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<document id="4EDE38FECA591013DD97EF62160DB267" ID-CLB-Dataset="38995" ID-DOI="10.11646/zootaxa.4114.4.9" ID-GBIF-Dataset="76649fac-7b9f-4baa-ab92-46dd8e33a5e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="271618" ID-ZooBank="7B68F221-7CB1-468D-8161-B64813B737B9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464160412348" checkinUser="plazi" docAuthor="Królik, Roman" docDate="2016" docId="513229017233156AFF2EFA59FD6CFF73" docLanguage="en" docName="zootaxa.4114.4.9.pdf.imf" docOrigin="Zootaxa 4114 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cis castaneus Herbst 1793" docType="treatment" docVersion="9" lastPageNumber="500" masterDocId="AD0B517972351562FFB9FFC3FF9FFF8B" masterDocTitle="Two new species and nomenclatural changes in the Cis castaneus species group (Coleoptera, Ciidae)" masterLastPageNumber="500" masterPageNumber="492" pageNumber="498" updateTime="1698662620247" updateUser="ExternalLinkService">
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<mods:title id="BC2DA91B88204820D2ED855D627A482B">Two new species and nomenclatural changes in the Cis castaneus species group (Coleoptera, Ciidae)</mods:title>
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<mods:namePart id="831904B39F6CC1F0667FAAEA8AE34B1A">Królik, Roman</mods:namePart>
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<mods:date id="AEC3700AC434F6B7616B460A9EF0AE72">2016</mods:date>
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<treatment id="513229017233156AFF2EFA59FD6CFF73" ID-DOI="http://doi.org/10.5281/zenodo.6059296" ID-GBIF-Taxon="120701497" ID-Zenodo-Dep="6059296" LSID="urn:lsid:plazi:treatment:513229017233156AFF2EFA59FD6CFF73" httpUri="http://treatment.plazi.org/id/513229017233156AFF2EFA59FD6CFF73" lastPageId="8" lastPageNumber="500" pageId="6" pageNumber="498">
<subSubSection id="9181CB9C72331564FF2EFA59FDF7F87A" pageId="6" pageNumber="498" type="nomenclature">
<paragraph id="D924981772331564FF2EFA59FD41FA3F" blockId="6.[151,734,1433,1460]" box="[151,734,1433,1460]" pageId="6" pageNumber="498">
<heading id="826C2F7B72331564FF2EFA59FD41FA3F" bold="true" box="[151,734,1433,1460]" fontSize="11" level="1" pageId="6" pageNumber="498" reason="1">
<emphasis id="EBEF440572331564FF2EFA59FD41FA3F" bold="true" box="[151,734,1433,1460]" pageId="6" pageNumber="498">
The
<taxonomicName id="1E9BE39472331564FF76FA5AFDB1FA3F" authority="Herbst, 1793" authorityName="Herbst" authorityYear="1793" box="[207,558,1433,1460]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="498" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EBEF440572331564FF76FA5AFEECFA38" bold="true" box="[207,371,1433,1459]" italics="true" pageId="6" pageNumber="498">Cis castaneus</emphasis>
(Herbst, 1793)
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="D924981772331564FF2EFA23FDF7F87A" blockId="6.[151,1437,1504,2033]" pageId="6" pageNumber="498">
The species belonging to the
<taxonomicName id="1E9BE39472331564FE53FA22FD19FA73" box="[490,646,1505,1528]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="498" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EBEF440572331564FE53FA22FD19FA73" box="[490,646,1505,1528]" italics="true" pageId="6" pageNumber="498">Cis castaneus</emphasis>
</taxonomicName>
group, previously known as
<taxonomicName id="1E9BE39472331564FC6EFA22FBCDFA73" box="[983,1106,1504,1528]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="498" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="EBEF440572331564FC6EFA22FBCDFA73" box="[983,1106,1504,1528]" italics="true" pageId="6" pageNumber="498">Cis nitidus</emphasis>
</taxonomicName>
are morphologically distinct from other species of the genus
<taxonomicName id="1E9BE39472331564FE42F9C6FD54F996" authority="Latreille, 1796" authorityName="Latreille" authorityYear="1796" box="[507,715,1540,1565]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="498" phylum="Arthropoda" rank="genus">
<emphasis id="EBEF440572331564FE42F9C6FDBFF997" box="[507,544,1541,1564]" italics="true" pageId="6" pageNumber="498">Cis</emphasis>
Latreille, 1796
</taxonomicName>
and seem to represent a monophyletic group (
<bibRefCitation id="BD0AE5E672331564FB77F9C7FAFCF996" author="Jelinek" box="[1230,1379,1540,1565]" pageId="6" pageNumber="498" refString="Jelinek, J. (2007) Nomenclatural changes in the family Ciidae (Coleoptera). Acta Entomologica Musei Nationalis Pragae, 47, 135 - 141." type="journal article" year="2007">Jelínek 2007</bibRefCitation>
). For this reason it was given a rank varying between the discrete genus
<emphasis id="EBEF440572331564FC39F9EBFC71F9CB" box="[896,1006,1576,1600]" italics="true" pageId="6" pageNumber="498">Eridaulus</emphasis>
Thomson, 1863 (
<bibRefCitation id="BD0AE5E672331564FB0AF9EAFAF9F9CA" author="Lawrence" box="[1203,1382,1576,1601]" pageId="6" pageNumber="498" refString="Lawrence, J. F. (1965) Comments on some recent changes in the classification of the Ciidae (Coleoptera). Bulletin of the Museum of Comparative Zoology, 133, 273 - 293." type="journal article" year="1965">Lawrence 1965</bibRefCitation>
), the subgenus
<emphasis id="EBEF440572331564FEBFF98EFEEBF9EE" box="[262,372,1613,1637]" italics="true" pageId="6" pageNumber="498">Eridaulus</emphasis>
(
<bibRefCitation id="BD0AE5E672331564FE3BF98FFD8EF9EE" author="Reitter" box="[386,529,1612,1637]" pageId="6" pageNumber="498" refString="Reitter, E. (1901) Byrrhidae (Anobiidae) und Cioidae. Bestimmungs-Tabelle der europaischen Coleopteren. XLVII. Heft 47. Edm. Reitter, Brunn, 64 pp." type="book" year="1901">Reitter 1901</bibRefCitation>
,
<bibRefCitation id="BD0AE5E672331564FDA4F98FFDC7F9EE" author="Reitter" box="[541,600,1612,1637]" pageId="6" pageNumber="498" refString="Reitter, E. (1915) Cis (Eridaulus) Mell. v. Matchanus nov. Wiener Entomologische Zeitung, 34, 66. http: // dx. doi. org / 10.5962 / bhl. part. 10591" type="journal article" year="1915">1915</bibRefCitation>
;
<bibRefCitation id="BD0AE5E672331564FDDDF98FFD73F9EE" author="Strand" box="[612,748,1612,1637]" pageId="6" pageNumber="498" refString="Strand, A. (1965) Uber die nordischen Arten der Gattung Cis Latr., Untergattung Eridaulus Thoms., mit Beschreibung einer neuen Art, Cis hanseni n. sp. (Col., Cisidae). Norsk Entomologisk Tidsskrift, 13, 61 - 65." type="journal article" year="1965">Strand 1965</bibRefCitation>
) and species group (
<bibRefCitation id="BD0AE5E672331564FC68F98FFBC6F9EE" author="Lohse" box="[977,1113,1612,1637]" pageId="6" pageNumber="498" refString="Lohse, G. A. (1965) Uber einige umstrittene Cis - Arten (2. Beitrag zur Kenntnis europaischer Cisidae). Entomologische Blatter, 60, 174 - 181." type="journal article" year="1965">Lohse 1965</bibRefCitation>
,
<bibRefCitation id="BD0AE5E672331564FBDDF98EFA8CF9EE" author="Lawrence" box="[1124,1299,1613,1637]" pageId="6" pageNumber="498" refString="Lawrence, J. F. (1971) Revision of the North American Ciidae (Coleoptera). Bulletin of the Museum of Comparative Zoology, 142, 419 - 522." type="journal article" year="1971">Lawrence 1971</bibRefCitation>
,
<bibRefCitation id="BD0AE5E672331564FAA6F98FFF4CF902" author="Kawanabe" pageId="6" pageNumber="498" refString="Kawanabe, M. (1997) The Japanese species related to the group of Cis nitidus (Coleoptera, Ciidae). Elytra, 25, 311 - 331." type="journal article" year="1997">Kawanabe, 1997</bibRefCitation>
). However, when this group is treated as a genus-group taxon,
<emphasis id="EBEF440572331564FC0CF9B3FBBCF903" box="[949,1059,1648,1672]" italics="true" pageId="6" pageNumber="498">Eridaulus</emphasis>
is not available as its name, for reasons mentioned by
<bibRefCitation id="BD0AE5E672331564FE28F957FDAAF926" author="Jelinek" box="[401,565,1684,1709]" pageId="6" pageNumber="498" refString="Jelinek, J. (2007) Nomenclatural changes in the family Ciidae (Coleoptera). Acta Entomologica Musei Nationalis Pragae, 47, 135 - 141." type="journal article" year="2007">Jelínek (2007)</bibRefCitation>
. These species are characterised by the following main features: body oblong, strongly convex with shining body surface; vestiture usually consisting of short, fine hairs; head moderately declined, partly covered by pronotum; fronto-clypeal ridge in male produced on each side forming two flat, rounded, triangular plates or forming two long horn-like projections; pronotum strongly convex, variously margined laterally, anterior angles rounded to produced and acute; anterior edge in male simple or produced and emarginate; elytra with dual punctation, consisting of larger nude macropunctures, and smaller setiferous micropunctures, the punctures uniform or seriate; prosternum relatively short, concave laterally and carinate medially; intercoxal process relatively narrow but not laminate, subacute at apex; protibia expanded at apex, outer apical angle produced and dentate; first abdominal ventrite of male with median pubescent fovea or patch (
<bibRefCitation id="BD0AE5E672331564FF27F81AFECFF87A" author="Lawrence" box="[158,336,2008,2033]" pageId="6" pageNumber="498" refString="Lawrence, J. F. (1965) Comments on some recent changes in the classification of the Ciidae (Coleoptera). Bulletin of the Museum of Comparative Zoology, 133, 273 - 293." type="journal article" year="1965">Lawrence 1965</bibRefCitation>
,
<bibRefCitation id="BD0AE5E672331564FEE5F81AFE08F87A" author="Lawrence" box="[348,407,2009,2033]" pageId="6" pageNumber="498" refString="Lawrence, J. F. (1971) Revision of the North American Ciidae (Coleoptera). Bulletin of the Museum of Comparative Zoology, 142, 419 - 522." type="journal article" year="1971">1971</bibRefCitation>
;
<bibRefCitation id="BD0AE5E672331564FE1BF81BFDC4F87A" author="Kawanabe" box="[418,603,2008,2033]" pageId="6" pageNumber="498" refString="Kawanabe, M. (1997) The Japanese species related to the group of Cis nitidus (Coleoptera, Ciidae). Elytra, 25, 311 - 331." type="journal article" year="1997">Kawanabe 1997</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="8DE4C89F72321565FF2EF8B7FE58F843" httpUri="https://zenodo.org/record/271622/files/figure.png" pageId="7" pageNumber="499" targetBox="[151,1434,221,1887]" targetPageId="7">
<paragraph id="D924981772321565FF2EF8B7FE58F843" blockId="7.[151,1436,1908,1992]" pageId="7" pageNumber="499">
<emphasis id="EBEF440572321565FF2EF8B7FECBF801" bold="true" box="[151,340,1908,1931]" pageId="7" pageNumber="499">FIGURES 2122.</emphasis>
Collecting sites. 21, Azerbaijan, Talysh Mts., Lerik rayonu, 2 km N of Peştətük (locus typicus of
<taxonomicName id="1E9BE39472321565FA8FF8B5FA03F800" box="[1334,1436,1909,1931]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="499" phylum="Arthropoda" rank="species" species="lasoni" status="sp. nov.">
<emphasis id="EBEF440572321565FA8FF8B5FA03F800" box="[1334,1436,1909,1931]" italics="true" pageId="7" pageNumber="499">Cis lasoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DCF97E72321565FF2EF857FF75F822" box="[151,234,1940,1961]" pageId="7" pageNumber="499" rank="species">
<emphasis id="EBEF440572321565FF2EF857FF75F822" bold="true" box="[151,234,1940,1961]" pageId="7" pageNumber="499">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="1E9BE39472321565FE99F856FE3BF821" box="[288,420,1940,1962]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="499" phylum="Arthropoda" rank="species" species="lugowoji" status="sp. nov.">
<emphasis id="EBEF440572321565FE99F856FE3BF821" box="[288,420,1940,1962]" italics="true" pageId="7" pageNumber="499">Cis lugowoji</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DCF97E72321565FE15F857FE60F822" box="[428,511,1940,1961]" pageId="7" pageNumber="499" rank="species">
<emphasis id="EBEF440572321565FE15F857FE60F822" bold="true" box="[428,511,1940,1961]" pageId="7" pageNumber="499">sp. nov.</emphasis>
</taxonomicNameLabel>
); 22, Iran, Alborz Mts., Golestān Prov., 11 km SE of Minudasht (the locality where
<taxonomicName id="1E9BE39472321565FAC3F856FF4AF843" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="499" phylum="Arthropoda" rank="species" species="lasoni" status="sp. nov.">
<emphasis id="EBEF440572321565FAC3F856FF4AF843" italics="true" pageId="7" pageNumber="499">Cis lasoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0DCF97E72321565FF62F870FEB4F843" box="[219,299,1971,1992]" pageId="7" pageNumber="499" rank="species">
<emphasis id="EBEF440572321565FF62F870FEB4F843" bold="true" box="[219,299,1971,1992]" pageId="7" pageNumber="499">sp. nov.</emphasis>
</taxonomicNameLabel>
was captured).
</paragraph>
</caption>
<subSubSection id="9181CB9C723D156AFF7EFF5BFD6CFF73" pageId="8" pageNumber="500" type="description">
<paragraph id="D9249817723D156AFF7EFF5BFD6CFF73" blockId="8.[151,1436,151,248]" pageId="8" pageNumber="500">
<taxonomicName id="1E9BE394723D156AFF7EFF5BFEA6FF3B" box="[199,313,152,176]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="500" phylum="Arthropoda" rank="species" species="lasoni">
<emphasis id="EBEF4405723D156AFF7EFF5BFEA6FF3B" box="[199,313,152,176]" italics="true" pageId="8" pageNumber="500">Cis lasoni</emphasis>
</taxonomicName>
and
<taxonomicName id="1E9BE394723D156AFEC8FF5BFD9FFF3B" box="[369,512,152,176]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="500" phylum="Arthropoda" rank="species" species="lugowoji">
<emphasis id="EBEF4405723D156AFEC8FF5BFD9FFF3B" box="[369,512,152,176]" italics="true" pageId="8" pageNumber="500">Cis lugowoji</emphasis>
</taxonomicName>
share most of the features of the
<taxonomicName id="1E9BE394723D156AFCC5FF5BFB88FF3B" box="[892,1047,152,176]" class="Insecta" family="Ciidae" genus="Cis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="500" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EBEF4405723D156AFCC5FF5BFB88FF3B" box="[892,1047,152,176]" italics="true" pageId="8" pageNumber="500">Cis castaneus</emphasis>
</taxonomicName>
group, although none of them has the modified fronto-clypeal ridge. The males of both species have the fronto-clypeal ridge devoid of any plates or projections, resembling the same structure in females.
</paragraph>
</subSubSection>
</treatment>
</document>