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<document id="57E4CD83128580E9864323E745BCB10E" ID-DOI="10.1111/j.1096-3642.2008.00466.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5445358" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630458879514" checkinUser="felipe" docAuthor="Biju, S. D. &amp; Bossuyt, Franky" docDate="2009" docId="03F5945FFF821E68FF77FCBFFCE8C0D8" docLanguage="en" docName="j.1096-3642.2008.00466.x.pdf" docOrigin="Zoological Journal of the Linnean Society 155 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00466.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Philautus graminirupes Biju &amp; Bossuyt 2005" docType="treatment" docVersion="4" lastPageNumber="415" masterDocId="FFCCEC27FFAB1E41FFE9FFFDFF8CC65A" masterDocTitle="Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species" masterLastPageNumber="444" masterPageNumber="374" pageNumber="415" updateTime="1699278541911" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="7ACAB0F1900B11107514006AC313615D">Systematics and phylogeny of Philautus Gistel, 1848 (Anura, Rhacophoridae) in the Western Ghats of India, with descriptions of 12 new species</mods:title>
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<mods:namePart id="ABAAE7D28EAEE20201CC2D20D2BC0E92">Biju, S. D.</mods:namePart>
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<mods:namePart id="5160D0E6E2B540B158B2BC6B3DCDB6F7">Bossuyt, Franky</mods:namePart>
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<mods:title id="0E402A2F341DF2D8D8983ED3225F2772">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="632D026DF410A8A210AC9CF6AB87ABA5">2009</mods:date>
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<mods:number id="A5D456E36F502890A3901DF1BFAD818B">2009-02-28</mods:number>
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<mods:identifier id="068D1463001436EABCD1DD5189A2799B" type="Zenodo-Dep">5445358</mods:identifier>
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<treatment id="03F5945FFF821E68FF77FCBFFCE8C0D8" ID-DOI="http://doi.org/10.5281/zenodo.5492423" ID-Zenodo-Dep="5492423" LSID="urn:lsid:plazi:treatment:03F5945FFF821E68FF77FCBFFCE8C0D8" httpUri="http://treatment.plazi.org/id/03F5945FFF821E68FF77FCBFFCE8C0D8" lastPageNumber="415" pageId="41" pageNumber="415">
<subSubSection id="C34676C2FF821E68FF77FCBFFD64C50D" box="[158,744,833,855]" pageId="41" pageNumber="415" type="nomenclature">
<paragraph id="8BE32549FF821E68FF77FCBFFD64C50D" blockId="41.[158,744,833,855]" box="[158,744,833,855]" pageId="41" pageNumber="415">
<heading id="D0AB9225FF821E68FF77FCBFFD64C50D" box="[158,744,833,855]" centered="true" fontSize="9" level="2" pageId="41" pageNumber="415" reason="2">
<taxonomicName id="4C5C5ECAFF821E68FF77FCBFFD64C50D" ID-CoL="4G5GD" authority="BIJU &amp; BOSSUYT, 2005" authorityName="Biju &amp; Bossuyt" authorityYear="2005" box="[158,744,833,855]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="379" phylum="Chordata" rank="species" species="graminirupes">
<emphasis id="B928F95BFF821E68FF77FCBFFE5CC50F" box="[158,464,834,855]" italics="true" pageId="41" pageNumber="415">PHILAUTUS GRAMINIRUPES</emphasis>
BIJU &amp; BOSSUYT, 2005
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34676C2FF821E68FECEFC95FCE8C0D8" pageId="41" pageNumber="415" type="description">
<paragraph id="8BE32549FF821E68FECEFC95FDD3C524" blockId="41.[295,607,872,894]" box="[295,607,872,894]" pageId="41" pageNumber="415">
(
<figureCitation id="136739CCFF821E68FEC6FC95FEF7C527" box="[303,379,872,893]" captionStart="Figure 2" captionStartId="6.[164,243,1531,1550]" captionTargetBox="[167,1442,195,1498]" captionTargetId="figure-103@6.[164,1444,193,1501]" captionTargetPageId="6" captionText="Figure 2. Maximum-likelihood phylogram (GTR + G + I; –Ln L = 14 763.97) for the mitochondrial DNA data set of 1488 bp of 34 taxa of Western Ghats Philautus and five outgroup species. Numbers above branches indicate nonparametric bootstrap values under maximum parsimony (MP). Numbers below branches indicate Bayesian posterior probabilities. *Values of less than 50." figureDoi="http://doi.org/10.5281/zenodo.5445362" httpUri="https://zenodo.org/record/5445362/files/figure.png" pageId="41" pageNumber="415">FIGS 2</figureCitation>
,
<figureCitation id="136739CCFF821E68FE63FC95FE30C527" box="[394,444,872,893]" captionStart="Figure 31" captionStartId="38.[164,243,1842,1861]" captionTargetBox="[185,1422,197,1810]" captionTargetId="figure-0@38.[182,1425,194,1813]" captionTargetPageId="38" captionText="Figure 31. Distribution map. A, Philautus dubois; B, map showing Malabar (shaded in grey), from which the type of Philautus flaviventris was collected; C, Philautus glandulosus; D, Philautus graminirupes." figureDoi="http://doi.org/10.5281/zenodo.5445420" httpUri="https://zenodo.org/record/5445420/files/figure.png" pageId="41" pageNumber="415">31D</figureCitation>
,
<figureCitation id="136739CCFF821E68FE22FC95FE66C524" box="[459,490,872,894]" captionStart="Figure 36" captionStartId="41.[144,223,1853,1872]" captionTargetBox="[148,757,1357,1823]" captionTargetId="figure-583@41.[148,757,1357,1823]" captionTargetPageId="41" captionText="Figure 36. Holotype of Philautus graminirupes (BNHS 4264) from Ponmudi." figureDoi="http://doi.org/10.5281/zenodo.5445430" httpUri="https://zenodo.org/record/5445430/files/figure.png" pageId="41" pageNumber="415">36</figureCitation>
;
<tableCitation id="C6DE10F2FF821E68FE11FC95FDDBC524" box="[504,599,872,894]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="41" pageNumber="415">TABLE 2</tableCitation>
)
</paragraph>
<paragraph id="8BE32549FF821E68FF79FC72FF72C264" blockId="41.[144,759,911,1086]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FF79FC72FEB8C5FE" box="[144,308,911,932]" italics="true" pageId="41" pageNumber="415">Type material:</emphasis>
<materialsCitation id="3B342F14FF821E68FED7FC72FF72C264" collectionCode="SVL" location="Thiruvananthapuram" municipality="Ponmudi" pageId="41" pageNumber="415" specimenCode="BNHS 4264, BNHS 4265-4266, BNHS 4587, TBGRI 2002.0055, BNHS 4267, FMNH 218118" specimenCount="2" specimenCount-adult="2" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="54E79BEBFF821E68FED7FC72FE2BC5FF" box="[318,423,911,933]" pageId="41" pageNumber="415" type="holotype">Holotype</typeStatus>
,
<specimenCode id="DBFA8D32FF821E68FE5DFC72FDCCC5FF" box="[436,576,911,933]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4264</specimenCode>
, an adult male,
<collectionCode id="ED4DBD8CFF821E68FF79FC50FF4EC598" box="[144,194,941,962]" pageId="41" pageNumber="415">SVL</collectionCode>
<quantity id="4CA488ACFF821E68FF22FC50FEBCC599" box="[203,304,941,963]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="41" pageNumber="415" unit="mm" value="22.6">22.6 mm</quantity>
, from
<collectingMunicipality id="6B87BF33FF821E68FE97FC50FE6BC599" box="[382,487,941,963]" pageId="41" pageNumber="415">Ponmudi</collectingMunicipality>
,
<location id="8E837392FF821E68FE1FFC50FD7EC599" LSID="urn:lsid:plazi:treatment:03F5945FFF821E68FF77FCBFFCE8C0D8:8E837392FF821E68FE1FFC50FD7EC599" box="[502,754,941,963]" municipality="Ponmudi" name="Thiruvananthapuram" pageId="41" pageNumber="415" stateProvince="Kerala">Thiruvananthapuram</location>
,
<collectingRegion id="4998EBABFF821E68FF79FC31FF6CC5B8" box="[144,224,972,994]" country="India" name="Kerala" pageId="41" pageNumber="415">Kerala</collectingRegion>
;
<typeStatus id="54E79BEBFF821E68FF02FC31FED3C5BB" box="[235,351,972,993]" pageId="41" pageNumber="415" type="paratype">paratypes</typeStatus>
,
<specimenCode id="DBFA8D32FF821E68FE83FC31FDBAC5B8" box="[362,566,972,994]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4265–4266</specimenCode>
and
<specimenCode id="DBFA8D32FF821E68FD86FC31FD7BC5B8" box="[623,759,972,994]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4587</specimenCode>
(ex
<specimenCode id="DBFA8D32FF821E68FF2BFC16FE10C25A" box="[194,412,1002,1024]" collectionCode="TBGRI" pageId="41" pageNumber="415">TBGRI 2002.0055</specimenCode>
),
<specimenCount id="9D5AEEC0FF821E68FE51FC17FDCDC25A" box="[440,577,1002,1024]" pageId="41" pageNumber="415" type="adult">three adult</specimenCount>
males;
<specimenCode id="DBFA8D32FF821E68FD44FC16FF4BC245" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4267</specimenCode>
and
<specimenCode id="DBFA8D32FF821E68FEE7FBF4FE4CC244" box="[270,448,1033,1054]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="41" pageNumber="415" type="Museum">FMNH 218118</specimenCode>
,
<specimenCount id="9D5AEEC0FF821E68FE3BFBF4FDCAC245" box="[466,582,1033,1055]" pageId="41" pageNumber="415" type="adult">two adult</specimenCount>
females, from Ponmudi.
</materialsCitation>
</paragraph>
<paragraph id="8BE32549FF821E68FF79FB95FE61C224" blockId="41.[144,493,1128,1150]" box="[144,493,1128,1150]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FF79FB95FE13C227" box="[144,415,1128,1149]" italics="true" pageId="41" pageNumber="415">Other material studied:</emphasis>
None.
</paragraph>
<paragraph id="8BE32549FF821E68FF79FB54FA02C72F" blockId="41.[144,759,1193,1307]" lastBlockId="41.[806,1422,198,465]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FF79FB54FDB9C2E4" box="[144,565,1193,1214]" italics="true" pageId="41" pageNumber="415">
Diagnosis:
<taxonomicName id="4C5C5ECAFF821E68FEF0FB54FDB9C2E4" authorityName="Biju &amp; Bossuyt" authorityYear="2005" box="[281,565,1193,1214]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="graminirupes">Philautus graminirupes</taxonomicName>
</emphasis>
can be distinguished from known congeners by the following combination of characters: (1) small male adult snout–vent length (SVL 22.0 ±
<quantity id="4CA488ACFF821E68FDE0FAF8FDECC340" box="[521,608,1285,1307]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="41" pageNumber="415" unit="mm" value="0.6">0.6 mm</quantity>
,
<emphasis id="B928F95BFF821E68FD9AFAF8FD0BC340" box="[627,647,1285,1306]" italics="true" pageId="41" pageNumber="415">N</emphasis>
= 4; SVL
<quantity id="4CA488ACFF821E68FCCFFF3BFC45C681" box="[806,969,198,219]" metricMagnitude="-2" metricUnit="m" metricValue="2.835" metricValueMax="2.94" metricValueMin="2.73" pageId="41" pageNumber="415" unit="mm" value="28.35" valueMax="29.4" valueMin="27.3">27.3–29.4 mm</quantity>
, female); (2) pointed snout in dorsal view; (3) canthus rostralis sharp; (4) posterior surface of thighs chocolate brown, vermiculated with bluish green; (5) dorsum shagreened with a few granular projections; (6) between the eyes a horny ridge from the snout to the middle of the body; (7) long hindlimbs.
</paragraph>
<paragraph id="8BE32549FF821E68FCD6FE83FB7CC78A" blockId="41.[806,1422,198,465]" pageId="41" pageNumber="415">
For comparison with the closely related species,
<taxonomicName id="4C5C5ECAFF821E68FCCFFE60FC6DC7EB" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" box="[806,993,412,434]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="chlorosomma" status="sp. nov.">
<emphasis id="B928F95BFF821E68FCCFFE60FC6DC7EB" box="[806,993,412,434]" italics="true" pageId="41" pageNumber="415">P. chlorosomma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FF821E68FC19FE61FBC1C7EB" box="[1008,1101,412,433]" pageId="41" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C5C5ECAFF821E68FB7FFE60FADFC7E8" authorityName="Biju &amp; Bossuyt" authorityYear="2005" box="[1174,1363,413,434]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="graminirupes">
<emphasis id="B928F95BFF821E68FB7FFE60FADFC7E8" box="[1174,1363,413,434]" italics="true" pageId="41" pageNumber="415">P. graminirupes</emphasis>
</taxonomicName>
, see ‘comparison’ of
<taxonomicName id="4C5C5ECAFF821E68FC30FE46FB02C78A" authority="Biju &amp; Bossuyt, 2009" authorityName="Biju &amp; Bossuyt" authorityYear="2009" box="[985,1166,443,464]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="chlorosomma" status="sp. nov.">
<emphasis id="B928F95BFF821E68FC30FE46FB02C78A" box="[985,1166,443,464]" italics="true" pageId="41" pageNumber="415">P. chlorosomma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21B4420FF821E68FB71FE46FB7CC78A" box="[1176,1264,443,464]" pageId="41" pageNumber="415" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BE32549FF821E68FCCFFE09FAA3C41C" blockId="41.[806,1421,499,582]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FCCFFE09FBFFC452" box="[806,1139,499,521]" italics="true" pageId="41" pageNumber="415">
Description of the
<typeStatus id="54E79BEBFF821E68FBE7FE0EFBE1C452" box="[1038,1133,499,520]" pageId="41" pageNumber="415" type="holotype">holotype</typeStatus>
:
</emphasis>
A detailed description and illustrations were published in
<bibRefCitation id="EFCD58B8FF821E68FB3FFDEFFCF3C41C" author="Biju SD &amp; Bossuyt F" pageId="41" pageNumber="415" pagination="29 - 37" refId="ref42861" refString="Biju SD, Bossuyt F. 2005 b. Two new Philautus (Anura: Ranidae, Rhacophorinae) from Ponmudi Hills in the Western Ghats of India. Copeia 2005: 29 - 37." type="journal article" year="2005">Biju &amp; Bossuyt (2005b)</bibRefCitation>
. Measurements are given in
<tableCitation id="C6DE10F2FF821E68FB3CFDCDFAA7C41C" box="[1237,1323,560,582]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="41" pageNumber="415">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE32549FF821E68FCCFFD94FC77C5EB" blockId="41.[806,1421,617,945]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FCCFFD94FC43C424" box="[806,975,617,639]" italics="true" pageId="41" pageNumber="415">Colour in life:</emphasis>
Dorsum greyish brown with various numbers of irregular black patches, a brownish black band between the eyes, loreal and tympanic regions dark brown, inguinal region vermiculated with brown–yellow, tinted with bluish green, both lips with between three and five narrow light bands, iris silvery brown with dark-brown horizontal bands (
<figureCitation id="136739CCFF821E68FCC7FCC2FCF3C50F" box="[814,895,831,853]" captionStart="Figure 36" captionStartId="41.[144,223,1853,1872]" captionTargetBox="[148,757,1357,1823]" captionTargetId="figure-583@41.[148,757,1357,1823]" captionTargetPageId="41" captionText="Figure 36. Holotype of Philautus graminirupes (BNHS 4264) from Ponmudi." figureDoi="http://doi.org/10.5281/zenodo.5445430" httpUri="https://zenodo.org/record/5445430/files/figure.png" pageId="41" pageNumber="415">Fig. 36</figureCitation>
); anterior surface of thighs vermiculated with brown–yellow, tinted with bluish green, posterior surface of thighs light-chocolate brown, vermiculated with bluish green.
</paragraph>
<paragraph id="8BE32549FF821E68FCCFFC29FBA9C2D9" blockId="41.[806,1422,980,1155]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FCCFFC29FC1BC5B3" box="[806,919,980,1001]" italics="true" pageId="41" pageNumber="415">Variation:</emphasis>
<materialsCitation id="3B342F14FF821E68FC4BFC29FBADC2D9" collectionCode="BNHS" pageId="41" pageNumber="415" specimenCode="BNHS 4265, BNHS 4266, BNHS 4587" specimenCount="1">
Measurements of
<specimenCount id="9D5AEEC0FF821E68FB9FFC29FA9BC5B0" box="[1142,1303,980,1002]" pageId="41" pageNumber="415" type="generic">six specimens</specimenCount>
including
<typeStatus id="54E79BEBFF821E68FCCFFC0EFCDBC252" box="[806,855,1011,1032]" pageId="41" pageNumber="415">type</typeStatus>
series are given in
<tableCitation id="C6DE10F2FF821E68FBDFFC0FFB00C252" box="[1078,1164,1010,1032]" captionStart="Table 2" captionStartId="10.[113,177,400,421]" captionText="Table 2. Morphometric measurements (in mm) of the specimens used in this study" pageId="41" pageNumber="415">Table 2</tableCitation>
.
<specimenCode id="DBFA8D32FF821E68FB7EFC0EFA93C252" box="[1175,1311,1010,1032]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4265</specimenCode>
is darker brown,
<specimenCode id="DBFA8D32FF821E68FC96FBECFB86C27D" box="[895,1034,1041,1063]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4266</specimenCode>
has fewer markings, and
<specimenCode id="DBFA8D32FF821E68FAAAFBECFCD1C21F" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="41" pageNumber="415">BNHS 4587</specimenCode>
has a discontinuous cross marking on the back. In captivity, the colour changes from light-greyish brown to dark brown
</materialsCitation>
.
</paragraph>
<paragraph id="8BE32549FF821E68FCCFFB58FB08C395" blockId="41.[806,1421,1189,1487]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FCCFFB58FCD5C280" italics="true" pageId="41" pageNumber="415">
Distribution and natural history:
<taxonomicName id="4C5C5ECAFF821E68FB4EFB58FCD5C280" authorityName="Biju &amp; Bossuyt" authorityYear="2005" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="graminirupes">Philautus graminirupes</taxonomicName>
</emphasis>
is known only from the type locality, Ponmudi (
<figureCitation id="136739CCFF821E68FCC7FB1EFC1FC2A2" box="[814,915,1251,1273]" captionStart="Figure 31" captionStartId="38.[164,243,1842,1861]" captionTargetBox="[185,1422,197,1810]" captionTargetId="figure-0@38.[182,1425,194,1813]" captionTargetPageId="38" captionText="Figure 31. Distribution map. A, Philautus dubois; B, map showing Malabar (shaded in grey), from which the type of Philautus flaviventris was collected; C, Philautus glandulosus; D, Philautus graminirupes." figureDoi="http://doi.org/10.5281/zenodo.5445420" httpUri="https://zenodo.org/record/5445420/files/figure.png" pageId="41" pageNumber="415">Fig. 31D</figureCitation>
,
<tableCitation id="C6DE10F2FF821E68FC41FB1EFC77C2A3" box="[936,1019,1251,1273]" captionStart="Table 1" captionStartId="2.[164,228,200,219]" captionText="Table 1. Collection localities that are discussed in the text. Localities are arranged by state" pageId="41" pageNumber="415">Table 1</tableCitation>
). The
<typeStatus id="54E79BEBFF821E68FBB0FB1EFB36C2A3" box="[1113,1210,1251,1273]" pageId="41" pageNumber="415" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D5AEEC0FF821E68FAE1FB1EFA01C2A3" box="[1288,1421,1251,1273]" pageId="41" pageNumber="415" type="male">three male</specimenCount>
<typeStatus id="54E79BEBFF821E68FCCFFAFFFC1BC34D" box="[806,919,1282,1303]" pageId="41" pageNumber="415" type="paratype">paratypes</typeStatus>
were collected from leaf blades in grassland near evergreen forest patches. The female
<typeStatus id="54E79BEBFF821E68FACEFADDFA01C36F" box="[1319,1421,1312,1333]" pageId="41" pageNumber="415" type="paratype">paratype</typeStatus>
BNHS 4267 was collected from a 2-m high mosscovered rock in grassland. Breeding was observed at the type locality. The eggs underwent direct development, and hatching of froglets took place after 24 days (
<bibRefCitation id="EFCD58B8FF821E68FC85FA44FBF4C395" author="Biju SD &amp; Bossuyt F" box="[876,1144,1465,1487]" pageId="41" pageNumber="415" pagination="29 - 37" refId="ref42861" refString="Biju SD, Bossuyt F. 2005 b. Two new Philautus (Anura: Ranidae, Rhacophorinae) from Ponmudi Hills in the Western Ghats of India. Copeia 2005: 29 - 37." type="journal article" year="2005">Biju &amp; Bossuyt, 2005b</bibRefCitation>
).
</paragraph>
<paragraph id="8BE32549FF821E68FCCFFA0CFCE8C0D8" blockId="41.[806,1421,1521,1666]" pageId="41" pageNumber="415">
<emphasis id="B928F95BFF821E68FCCFFA0CFC1EC05C" box="[806,914,1521,1542]" italics="true" pageId="41" pageNumber="415">Remarks:</emphasis>
This species was confused with
<taxonomicName id="4C5C5ECAFF821E68FAF9FA0FFA01C05D" baseAuthorityName="BOULENGER" baseAuthorityYear="1882" box="[1296,1421,1522,1543]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="41" pageNumber="415" phylum="Chordata" rank="species" species="signatus">
<emphasis id="B928F95BFF821E68FAF9FA0FFA01C05D" box="[1296,1421,1522,1543]" italics="true" pageId="41" pageNumber="415">P. signatus</emphasis>
</taxonomicName>
(FMNH 218123 and FMNH 218118) by
<bibRefCitation id="EFCD58B8FF821E68FAE2F9ECFCFDC01F" author="Inger RF &amp; Shaffer HB &amp; Koshy M &amp; Bakde R" pageId="41" pageNumber="415" pagination="551 - 570" refId="ref43700" refString="Inger RF, Shaffer HB, Koshy M, Bakde R. 1984. A report on a collection of amphibians and reptiles from the Ponmudi, Kerala, South India. Journal of Bombay Natural History Society 81: 551 - 570." type="journal article" year="1984">
Inger
<emphasis id="B928F95BFF821E68FABCF9ECFA01C07F" box="[1365,1421,1552,1574]" italics="true" pageId="41" pageNumber="415">et al.</emphasis>
(1984)
</bibRefCitation>
. The
<typeStatus id="54E79BEBFF821E68FC2EF9D2FBA1C01E" box="[967,1069,1583,1604]" pageId="41" pageNumber="415" type="paratype">paratype</typeStatus>
TBGRI 2002.0055 (
<bibRefCitation id="EFCD58B8FF821E68FADDF9D2FC54C039" author="Biju SD &amp; Bossuyt F" pageId="41" pageNumber="415" pagination="29 - 37" refId="ref42861" refString="Biju SD, Bossuyt F. 2005 b. Two new Philautus (Anura: Ranidae, Rhacophorinae) from Ponmudi Hills in the Western Ghats of India. Copeia 2005: 29 - 37." type="journal article" year="2005">Biju &amp; Bossuyt, 2005b</bibRefCitation>
) was transferred to BNHS as BNHS 4587.
</paragraph>
</subSubSection>
</treatment>
</document>
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