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<document id="A30077670F9F9AF9C1E39CBB389C6529" ID-DOI="10.5852/ejt.2025.976.2775" ID-ISSN="2118-9773" ID-Zenodo-Dep="14771799" ID-ZooBank="58ACBB9C-CB70-43A5-8261-11FC8AF8E054" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1738237399256" checkinUser="plazi" docAuthor="Zhang, Yu-Hao, Li, Hai-Tao, Hu, Zhan-Yu &amp; Guo, Jian-Jun" docDate="2025" docId="0396422EFFEFFF91FE5A85478330F13F" docLanguage="en" docName="EJT.2025.976.47-76.pdf" docOrigin="European Journal of Taxonomy 976" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2775/12685" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Atractides (Atractides) fodingensis Zhang &amp; Guo 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="FFAF3A56FFE0FF83FFE5844F8646F47F" masterDocTitle="Two new species in the genus Atractides Koch, 1837 (Acari: Hydrachnidiae: Hygrobatidae), with the first descriptions of the female of two species from China" masterLastPageNumber="76" masterPageNumber="47" pageNumber="62" updateTime="1738328309856" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2E2FD02DFD639FB28B740C3CCEE5ECA7">Two new species in the genus Atractides Koch, 1837 (Acari: Hydrachnidiae: Hygrobatidae), with the first descriptions of the female of two species from China</mods:title>
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<mods:namePart id="5E0514249FB921EFA1468986E034064C">Zhang, Yu-Hao</mods:namePart>
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<mods:affiliation id="A41EE5772C4D27956B89C61F23339E68">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="14245A465CDE732B9BCCC89658D8F476" type="email">1289568720@qq.com</mods:nameIdentifier>
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<mods:name id="2E7E0FC5B99530D72FAAC050B1A6A2A2" type="personal">
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<mods:namePart id="DCD8E1C0F005EAE6C2B40CC4A6E74807">Li, Hai-Tao</mods:namePart>
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<mods:affiliation id="16E113A006883D75B61FADA6423331EF">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="1424842BDAC5A5123B240C53D11DA002" type="email">3221448736@qq.com</mods:nameIdentifier>
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<mods:name id="D352E867D4E82DA0CEC2888164255093" type="personal">
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<mods:namePart id="060221DA20639704BF606C8F0573C5E3">Hu, Zhan-Yu</mods:namePart>
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<mods:affiliation id="A1772B4CA4C1AF8ECD13078B02C14ED7">Sanqi Medical College, Wenshan University, Wenshan 663099, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="F0F017CA8EFACDC7EEB441DBD5682F7B" type="email">zhanyu3599@126.com</mods:nameIdentifier>
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<mods:namePart id="22F7B7473F314C3CC038111207A5F7D7">Guo, Jian-Jun</mods:namePart>
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<mods:affiliation id="DCEE0C4F76B7738DB650B8D7F5018F88">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="9C380B4A2E2FA4AE8D80722AEA365C9C" type="email">jjguo@gzu.edu.cn</mods:nameIdentifier>
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<mods:title id="5433750982C20FBA9A51ADBE3B4C6336">European Journal of Taxonomy</mods:title>
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<mods:date id="AE9C670E18BA1B66B354DA2E9F36D3E2">2025</mods:date>
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<subSubSection id="C325A0B3FFEFFF8CFE5A85478233F55C" box="[447,1141,264,291]" pageId="15" pageNumber="62" type="nomenclature">
<paragraph id="8B80F338FFEFFF8CFE5A85478233F55C" blockId="15.[364,1224,264,361]" box="[447,1141,264,291]" pageId="15" pageNumber="62">
<heading id="D0C84454FFEFFF8CFE5A85478233F55C" box="[447,1141,264,291]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="62" reason="2">
<taxonomicName id="4C3F88BBFFEFFF8CFE5A85478254F55C" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[447,1042,264,291]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="62" phylum="Arthropoda" rank="species" species="fodingensis" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFEFFF8CFE5A85478523F55C" bold="true" box="[447,869,264,291]" pageId="15" pageNumber="62">
<emphasis id="B94B2F2AFFEFFF8CFE5A8547847DF55C" bold="true" box="[447,571,264,291]" italics="true" pageId="15" pageNumber="62">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFEFFF8CFDA98547848EF55C" bold="true" box="[588,712,264,291]" italics="true" pageId="15" pageNumber="62">Atractides</emphasis>
)
<emphasis id="B94B2F2AFFEFFF8CFD3C85478523F55C" bold="true" box="[729,869,264,291]" italics="true" pageId="15" pageNumber="62">fodingensis</emphasis>
</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFEFFF8CFBFC85468233F55C" box="[1049,1141,265,291]" pageId="15" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C325A0B3FFEFFF8CFE898563851DF516" pageId="15" pageNumber="62" type="description">
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<uri id="FFAEFF3AFFEFFF8CFE898563828EF538" box="[364,1224,300,327]" pageId="15" pageNumber="62">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8B80F338FFEFFF8CFD3D8500851DF516" blockId="15.[364,1224,264,361]" box="[728,859,335,361]" pageId="15" pageNumber="62">
<figureCitation id="1304EFBDFFEFFF8CFD3D8500851DF516" box="[728,859,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="15.[189,232,1928,1954]" captionStartId-1="16.[189,232,1963,1989]" captionStartId-2="17.[189,232,1928,1954]" captionTargetBox-0="[302,1271,439,1872]" captionTargetBox-1="[214,1377,651,1903]" captionTargetBox-2="[189,1398,786,1889]" captionTargetId-0="figure-25@15.[276,1312,436,1888]" captionTargetId-1="figure-87@16.[189,1398,633,1923]" captionTargetId-2="figure-142@17.[189,1398,786,1889]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Fig. 16. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." captionText-1="Fig. 17. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC). A–D. I-L– IV-L. Scale bar = 100 μm." captionText-2="Fig. 18. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC), light microscope photographs.A. Idiosoma, dorsal view. B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.14771837" figureDoi-1="http://doi.org/10.5281/zenodo.14771839" figureDoi-2="http://doi.org/10.5281/zenodo.14771841" httpUri-0="https://zenodo.org/record/14771837/files/figure.png" httpUri-1="https://zenodo.org/record/14771839/files/figure.png" httpUri-2="https://zenodo.org/record/14771841/files/figure.png" pageId="15" pageNumber="62">Figs 16–18</figureCitation>
</paragraph>
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<caption id="DF40A3B0FFEFFF8CFF5883C78410F396" ID-DOI="http://doi.org/10.5281/zenodo.14771837" ID-Zenodo-Dep="14771837" httpUri="https://zenodo.org/record/14771837/files/figure.png" pageId="15" pageNumber="62" startId="15.[189,232,1928,1954]" targetBox="[302,1271,439,1872]" targetPageId="15">
<paragraph id="8B80F338FFEFFF8CFF5883C78410F396" blockId="15.[187,1399,1928,2025]" pageId="15" pageNumber="62">
<emphasis id="B94B2F2AFFEFFF8CFF5883C78764F3DD" bold="true" box="[189,290,1928,1954]" pageId="15" pageNumber="62">Fig. 16.</emphasis>
<taxonomicName id="4C3F88BBFFEFFF8CFED583C785DAF3DD" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[304,924,1928,1954]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="15" pageNumber="62" phylum="Arthropoda" rank="species" species="fodingensis" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFEFFF8CFED583C787ECF3DD" box="[304,426,1928,1954]" italics="true" pageId="15" pageNumber="62">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFEFFF8CFE2783C7847AF3DD" box="[450,572,1928,1954]" italics="true" pageId="15" pageNumber="62">Atractides</emphasis>
)
<emphasis id="B94B2F2AFFEFFF8CFDB083C7849AF3DD" box="[597,732,1928,1954]" italics="true" pageId="15" pageNumber="62">fodingensis</emphasis>
Zhang &amp; Guo
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<taxonomicNameLabel id="A2789251FFEFFF8CFC4F83C78249F3DD" box="[938,1039,1928,1954]" pageId="15" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (slide no. GZHY-2023061901, GUGC).
<emphasis id="B94B2F2AFFEFFF8CFE1183E4844FF3BA" bold="true" box="[500,521,1963,1989]" pageId="15" pageNumber="62">A</emphasis>
. Idiosoma, dorsal view.
<emphasis id="B94B2F2AFFEFFF8CFCC583E48572F3BA" bold="true" box="[800,820,1963,1989]" pageId="15" pageNumber="62">B</emphasis>
. Idiosoma, ventral view.
<emphasis id="B94B2F2AFFEFFF8CFBB183E4822FF3BA" bold="true" box="[1108,1129,1963,1989]" pageId="15" pageNumber="62">C</emphasis>
. Palp.
<emphasis id="B94B2F2AFFEFFF8CFB5283E4828AF3BA" bold="true" box="[1207,1228,1963,1989]" pageId="15" pageNumber="62">D</emphasis>
. Gnathosoma.
<emphasis id="B94B2F2AFFEFFF8CFF5E83818689F397" bold="true" box="[187,207,1998,2024]" pageId="15" pageNumber="62">E</emphasis>
. Chelicera. Scale bars = 100 μm.
</paragraph>
</caption>
<subSubSection id="C325A0B3FFF0FF93FF58854782E8F5E8" pageId="16" pageNumber="63" type="diagnosis">
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<heading id="D0C84454FFF0FF93FF5885478771F55D" bold="true" box="[189,311,264,290]" fontSize="11" level="3" pageId="16" pageNumber="63" reason="3">
<emphasis id="B94B2F2AFFF0FF93FF5885478771F55D" bold="true" box="[189,311,264,290]" pageId="16" pageNumber="63">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFF0FF93FF58857882E8F5E8" blockId="16.[188,1399,311,408]" pageId="16" pageNumber="63">
The apodemes of ACG almost transverse. Pregenital sclerite relatively large,
<emphasis id="B94B2F2AFFF0FF93FBAA8577822DF527" box="[1103,1131,312,344]" italics="true" pageId="16" pageNumber="63">
V
<subScript id="17BBF17DFFF0FF93FB878505822DF527" attach="left" box="[1122,1131,330,344]" fontSize="6" pageId="16" pageNumber="63">1</subScript>
</emphasis>
separated fro
<emphasis id="B94B2F2AFFF0FF93FAF085778369F527" box="[1301,1327,312,344]" italics="true" pageId="16" pageNumber="63">
V
<subScript id="17BBF17DFFF0FF93FAC385058369F527" attach="left" box="[1318,1327,330,344]" fontSize="6" pageId="16" pageNumber="63">2</subScript>
</emphasis>
,
<emphasis id="B94B2F2AFFF0FF93FADB8577831FF527" box="[1342,1369,312,344]" italics="true" pageId="16" pageNumber="63">
V
<subScript id="17BBF17DFFF0FF93FAB58505831FF527" attach="left" box="[1360,1369,330,344]" fontSize="6" pageId="16" pageNumber="63">3</subScript>
</emphasis>
at same level of pregenital sclerite and near Cx-IV, excretory pore smooth. P-4 with straight ventral margin and two ventral hairs dividing it in 2:1:2, sword seta at same level of distoventral hair.
</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFF0FF93FF5885F082AEF678" box="[189,1256,447,520]" pageId="16" pageNumber="63" type="etymology">
<paragraph id="8B80F338FFF0FF93FF5885F08703F5A6" blockId="16.[189,325,447,473]" box="[189,325,447,473]" pageId="16" pageNumber="63">
<heading id="D0C84454FFF0FF93FF5885F08703F5A6" bold="true" box="[189,325,447,473]" fontSize="11" level="3" pageId="16" pageNumber="63" reason="3">
<emphasis id="B94B2F2AFFF0FF93FF5885F08703F5A6" bold="true" box="[189,325,447,473]" pageId="16" pageNumber="63">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFF0FF93FF5885A282AEF678" blockId="16.[189,1256,493,520]" box="[189,1256,493,520]" pageId="16" pageNumber="63">The new species is named after the Foding Mountain, where the specimens were collected.</paragraph>
</subSubSection>
<caption id="DF40A3B0FFF0FF93FF5883E485F7F396" ID-DOI="http://doi.org/10.5281/zenodo.14771839" ID-Zenodo-Dep="14771839" httpUri="https://zenodo.org/record/14771839/files/figure.png" pageId="16" pageNumber="63" startId="16.[189,232,1963,1989]" targetBox="[214,1377,651,1903]" targetPageId="16">
<paragraph id="8B80F338FFF0FF93FF5883E485F7F396" blockId="16.[189,1399,1963,2025]" pageId="16" pageNumber="63">
<emphasis id="B94B2F2AFFF0FF93FF5883E48764F3BA" bold="true" box="[189,290,1963,1989]" pageId="16" pageNumber="63">Fig. 17.</emphasis>
<taxonomicName id="4C3F88BBFFF0FF93FED583E485DAF3BA" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[304,924,1963,1989]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="63" phylum="Arthropoda" rank="species" species="fodingensis" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFF0FF93FED583E487ECF3BA" box="[304,426,1963,1989]" italics="true" pageId="16" pageNumber="63">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFF0FF93FE2783E4847AF3BA" box="[450,572,1963,1989]" italics="true" pageId="16" pageNumber="63">Atractides</emphasis>
)
<emphasis id="B94B2F2AFFF0FF93FDB083E4849AF3BA" box="[597,732,1963,1989]" italics="true" pageId="16" pageNumber="63">fodingensis</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFF0FF93FC4F83E38249F3B9" box="[938,1039,1964,1990]" pageId="16" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (slide no. GZHY-2023061901, GUGC).
<emphasis id="B94B2F2AFFF0FF93FE1E83818472F397" bold="true" box="[507,564,1998,2025]" pageId="16" pageNumber="63">A–D</emphasis>
. I-L– IV-L. Scale bar = 100 μm.
</paragraph>
</caption>
<subSubSection id="C325A0B3FFF1FF92FF58854682EEF660" pageId="17" pageNumber="64" type="materials_examined">
<paragraph id="8B80F338FFF1FF92FF588546872BF55D" blockId="17.[189,365,264,291]" box="[189,365,264,291]" pageId="17" pageNumber="64">
<heading id="D0C84454FFF1FF92FF588546872BF55D" bold="true" box="[189,365,264,291]" fontSize="11" level="3" pageId="17" pageNumber="64" reason="3">
<emphasis id="B94B2F2AFFF1FF92FF588546872BF55D" bold="true" box="[189,365,264,291]" pageId="17" pageNumber="64">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFF1FF92FF0F85788219F5C4" blockId="17.[189,1399,311,443]" pageId="17" pageNumber="64">
<materialsCitation id="3B57F965FFF1FF92FF0F85788219F5C4" collectingDate="2023-06-19" collectionCode="GUGC" collectorName="Ri-Xin Jiang &amp; Ping Li &amp; Lan Jia" country="China" elevation="693" latitude="27.3113" location="Foding Mountain" longLatPrecision="6" longitude="108.1363" municipality="Tongren City" pageId="17" pageNumber="64" specimenCode="GZ-HY-2023061901" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou Province" typeStatus="holotype">
<emphasis id="B94B2F2AFFF1FF92FF0F8578871AF52E" bold="true" box="[234,348,311,337]" pageId="17" pageNumber="64">
<typeStatus id="54844D9AFFF1FF92FF0F8578871AF52E" box="[234,348,311,337]" pageId="17" pageNumber="64" type="holotype">Holotype</typeStatus>
</emphasis>
P.R.
<collectingCountry id="F328B3A8FFF1FF92FF1185158709F50B" box="[244,335,346,372]" name="China" pageId="17" pageNumber="64">CHINA</collectingCountry>
•
<specimenCount id="9D3938B1FFF1FF92FE8D85148739F50A" box="[360,383,347,373]" count="1" pageId="17" pageNumber="64" type="female">♀</specimenCount>
;
<collectingRegion id="49FB3DDAFFF1FF92FE6885158423F50B" box="[397,613,346,372]" country="China" name="Guizhou" pageId="17" pageNumber="64">Guizhou Province</collectingRegion>
,
<collectingMunicipality id="6BE46942FFF1FF92FD958514854BF50B" box="[624,781,346,373]" pageId="17" pageNumber="64">Tongren City</collectingMunicipality>
,
<location id="8EE0A5E3FFF1FF92FCFD851585AFF50B" LSID="urn:lsid:plazi:treatment:0396422EFFEFFF91FE5A85478330F13F:8EE0A5E3FFF1FF92FCFD851585AFF50B" box="[792,1001,346,372]" country="China" latitude="27.3113" longLatPrecision="6" longitude="108.1363" municipality="Tongren City" name="Foding Mountain" pageId="17" pageNumber="64" stateProvince="Guizhou Province">Foding Mountain</location>
;
<geoCoordinate id="EE0B95FFFFF1FF92FC1185158238F50A" box="[1012,1150,346,373]" degrees="27.3113" direction="north" orientation="latitude" pageId="17" pageNumber="64" precision="5" value="27.3113">27.3113° N</geoCoordinate>
,
<geoCoordinate id="EE0B95FFFFF1FF92FB6F85158366F50A" box="[1162,1312,346,373]" degrees="108.1363" direction="east" orientation="longitude" pageId="17" pageNumber="64" precision="5" value="108.1363">108.1363° E</geoCoordinate>
;
<elevation id="0012140BFFF1FF92FAC9851586B0F5E8" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="17" pageNumber="64" unit="m" value="693.0">
<quantity id="4CC75EDDFFF1FF92FAC985158331F50A" box="[1324,1399,346,373]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="17" pageNumber="64" unit="m" value="693.0">693 m</quantity>
a.s.l.
</elevation>
;
<date id="FF81D5F8FFF1FF92FEE6853187DBF5E7" box="[259,413,382,408]" pageId="17" pageNumber="64" value="2023-06-19">
<collectingDate id="EFC52C10FFF1FF92FEE6853187DBF5E7" box="[259,413,382,408]" pageId="17" pageNumber="64" value="2023-06-19">19 Jun. 2023</collectingDate>
</date>
;
<collectorName id="26CA96EEFFF1FF92FE4F85328405F5E8" box="[426,579,381,407]" pageId="17" pageNumber="64">Ri-Xin Jiang</collectorName>
,
<collectorName id="26CA96EEFFF1FF92FDAA853284E0F5E8" box="[591,678,381,407]" pageId="17" pageNumber="64">Ping Li</collectorName>
and
<collectorName id="26CA96EEFFF1FF92FD0585318570F5E8" box="[736,822,381,408]" pageId="17" pageNumber="64">Lan Jia</collectorName>
leg.; waterfall in the mountain, with fast running water, and many plants along the shore; slide no.
<specimenCode id="DB995B43FFF1FF92FCE485EF85BFF5C5" box="[769,1017,416,442]" pageId="17" pageNumber="64">GZ-HY-2023061901</specimenCode>
;
<collectionCode id="ED2E6BFDFFF1FF92FBE085EE821AF5C4" box="[1029,1116,417,443]" pageId="17" pageNumber="64">GUGC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B80F338FFF1FF92FF0F85AD82EEF660" blockId="17.[189,1192,482,544]" pageId="17" pageNumber="64">
<materialsCitation id="3B57F965FFF1FF92FF0F85AD82E2F660" collectingDate="2023-06-19" collectionCode="GUGC" collectorName="Ri-Xin Jiang &amp; Ping Li &amp; Lan Jia" country="China" elevation="693" latitude="27.3113" location="CHINA" longLatPrecision="6" longitude="108.1363" municipality="Tongren City" pageId="17" pageNumber="64" specimenCode="GZ-HY-2023061902" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou" typeStatus="paratype">
<emphasis id="B94B2F2AFFF1FF92FF0F85AD871AF583" bold="true" box="[234,348,482,508]" pageId="17" pageNumber="64">
<typeStatus id="54844D9AFFF1FF92FF0F85AD871AF583" box="[234,348,482,508]" pageId="17" pageNumber="64" type="paratype">Paratype</typeStatus>
</emphasis>
P.R.
<collectingCountry id="F328B3A8FFF1FF92FF11864A8709F660" box="[244,335,517,543]" name="China" pageId="17" pageNumber="64">CHINA</collectingCountry>
•
<specimenCount id="9D3938B1FFF1FF92FE8D864A87D3F65F" box="[360,405,517,544]" count="1" pageId="17" pageNumber="64" type="female">1 ♀</specimenCount>
; same data as for holotype; slide no.
<specimenCode id="DB995B43FFF1FF92FCAC864A8207F660" box="[841,1089,517,543]" pageId="17" pageNumber="64">GZ-HY-2023061902</specimenCode>
;
<collectionCode id="ED2E6BFDFFF1FF92FBA8864A82E2F660" box="[1101,1188,517,543]" pageId="17" pageNumber="64">GUGC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFF1FF91FF5886098294F783" lastPageId="18" lastPageNumber="65" pageId="17" pageNumber="64" type="description">
<paragraph id="8B80F338FFF1FF92FF5886098709F61F" blockId="17.[189,335,582,608]" box="[189,335,582,608]" pageId="17" pageNumber="64">
<heading id="D0C84454FFF1FF92FF5886098709F61F" bold="true" box="[189,335,582,608]" fontSize="11" level="3" pageId="17" pageNumber="64" reason="3">
<emphasis id="B94B2F2AFFF1FF92FF5886098709F61F" bold="true" box="[189,335,582,608]" pageId="17" pageNumber="64">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFF1FF92FF0F863A87DBF6F0" blockId="17.[188,1398,629,726]" box="[234,413,629,656]" pageId="17" pageNumber="64">
<emphasis id="B94B2F2AFFF1FF92FF0F863A8703F6F0" bold="true" box="[234,325,629,655]" pageId="17" pageNumber="64">Female</emphasis>
(n = 2)
</paragraph>
<paragraph id="8B80F338FFF1FF92FF5886D78331F6A2" blockId="17.[188,1398,629,726]" pageId="17" pageNumber="64">
Idiosoma soft and oval, frontal edge slightly concave;
<emphasis id="B94B2F2AFFF1FF92FCD186D68515F6C6" box="[820,851,665,697]" italics="true" pageId="17" pageNumber="64">
O
<subScript id="17BBF17DFFF1FF92FCAF86E48515F6C6" attach="left" box="[842,851,683,697]" fontSize="6" pageId="17" pageNumber="64">1</subScript>
</emphasis>
between
<emphasis id="B94B2F2AFFF1FF92FC2486D6859AF6C6" box="[961,988,665,697]" italics="true" pageId="17" pageNumber="64">
A
<subScript id="17BBF17DFFF1FF92FC3686E4859AF6C6" attach="left" box="[979,988,683,697]" fontSize="6" pageId="17" pageNumber="64">1</subScript>
</emphasis>
and
<emphasis id="B94B2F2AFFF1FF92FBF686D68269F6C6" box="[1043,1071,665,697]" italics="true" pageId="17" pageNumber="64">
A
<subScript id="17BBF17DFFF1FF92FBC386E48269F6C6" attach="right" box="[1062,1071,683,697]" fontSize="6" pageId="17" pageNumber="64">2</subScript>
</emphasis>
,
<emphasis id="B94B2F2AFFF1FF92FBD986D6821CF6C6" box="[1084,1114,665,697]" italics="true" pageId="17" pageNumber="64">
O
<subScript id="17BBF17DFFF1FF92FBB486E4821CF6C6" attach="left" box="[1105,1114,683,697]" fontSize="6" pageId="17" pageNumber="64">2</subScript>
</emphasis>
between
<emphasis id="B94B2F2AFFF1FF92FB2D86D682A1F6C6" box="[1224,1255,665,697]" italics="true" pageId="17" pageNumber="64">
D
<subScript id="17BBF17DFFF1FF92FB3B86E482A1F6C6" attach="left" box="[1246,1255,683,697]" fontSize="6" pageId="17" pageNumber="64">1</subScript>
</emphasis>
and slightly front of
<emphasis id="B94B2F2AFFF1FF92FEFE86F38776F6AA" box="[283,304,700,725]" italics="true" pageId="17" pageNumber="64">D</emphasis>
<emphasis id="B94B2F2AFFF1FF92FED48680877CF6A2" box="[305,314,719,733]" italics="true" pageId="17" pageNumber="64">1</emphasis>
; all slit organs visible,
<emphasis id="B94B2F2AFFF1FF92FDA286F38422F6A9" box="[583,612,700,726]" italics="true" pageId="17" pageNumber="64">So</emphasis>
<emphasis id="B94B2F2AFFF1FF92FD8386808429F6A2" box="[614,623,719,733]" italics="true" pageId="17" pageNumber="64">1</emphasis>
near
<emphasis id="B94B2F2AFFF1FF92FD4886F384F9F6A9" box="[685,703,700,726]" italics="true" pageId="17" pageNumber="64">A</emphasis>
<emphasis id="B94B2F2AFFF1FF92FD258680848FF6A2" box="[704,713,719,733]" italics="true" pageId="17" pageNumber="64">2</emphasis>
,
<emphasis id="B94B2F2AFFF1FF92FD3286F384B2F6A9" box="[727,756,700,726]" italics="true" pageId="17" pageNumber="64">So</emphasis>
<emphasis id="B94B2F2AFFF1FF92FD11868084BBF6A2" box="[756,765,719,733]" italics="true" pageId="17" pageNumber="64">2</emphasis>
at the middle of
<emphasis id="B94B2F2AFFF1FF92FC2486F38597F6AA" box="[961,977,700,725]" italics="true" pageId="17" pageNumber="64">L</emphasis>
<emphasis id="B94B2F2AFFF1FF92FC378680859DF6A2" box="[978,987,719,733]" italics="true" pageId="17" pageNumber="64">1</emphasis>
and
<emphasis id="B94B2F2AFFF1FF92FBF786F38264F6AA" box="[1042,1058,700,725]" italics="true" pageId="17" pageNumber="64">L</emphasis>
<emphasis id="B94B2F2AFFF1FF92FBC68680826AF6A2" box="[1059,1068,719,733]" italics="true" pageId="17" pageNumber="64">2</emphasis>
,
<emphasis id="B94B2F2AFFF1FF92FBDF86F38211F6A9" box="[1082,1111,700,726]" italics="true" pageId="17" pageNumber="64">So</emphasis>
<emphasis id="B94B2F2AFFF1FF92FBB286808226F6A2" box="[1111,1120,719,733]" italics="true" pageId="17" pageNumber="64">3</emphasis>
at same level of
<emphasis id="B94B2F2AFFF1FF92FAC186F3837FF6AA" box="[1316,1337,700,725]" italics="true" pageId="17" pageNumber="64">D</emphasis>
<emphasis id="B94B2F2AFFF1FF92FADF86808305F6A2" box="[1338,1347,719,733]" italics="true" pageId="17" pageNumber="64">2</emphasis>
,
<emphasis id="B94B2F2AFFF1FF92FAB586F3832BF6A9" box="[1360,1389,700,726]" italics="true" pageId="17" pageNumber="64">So</emphasis>
<emphasis id="B94B2F2AFFF1FF92FA8B86808331F6A2" box="[1390,1399,719,733]" italics="true" pageId="17" pageNumber="64">4</emphasis>
</paragraph>
<caption id="DF40A3B0FFF1FF92FF5883C7856FF396" ID-DOI="http://doi.org/10.5281/zenodo.14771841" ID-Zenodo-Dep="14771841" httpUri="https://zenodo.org/record/14771841/files/figure.png" pageId="17" pageNumber="64" startId="17.[189,232,1928,1954]" targetBox="[189,1398,786,1889]" targetPageId="17">
<paragraph id="8B80F338FFF1FF92FF5883C7856FF396" blockId="17.[187,1399,1928,2025]" pageId="17" pageNumber="64">
<emphasis id="B94B2F2AFFF1FF92FF5883C78767F3DD" bold="true" box="[189,289,1928,1954]" pageId="17" pageNumber="64">Fig. 18.</emphasis>
<taxonomicName id="4C3F88BBFFF1FF92FED583C785DAF3DD" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[304,924,1928,1954]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="64" phylum="Arthropoda" rank="species" species="fodingensis" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFF1FF92FED583C787ECF3DD" box="[304,426,1928,1954]" italics="true" pageId="17" pageNumber="64">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFF1FF92FE2783C7847AF3DD" box="[450,572,1928,1954]" italics="true" pageId="17" pageNumber="64">Atractides</emphasis>
)
<emphasis id="B94B2F2AFFF1FF92FDB083C7849AF3DD" box="[597,732,1928,1954]" italics="true" pageId="17" pageNumber="64">fodingensis</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFF1FF92FC4F83C78249F3DD" box="[938,1039,1928,1954]" pageId="17" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (slide no. GZHY-2023061901, GUGC), light microscope photographs.
<emphasis id="B94B2F2AFFF1FF92FCBE83E48536F3BA" bold="true" box="[859,880,1963,1989]" pageId="17" pageNumber="64">A</emphasis>
. Idiosoma, dorsal view.
<emphasis id="B94B2F2AFFF1FF92FB6C83E482DBF3BA" bold="true" box="[1161,1181,1963,1989]" pageId="17" pageNumber="64">B</emphasis>
. Idiosoma, ventral view.
<emphasis id="B94B2F2AFFF1FF92FEE68381875EF397" bold="true" box="[259,280,1998,2024]" pageId="17" pageNumber="64">C</emphasis>
. Idiosoma lateral view. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8B80F338FFF2FF91FF58854587E2F5CF" blockId="18.[189,1399,265,432]" pageId="18" pageNumber="65">
near
<emphasis id="B94B2F2AFFF2FF91FF1D85458757F555" box="[248,273,266,298]" italics="true" pageId="18" pageNumber="65">
L
<subScript id="17BBF17DFFF2FF91FEED85538757F555" attach="left" box="[264,273,284,298]" fontSize="6" pageId="18" pageNumber="65">4</subScript>
</emphasis>
,
<emphasis id="B94B2F2AFFF2FF91FEC585468701F555" box="[288,327,265,298]" italics="true" pageId="18" pageNumber="65">
So
<subScript id="17BBF17DFFF2FF91FEDB85538701F555" attach="left" box="[318,327,284,298]" fontSize="6" pageId="18" pageNumber="65">5</subScript>
</emphasis>
behind of
<emphasis id="B94B2F2AFFF2FF91FE2D854587A3F555" box="[456,485,266,298]" italics="true" pageId="18" pageNumber="65">
D
<subScript id="17BBF17DFFF2FF91FE39855387A3F555" attach="left" box="[476,485,284,298]" fontSize="6" pageId="18" pageNumber="65">4</subScript>
</emphasis>
(
<figureCitation id="1304EFBDFFF2FF91FE1085468421F55C" box="[501,615,265,291]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[302,1271,439,1872]" captionTargetId="figure-25@15.[276,1312,436,1888]" captionTargetPageId="15" captionText="Fig. 16. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771837" httpUri="https://zenodo.org/record/14771837/files/figure.png" pageId="18" pageNumber="65">Figs 16A</figureCitation>
,
<figureCitation id="1304EFBDFFF2FF91FD91854684EDF55C" box="[628,683,265,291]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,786,1889]" captionTargetId="figure-142@17.[189,1398,786,1889]" captionTargetPageId="17" captionText="Fig. 18. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC), light microscope photographs.A. Idiosoma, dorsal view. B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771841" httpUri="https://zenodo.org/record/14771841/files/figure.png" pageId="18" pageNumber="65">18A</figureCitation>
). ACG fused and with suture line, apodemes of ACG almost transverse; PCG separated. Pregenital sclerite relatively large, three pairs of acetabula in weakly curved line, Ac3 largest.
<emphasis id="B94B2F2AFFF2FF91FE71851F87E9F50F" box="[404,431,336,368]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FE43852D87E9F50F" attach="left" box="[422,431,354,368]" fontSize="6" pageId="18" pageNumber="65">1</subScript>
</emphasis>
separated from
<emphasis id="B94B2F2AFFF2FF91FD90851F84D6F50F" box="[629,656,336,368]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FD62852D84D6F50F" attach="left" box="[647,656,354,368]" fontSize="6" pageId="18" pageNumber="65">2</subScript>
</emphasis>
,
<emphasis id="B94B2F2AFFF2FF91FD47851F84FBF50F" box="[674,701,336,368]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FD51852D84FBF50F" attach="left" box="[692,701,354,368]" fontSize="6" pageId="18" pageNumber="65">3</subScript>
</emphasis>
at same level of pregenital sclerite and near Cx-IV,
<emphasis id="B94B2F2AFFF2FF91FAD9851F8311F50F" box="[1340,1367,336,368]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FAAB852D8311F50F" attach="left" box="[1358,1367,354,368]" fontSize="6" pageId="18" pageNumber="65">4</subScript>
</emphasis>
at horizontal line between Ac1 and Ac2,
<emphasis id="B94B2F2AFFF2FF91FD98853C84DEF5EC" box="[637,664,371,403]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FD6A85CA84DEF5EC" attach="left" box="[655,664,389,403]" fontSize="6" pageId="18" pageNumber="65">3</subScript>
</emphasis>
and
<emphasis id="B94B2F2AFFF2FF91FD35853C84ADF5EC" box="[720,747,371,403]" italics="true" pageId="18" pageNumber="65">
V
<subScript id="17BBF17DFFF2FF91FD0785CA84ADF5EC" attach="left" box="[738,747,389,403]" fontSize="6" pageId="18" pageNumber="65">4</subScript>
</emphasis>
forming trapezoid; excretory pore smooth and between
<emphasis id="B94B2F2AFFF2FF91FF5885D88689F5CF" box="[189,207,407,432]" italics="true" pageId="18" pageNumber="65">V</emphasis>
<emphasis id="B94B2F2AFFF2FF91FF2A85E6869EF5C8" box="[207,216,425,439]" italics="true" pageId="18" pageNumber="65">2</emphasis>
(
<figureCitation id="1304EFBDFFF2FF91FF0285D98710F5CF" box="[231,342,406,432]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[302,1271,439,1872]" captionTargetId="figure-25@15.[276,1312,436,1888]" captionTargetPageId="15" captionText="Fig. 16. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771837" httpUri="https://zenodo.org/record/14771837/files/figure.png" pageId="18" pageNumber="65">Figs 16B</figureCitation>
,
<figureCitation id="1304EFBDFFF2FF91FE8785D987D0F5CF" box="[354,406,406,432]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[189,1398,786,1889]" captionTargetId="figure-142@17.[189,1398,786,1889]" captionTargetPageId="17" captionText="Fig. 18. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC), light microscope photographs.A. Idiosoma, dorsal view. B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771841" httpUri="https://zenodo.org/record/14771841/files/figure.png" pageId="18" pageNumber="65">18B</figureCitation>
).
</paragraph>
<paragraph id="8B80F338FFF2FF91FF5885988787F601" blockId="18.[189,1400,471,638]" pageId="18" pageNumber="65">
Palp five-segmented; P-1 short, P-2 and P-3 without projection, P-4 with straight ventral margin and two ventral hairs, dividing P-
<quantity id="4CC75EDDFFF2FF91FDF685B48406F66B" box="[531,576,506,533]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="18" pageNumber="65" unit="in" value="4.0">4 in</quantity>
2:1:2, sword seta at same level of distoventral hair (
<figureCitation id="1304EFBDFFF2FF91FB4685B58348F66B" box="[1187,1294,506,532]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[302,1271,439,1872]" captionTargetId="figure-25@15.[276,1312,436,1888]" captionTargetPageId="15" captionText="Fig. 16. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771837" httpUri="https://zenodo.org/record/14771837/files/figure.png" pageId="18" pageNumber="65">Fig. 16C</figureCitation>
). Ventral margin of I-L-5 as long as dorsal margin, S-1 close to S-2, S-2 longer and thicker than S-1, ventrally curved whip-like seta at end of I-L-5, and near dorsal slide; I-L-6 stout, thick at its base, tapering towards end (
<figureCitation id="1304EFBDFFF2FF91FEB9862B87F4F601" box="[348,434,612,638]" captionStart="Fig" captionStartId="16.[189,232,1963,1989]" captionTargetBox="[214,1377,651,1903]" captionTargetId="figure-87@16.[189,1398,633,1923]" captionTargetPageId="16" captionText="Fig. 17. Atractides (Atractides) fodingensis Zhang &amp; Guo sp. nov., holotype, ♀ (slide no. GZ- HY-2023061901, GUGC). A–D. I-L– IV-L. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771839" httpUri="https://zenodo.org/record/14771839/files/figure.png" pageId="18" pageNumber="65">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="8B80F338FFF2FF91FF5886E98294F783" blockId="18.[188,1400,677,1021]" pageId="18" pageNumber="65">
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(measurements of
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in parantheses). Idiosoma L 495 (443), W 408 (368); coxal field L 295 (264), Cx-III W 252 (289), ACG IL 161 (179), mL 104 (116), W 210 (226); gnathosoma bay L 80 (86); gonopore L 95 (95), pregenital sclerite L 76 (65), postgenital sclerite L 37 (30); Ac1–3 L 31 (33), 23 (31), 29 (34); chelicera L 125 (114), cheliceral claw L 37 (39), basal segment L 94 (96); gnathosoma L 107 (131); palp dL: P-1 22 (25), P-2 46 (49), P-3 54 (59), P-4 57 (63), P-5 23 (23); leg segments: I-L-1 dL 42 (49), I-L-2 dL 60 (68), I-L-3 dL 78 (83), I-L-4 dL 106 (113), I-L-5 dL 117 (124), HB 31 (35), I-L-6 dL 61 (62), HB 12 (14), S-1 L 46 (53), W 6 (8), S-2 L 51 (51), W 7 (12); dL: II-L-1 40 (45), II-L-2 50 (56), II-L-3 62 (66), II-L-4 81 (85), II-L-5 84 (96), II-L-6 83 (88); dL: III-L-1 44 (44), III-L-2 60 (65), III-L-3 69 (76), III-L-4 97 (111), III-L-5 102 (112), III-L-6 90 (105); dL: IV-L-1 85 (93), IV-L-2 79 (78), IV-L-3 110 (117), IV-L-4 129 (137), IV-L-5 132 (143), IV-L-6 115 (130).
</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFF2FF91FF58806B8330F13F" pageId="18" pageNumber="65" type="discussion">
<paragraph id="8B80F338FFF2FF91FF58806B8776F041" blockId="18.[189,304,1060,1086]" box="[189,304,1060,1086]" pageId="18" pageNumber="65">
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<emphasis id="B94B2F2AFFF2FF91FF58806B8776F041" bold="true" box="[189,304,1060,1086]" pageId="18" pageNumber="65">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFF2FF91FF58801D8330F13F" blockId="18.[187,1399,1106,1350]" pageId="18" pageNumber="65">
This new species is similar to
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<emphasis id="B94B2F2AFFF2FF91FDFB801D84DEF013" box="[542,664,1106,1132]" italics="true" pageId="18" pageNumber="65">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFF2FF91FD4C801D8565F013" box="[681,803,1106,1132]" italics="true" pageId="18" pageNumber="65">Atractides</emphasis>
)
<emphasis id="B94B2F2AFFF2FF91FCD1801D85EEF013" box="[820,936,1106,1132]" italics="true" pageId="18" pageNumber="65">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFF2FF91FBBF801C82F2F012" box="[1114,1204,1107,1133]" pageId="18" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
in the following characters: (1)
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V
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separated from
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V
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</emphasis>
; (2) excretory pore smooth; (3) I-L-6 stout, thick at its base, tapering towards the end. However, the new species differs from
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<emphasis id="B94B2F2AFFF2FF91FC9480D685CDF0CC" box="[881,907,1177,1203]" italics="true" pageId="18" pageNumber="65">A.</emphasis>
(
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)
<emphasis id="B94B2F2AFFF2FF91FC2B80D68204F0CC" box="[974,1090,1177,1203]" italics="true" pageId="18" pageNumber="65">cardiacus</emphasis>
</taxonomicName>
in the following aspects: (1) sword seta of P-4 at the same level of distoventral hair in the new species, but at the same level of proximoventral hair in
<emphasis id="B94B2F2AFFF2FF91FE2C8090879DF086" box="[457,475,1247,1273]" italics="true" pageId="18" pageNumber="65">A</emphasis>
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(
<emphasis id="B94B2F2AFFF2FF91FE09809087B8F086" box="[492,510,1247,1273]" italics="true" pageId="18" pageNumber="65">A</emphasis>
.)
<emphasis id="B94B2F2AFFF2FF91FDF0809084CFF086" box="[533,649,1247,1273]" italics="true" pageId="18" pageNumber="65">cardiacus</emphasis>
</taxonomicName>
; (2) ACG and PCG occupy half of the venter in the new species, but ACG and PCG of the female
<emphasis id="B94B2F2AFFF2FF91FDD1814C8400F162" box="[564,582,1283,1309]" italics="true" pageId="18" pageNumber="65">A</emphasis>
<taxonomicName id="4C3F88BBFFF2FF91FDA8814D84B2F163" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[589,756,1282,1309]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="65" phylum="Arthropoda" rank="species" species="cardiacus" subGenus="Atractides">
(
<emphasis id="B94B2F2AFFF2FF91FDB2814C842FF162" box="[599,617,1283,1309]" italics="true" pageId="18" pageNumber="65">A</emphasis>
.)
<emphasis id="B94B2F2AFFF2FF91FD65814D84B2F163" box="[640,756,1282,1308]" italics="true" pageId="18" pageNumber="65">cardiacus</emphasis>
</taxonomicName>
occupy one third of the ventral view; (3) excretory pore between
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V
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</emphasis>
in the new species, but slightly behind the parallel line of
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V
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</emphasis>
in the female
<emphasis id="B94B2F2AFFF2FF91FB4B81698286F13F" box="[1198,1216,1318,1344]" italics="true" pageId="18" pageNumber="65">A</emphasis>
<taxonomicName id="4C3F88BBFFF2FF91FB2281698329F140" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[1223,1391,1317,1344]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="18" pageNumber="65" phylum="Arthropoda" rank="species" species="cardiacus" subGenus="Atractides">
(
<emphasis id="B94B2F2AFFF2FF91FB34816982A5F13F" box="[1233,1251,1318,1344]" italics="true" pageId="18" pageNumber="65">A</emphasis>
.)
<emphasis id="B94B2F2AFFF2FF91FB1E816A8329F140" box="[1275,1391,1317,1343]" italics="true" pageId="18" pageNumber="65">cardiacus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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