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<document id="25FB86E1BA431F07EB5117280F6C4128" ID-DOI="10.5852/ejt.2025.976.2775" ID-ISSN="2118-9773" ID-Zenodo-Dep="14771799" ID-ZooBank="58ACBB9C-CB70-43A5-8261-11FC8AF8E054" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1738237399256" checkinUser="plazi" docAuthor="Zhang, Yu-Hao, Li, Hai-Tao, Hu, Zhan-Yu &amp; Guo, Jian-Jun" docDate="2025" docId="0396422EFFE2FF89FE29871A827AF68C" docLanguage="en" docName="EJT.2025.976.47-76.pdf" docOrigin="European Journal of Taxonomy 976" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2775/12685" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Atractides (Atractides) cardiacus Zhang &amp; Guo 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="57" masterDocId="FFAF3A56FFE0FF83FFE5844F8646F47F" masterDocTitle="Two new species in the genus Atractides Koch, 1837 (Acari: Hydrachnidiae: Hygrobatidae), with the first descriptions of the female of two species from China" masterLastPageNumber="76" masterPageNumber="47" pageNumber="49" updateTime="1738328309856" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="576DBA15140B098906815F55B9DB00BD">Two new species in the genus Atractides Koch, 1837 (Acari: Hydrachnidiae: Hygrobatidae), with the first descriptions of the female of two species from China</mods:title>
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<mods:namePart id="0E6E829A01097688B2B164E918BBEADB">Zhang, Yu-Hao</mods:namePart>
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<mods:affiliation id="7A4DB43C93065B7EE5A9BC94C1245C12">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
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<mods:namePart id="EB1776173B087CB8DF7179E37CA2C691">Li, Hai-Tao</mods:namePart>
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<mods:affiliation id="0584AFEA40ADF821E21496A6805230A9">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
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<mods:affiliation id="13D3C267CF2E8D140C2882B358978B99">Institute of Entomology, Guizhou University, Scientific Observing and Experimental Station of Crop Pest in Guiyang, Ministry of Agriculture and Rural Affairs of the P. R. China, Guiyang 550025, P. R. China.</mods:affiliation>
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<mods:date id="092D20E1D75B4B6471BDB6990EADEF33">2025</mods:date>
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<taxonomicName id="4C3F88BBFFE2FF81FE29871A8241F710" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[460,1031,853,880]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
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Zhang &amp; Guo
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<figureCitation id="1304EFBDFFE2FF81FD2C87D48506F7C9" box="[713,832,923,950]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-12="Fig" captionStart-13="Fig" captionStart-14="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="3.[188,231,1891,1917]" captionStartId-1="4.[189,232,1927,1953]" captionStartId-10="11.[189,232,1963,1989]" captionStartId-11="12.[189,232,1024,1050]" captionStartId-12="12.[189,232,1963,1989]" captionStartId-13="13.[189,232,1963,1989]" captionStartId-14="14.[189,232,1963,1989]" captionStartId-2="5.[189,232,1962,1988]" captionStartId-3="6.[189,232,1928,1954]" captionStartId-4="7.[189,232,1962,1988]" captionStartId-5="8.[190,233,1928,1954]" captionStartId-6="9.[189,232,1325,1351]" captionStartId-7="9.[189,232,1963,1989]" captionStartId-8="10.[189,232,1928,1954]" captionStartId-9="11.[189,232,1078,1104]" captionTargetBox-0="[189,1398,662,1853]" captionTargetBox-1="[258,1346,486,1872]" captionTargetBox-10="[303,1285,1187,1924]" captionTargetBox-11="[315,1272,265,983]" captionTargetBox-12="[289,1298,1168,1925]" captionTargetBox-13="[189,1398,1016,1923]" captionTargetBox-14="[189,1398,561,1922]" captionTargetBox-2="[287,1304,594,1883]" captionTargetBox-3="[322,1241,445,1857]" captionTargetBox-4="[205,1385,667,1913]" captionTargetBox-5="[210,1378,731,1888]" captionTargetBox-6="[246,1342,265,1285]" captionTargetBox-7="[189,1398,1470,1924]" captionTargetBox-8="[189,1398,983,1890]" captionTargetBox-9="[277,1311,265,1041]" captionTargetId-0="figure-155@3.[189,1398,662,1853]" captionTargetId-1="figure-66@4.[205,1383,465,1892]" captionTargetId-10="figure-46@11.[303,1285,1187,1924]" captionTargetId-11="figure-19@12.[315,1272,265,983]" captionTargetId-12="figure-63@12.[289,1298,1168,1925]" captionTargetId-13="figure-11@13.[189,1398,1016,1923]" captionTargetId-14="figure-18@14.[189,1398,561,1922]" captionTargetId-2="figure-104@5.[241,1346,559,1922]" captionTargetId-3="figure-70@6.[298,1290,416,1890]" captionTargetId-4="figure-158@7.[189,1398,642,1925]" captionTargetId-5="figure-182@8.[210,1378,731,1888]" captionTargetId-6="figure-11@9.[246,1342,265,1285]" captionTargetId-7="figure-56@9.[189,1398,1470,1924]" captionTargetId-8="figure-222@10.[189,1398,983,1890]" captionTargetId-9="figure-11@11.[277,1311,265,1041]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="11" captionTargetPageId-11="12" captionTargetPageId-12="12" captionTargetPageId-13="13" captionTargetPageId-14="14" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="Fig. 1. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov. A–B. Paratype, ♂ (slide no. HNHY-2023041102, GUGC). C–D. Paratype, ♀ (slide no. HN-HY-2023041106, GUGC). A. Idiosoma, dorsal view. B. Idiosoma, ventral view. C. Idiosoma, dorsal view. D. Idiosoma, ventral view. Scale bars = 200 μm." captionText-1="Fig. 2. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HNHY-2023041101, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." captionText-10="Fig. 11. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HNHY-2023041103, GUGC), SEM photographs. A. II-L. B. II-L. C. III-L. D. IV-L." captionText-11="Fig. 12. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. Gnathosoma. B. ACG and PCG. C. Genital field. D. V 1, V 2 and excretory pore." captionText-12="Fig. 13. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. Palp. B. P-4 and P-5. C. palp. D. Sword seta of P-4." captionText-13="Fig. 14. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. I-L. B. I-L-5–6. C. I-L. D. I-L-5–6." captionText-14="Fig. 15. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. II-L. B. II-L. C. III-L. D. III-L. E. IV-L. F. IV-L." captionText-2="Fig. 3. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HNHY-2023041101, GUGC), I-L– IV-L. Scale bar = 100 μm." captionText-3="Fig. 4. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041105, GUGC). A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp. D. Pnathosoma. E. Chelicera. Scale bars = 100 μm." captionText-4="Fig. 5. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041105, GUGC). A–D. I-L– IV-L. Scale bar = 100 μm." captionText-5="Fig. 6. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HNHY-2023041101, GUGC), light microscope photographs. A. Idiosoma, dorsal view.B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." captionText-6="Fig. 7. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041105, GUGC), light microscope photographs. A. Idiosoma, dorsal view.B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." captionText-7="Fig. 8. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HNHY-2023041103, GUGC), SEM photographs. A. ACG and PCG. B. V 1, V 2 and excretory pore." captionText-8="Fig. 9. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HNHY-2023041103, GUGC), SEM photographs. A. Palp. B. Sword seta of P-4. C. Gnathosoma. D. Genital field." captionText-9="Fig. 10. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HNHY-2023041103, GUGC), SEM photographs. A. I-L. B. I-L-5–6. C. I-L. D. I-L-5–6." figureDoi-0="http://doi.org/10.5281/zenodo.14771801" figureDoi-1="http://doi.org/10.5281/zenodo.14771803" figureDoi-10="http://doi.org/10.5281/zenodo.14771825" figureDoi-11="http://doi.org/10.5281/zenodo.14771827" figureDoi-12="http://doi.org/10.5281/zenodo.14771829" figureDoi-13="http://doi.org/10.5281/zenodo.14771833" figureDoi-14="http://doi.org/10.5281/zenodo.14771835" figureDoi-2="http://doi.org/10.5281/zenodo.14771805" figureDoi-3="http://doi.org/10.5281/zenodo.14771807" figureDoi-4="http://doi.org/10.5281/zenodo.14771809" figureDoi-5="http://doi.org/10.5281/zenodo.14771811" figureDoi-6="http://doi.org/10.5281/zenodo.14771813" figureDoi-7="http://doi.org/10.5281/zenodo.14771815" figureDoi-8="http://doi.org/10.5281/zenodo.14771819" figureDoi-9="http://doi.org/10.5281/zenodo.14771823" httpUri-0="https://zenodo.org/record/14771801/files/figure.png" httpUri-1="https://zenodo.org/record/14771803/files/figure.png" httpUri-10="https://zenodo.org/record/14771825/files/figure.png" httpUri-11="https://zenodo.org/record/14771827/files/figure.png" httpUri-12="https://zenodo.org/record/14771829/files/figure.png" httpUri-13="https://zenodo.org/record/14771833/files/figure.png" httpUri-14="https://zenodo.org/record/14771835/files/figure.png" httpUri-2="https://zenodo.org/record/14771805/files/figure.png" httpUri-3="https://zenodo.org/record/14771807/files/figure.png" httpUri-4="https://zenodo.org/record/14771809/files/figure.png" httpUri-5="https://zenodo.org/record/14771811/files/figure.png" httpUri-6="https://zenodo.org/record/14771813/files/figure.png" httpUri-7="https://zenodo.org/record/14771815/files/figure.png" httpUri-8="https://zenodo.org/record/14771819/files/figure.png" httpUri-9="https://zenodo.org/record/14771823/files/figure.png" pageId="2" pageNumber="49">Figs 1–15</figureCitation>
, 
<figureCitation id="1304EFBDFFE2FF81FCA887D4852CF7C9" box="[845,874,923,950]" captionStart="Fig" captionStartId="26.[189,232,1857,1883]" captionTargetBox="[189,1398,1364,1818]" captionTargetId="figure-86@26.[189,1398,1364,1818]" captionTargetPageId="26" captionText="Fig. 25. Photographs of selected sampling sites. A. Longjiang Town, Qionghai City, Hainan Province, Wanquan River. (sampling sites of Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov. and A. (A.) bitergumus Zhang &amp; Guo, 2023). B. Diao Luo Mountain National Forest Park, Lingshui Li Autonomous County, Hainan Province, Diaoluo River (sampling site of A. (Tympanomegapus) tergumus Zhang &amp; Guo, 2023)." figureDoi="http://doi.org/10.5281/zenodo.14771858" httpUri="https://zenodo.org/record/14771858/files/figure.png" pageId="2" pageNumber="49">25</figureCitation>
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<emphasis id="B94B2F2AFFE2FF81FF5887928771F788" bold="true" box="[189,311,989,1015]" pageId="2" pageNumber="49">Diagnosis</emphasis>
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<emphasis id="B94B2F2AFFE2FF81FF0F8043876CF059" bold="true" box="[234,298,1036,1062]" pageId="2" pageNumber="49">Male</emphasis>
</paragraph>
<paragraph id="8B80F338FFE2FF81FF5880608518F0CD" blockId="2.[188,1399,1036,1203]" pageId="2" pageNumber="49">
Apodemes of ACG curved hook-shaped, and posterior margin of Cx-IV almost straight. Anterior margin of genital plate slightly indented, posterior margin slightly convex, three pairs of acetabula forming an obtuse triangle. 
<emphasis id="B94B2F2AFFE2FF81FE41803987F9F0E9" box="[420,447,1142,1174]" italics="true" pageId="2" pageNumber="49">
V 
<subScript id="17BBF17DFFE2FF81FE5380C787F9F0E9" attach="left" box="[438,447,1160,1174]" fontSize="6" pageId="2" pageNumber="49">1</subScript>
</emphasis>
separated from 
<emphasis id="B94B2F2AFFE2FF81FD61803984D9F0E9" box="[644,671,1142,1174]" italics="true" pageId="2" pageNumber="49">
V 
<subScript id="17BBF17DFFE2FF81FD7380C784D9F0E9" attach="left" box="[662,671,1160,1174]" fontSize="6" pageId="2" pageNumber="49">2</subScript>
</emphasis>
, excretory pore smooth. Sword seta on P-4 at same level of proximoventral hair. I-L-6 thick at base, tapering distally.
</paragraph>
<paragraph id="8B80F338FFE2FF81FF0F80958700F08B" blockId="2.[189,1399,1242,1338]" box="[234,326,1242,1268]" pageId="2" pageNumber="49">
<emphasis id="B94B2F2AFFE2FF81FF0F80958700F08B" bold="true" box="[234,326,1242,1268]" pageId="2" pageNumber="49">Female</emphasis>
</paragraph>
<paragraph id="8B80F338FFE2FF81FF5880B28746F145" blockId="2.[189,1399,1242,1338]" pageId="2" pageNumber="49">Similar to male. Ac in weakly curved line, Ac2 at middle of Ac1 and Ac3. Palp more slender than in male.</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFE2FF81FF58812D85BCF1D5" box="[189,1018,1378,1451]" pageId="2" pageNumber="49" type="etymology">
<paragraph id="8B80F338FFE2FF81FF58812D8703F103" blockId="2.[189,325,1378,1404]" box="[189,325,1378,1404]" pageId="2" pageNumber="49">
<heading id="D0C84454FFE2FF81FF58812D8703F103" bold="true" box="[189,325,1378,1404]" fontSize="11" level="3" pageId="2" pageNumber="49" reason="3">
<emphasis id="B94B2F2AFFE2FF81FF58812D8703F103" bold="true" box="[189,325,1378,1404]" pageId="2" pageNumber="49">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFE2FF81FF5881DF85BCF1D5" blockId="2.[189,1018,1424,1451]" box="[189,1018,1424,1451]" pageId="2" pageNumber="49">The new species is named after the heart-like genital plate of the male.</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFE2FF81FF58819D875EF373" pageId="2" pageNumber="49" type="materials_examined">
<paragraph id="8B80F338FFE2FF81FF58819D872BF194" blockId="2.[189,365,1489,1516]" box="[189,365,1489,1516]" pageId="2" pageNumber="49">
<heading id="D0C84454FFE2FF81FF58819D872BF194" bold="true" box="[189,365,1489,1516]" fontSize="11" level="3" pageId="2" pageNumber="49" reason="3">
<emphasis id="B94B2F2AFFE2FF81FF58819D872BF194" bold="true" box="[189,365,1489,1516]" pageId="2" pageNumber="49">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFE2FF81FF0F824F8337F2FB" blockId="2.[188,1398,1536,1668]" pageId="2" pageNumber="49">
<materialsCitation id="3B57F965FFE2FF81FF0F824F8337F2FB" collectingDate="2023-04-11" collectionCode="GUGC" collectorName="Hai-Tao Li &amp; Yu-Lin Zheng &amp; Yu-Hao Zhang" country="China" county="Qionghai City" elevation="4" latitude="19.173" location="Longjiang Town" longLatPrecision="7" longitude="110.3459" municipality="Longjiang Town" pageId="2" pageNumber="49" specimenCode="HN-HY-2023041101" specimenCount="1" specimenCount-male="1" stateProvince="Hainan Province" typeStatus="holotype">
<emphasis id="B94B2F2AFFE2FF81FF0F824F871AF265" bold="true" box="[234,348,1536,1562]" pageId="2" pageNumber="49">
<typeStatus id="54844D9AFFE2FF81FF0F824F871AF265" box="[234,348,1536,1562]" pageId="2" pageNumber="49" type="holotype">Holotype</typeStatus>
</emphasis>
P.R. 
<collectingCountry id="F328B3A8FFE2FF81FF17826C870BF242" box="[242,333,1571,1597]" name="China" pageId="2" pageNumber="49">CHINA</collectingCountry>
• 
<specimenCount id="9D3938B1FFE2FF81FE86826B873AF241" box="[355,380,1572,1598]" count="1" pageId="2" pageNumber="49" type="male">♂</specimenCount>
; 
<collectingRegion id="49FB3DDAFFE2FF81FE62826C840AF242" box="[391,588,1571,1597]" country="China" name="Hainan" pageId="2" pageNumber="49">Hainan Province</collectingRegion>
, 
<collectingCounty id="62E18BB4FFE2FF81FDB2826C84B8F242" box="[599,766,1571,1597]" pageId="2" pageNumber="49">Qionghai City</collectingCounty>
, 
<collectingMunicipality id="6BE46942FFE2FF81FCE2826C858DF241" box="[775,971,1571,1598]" pageId="2" pageNumber="49">Longjiang Town</collectingMunicipality>
; 
<geoCoordinate id="EE0B95FFFFE2FF81FC30826C8218F241" box="[981,1118,1571,1598]" degrees="19.1730" direction="north" orientation="latitude" pageId="2" pageNumber="49" precision="5" value="19.173">19.1730° N</geoCoordinate>
, 
<geoCoordinate id="EE0B95FFFFE2FF81FB8C826C82BAF241" box="[1129,1276,1571,1598]" degrees="110.3459" direction="east" orientation="longitude" pageId="2" pageNumber="49" precision="5" value="110.3459">110.3459° E</geoCoordinate>
; 
<elevation id="0012140BFFE2FF81FAE2826C8336F242" box="[1287,1392,1571,1598]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="49" unit="m" value="4.0">
<quantity id="4CC75EDDFFE2FF81FAE2826C8374F241" box="[1287,1330,1571,1598]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="49" unit="m" value="4.0">4 m</quantity>
a.s.l.
</elevation>
; 
<date id="FF81D5F8FFE2FF81FF5882088710F21E" box="[189,342,1607,1633]" pageId="2" pageNumber="49" value="2023-04-11">
<collectingDate id="EFC52C10FFE2FF81FF5882088710F21E" box="[189,342,1607,1633]" pageId="2" pageNumber="49" value="2023-04-11">11 Apr. 2023</collectingDate>
</date>
; 
<collectorName id="26CA96EEFFE2FF81FE84820987A2F21F" box="[353,484,1606,1632]" pageId="2" pageNumber="49">Hai-Tao Li</collectorName>
, 
<collectorName id="26CA96EEFFE2FF81FE0B820984D4F21F" box="[494,658,1606,1632]" pageId="2" pageNumber="49">Yu-Lin Zheng</collectorName>
and 
<collectorName id="26CA96EEFFE2FF81FD2D82088533F21F" box="[712,885,1606,1633]" pageId="2" pageNumber="49">Yu-Hao Zhang</collectorName>
leg.; main river (Wanquan River) with wide surface, slowly flowing, with sandy soil and humus on bottom; slide no. 
<specimenCode id="DB995B43FFE2FF81FBEA8225834DF2FB" box="[1039,1291,1642,1668]" pageId="2" pageNumber="49">HN-HY-2023041101</specimenCode>
; 
<collectionCode id="ED2E6BFDFFE2FF81FAF38225832BF2FB" box="[1302,1389,1642,1668]" pageId="2" pageNumber="49">GUGC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B80F338FFE2FF81FF0F82E4875EF373" blockId="2.[189,1398,1707,1804]" pageId="2" pageNumber="49">
<materialsCitation id="3B57F965FFE2FF81FF0F82E48752F373" collectingDate="2023-04-11" collectionCode="GUGC" collectorName="Hai-Tao Li &amp; Yu-Lin Zheng &amp; Yu-Hao Zhang" country="China" county="Qionghai City" elevation="4" latitude="19.173" location="CHINA" longLatPrecision="7" longitude="110.3459" municipality="Longjiang Town" pageId="2" pageNumber="49" specimenCode="HN-HY-2023041102 to 2023041107" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hainan" typeStatus="paratype">
<emphasis id="B94B2F2AFFE2FF81FF0F82E48721F2BA" bold="true" box="[234,359,1707,1733]" pageId="2" pageNumber="49">
<typeStatus id="54844D9AFFE2FF81FF0F82E48721F2BA" box="[234,359,1707,1733]" pageId="2" pageNumber="49" type="paratype">Paratypes</typeStatus>
</emphasis>
P.R. 
<collectingCountry id="F328B3A8FFE2FF81FF1682818708F297" box="[243,334,1742,1768]" name="China" pageId="2" pageNumber="49">CHINA</collectingCountry>
• 
<specimenCount id="9D3938B1FFE2FF81FE82828187ECF296" box="[359,426,1742,1769]" count="3" pageId="2" pageNumber="49" type="male">3 ♂♂</specimenCount>
, 
<specimenCount id="9D3938B1FFE2FF81FE52828187BFF296" box="[439,505,1742,1769]" count="3" pageId="2" pageNumber="49" type="female">3 ♀♀</specimenCount>
; same data as for holotype; slides nos. 
<specimenCode id="DB995B43FFE2FF81FC2482818335F297" box="[961,1395,1742,1769]" pageId="2" pageNumber="49">HN-HY-2023041102 to 2023041107</specimenCode>
; 
<collectionCode id="ED2E6BFDFFE2FF81FF5882BD8752F373" box="[189,276,1778,1804]" pageId="2" pageNumber="49">GUGC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFE3FF89FF588547878CF588" lastPageId="10" lastPageNumber="57" pageId="3" pageNumber="50" type="description">
<paragraph id="8B80F338FFE3FF80FF5885478709F55D" blockId="3.[189,335,264,290]" box="[189,335,264,290]" pageId="3" pageNumber="50">
<heading id="D0C84454FFE3FF80FF5885478709F55D" bold="true" box="[189,335,264,290]" fontSize="11" level="3" pageId="3" pageNumber="50" reason="3">
<emphasis id="B94B2F2AFFE3FF80FF5885478709F55D" bold="true" box="[189,335,264,290]" pageId="3" pageNumber="50">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFE3FF80FF0F857887C4F52E" blockId="3.[188,1400,311,549]" box="[234,386,311,338]" pageId="3" pageNumber="50">
<emphasis id="B94B2F2AFFE3FF80FF0F8578876CF52E" bold="true" box="[234,298,311,337]" pageId="3" pageNumber="50">Male</emphasis>
(n = 4)
</paragraph>
<paragraph id="8B80F338FFE3FF87FF5885158310F539" blockId="3.[188,1400,311,549]" lastBlockId="4.[189,1398,265,333]" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50">
Idiosoma soft and oval; 
<emphasis id="B94B2F2AFFE3FF80FE34851487B7F504" box="[465,497,347,379]" italics="true" pageId="3" pageNumber="50">
O 
<subScript id="17BBF17DFFE3FF80FE0D852287B7F504" attach="left" box="[488,497,365,379]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
between 
<emphasis id="B94B2F2AFFE3FF80FDB88514843FF504" box="[605,633,347,379]" italics="true" pageId="3" pageNumber="50">
A 
<subScript id="17BBF17DFFE3FF80FD958522843FF504" attach="left" box="[624,633,365,379]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
and 
<emphasis id="B94B2F2AFFE3FF80FD4B8514848CF504" box="[686,714,347,379]" italics="true" pageId="3" pageNumber="50">
A 
<subScript id="17BBF17DFFE3FF80FD248522848CF504" attach="right" box="[705,714,365,379]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
, but closer to 
<emphasis id="B94B2F2AFFE3FF80FC8F851485C0F504" box="[874,902,347,379]" italics="true" pageId="3" pageNumber="50">
A 
<subScript id="17BBF17DFFE3FF80FC98852285C0F504" attach="right" box="[893,902,365,379]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE3FF80FC76851485F7F504" box="[915,945,347,379]" italics="true" pageId="3" pageNumber="50">
O 
<subScript id="17BBF17DFFE3FF80FC4D852285F7F504" attach="left" box="[936,945,365,379]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
between 
<emphasis id="B94B2F2AFFE3FF80FBF88514827BF504" box="[1053,1085,347,379]" italics="true" pageId="3" pageNumber="50">
D 
<subScript id="17BBF17DFFE3FF80FBD18522827BF504" attach="left" box="[1076,1085,365,379]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
and at same level of 
<emphasis id="B94B2F2AFFE3FF80FAC98514830DF504" box="[1324,1355,347,379]" italics="true" pageId="3" pageNumber="50">
D 
<subScript id="17BBF17DFFE3FF80FAA78522830DF504" attach="right" box="[1346,1355,365,379]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
; all slit organs visible, 
<emphasis id="B94B2F2AFFE3FF80FE70853187FBF5E1" box="[405,445,382,414]" italics="true" pageId="3" pageNumber="50">
So 
<subScript id="17BBF17DFFE3FF80FE5185DF87FBF5E1" attach="left" box="[436,445,400,414]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
near 
<emphasis id="B94B2F2AFFE3FF80FE1F85318450F5E1" box="[506,534,382,414]" italics="true" pageId="3" pageNumber="50">
A 
<subScript id="17BBF17DFFE3FF80FDE885DF8450F5E1" attach="right" box="[525,534,400,414]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE3FF80FDC68531840CF5E1" box="[547,586,382,414]" italics="true" pageId="3" pageNumber="50">
So 
<subScript id="17BBF17DFFE3FF80FDA485DF840CF5E1" attach="left" box="[577,586,400,414]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
at same level of 
<emphasis id="B94B2F2AFFE3FF80FCEF8531856FF5E1" box="[778,809,382,414]" italics="true" pageId="3" pageNumber="50">
D 
<subScript id="17BBF17DFFE3FF80FCC585DF856FF5E1" attach="left" box="[800,809,400,414]" fontSize="6" pageId="3" pageNumber="50">1</subScript>
</emphasis>
and 
<emphasis id="B94B2F2AFFE3FF80FCBB85318538F5E1" box="[862,894,382,414]" italics="true" pageId="3" pageNumber="50">
O 
<subScript id="17BBF17DFFE3FF80FC9085DF8538F5E1" attach="right" box="[885,894,400,414]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE3FF80FC6E853185F4F5E1" box="[907,946,382,414]" italics="true" pageId="3" pageNumber="50">
So 
<subScript id="17BBF17DFFE3FF80FC4C85DF85F4F5E1" attach="left" box="[937,946,400,414]" fontSize="6" pageId="3" pageNumber="50">3</subScript>
</emphasis>
at same level of 
<emphasis id="B94B2F2AFFE3FF80FB94853182D7F5E1" box="[1137,1169,382,414]" italics="true" pageId="3" pageNumber="50">
D 
<subScript id="17BBF17DFFE3FF80FB6D85DF82D7F5E1" attach="right" box="[1160,1169,400,414]" fontSize="6" pageId="3" pageNumber="50">2</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE3FF80FB7B85318283F5E1" box="[1182,1221,382,414]" italics="true" pageId="3" pageNumber="50">
So 
<subScript id="17BBF17DFFE3FF80FB5985DF8283F5E1" attach="left" box="[1212,1221,400,414]" fontSize="6" pageId="3" pageNumber="50">4</subScript>
</emphasis>
close to 
<emphasis id="B94B2F2AFFE3FF80FACD85318305F5E1" box="[1320,1347,382,414]" italics="true" pageId="3" pageNumber="50">
L 
<subScript id="17BBF17DFFE3FF80FADF85DF8305F5E1" attach="right" box="[1338,1347,400,414]" fontSize="6" pageId="3" pageNumber="50">4</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE3FF80FAB585318331F5E1" box="[1360,1399,382,414]" italics="true" pageId="3" pageNumber="50">
So 
<subScript id="17BBF17DFFE3FF80FA8B85DF8331F5E1" attach="left" box="[1390,1399,400,414]" fontSize="6" pageId="3" pageNumber="50">5</subScript>
</emphasis>
behind of 
<emphasis id="B94B2F2AFFE3FF80FED285EE8712F5BD" box="[311,340,417,450]" italics="true" pageId="3" pageNumber="50">D 4</emphasis>
(
<figureCitation id="1304EFBDFFE3FF80FE8185EE8781F5C4" box="[356,455,417,443]" captionStart="Fig" captionStartId="4.[189,232,1927,1953]" captionTargetBox="[258,1346,486,1872]" captionTargetId="figure-66@4.[205,1383,465,1892]" captionTargetPageId="4" captionText="Fig. 2. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771803" httpUri="https://zenodo.org/record/14771803/files/figure.png" pageId="3" pageNumber="50">Figs 2A</figureCitation>
, 
<figureCitation id="1304EFBDFFE3FF80FE3185EF87BAF5C5" box="[468,508,416,442]" captionStart="Fig" captionStartId="8.[190,233,1928,1954]" captionTargetBox="[210,1378,731,1888]" captionTargetId="figure-182@8.[210,1378,731,1888]" captionTargetPageId="8" captionText="Fig. 6. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC), light microscope photographs. A. Idiosoma, dorsal view.B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771811" httpUri="https://zenodo.org/record/14771811/files/figure.png" pageId="3" pageNumber="50">6A</figureCitation>
). ACG fused together and with suture line, apodemes of ACG curved hook shape; PCG separated, posterior margin of Cx-IV almost straight, 
<emphasis id="B94B2F2AFFE3FF80FC3E858B85B0F59A" box="[987,1014,452,485]" italics="true" pageId="3" pageNumber="50">C 4</emphasis>
near suture line between Cx-III and Cx-IV, suture line of Cx-III and Cx-IV in medial part nearly straight (
<figureCitation id="1304EFBDFFE3FF80FBCD85A882C1F67E" box="[1064,1159,487,513]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[189,1398,1470,1924]" captionTargetId="figure-56@9.[189,1398,1470,1924]" captionTargetPageId="9" captionText="Fig. 8. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HN- HY-2023041103, GUGC), SEM photographs. A. ACG and PCG. B. V 1, V 2 and excretory pore." figureDoi="http://doi.org/10.5281/zenodo.14771815" httpUri="https://zenodo.org/record/14771815/files/figure.png" pageId="3" pageNumber="50">Fig. 8A</figureCitation>
). Anterior margin of genital plate slightly indented, posterior margin slightly convex, three pairs of acetabula forming obtuse triangle, Ac2 near Ac1 (
<figureCitation id="1304EFBDFFE4FF87FE3385468473F55C" box="[470,565,265,291]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,983,1890]" captionTargetId="figure-222@10.[189,1398,983,1890]" captionTargetPageId="10" captionText="Fig. 9. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HN- HY-2023041103, GUGC), SEM photographs. A. Palp. B. Sword seta of P-4. C. Gnathosoma. D. Genital field." figureDoi="http://doi.org/10.5281/zenodo.14771819" httpUri="https://zenodo.org/record/14771819/files/figure.png" pageId="4" pageNumber="51">Fig. 9D</figureCitation>
). 
<emphasis id="B94B2F2AFFE4FF87FDAF85458423F555" box="[586,613,266,298]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FDB985538423F555" attach="left" box="[604,613,284,298]" fontSize="6" pageId="4" pageNumber="51">1</subScript>
</emphasis>
separated from 
<emphasis id="B94B2F2AFFE4FF87FCC685458578F555" box="[803,830,266,298]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FCD085538578F555" attach="left" box="[821,830,284,298]" fontSize="6" pageId="4" pageNumber="51">2</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE4FF87FCA88545852EF555" box="[845,872,266,298]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FCBA8553852EF555" attach="left" box="[863,872,284,298]" fontSize="6" pageId="4" pageNumber="51">2</subScript>
</emphasis>
posterior to 
<emphasis id="B94B2F2AFFE4FF87FC1B8545825EF555" box="[1022,1048,266,298]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FBEA8553825EF555" attach="left" box="[1039,1048,284,298]" fontSize="6" pageId="4" pageNumber="51">1</subScript>
</emphasis>
(
<figureCitation id="1304EFBDFFE4FF87FBCD854682C2F55C" box="[1064,1156,265,291]" captionStart="Fig" captionStartId="9.[189,232,1963,1989]" captionTargetBox="[189,1398,1470,1924]" captionTargetId="figure-56@9.[189,1398,1470,1924]" captionTargetPageId="9" captionText="Fig. 8. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HN- HY-2023041103, GUGC), SEM photographs. A. ACG and PCG. B. V 1, V 2 and excretory pore." figureDoi="http://doi.org/10.5281/zenodo.14771815" httpUri="https://zenodo.org/record/14771815/files/figure.png" pageId="4" pageNumber="51">Fig. 8B</figureCitation>
), 
<emphasis id="B94B2F2AFFE4FF87FB7E854582F3F555" box="[1179,1205,266,298]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FB49855382F3F555" attach="left" box="[1196,1205,284,298]" fontSize="6" pageId="4" pageNumber="51">4</subScript>
</emphasis>
at same level of Ac2, 
<emphasis id="B94B2F2AFFE4FF87FF188562875EF532" box="[253,280,301,333]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FEEA8570875EF532" attach="left" box="[271,280,319,333]" fontSize="6" pageId="4" pageNumber="51">3</subScript>
</emphasis>
and 
<emphasis id="B94B2F2AFFE4FF87FEB58562872DF532" box="[336,363,301,333]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FE878570872DF532" attach="left" box="[354,363,319,333]" fontSize="6" pageId="4" pageNumber="51">4</subScript>
</emphasis>
arranged in inverted trapezoid; excretory pore smooth and between 
<emphasis id="B94B2F2AFFE4FF87FB69856282E1F532" box="[1164,1191,301,333]" italics="true" pageId="4" pageNumber="51">
V 
<subScript id="17BBF17DFFE4FF87FB7B857082E1F532" attach="left" box="[1182,1191,319,333]" fontSize="6" pageId="4" pageNumber="51">2</subScript>
</emphasis>
(
<figureCitation id="1304EFBDFFE4FF87FB5285638351F539" box="[1207,1303,300,326]" captionStart="Fig" captionStartId="4.[189,232,1927,1953]" captionTargetBox="[258,1346,486,1872]" captionTargetId="figure-66@4.[205,1383,465,1892]" captionTargetPageId="4" captionText="Fig. 2. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771803" httpUri="https://zenodo.org/record/14771803/files/figure.png" pageId="4" pageNumber="51">Figs 2B</figureCitation>
, 
<figureCitation id="1304EFBDFFE4FF87FAC68563830EF539" box="[1315,1352,300,326]" captionStart="Fig" captionStartId="8.[190,233,1928,1954]" captionTargetBox="[210,1378,731,1888]" captionTargetId="figure-182@8.[210,1378,731,1888]" captionTargetPageId="8" captionText="Fig. 6. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC), light microscope photographs. A. Idiosoma, dorsal view.B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771811" httpUri="https://zenodo.org/record/14771811/files/figure.png" pageId="4" pageNumber="51">6B</figureCitation>
).
</paragraph>
<caption id="DF40A3B0FFE3FF80FF59832C8728F398" ID-DOI="http://doi.org/10.5281/zenodo.14771801" ID-Zenodo-Dep="14771801" httpUri="https://zenodo.org/record/14771801/files/figure.png" pageId="3" pageNumber="50" startId="3.[188,231,1891,1917]" targetBox="[189,1398,662,1853]" targetPageId="3">
<paragraph id="8B80F338FFE3FF80FF59832C8728F398" blockId="3.[188,1399,1891,2023]" pageId="3" pageNumber="50">
<emphasis id="B94B2F2AFFE3FF80FF59832C8756F302" bold="true" box="[188,272,1891,1917]" pageId="3" pageNumber="50">Fig. 1.</emphasis>
<taxonomicName id="4C3F88BBFFE3FF80FEF9832C852EF302" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[284,872,1891,1917]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE3FF80FEF9832C87D0F302" box="[284,406,1891,1917]" italics="true" pageId="3" pageNumber="50">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE3FF80FE49832C8460F302" box="[428,550,1891,1917]" italics="true" pageId="3" pageNumber="50">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE3FF80FDD9832C84F6F302" box="[572,688,1891,1917]" italics="true" pageId="3" pageNumber="50">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE3FF80FC96832B8595F301" box="[883,979,1892,1918]" pageId="3" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94B2F2AFFE3FF80FC3B832C8250F302" bold="true" box="[990,1046,1891,1918]" pageId="3" pageNumber="50">A–B</emphasis>
. Paratype, ♂ (slide no. HNHY-2023041102, GUGC). 
<emphasis id="B94B2F2AFFE3FF80FDE483C9847CF3DF" bold="true" box="[513,570,1926,1953]" pageId="3" pageNumber="50">C–D</emphasis>
. Paratype, ♀ (slide no. HN-HY-2023041106, GUGC). 
<emphasis id="B94B2F2AFFE3FF80FB3F83C982A9F3DF" bold="true" box="[1242,1263,1926,1952]" pageId="3" pageNumber="50">A</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE3FF80FEBC83E5872BF3BB" bold="true" box="[345,365,1962,1988]" pageId="3" pageNumber="50">B</emphasis>
. Idiosoma, ventral view. 
<emphasis id="B94B2F2AFFE3FF80FD4083E684FCF3BC" bold="true" box="[677,698,1961,1987]" pageId="3" pageNumber="50">C</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE3FF80FC0C83E585B8F3BB" bold="true" box="[1001,1022,1962,1988]" pageId="3" pageNumber="50">D</emphasis>
. Idiosoma, ventral view. Scale bars = 200 μm.
</paragraph>
</caption>
<caption id="DF40A3B0FFE4FF87FF5883C88410F397" ID-DOI="http://doi.org/10.5281/zenodo.14771803" ID-Zenodo-Dep="14771803" httpUri="https://zenodo.org/record/14771803/files/figure.png" pageId="4" pageNumber="51" startId="4.[189,232,1927,1953]" targetBox="[258,1346,486,1872]" targetPageId="4">
<paragraph id="8B80F338FFE4FF87FF5883C88410F397" blockId="4.[187,1399,1927,2024]" pageId="4" pageNumber="51">
<emphasis id="B94B2F2AFFE4FF87FF5883C88753F3DE" bold="true" box="[189,277,1927,1953]" pageId="4" pageNumber="51">Fig. 2.</emphasis>
<taxonomicName id="4C3F88BBFFE4FF87FEC383C885CCF3DE" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,906,1927,1953]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="51" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE4FF87FEC383C887E6F3DE" box="[294,416,1927,1953]" italics="true" pageId="4" pageNumber="51">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE4FF87FE5F83C88472F3DE" box="[442,564,1927,1953]" italics="true" pageId="4" pageNumber="51">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE4FF87FDAA83C88485F3DE" box="[591,707,1927,1953]" italics="true" pageId="4" pageNumber="51">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE4FF87FC7F83C78246F3DD" box="[922,1024,1928,1954]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (slide no. HNHY-2023041101, GUGC). 
<emphasis id="B94B2F2AFFE4FF87FE1683E5844EF3BB" bold="true" box="[499,520,1962,1988]" pageId="4" pageNumber="51">A</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE4FF87FCC583E48572F3BA" bold="true" box="[800,820,1963,1989]" pageId="4" pageNumber="51">B</emphasis>
. Idiosoma, ventral view. 
<emphasis id="B94B2F2AFFE4FF87FBB183E5822FF3BB" bold="true" box="[1108,1129,1962,1988]" pageId="4" pageNumber="51">C</emphasis>
. Palp. 
<emphasis id="B94B2F2AFFE4FF87FB5283E4828AF3BA" bold="true" box="[1207,1228,1963,1989]" pageId="4" pageNumber="51">D</emphasis>
. Gnathosoma. 
<emphasis id="B94B2F2AFFE4FF87FF5E83818689F397" bold="true" box="[187,207,1998,2024]" pageId="4" pageNumber="51">E</emphasis>
. Chelicera. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8B80F338FFE5FF86FF5885468524F5AC" blockId="5.[188,1400,265,467]" pageId="5" pageNumber="52">
Palp five-segmented, without obvious sexual dimorphism; P-1 short, P-2 and P-3 short with straight ventral margin, P-4 with some dorsal hairs and two ventral hairs, dividing P- 
<quantity id="4CC75EDDFFE5FF86FBB4856382C6F539" box="[1105,1152,300,326]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="52" unit="in" value="4.0">4 in</quantity>
sectors 3:2:2, sword seta at same level of proximoventral hair (
<figureCitation id="1304EFBDFFE5FF86FD4B85008548F516" box="[686,782,335,361]" captionStart="Fig" captionStartId="4.[189,232,1927,1953]" captionTargetBox="[258,1346,486,1872]" captionTargetId="figure-66@4.[205,1383,465,1892]" captionTargetPageId="4" captionText="Fig. 2. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC).A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp.D. Gnathosoma. E. Chelicera. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771803" httpUri="https://zenodo.org/record/14771803/files/figure.png" pageId="5" pageNumber="52">Figs 2C</figureCitation>
, 
<figureCitation id="1304EFBDFFE5FF86FCFF85008523F516" box="[794,869,335,361]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1398,983,1890]" captionTargetId="figure-222@10.[189,1398,983,1890]" captionTargetPageId="10" captionText="Fig. 9. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HN- HY-2023041103, GUGC), SEM photographs. A. Palp. B. Sword seta of P-4. C. Gnathosoma. D. Genital field." figureDoi="http://doi.org/10.5281/zenodo.14771819" httpUri="https://zenodo.org/record/14771819/files/figure.png" pageId="5" pageNumber="52">9A–B</figureCitation>
). Ventral margin of I-L-5 slightly longer than dorsal margin, S-1 close to S-2 and S-2 thickens at three-fifth position and then narrows immediately, whip-like seta at anterior distal margin; I-L-6 thick at base, distally tapering. IV-L-5 with swimming seta at distal margin and almost as long as IV-L-6 (
<figureCitation id="1304EFBDFFE5FF86FD3E85F68561F5AC" box="[731,807,441,467]" captionStart="Fig" captionStartId="5.[189,232,1962,1988]" captionTargetBox="[287,1304,594,1883]" captionTargetId="figure-104@5.[241,1346,559,1922]" captionTargetPageId="5" captionText="Fig. 3. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., holotype, ♂ (slide no. HN- HY-2023041101, GUGC), I-L– IV-L. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771805" httpUri="https://zenodo.org/record/14771805/files/figure.png" pageId="5" pageNumber="52">Figs 3</figureCitation>
, 
<figureCitation id="1304EFBDFFE5FF86FCD185F68513F5AC" box="[820,853,441,467]" captionStart="Fig" captionStartId="11.[189,232,1078,1104]" captionTargetBox="[277,1311,265,1041]" captionTargetId="figure-11@11.[277,1311,265,1041]" captionTargetPageId="11" captionText="Fig. 10. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♂ (slide no. HN- HY-2023041103, GUGC), SEM photographs. A. I-L. B. I-L-5–6. C. I-L. D. I-L-5–6." figureDoi="http://doi.org/10.5281/zenodo.14771823" httpUri="https://zenodo.org/record/14771823/files/figure.png" pageId="5" pageNumber="52">10</figureCitation>
).
</paragraph>
<caption id="DF40A3B0FFE5FF86FF5883E5852EF397" ID-DOI="http://doi.org/10.5281/zenodo.14771805" ID-Zenodo-Dep="14771805" httpUri="https://zenodo.org/record/14771805/files/figure.png" pageId="5" pageNumber="52" startId="5.[189,232,1962,1988]" targetBox="[287,1304,594,1883]" targetPageId="5">
<paragraph id="8B80F338FFE5FF86FF5883E5852EF397" blockId="5.[189,1398,1962,2024]" pageId="5" pageNumber="52">
<emphasis id="B94B2F2AFFE5FF86FF5883E58753F3BB" bold="true" box="[189,277,1962,1988]" pageId="5" pageNumber="52">Fig. 3.</emphasis>
<taxonomicName id="4C3F88BBFFE5FF86FEC383E585CCF3BB" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,906,1962,1988]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="52" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE5FF86FEC383E587E6F3BB" box="[294,416,1962,1988]" italics="true" pageId="5" pageNumber="52">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE5FF86FE5F83E58472F3BB" box="[442,564,1962,1988]" italics="true" pageId="5" pageNumber="52">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE5FF86FDAA83E58485F3BB" box="[591,707,1962,1988]" italics="true" pageId="5" pageNumber="52">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE5FF86FC7F83E58246F3BB" box="[922,1024,1962,1988]" pageId="5" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (slide no. HNHY-2023041101, GUGC), I-L– IV-L. Scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="8B80F338FFE6FF85FF5885468330F516" blockId="6.[189,1399,265,362]" pageId="6" pageNumber="53">
<smallCapsWord id="8D6665E4FFE6FF85FF5885468728F55E" baselines="284,284" box="[189,366,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="6" pageNumber="53">MEASUREMENTS</smallCapsWord>
(measurements of 
<typeStatus id="54844D9AFFE6FF85FDB685468483F55C" box="[595,709,265,291]" pageId="6" pageNumber="53" type="paratype">paratypes</typeStatus>
in parentheses). Idiosoma L 450 (420–460), W 385 (371– 386); coxal field L 245 (216–245), Cx-III W 263 (238–263), ACG IL 180 (163–180), mL 116 (112–123), W 192 (183–192); gnathosoma bay L 83 (68–83); genital field L 81 (70–82), W 109 (92–109); Ac1–3
</paragraph>
<caption id="DF40A3B0FFE6FF85FF5883C78410F396" ID-DOI="http://doi.org/10.5281/zenodo.14771807" ID-Zenodo-Dep="14771807" httpUri="https://zenodo.org/record/14771807/files/figure.png" pageId="6" pageNumber="53" startId="6.[189,232,1928,1954]" targetBox="[322,1241,445,1857]" targetPageId="6">
<paragraph id="8B80F338FFE6FF85FF5883C78410F396" blockId="6.[187,1399,1928,2025]" pageId="6" pageNumber="53">
<emphasis id="B94B2F2AFFE6FF85FF5883C78753F3DD" bold="true" box="[189,277,1928,1954]" pageId="6" pageNumber="53">Fig. 4.</emphasis>
<taxonomicName id="4C3F88BBFFE6FF85FEC383C785CCF3DD" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,906,1928,1954]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE6FF85FEC383C787E6F3DD" box="[294,416,1928,1954]" italics="true" pageId="6" pageNumber="53">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE6FF85FE5F83C78472F3DD" box="[442,564,1928,1954]" italics="true" pageId="6" pageNumber="53">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE6FF85FDAA83C78485F3DD" box="[591,707,1928,1954]" italics="true" pageId="6" pageNumber="53">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE6FF85FC7E83C78247F3DD" box="[923,1025,1928,1954]" pageId="6" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (slide no. HNHY-2023041105, GUGC). 
<emphasis id="B94B2F2AFFE6FF85FE1083E4844CF3BA" bold="true" box="[501,522,1963,1989]" pageId="6" pageNumber="53">A</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE6FF85FCC783E48570F3BA" bold="true" box="[802,822,1963,1989]" pageId="6" pageNumber="53">B</emphasis>
. Idiosoma, ventral view. 
<emphasis id="B94B2F2AFFE6FF85FBBD83E4822BF3BA" bold="true" box="[1112,1133,1963,1989]" pageId="6" pageNumber="53">C</emphasis>
. Palp. 
<emphasis id="B94B2F2AFFE6FF85FB5983E48297F3BA" bold="true" box="[1212,1233,1963,1989]" pageId="6" pageNumber="53">D</emphasis>
. Pnathosoma. 
<emphasis id="B94B2F2AFFE6FF85FF5E83818689F397" bold="true" box="[187,207,1998,2024]" pageId="6" pageNumber="53">E</emphasis>
. Chelicera. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8B80F338FFE7FF84FF5885468520F63F" blockId="7.[188,1401,265,577]" pageId="7" pageNumber="54">L 20 (19–21), 22 (20–22), 24 (21–24); chelicera L 166 (155–171), cheliceral claw L 43 (42–47), basal segment L 126 (123–138); gnathosoma L 136 (131–139); palp dL: P-1 22 (19–22), P-2 51 (48–54), P-3 49 (45–50), P-4 73 (67–74), P-5 27 (24–28); leg segments: I-L-1 dL 39 (36–41), I-L-2 dL 66 (65–73), I-L-3 dL 80 (74–84), I-L-4 dL 125 (115–128), I-L-5 dL 117 (103–117), HB 37 (33–37), I-L-6 dL 65 (65–73), HB 15 (11–15), S-1 L 61 (57–61), W 6 (5–6), S-2 L 57 (55–59), W 7 (6–8); dL: II-L-1 39 (37– 43), II-L-2 62 (55–71), II-L-3 68 (60–69), II-L-4 89 (76–90), II-L-5 98 (86–104), II-L-6 85 (82–91); dL: III-L-1 42 (36–46), III-L-2 62 (61–66), III-L-3 67 (64–69), III-L-4 96 (87–96), III-L-5 112 (101–112), III-L-6 108 (95–108); dL: IV-L-1 88 (78–88), IV-L-2 85 (71–85), IV-L-3 106 (102–109), IV-L-4 122 (118–127), IV-L-5 137 (125–139), IV-L-6 134 (118–134).</paragraph>
<caption id="DF40A3B0FFE7FF84FF5883E585F6F397" ID-DOI="http://doi.org/10.5281/zenodo.14771809" ID-Zenodo-Dep="14771809" httpUri="https://zenodo.org/record/14771809/files/figure.png" pageId="7" pageNumber="54" startId="7.[189,232,1962,1988]" targetBox="[205,1385,667,1913]" targetPageId="7">
<paragraph id="8B80F338FFE7FF84FF5883E585F6F397" blockId="7.[189,1398,1962,2024]" pageId="7" pageNumber="54">
<emphasis id="B94B2F2AFFE7FF84FF5883E58753F3BB" bold="true" box="[189,277,1962,1988]" pageId="7" pageNumber="54">Fig. 5.</emphasis>
<taxonomicName id="4C3F88BBFFE7FF84FEC383E585CDF3BB" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,907,1962,1988]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE7FF84FEC383E587E6F3BB" box="[294,416,1962,1988]" italics="true" pageId="7" pageNumber="54">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE7FF84FE5E83E58473F3BB" box="[443,565,1962,1988]" italics="true" pageId="7" pageNumber="54">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE7FF84FDAA83E58485F3BB" box="[591,707,1962,1988]" italics="true" pageId="7" pageNumber="54">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE7FF84FC7E83E58244F3BB" box="[923,1026,1962,1988]" pageId="7" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (slide no. HNHY-2023041105, GUGC). 
<emphasis id="B94B2F2AFFE7FF84FE1F83828475F398" bold="true" box="[506,563,1997,2024]" pageId="7" pageNumber="54">A–D</emphasis>
. I-L– IV-L. Scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="8B80F338FFE8FF8BFF0F854687DBF55C" blockId="8.[189,1399,265,433]" box="[234,413,265,291]" pageId="8" pageNumber="55">
<emphasis id="B94B2F2AFFE8FF8BFF0F85468703F55C" bold="true" box="[234,325,265,291]" pageId="8" pageNumber="55">Female</emphasis>
(n = 3)
</paragraph>
<paragraph id="8B80F338FFE8FF8BFF588563875EF5CE" blockId="8.[189,1399,265,433]" pageId="8" pageNumber="55">
Similar to male. Ac in weakly curved line, Ac2 at middle of Ac1 and Ac3, 
<emphasis id="B94B2F2AFFE8FF8BFBC58562827AF532" box="[1056,1084,301,333]" italics="true" pageId="8" pageNumber="55">
V 
<subScript id="17BBF17DFFE8FF8BFBD68570827AF532" attach="left" box="[1075,1084,319,333]" fontSize="6" pageId="8" pageNumber="55">3</subScript>
</emphasis>
at same level of pregenital sclerite, distance between pregenital sclerite and postgenital sclerite longer, 
<emphasis id="B94B2F2AFFE8FF8BFB82851F82C4F50F" box="[1127,1154,336,368]" italics="true" pageId="8" pageNumber="55">V 3</emphasis>
and 
<emphasis id="B94B2F2AFFE8FF8BFB20851F82A6F50F" box="[1221,1248,336,368]" italics="true" pageId="8" pageNumber="55">V 4</emphasis>
arranged in trapezoid; excretory pore slightly posterior to level of 
<emphasis id="B94B2F2AFFE8FF8BFCDA853C851FF5EC" box="[831,857,371,403]" italics="true" pageId="8" pageNumber="55">
V 
<subScript id="17BBF17DFFE8FF8BFCB585CA851FF5EC" attach="left" box="[848,857,389,403]" fontSize="6" pageId="8" pageNumber="55">2</subScript>
</emphasis>
. Palp more slender than in male (
<figureCitation id="1304EFBDFFE8FF8BFB0C853D8312F5F2" box="[1257,1364,370,397]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1928,1954]" captionStartId-1="7.[189,232,1962,1988]" captionTargetBox-0="[322,1241,445,1857]" captionTargetBox-1="[205,1385,667,1913]" captionTargetId-0="figure-70@6.[298,1290,416,1890]" captionTargetId-1="figure-158@7.[189,1398,642,1925]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 4. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HN- HY-2023041105, GUGC). A. Idiosoma, dorsal view.B. Idiosoma, ventral view.C. Palp. D. Pnathosoma. E. Chelicera. Scale bars = 100 μm." captionText-1="Fig. 5. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HN- HY-2023041105, GUGC). A–D. I-L– IV-L. Scale bar = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.14771807" figureDoi-1="http://doi.org/10.5281/zenodo.14771809" httpUri-0="https://zenodo.org/record/14771807/files/figure.png" httpUri-1="https://zenodo.org/record/14771809/files/figure.png" pageId="8" pageNumber="55">Figs 4–5</figureCitation>
, 
<figureCitation id="1304EFBDFFE8FF8BFA85853C8337F5F2" box="[1376,1393,371,397]" captionStart="Fig" captionStartId="9.[189,232,1325,1351]" captionTargetBox="[246,1342,265,1285]" captionTargetId="figure-11@9.[246,1342,265,1285]" captionTargetPageId="9" captionText="Fig. 7. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HN- HY-2023041105, GUGC), light microscope photographs. A. Idiosoma, dorsal view.B. Idiosoma, ventral view. C. Idiosoma lateral view. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.14771813" httpUri="https://zenodo.org/record/14771813/files/figure.png" pageId="8" pageNumber="55">7</figureCitation>
, 
<figureCitation id="1304EFBDFFE8FF8BFF5B85D9874DF5CE" box="[190,267,406,433]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="12.[189,232,1024,1050]" captionStartId-1="12.[189,232,1963,1989]" captionStartId-2="13.[189,232,1963,1989]" captionStartId-3="14.[189,232,1963,1989]" captionTargetBox-0="[315,1272,265,983]" captionTargetBox-1="[289,1298,1168,1925]" captionTargetBox-2="[189,1398,1016,1923]" captionTargetBox-3="[189,1398,561,1922]" captionTargetId-0="figure-19@12.[315,1272,265,983]" captionTargetId-1="figure-63@12.[289,1298,1168,1925]" captionTargetId-2="figure-11@13.[189,1398,1016,1923]" captionTargetId-3="figure-18@14.[189,1398,561,1922]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="Fig. 12. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. Gnathosoma. B. ACG and PCG. C. Genital field. D. V 1, V 2 and excretory pore." captionText-1="Fig. 13. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. Palp. B. P-4 and P-5. C. palp. D. Sword seta of P-4." captionText-2="Fig. 14. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. I-L. B. I-L-5–6. C. I-L. D. I-L-5–6." captionText-3="Fig. 15. Atractides (Atractides) cardiacus Zhang &amp; Guo sp. nov., paratype, ♀ (slide no. HNHY-2023041107, GUGC), SEM photographs. A. II-L. B. II-L. C. III-L. D. III-L. E. IV-L. F. IV-L." figureDoi-0="http://doi.org/10.5281/zenodo.14771827" figureDoi-1="http://doi.org/10.5281/zenodo.14771829" figureDoi-2="http://doi.org/10.5281/zenodo.14771833" figureDoi-3="http://doi.org/10.5281/zenodo.14771835" httpUri-0="https://zenodo.org/record/14771827/files/figure.png" httpUri-1="https://zenodo.org/record/14771829/files/figure.png" httpUri-2="https://zenodo.org/record/14771833/files/figure.png" httpUri-3="https://zenodo.org/record/14771835/files/figure.png" pageId="8" pageNumber="55">12–15</figureCitation>
).
</paragraph>
<paragraph id="8B80F338FFE8FF8BFF5885978331F6DE" blockId="8.[189,1400,471,674]" pageId="8" pageNumber="55">
<smallCapsWord id="8D6665E4FFE8FF8BFF5885978728F58F" baselines="491,491" box="[189,366,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="8" pageNumber="55">MEASUREMENTS</smallCapsWord>
(measurements of 
<typeStatus id="54844D9AFFE8FF8BFDB685978483F58D" box="[595,709,472,498]" pageId="8" pageNumber="55" type="paratype">paratypes</typeStatus>
in parentheses). Idiosoma L 783 (722–807), W 621 (621– 678); coxal field L 254 (254–293), Cx-III W 327 (327–363),ACG IL 176 (176–212), mL 101 (101–115), W 223 (223–259); gnathosoma bay L 97 (97–120); gonopore L 141 (99–141), pregenital sclerite L 79 (61–79), postgenital sclerite L 37 (33–40); Ac1–3 L 22 (22–26), 28 (23–28), 25 (25–28); chelicera L 219 (219–240), cheliceral claw L 58 (58–66), basal segment L 175 (175–186); gnathosoma L 146 (146–184); palp dL: P-1 26 (25–26), P-2 64 (64–77), P-3 65 (65–74), P-4 85 (85–96), P-5 31 (31–34);
</paragraph>
<caption id="DF40A3B0FFE8FF8BFF5B83C7856FF396" ID-DOI="http://doi.org/10.5281/zenodo.14771811" ID-Zenodo-Dep="14771811" httpUri="https://zenodo.org/record/14771811/files/figure.png" pageId="8" pageNumber="55" startId="8.[190,233,1928,1954]" targetBox="[210,1378,731,1888]" targetPageId="8">
<paragraph id="8B80F338FFE8FF8BFF5B83C7856FF396" blockId="8.[187,1398,1928,2025]" pageId="8" pageNumber="55">
<emphasis id="B94B2F2AFFE8FF8BFF5B83C78753F3DD" bold="true" box="[190,277,1928,1954]" pageId="8" pageNumber="55">Fig. 6.</emphasis>
<taxonomicName id="4C3F88BBFFE8FF8BFEC383C785CCF3DD" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,906,1928,1954]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE8FF8BFEC383C787E6F3DD" box="[294,416,1928,1954]" italics="true" pageId="8" pageNumber="55">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE8FF8BFE5F83C78472F3DD" box="[442,564,1928,1954]" italics="true" pageId="8" pageNumber="55">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE8FF8BFDAA83C78485F3DD" box="[591,707,1928,1954]" italics="true" pageId="8" pageNumber="55">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE8FF8BFC7F83C78246F3DD" box="[922,1024,1928,1954]" pageId="8" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (slide no. HNHY-2023041101, GUGC), light microscope photographs. 
<emphasis id="B94B2F2AFFE8FF8BFCBE83E48536F3BA" bold="true" box="[859,880,1963,1989]" pageId="8" pageNumber="55">A</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE8FF8BFB6D83E482DAF3BA" bold="true" box="[1160,1180,1963,1989]" pageId="8" pageNumber="55">B</emphasis>
. Idiosoma, ventral view. 
<emphasis id="B94B2F2AFFE8FF8BFEE68381875EF397" bold="true" box="[259,280,1998,2024]" pageId="8" pageNumber="55">C</emphasis>
. Idiosoma lateral view. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="DF40A3B0FFE9FF8AFF588162856FF1F0" ID-DOI="http://doi.org/10.5281/zenodo.14771813" ID-Zenodo-Dep="14771813" httpUri="https://zenodo.org/record/14771813/files/figure.png" pageId="9" pageNumber="56" startId="9.[189,232,1325,1351]" targetBox="[246,1342,265,1285]" targetPageId="9">
<paragraph id="8B80F338FFE9FF8AFF588162856FF1F0" blockId="9.[187,1398,1325,1423]" pageId="9" pageNumber="56">
<emphasis id="B94B2F2AFFE9FF8AFF5881628753F137" bold="true" box="[189,277,1325,1352]" pageId="9" pageNumber="56">Fig. 7.</emphasis>
<taxonomicName id="4C3F88BBFFE9FF8AFEC3816185CDF137" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,907,1326,1352]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE9FF8AFEC3816187E6F137" box="[294,416,1326,1352]" italics="true" pageId="9" pageNumber="56">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE9FF8AFE5E81618473F137" box="[443,565,1326,1352]" italics="true" pageId="9" pageNumber="56">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE9FF8AFDAA81618485F137" box="[591,707,1326,1352]" italics="true" pageId="9" pageNumber="56">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE9FF8AFC7E81618244F137" box="[923,1026,1326,1352]" pageId="9" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♀ (slide no. HNHY-2023041105, GUGC), light microscope photographs. 
<emphasis id="B94B2F2AFFE9FF8AFCBE811E8536F114" bold="true" box="[859,880,1361,1387]" pageId="9" pageNumber="56">A</emphasis>
. Idiosoma, dorsal view. 
<emphasis id="B94B2F2AFFE9FF8AFB6D811E82DAF114" bold="true" box="[1160,1180,1361,1387]" pageId="9" pageNumber="56">B</emphasis>
. Idiosoma, ventral view. 
<emphasis id="B94B2F2AFFE9FF8AFEE6813B875EF1F1" bold="true" box="[259,280,1396,1422]" pageId="9" pageNumber="56">C</emphasis>
. Idiosoma lateral view. Scale bars = 100 μm.
</paragraph>
</caption>
<caption id="DF40A3B0FFE9FF8AFF5883E48356F396" ID-DOI="http://doi.org/10.5281/zenodo.14771815" ID-Zenodo-Dep="14771815" httpUri="https://zenodo.org/record/14771815/files/figure.png" pageId="9" pageNumber="56" startId="9.[189,232,1963,1989]" targetBox="[189,1398,1470,1924]" targetPageId="9">
<paragraph id="8B80F338FFE9FF8AFF5883E48356F396" blockId="9.[189,1398,1963,2031]" pageId="9" pageNumber="56">
<emphasis id="B94B2F2AFFE9FF8AFF5883E48753F3BA" bold="true" box="[189,277,1963,1989]" pageId="9" pageNumber="56">Fig. 8.</emphasis>
<taxonomicName id="4C3F88BBFFE9FF8AFEC383E485CDF3BA" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[294,907,1963,1989]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFE9FF8AFEC383E487E6F3BA" box="[294,416,1963,1989]" italics="true" pageId="9" pageNumber="56">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFE9FF8AFE5E83E48473F3BA" box="[443,565,1963,1989]" italics="true" pageId="9" pageNumber="56">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFE9FF8AFDAA83E48485F3BA" box="[591,707,1963,1989]" italics="true" pageId="9" pageNumber="56">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFE9FF8AFC7E83E38244F3B9" box="[923,1026,1964,1990]" pageId="9" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, ♂ (slide no. HNHY-2023041103, GUGC), SEM photographs. 
<emphasis id="B94B2F2AFFE9FF8AFD38838184B4F397" bold="true" box="[733,754,1998,2024]" pageId="9" pageNumber="56">A</emphasis>
. ACG and PCG. 
<emphasis id="B94B2F2AFFE9FF8AFC5B83818594F397" bold="true" box="[958,978,1998,2024]" pageId="9" pageNumber="56">B</emphasis>
. 
<emphasis id="B94B2F2AFFE9FF8AFC05838085BDF390" box="[992,1019,1999,2031]" italics="true" pageId="9" pageNumber="56">
V 
<subScript id="17BBF17DFFE9FF8AFC1783AE85BDF390" attach="left" box="[1010,1019,2017,2031]" fontSize="6" pageId="9" pageNumber="56">1</subScript>
</emphasis>
, 
<emphasis id="B94B2F2AFFE9FF8AFBEF83808263F390" box="[1034,1061,1999,2031]" italics="true" pageId="9" pageNumber="56">
V 
<subScript id="17BBF17DFFE9FF8AFBF983AE8263F390" attach="left" box="[1052,1061,2017,2031]" fontSize="6" pageId="9" pageNumber="56">2</subScript>
</emphasis>
and excretory pore.
</paragraph>
</caption>
<paragraph id="8B80F338FFEAFF89FF588546878CF588" blockId="10.[188,1400,265,503]" pageId="10" pageNumber="57">leg segments: I-L-1 dL 43 (43–48), I-L-2 dL 80 (80–92), I-L-3 dL 95 (95–104), I-L-4 dL 148 (148–168), I-L-5 dL 131 (131–143), HB 44 (44–49), I-L-6 dL 78 (77–85), HB 12 (11–13), S-1 L 63 (63–71), W 6 (6–7), S-2 L 59 (59–68), W 8 (8–10); dL: II-L-1 48 (48–56), II-L-2 70 (70–88), II-L-3 78 (78–87), II-L-4 107 (107–116), II-L-5 115 (115–126), II-L-6 94 (94–104); dL: III-L-1 48 (48–57), III-L-2 71 (71–91), III-L-3 79 (79–87), III-L-4 109 (109–125), III-L-5 125 (125–140), III-L-6 109 (109–122); dL: IV-L-1 87 (87–106), IV-L-2 84 (84–98), IV-L-3 121 (121–133), IV-L-4 146 (146–163), IV-L-5 156 (156–177), IV-L-6 151 (151–154).</paragraph>
</subSubSection>
<subSubSection id="C325A0B3FFEAFF89FF588651827AF68C" pageId="10" pageNumber="57" type="discussion">
<paragraph id="8B80F338FFEAFF89FF5886518776F647" blockId="10.[189,304,542,568]" box="[189,304,542,568]" pageId="10" pageNumber="57">
<heading id="D0C84454FFEAFF89FF5886518776F647" bold="true" box="[189,304,542,568]" fontSize="11" level="3" pageId="10" pageNumber="57" reason="3">
<emphasis id="B94B2F2AFFEAFF89FF5886518776F647" bold="true" box="[189,304,542,568]" pageId="10" pageNumber="57">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B80F338FFEAFF89FF588603827AF68C" blockId="10.[187,1399,588,755]" pageId="10" pageNumber="57">
The new species 
<taxonomicName id="4C3F88BBFFEAFF89FE60860385FEF619" authority="Zhang &amp; Guo" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[389,952,588,614]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="cardiacus" status="sp. nov." subGenus="Atractides">
<emphasis id="B94B2F2AFFEAFF89FE60860387B9F619" box="[389,511,588,614]" italics="true" pageId="10" pageNumber="57">Atractides</emphasis>
(
<emphasis id="B94B2F2AFFEAFF89FDF5860384CCF619" box="[528,650,588,614]" italics="true" pageId="10" pageNumber="57">Atractides</emphasis>
) 
<emphasis id="B94B2F2AFFEAFF89FD7E86038549F619" box="[667,783,588,614]" italics="true" pageId="10" pageNumber="57">cardiacus</emphasis>
Zhang &amp; Guo
</taxonomicName>
<taxonomicNameLabel id="A2789251FFEAFF89FC5A8602825FF618" box="[959,1049,589,615]" pageId="10" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
is similar to 
<taxonomicName id="4C3F88BBFFEAFF89FB558603870CF6F6" authority="Wiles, 1991" authorityName="Wiles" authorityYear="1991" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B94B2F2AFFEAFF89FB5586038330F619" box="[1200,1398,588,614]" italics="true" pageId="10" pageNumber="57">Atractides putihi</emphasis>
Wiles, 
<quantity id="4CC75EDDFFEAFF89FEEA8620870CF6F6" box="[271,330,623,649]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="10" pageNumber="57" unit="in" value="1991.0">1991</quantity>
</taxonomicName>
in the following points: (1) the posterior margin of Cx-IV almost straight; (2) 
<emphasis id="B94B2F2AFFEAFF89FB03863F8346F6EF" box="[1254,1280,624,656]" italics="true" pageId="10" pageNumber="57">V 1</emphasis>
separated from 
<emphasis id="B94B2F2AFFEAFF89FF1886DC875EF6CC" box="[253,280,659,691]" italics="true" pageId="10" pageNumber="57">
V 
<subScript id="17BBF17DFFEAFF89FEEA86EA875EF6CC" attach="left" box="[271,280,677,691]" fontSize="6" pageId="10" pageNumber="57">2</subScript>
</emphasis>
; (3) excretory pore smooth. But 
<taxonomicName id="4C3F88BBFFEAFF89FD7386DC8519F6D3" authorityName="Zhang &amp; Guo" authorityYear="2025" box="[662,863,658,685]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="65" phylum="Arthropoda" rank="species" species="cardiacus" subGenus="Atractides">
<emphasis id="B94B2F2AFFEAFF89FD7386DC84EEF6D2" box="[662,680,659,685]" italics="true" pageId="10" pageNumber="57">A</emphasis>
. (
<emphasis id="B94B2F2AFFEAFF89FD2486DC8495F6D2" box="[705,723,659,685]" italics="true" pageId="10" pageNumber="57">A</emphasis>
.) 
<emphasis id="B94B2F2AFFEAFF89FD0E86DD8519F6D3" box="[747,863,658,684]" italics="true" pageId="10" pageNumber="57">cardiacus</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C3F88BBFFEAFF89FC1D86DC8218F6D3" authorityName="Wiles" authorityYear="1991" box="[1016,1118,658,685]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B94B2F2AFFEAFF89FC1D86DC824CF6D2" box="[1016,1034,659,685]" italics="true" pageId="10" pageNumber="57">A</emphasis>
. 
<emphasis id="B94B2F2AFFEAFF89FBFC86DD8218F6D3" box="[1049,1118,658,684]" italics="true" pageId="10" pageNumber="57">putihi</emphasis>
</taxonomicName>
in the following points: (1) the posterior margin of the new species genital field is narrow, but wider in 
<taxonomicName id="4C3F88BBFFEAFF89FBA186F982EEF6AF" authorityName="Wiles" authorityYear="1991" box="[1092,1192,694,720]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B94B2F2AFFEAFF89FBA186F98210F6AF" box="[1092,1110,694,720]" italics="true" pageId="10" pageNumber="57">A</emphasis>
. 
<emphasis id="B94B2F2AFFEAFF89FB8686F982EEF6AF" box="[1123,1192,694,720]" italics="true" pageId="10" pageNumber="57">putihi</emphasis>
</taxonomicName>
; (2) the gonopore of the male new species is short but longer in 
<taxonomicName id="4C3F88BBFFEAFF89FD328696827EF68C" authority="(Pesic &amp; Smit 2009)" baseAuthorityName="Pesic &amp; Smit" baseAuthorityYear="2009" box="[727,1080,729,755]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="57" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="B94B2F2AFFEAFF89FD32869684AFF68C" box="[727,745,729,755]" italics="true" pageId="10" pageNumber="57">A</emphasis>
. 
<emphasis id="B94B2F2AFFEAFF89FD128696857AF68C" box="[759,828,729,755]" italics="true" pageId="10" pageNumber="57">putihi</emphasis>
(
<bibRefCitation id="EFAE8EC9FFEAFF89FCA986968276F68C" author="Pesic V. &amp; Smit H." box="[844,1072,729,755]" pageId="10" pageNumber="57" pagination="1 - 30" refId="ref8217" refString="Pesic V. &amp; Smit H. 2009. New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species. Zootaxa 2240 (1): 1 - 30. https: // doi. org / 10.11646 / zootaxa. 2240.1.1" type="journal article" year="2009">Pešić &amp; Smit 2009</bibRefCitation>
)
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.
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