="Okada" pageId="6" pageNumber="38" refString="Okada, S., Izawa, M. & Ota, H. (2002) Growth and Reproduction of Gekko hokouensis (Reptilia: Squamata) on Okinawajima Island of the Ryukyu Archipelago, Japan. Journal of Herpetology, 36, 473 - 479." type="journal article" year="2002">
|
||
Okada
|
||
<emphasis id="B943B44EBC272D31FAEDFDE32C81FD2F" box="[1378,1429,620,644]" italics="true" pageId="6" pageNumber="38">et al</emphasis>
|
||
. 2002
|
||
</bibRefCitation>
|
||
) although there are some genera in which sexual maturity may be reached at smaller relative sizes (e.g.,
|
||
<taxonomicName id="4C3713DFBC272D31FF18FD3B2B3AFD67" authority="Gray, Hibbitts et al. 2005" authorityName="Gray, Hibbitts et al." authorityYear="2005" box="[151,558,692,716]" class="Reptilia" family="Gekkonidae" genus="Ptenopus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC272D31FF18FD3B29EBFD67" box="[151,255,693,716]" italics="true" pageId="6" pageNumber="38">Ptenopus</emphasis>
|
||
Gray,
|
||
<bibRefCitation id="EFA615ADBC272D31FEC2FD3A2B3AFD67" author="Hibbitts" box="[333,558,692,716]" pageId="6" pageNumber="38" refString="Hibbitts, T. J., Pianka, E. R., Huey, R. B. & Whiting, M. J. (2005) Ecology of the common barking gecko (Ptenopus garrulus) in southern Africa. Journal of Herpetology, 39, 509 - 515." type="journal article" year="2005">
|
||
Hibbitts
|
||
<emphasis id="B943B44EBC272D31FE3FFD3B28F6FD67" box="[432,482,692,716]" italics="true" pageId="6" pageNumber="38">et al</emphasis>
|
||
. 2005
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C3713DFBC272D31FDB4FD3B2D61FD67" authority="Gray, Ibarguengoytia & Casalins 2007" authorityName="Gray, Ibarguengoytia & Casalins" authorityYear="2007" box="[571,1141,692,716]" class="Reptilia" family="Phyllodactylidae" genus="Homonota" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC272D31FDB4FD3B2BA5FD67" box="[571,689,693,716]" italics="true" pageId="6" pageNumber="38">Homonota</emphasis>
|
||
Gray,
|
||
<bibRefCitation id="EFA615ADBC272D31FC8FFD3A2D61FD67" author="Ibarguengoytia" box="[768,1141,692,716]" pageId="6" pageNumber="38" refString="Ibarguengoytia, N. R. & Casalins, L. M. (2007) Reproductive biology of the southernmost gecko Homonota darwini: convergent life-history patterns among Southern Hemisphere reptiles living in harsh environments. Journal of Herpetology, 41, 72 - 80." type="journal article" year="2007">Ibargüengoytía & Casalins 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
). These values may, however, be influenced by the criteria used to determine maturity. For example, in
|
||
<taxonomicName id="4C3713DFBC272D31FBCEFD562C88FD44" authority="Smith" authorityName="Smith" box="[1089,1436,727,752]" class="Reptilia" family="Gekkonidae" genus="Homopholis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="wahlbergii">
|
||
<emphasis id="B943B44EBC272D31FBCEFD562C5AFD5B" box="[1089,1358,728,752]" italics="true" pageId="6" pageNumber="38">Homopholis wahlbergii</emphasis>
|
||
Smith
|
||
</taxonomicName>
|
||
females had muscular oviducts at <50% of maximum size, but fully developed eggs were only found in females>71.4% of maximum size (
|
||
<bibRefCitation id="EFA615ADBC272D31FE48FC912BB3FC9C" author="Whiting" box="[455,679,799,824]" pageId="6" pageNumber="38" refString="Whiting, M. J., Reaney, L. T. & Keogh, J. S. (2007) Ecology of Wahlberg's velvet gecko, Homopholis wahlbergii, in southern Africa. African Zoology, 42, 38 - 44." type="journal article" year="2007">
|
||
Whiting
|
||
<emphasis id="B943B44EBC272D31FDA4FCAF2B4FFC93" box="[555,603,800,824]" italics="true" pageId="6" pageNumber="38">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4838D4BC272D31FF18F8C52820F82B" httpUri="https://zenodo.org/record/215284/files/figure.png" pageId="6" pageNumber="38" targetBox="[151,1436,879,1843]" targetPageId="6">
|
||
<paragraph id="8B88685CBC272D31FF18F8C52820F82B" blockId="6.[151,1436,1867,1920]" pageId="6" pageNumber="38">
|
||
<emphasis id="B943B44EBC272D31FF18F8C52807F8CA" bold="true" box="[151,275,1867,1890]" pageId="6" pageNumber="38">FIGURE 6.</emphasis>
|
||
Vine thicket habitat of
|
||
<taxonomicName id="4C3713DFBC272D31FD8DF8C22A38F8C9" box="[514,812,1868,1890]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC272D31FD8DF8C22A38F8C9" box="[514,812,1868,1890]" italics="true" pageId="6" pageNumber="38">Cyrtodactylus kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC272D31FCBBF8C22A91F8CA" bold="true" box="[820,901,1868,1889]" pageId="6" pageNumber="38">
|
||
<taxonomicNameLabel id="A2700935BC272D31FCBBF8C22A91F8CA" box="[820,901,1868,1889]" pageId="6" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
on East Montalivet Island, Western Australia. Photo —M. Bamford.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B88685CBC272D30FF48F8232C1CFAA7" blockId="6.[151,1436,1965,2025]" lastBlockId="7.[151,1437,151,1292]" lastPageId="7" lastPageNumber="39" pageId="6" pageNumber="38">
|
||
Based on data from other geckos one may thus conservatively estimate that the
|
||
<typeStatus id="548CD6FEBC272D31FBC3F8232DBAF86E" box="[1100,1198,1965,1989]" pageId="6" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C3713DFBC272D31FB5BF8202C88F86E" box="[1236,1436,1965,1989]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="38" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC272D31FB5BF8202C88F86E" box="[1236,1436,1965,1989]" italics="true" pageId="6" pageNumber="38">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC272D31FF18F85F29E4F842" bold="true" box="[151,240,2001,2025]" pageId="6" pageNumber="38">
|
||
<taxonomicNameLabel id="A2700935BC272D31FF18F85F29E4F842" box="[151,240,2001,2025]" pageId="6" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is at least 70% of the maximum size the species might attain. Of named species summarized by
|
||
<bibRefCitation id="EFA615ADBC272D30FAABF85F283CFF04" author="Rosler" lastPageId="7" lastPageNumber="39" pageId="6" pageNumber="38" refString="Rosler, H. & Glaw, F. (2008) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa, 1729, 8 - 22." type="journal article" year="2008">Rösler and Glaw (2008)</bibRefCitation>
|
||
, only the following species had maximum SVLs of less than
|
||
<quantity id="4CCFC5B9BC262D30FC59FF192D32FF04" box="[982,1062,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="39" unit="mm" value="65.0">65 mm</quantity>
|
||
, the maximum size of
|
||
<taxonomicName id="4C3713DFBC262D30FAAAFF1629E4FF7F" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC262D30FAAAFF1629E4FF7F" italics="true" pageId="7" pageNumber="39">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC262D30FF78FF322846FF7F" bold="true" box="[247,338,188,212]" pageId="7" pageNumber="39">
|
||
<taxonomicNameLabel id="A2700935BC262D30FF78FF322846FF7F" box="[247,338,188,212]" pageId="7" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
so predicted:
|
||
<taxonomicName id="4C3713DFBC262D30FE7DFF332A42FF7F" authority="Annandale" authorityName="Annandale" box="[498,854,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="consobrinoides">
|
||
<emphasis id="B943B44EBC262D30FE7DFF332BABFF7F" box="[498,703,188,212]" italics="true" pageId="7" pageNumber="39">C. consobrinoides</emphasis>
|
||
(Annandale)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FCEDFF332AA6FF7F" box="[866,946,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="feae">
|
||
<emphasis id="B943B44EBC262D30FCEDFF332AA6FF7F" box="[866,946,188,212]" italics="true" pageId="7" pageNumber="39">C. feae</emphasis>
|
||
</taxonomicName>
|
||
(Boulenger,
|
||
<taxonomicName id="4C3713DFBC262D30FBCAFF332C26FF7F" authority="Bauer" authorityName="Bauer" box="[1093,1330,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="annandalei">
|
||
<emphasis id="B943B44EBC262D30FBCAFF332DF1FF7F" box="[1093,1253,188,212]" italics="true" pageId="7" pageNumber="39">C. annandalei</emphasis>
|
||
Bauer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FAB2FF3329C8FF5C" authority="Bauer" authorityName="Bauer" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="gansi">
|
||
<emphasis id="B943B44EBC262D30FAB2FF332C88FF7F" box="[1341,1436,189,212]" italics="true" pageId="7" pageNumber="39">C. gansi</emphasis>
|
||
Bauer
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3713DFBC262D30FE97FF6E28BAFF53" box="[280,430,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="wakeorum">
|
||
<emphasis id="B943B44EBC262D30FE97FF6E28BAFF53" box="[280,430,224,248]" italics="true" pageId="7" pageNumber="39">C. wakeorum</emphasis>
|
||
</taxonomicName>
|
||
Bauer—all from
|
||
<collectingCountry id="F32028CCBC262D30FDF8FF6E2BF2FF53" box="[631,742,224,248]" name="Myanmar" pageId="7" pageNumber="39">Myanmar</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FD7DFF6E2AEFFF5C" authority="Bleeker" authorityName="Bleeker" box="[754,1019,223,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="B943B44EBC262D30FD7DFF6E2A9DFF53" box="[754,905,224,248]" italics="true" pageId="7" pageNumber="39">C. agamensis</emphasis>
|
||
(Bleeker)
|
||
</taxonomicName>
|
||
from Sumatra,
|
||
<taxonomicName id="4C3713DFBC262D30FB21FF6E29A0FEB7" authority="David et al." authorityName="David et al." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="buchardi">
|
||
<emphasis id="B943B44EBC262D30FB21FF6E2C27FF53" box="[1198,1331,224,248]" italics="true" pageId="7" pageNumber="39">C. buchardi</emphasis>
|
||
David
|
||
<emphasis id="B943B44EBC262D30FA07FF6F29A0FEB7" italics="true" pageId="7" pageNumber="39">et al.</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F32028CCBC262D30FF78FE8B2824FEB7" box="[247,304,261,284]" name="Laos" pageId="7" pageNumber="39">Laos</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FEB5FE8B2B69FEB7" authority="Annandale" authorityName="Annandale" box="[314,637,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="gubernatoris">
|
||
<emphasis id="B943B44EBC262D30FEB5FE8B28FFFEB7" box="[314,491,260,284]" italics="true" pageId="7" pageNumber="39">C. gubernatoris</emphasis>
|
||
(Annandale)
|
||
</taxonomicName>
|
||
from the eastern Himalayas,
|
||
<taxonomicName id="4C3713DFBC262D30FC4BFE8B2DDCFEB7" authority="Darevsky" authorityName="Darevsky" box="[964,1224,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="laevigatus">
|
||
<emphasis id="B943B44EBC262D30FC4BFE8B2D43FEB7" box="[964,1111,260,284]" italics="true" pageId="7" pageNumber="39">C. laevigatus</emphasis>
|
||
Darevsky
|
||
</taxonomicName>
|
||
from Komodo and
|
||
<collectingCountry id="F32028CCBC262D30FF18FEA929F4FE94" box="[151,224,295,319]" name="Indonesia" pageId="7" pageNumber="39">Flores</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FF65FEA62B2DFE94" authority="Constable" authorityName="Constable" box="[234,569,295,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="malcolmsmithi">
|
||
<emphasis id="B943B44EBC262D30FF65FEA628A4FEEB" box="[234,432,296,320]" italics="true" pageId="7" pageNumber="39">C. malcolmsmithi</emphasis>
|
||
(Constable)
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingCountry id="F32028CCBC262D30FD6EFEA92A09FE94" box="[737,797,295,319]" name="India" pageId="7" pageNumber="39">India</collectingCountry>
|
||
, and several species in the subgenus
|
||
<taxonomicName id="4C3713DFBC262D30FB36FEA62C3FFEEB" box="[1209,1323,296,320]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC262D30FB36FEA62C3FFEEB" box="[1209,1323,296,320]" italics="true" pageId="7" pageNumber="39">Geckoella</emphasis>
|
||
</taxonomicName>
|
||
occurring in peninsular
|
||
<collectingCountry id="F32028CCBC262D30FEA1FEC2287CFECF" box="[302,360,332,356]" name="India" pageId="7" pageNumber="39">India</collectingCountry>
|
||
and
|
||
<collectingCountry id="F32028CCBC262D30FE12FEC22B1AFECF" box="[413,526,332,356]" name="Sri Lanka" pageId="7" pageNumber="39">Sri Lanka</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C3713DFBC262D30FD97FEC22AA7FECF" authority="Boulenger" authorityName="Boulenger" box="[536,947,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="jellesmae">
|
||
<emphasis id="B943B44EBC262D30FD97FEC22A31FECF" box="[536,805,332,356]" italics="true" pageId="7" pageNumber="39">Cyrtodactylus jellesmae</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
from Sulawesi was reported to have a maximum SVL of
|
||
<quantity id="4CCFC5B9BC262D30FEB6FEE1289CFE2C" box="[313,392,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="39" unit="mm" value="63.0">63 mm</quantity>
|
||
by
|
||
<bibRefCitation id="EFA615ADBC262D30FE3EFEE12BB8FE2C" author="Rosler" box="[433,684,367,391]" pageId="7" pageNumber="39" refString="Rosler, H. & Glaw, F. (2008) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa, 1729, 8 - 22." type="journal article" year="2008">Rösler & Glaw (2008)</bibRefCitation>
|
||
but adults are now known to reach at least
|
||
<quantity id="4CCFC5B9BC262D30FB00FEE12DCAFE2C" box="[1167,1246,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="7" pageNumber="39" unit="mm" value="75.0">75 mm</quantity>
|
||
SVL (Linkem
|
||
<emphasis id="B943B44EBC262D30FA07FEFF29B9FE07" italics="true" pageId="7" pageNumber="39">et al</emphasis>
|
||
. 2008). All descriptions published since the summary of
|
||
<bibRefCitation id="EFA615ADBC262D30FCB1FE1A2D46FE07" author="Rosler" box="[830,1106,404,428]" pageId="7" pageNumber="39" refString="Rosler, H. & Glaw, F. (2008) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa, 1729, 8 - 22." type="journal article" year="2008">Rösler and Glaw (2008)</bibRefCitation>
|
||
were consulted and the only additional taxon with a mature SVL of less than
|
||
<quantity id="4CCFC5B9BC262D30FD58FE392A3EFE64" box="[727,810,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="39" unit="mm" value="65.0">65 mm</quantity>
|
||
subsequently described is
|
||
<taxonomicName id="4C3713DFBC262D30FBEBFE3629EBFE5F" authority="Mahony. However" authorityName="Mahony. However" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="mandalayensis">
|
||
<emphasis id="B943B44EBC262D30FBEBFE362C3AFE7B" box="[1124,1326,440,464]" italics="true" pageId="7" pageNumber="39">C. mandalayensis</emphasis>
|
||
Mahony. However
|
||
</taxonomicName>
|
||
, this description was based on a single, apparently immature specimen of
|
||
<quantity id="4CCFC5B9BC262D30FBB7FE522D8FFE5F" box="[1080,1179,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="6.17" pageId="7" pageNumber="39" unit="mm" value="61.7">61.7 mm</quantity>
|
||
SVL, implying a maximum adult size of>
|
||
<quantity id="4CCFC5B9BC262D30FEF3FE7128E0FDBC" box="[380,500,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="7" pageNumber="39" unit="mm" value="80.0" valueMax="85.0" valueMin="75.0">75–85 mm</quantity>
|
||
SVL. The new species may be distinguished from
|
||
<taxonomicName id="4C3713DFBC262D30FBA4FD8E2D89FDB3" box="[1067,1181,512,536]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="genus">
|
||
<emphasis id="B943B44EBC262D30FBA4FD8E2D89FDB3" box="[1067,1181,512,536]" italics="true" pageId="7" pageNumber="39">Geckoella</emphasis>
|
||
</taxonomicName>
|
||
spp. by its longer toes, more depressed habitus, tail longer than SVL, and tubercular dorsum. By the absence of enlarged caudal midventral plates it may be distinguished from
|
||
<taxonomicName id="4C3713DFBC262D30FDDEFDC62BE4FDCB" box="[593,752,584,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="annandalei">
|
||
<emphasis id="B943B44EBC262D30FDDEFDC62BE4FDCB" box="[593,752,584,608]" italics="true" pageId="7" pageNumber="39">C. annandalei</emphasis>
|
||
</taxonomicName>
|
||
and from this species and
|
||
<taxonomicName id="4C3713DFBC262D30FB97FDC62DBAFDCB" box="[1048,1198,584,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="agamensis">
|
||
<emphasis id="B943B44EBC262D30FB97FDC62DBAFDCB" box="[1048,1198,584,608]" italics="true" pageId="7" pageNumber="39">C. agamensis</emphasis>
|
||
</taxonomicName>
|
||
in having fewer ventral scales across midbody (36
|
||
<emphasis id="B943B44EBC262D30FE7BFDE32B28FD2F" box="[500,572,621,644]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
43 and 67, respectively) [Note: The ventral scale count in
|
||
<taxonomicName id="4C3713DFBC262D30FB5BFDE32C88FD2F" box="[1236,1436,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC262D30FB5BFDE32C88FD2F" box="[1236,1436,620,644]" italics="true" pageId="7" pageNumber="39">C. kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2700935BC262D30FF18FD1E29E6FD0C" box="[151,242,656,680]" pageId="7" pageNumber="39" rank="species">
|
||
<emphasis id="B943B44EBC262D30FF18FD1E29F8FD03" bold="true" box="[151,236,656,680]" pageId="7" pageNumber="39">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, which lacks ventrolateral folds, was made between the lowest rows of lateral tubercles and is thus not strictly comparable to counts of species with folds. However, as tubercles are never present ventral to such folds, and the ventralmost tubercles are often several scale rows above the folds, the ventral count in the new species may be assumed to be equal to or greater than (but not less than) corresponding inter-fold counts. Inter-fold scale counts of greater magnitude than the inter-tubercle count of the new species may thus be conservatively regarded as putatively diagnostic]. It differs from
|
||
<taxonomicName id="4C3713DFBC262D30FD9CFCCB2BCCFCF7" box="[531,728,836,860]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="malcolmsmithi">
|
||
<emphasis id="B943B44EBC262D30FD9CFCCB2BCCFCF7" box="[531,728,836,860]" italics="true" pageId="7" pageNumber="39">C. malcolmsmithi</emphasis>
|
||
</taxonomicName>
|
||
in having small caudal tubercles restricted to the proximal portion of the tail (versus prominent tubercles along the entire tail), from
|
||
<taxonomicName id="4C3713DFBC262D30FC76FCE62D90FC2B" box="[1017,1156,872,896]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="buchardi">
|
||
<emphasis id="B943B44EBC262D30FC76FCE62D90FC2B" box="[1017,1156,872,896]" italics="true" pageId="7" pageNumber="39">C. buchardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FB16FCE62C2AFC2B" box="[1177,1342,872,896]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="annandalei">
|
||
<emphasis id="B943B44EBC262D30FB16FCE62C2AFC2B" box="[1177,1342,872,896]" italics="true" pageId="7" pageNumber="39">C. annandalei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3713DFBC262D30FA0DFCE62856FC0F" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="consobrinoides">
|
||
<emphasis id="B943B44EBC262D30FA0DFCE62856FC0F" italics="true" pageId="7" pageNumber="39">C. consobrinoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3713DFBC262D30FEDDFC0328A6FC0F" box="[338,434,909,932]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="gansi">
|
||
<emphasis id="B943B44EBC262D30FEDDFC0328A6FC0F" box="[338,434,909,932]" italics="true" pageId="7" pageNumber="39">C. gansi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3713DFBC262D30FE63FC032B90FC0F" box="[492,644,908,932]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="wakeorum">
|
||
<emphasis id="B943B44EBC262D30FE63FC032B90FC0F" box="[492,644,908,932]" italics="true" pageId="7" pageNumber="39">C. wakeorum</emphasis>
|
||
</taxonomicName>
|
||
in lacking a banded dorsal pattern and from the last two of these, as well as
|
||
<taxonomicName id="4C3713DFBC262D30FF64FC3E287BFC63" box="[235,367,944,968]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="buchardi">
|
||
<emphasis id="B943B44EBC262D30FF64FC3E287BFC63" box="[235,367,944,968]" italics="true" pageId="7" pageNumber="39">C. buchardi</emphasis>
|
||
</taxonomicName>
|
||
, in having a lower number of dorsal tubercle rows (16–18
|
||
<emphasis id="B943B44EBC262D30FC73FC3F2D57FC63" box="[1020,1091,945,968]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
20–25 [
|
||
<quantity id="4CCFC5B9BC262D30FB12FC212DCCFC6C" box="[1181,1240,943,967]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" pageId="7" pageNumber="39" unit="in" value="24.0">24 in</quantity>
|
||
<taxonomicName id="4C3713DFBC262D30FB51FC3E2C67FC63" box="[1246,1395,944,968]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="wakeorum">
|
||
<emphasis id="B943B44EBC262D30FB51FC3E2C67FC63" box="[1246,1395,944,968]" italics="true" pageId="7" pageNumber="39">C. wakeorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CCFC5B9BC262D30FA0FFC2129B9FC47" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="7" pageNumber="39" unit="in" value="25.0">25 in</quantity>
|
||
<taxonomicName id="4C3713DFBC262D30FF3BFC5B282CFC47" box="[180,312,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="buchardi">
|
||
<emphasis id="B943B44EBC262D30FF3BFC5B282CFC47" box="[180,312,980,1004]" italics="true" pageId="7" pageNumber="39">C. buchardi</emphasis>
|
||
</taxonomicName>
|
||
]). It is distinguished from
|
||
<taxonomicName id="4C3713DFBC262D30FDEAFC5B2BA1FC47" box="[613,693,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="feae">
|
||
<emphasis id="B943B44EBC262D30FDEAFC5B2BA1FC47" box="[613,693,980,1004]" italics="true" pageId="7" pageNumber="39">C. feae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3713DFBC262D30FD64FC5B2A89FC47" box="[747,925,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="gubernatoris">
|
||
<emphasis id="B943B44EBC262D30FD64FC5B2A89FC47" box="[747,925,980,1004]" italics="true" pageId="7" pageNumber="39">C. gubernatoris</emphasis>
|
||
</taxonomicName>
|
||
in having a weakly defined ventrolateral fold (
|
||
<emphasis id="B943B44EBC262D30FF2FFC7729FCFBBB" box="[160,232,1017,1040]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
a well-defined row of enlarged tubercles separating flank and venter) and further from the former species in lacking a reticulate pattern on the head and from the latter in lacking irregular blackish transverse markings on the dorsum.
|
||
<taxonomicName id="4C3713DFBC262D30FF74FBCE2B5CFBF3" box="[251,584,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="kimberleyensis" status="sp. nov.">
|
||
<emphasis id="B943B44EBC262D30FF74FBCE2B5CFBF3" box="[251,584,1088,1112]" italics="true" pageId="7" pageNumber="39">Cyrtodactylus kimberleyensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943B44EBC262D30FDDDFBCE2BBAFBF3" bold="true" box="[594,686,1088,1112]" pageId="7" pageNumber="39">
|
||
<taxonomicNameLabel id="A2700935BC262D30FDDDFBCE2BBAFBF3" box="[594,686,1088,1112]" pageId="7" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the same size as
|
||
<taxonomicName id="4C3713DFBC262D30FC16FBCE2D3BFBF3" box="[921,1071,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="39" phylum="Chordata" rank="species" species="laevigatus">
|
||
<emphasis id="B943B44EBC262D30FC16FBCE2D3BFBF3" box="[921,1071,1088,1112]" italics="true" pageId="7" pageNumber="39">C. laevigatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B943B44EBC262D30FBE4FBCE2C06FBF3" box="[1131,1298,1088,1112]" italics="true" pageId="7" pageNumber="39">C. l. uniformis</emphasis>
|
||
Auffenberg from
|
||
<collectingCountry id="F32028CCBC262D30FF5BFBEA280EFBD7" box="[212,282,1124,1148]" name="Indonesia" pageId="7" pageNumber="39">Flores</collectingCountry>
|
||
but differs from these in its more regularly arranged dorsal tubercles (16–18 discrete longitudinal rows
|
||
<emphasis id="B943B44EBC262D30FF18FB0729CBFB0B" box="[151,223,1161,1184]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
scattered or irregularly arranged tubercles) and in having at most a single transverse row of granules separating the proximal and distal series of subdigital lamellae (
|
||
<emphasis id="B943B44EBC262D30FCA5FB232A66FB6F" box="[810,882,1197,1220]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
multiple such rows; see
|
||
<bibRefCitation id="EFA615ADBC262D30FB1CFB222C56FB6F" author="Darevsky" box="[1171,1346,1196,1220]" pageId="7" pageNumber="39" refString="Darevsky, I. S. (1964) Two new species of gekkonid lizards from the Komodo Island in Lesser Sundas Archipelago. Zoologischer Anzeiger, 173, 169 - 174." type="journal article" year="1964">Darevsky 1964</bibRefCitation>
|
||
,
|
||
<figureCitation id="130C74D9BC262D30FADFFB222C83FB6F" box="[1360,1431,1196,1220]" captionStart="FIGURE 4" captionStartId="4.[151,250,1306,1329]" captionTargetBox="[369,1154,517,1267]" captionTargetId="figure@4.[354,1233,517,1285]" captionTargetPageId="4" captionText="FIGURE 4. Ventral view of right pes of Cyrtodactylus kimberleyensis sp. nov. showing disposition of the subdigital lamellae. Roman numerals indicate digit numbers. Scale bar = 2 mm." httpUri="https://zenodo.org/record/215282/files/figure.png" pageId="7" pageNumber="39">Fig. 4</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFA615ADBC262D30FF18FB41284CFB4C" author="Auffenberg" box="[151,344,1231,1255]" pageId="7" pageNumber="39" refString="Auffenberg, W. (1980) The herpetofauna of Komodo, with notes on adjacent areas. Bulletin of the Florida State Museum, Biological Sciences, 25, 39 - 156." type="journal article" year="1980">Auffenberg 1980</bibRefCitation>
|
||
, Fig. 14E). It also differs from these species in dorsal pattern (light and dark blotches
|
||
<emphasis id="B943B44EBC262D30FA80FB5F2C42FB43" box="[1295,1366,1233,1256]" italics="true" pageId="7" pageNumber="39">versus</emphasis>
|
||
short, sometimes interrupted transverse bars in the nominate form and mostly patternless in
|
||
<emphasis id="B943B44EBC262D30FBD9FB7B2DECFAA7" box="[1110,1272,1268,1292]" italics="true" pageId="7" pageNumber="39">C. l. uniformis</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |