287 lines
36 KiB
XML
287 lines
36 KiB
XML
<document id="1673D0314AE69CDAC486712C4410920E" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. & Kodada, Ján" docDate="2024" docId="038B5001DA5BE71AFDB9FE51FD2D204D" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus prosternalis Hinton 1935" docType="treatment" docVersion="2" lastPageNumber="86" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="84" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B241922CB16D72528DDA9A3224E513DC">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
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<mods:namePart id="98533E79AC201924ACC337A3E32C7B5B">Selnekovič, Dávid</mods:namePart>
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<mods:affiliation id="9AD686A988CED73F75179B557A34A7C7">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="FE63B1A6287E542D1AFB46E0A4F39054" type="email">david.selnekovic@uniba.sk</mods:nameIdentifier>
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<mods:namePart id="496BC6B145550434E6E3DDCDCB6DDD2E">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="8352F7CDC19AE5194FB4A24EA9086240">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
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<mods:nameIdentifier id="E3D2C270895510FBC567F5B4E4ADBC3F" type="email">manfred.jaech@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="ED2FC46BC6D69FB8CEE36A42EAEFD7BB">Kodada, Ján</mods:namePart>
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<mods:affiliation id="3DD7C026C4BB17D9384D7C50E28FEE5D">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:nameIdentifier id="07402E825499172786D565F0AABDD233" type="email">jan.kodada@uniba.sk</mods:nameIdentifier>
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<mods:title id="BF71E47DA03DEFC06338ABBE4DBA0789">European Journal of Taxonomy</mods:title>
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<mods:part id="27384085BA41248FF18A9E393B325C49">
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<mods:date id="C098CD747B4006D54D95177A2BB028E7">2024</mods:date>
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<mods:number id="D9E74DA8CE9B4F0139E372A0E817AF8E">2024-09-11</mods:number>
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<mods:number id="2DA69B6EA927F45530FFDDFA107355BF">957</mods:number>
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<mods:identifier id="5EA47CF1FBFFB072E034CF9977FC2C62" type="DOI">10.5852/ejt.2024.957.2651</mods:identifier>
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<mods:identifier id="C4269F494420A506329293B091C2856B" type="ISSN">2118-9773</mods:identifier>
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<subSubSection id="C338B29CDA5BE71CFDB9FE51FBEE2646" box="[551,1037,491,518]" pageId="83" pageNumber="84" type="nomenclature">
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<paragraph id="8B9DE117DA5BE71CFDB9FE51FBEE2646" blockId="83.[551,1037,491,550]" box="[551,1037,491,518]" pageId="83" pageNumber="84">
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<heading id="D0D5567BDA5BE71CFDB9FE51FBEE2646" box="[551,1037,491,518]" centered="true" fontSize="11" level="2" pageId="83" pageNumber="84" reason="2">
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<taxonomicName id="4C229A94DA5BE71CFDB9FE51FBEE2646" authority="Hinton, 1935" authorityName="Hinton" authorityYear="1935" box="[551,1037,491,518]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
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<emphasis id="B9563D05DA5BE71CFDB9FE51FC892646" bold="true" box="[551,874,491,518]" italics="true" pageId="83" pageNumber="84">Elmomorphus prosternalis</emphasis>
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<bibRefCitation id="EFB39CE6DA5BE71CFCECFE51FBEE2646" author="Hinton H. E." box="[882,1037,491,518]" pageId="83" pageNumber="84" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C338B29CDA5BE71CFD20FDB6FC962666" box="[702,885,524,550]" pageId="83" pageNumber="84" type="description">
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<paragraph id="8B9DE117DA5BE71CFD20FDB6FC962666" blockId="83.[551,1037,491,550]" box="[702,885,524,550]" pageId="83" pageNumber="84">
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<figureCitation id="1319FD92DA5BE71CFD20FDB6FCCE2666" box="[702,813,524,550]" captionStart="Fig" captionStartId="84.[189,232,1811,1837]" captionTargetBox="[196,1393,285,1746]" captionTargetId="figure-62@84.[189,1399,265,1759]" captionTargetPageId="84" captionText="Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="83" pageNumber="84">Figs 39A</figureCitation>
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,
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<figureCitation id="1319FD92DA5BE71CFCA7FDB6FC962666" box="[825,885,524,550]" captionStart="Fig" captionStartId="218.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@218.[189,1399,265,1830]" captionTargetPageId="218" captionText="Fig. 110. Distribution maps.A. Elmomorphus corpulentus sp. nov.B. E. ellipticus sp. nov.C. E. mazzoldii sp. nov. D. E. bryanti Hinton, 1935. E. E. prosternalis Hinton, 1935. F. E. montanus (Grouvelle, 1913)." figureDoi="http://doi.org/10.5281/zenodo.13773714" httpUri="https://zenodo.org/record/13773714/files/figure.png" pageId="83" pageNumber="84">110E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C338B29CDA5BE71CFF23FDF1FCAB26E4" pageId="83" pageNumber="84" type="reference_group">
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<paragraph id="8B9DE117DA5BE71CFF23FDF1FC382625" blockId="83.[189,987,586,613]" box="[189,987,586,613]" pageId="83" pageNumber="84">
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<treatmentCitationGroup id="AB32C639DA5BE71CFF23FDF1FC382625" box="[189,987,586,613]" pageId="83" pageNumber="84">
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<treatmentCitation id="0A83C706DA5BE71CFF23FDF1FD302625" author="Hinton H. E." box="[189,723,586,613]" page="171" pageId="83" pageNumber="84" year="1935">
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<taxonomicName id="4C229A94DA5BE71CFF23FDF1FD302625" authority="Hinton, 1935: 171" authorityName="Hinton" authorityPageNumber="171" authorityYear="1935" box="[189,723,586,613]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
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<emphasis id="B9563D05DA5BE71CFF23FDF1FE152625" box="[189,502,587,613]" italics="true" pageId="83" pageNumber="84">Elmomorphus prosternalis</emphasis>
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<bibRefCitation id="EFB39CE6DA5BE71CFE63FDF1FD302625" author="Hinton H. E." box="[509,723,586,613]" pageId="83" pageNumber="84" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935: 171</bibRefCitation>
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</taxonomicName>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9DE117DA5BE71CFF23FD33FCAB26E4" blockId="83.[189,840,649,676]" box="[189,840,649,676]" pageId="83" pageNumber="84">
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<treatmentCitationGroup id="AB32C639DA5BE71CFF23FD33FCAB26E4" box="[189,840,649,676]" pageId="83" pageNumber="84">
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<taxonomicName id="4C229A94DA5BE71CFF23FD33FE1526E3" authorityName="Hinton" authorityYear="1935" box="[189,502,649,675]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
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<emphasis id="B9563D05DA5BE71CFF23FD33FE1526E3" box="[189,502,649,675]" italics="true" pageId="83" pageNumber="84">Elmomorphus prosternalis</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A83C706DA5BE71CFD8DFD33FCA726E4" author="Shepard W. D. & Sites R. W." box="[531,836,649,676]" page="93" pageId="83" pageNumber="84" year="2016">
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<bibRefCitation id="EFB39CE6DA5BE71CFD8DFD33FCA726E4" author="Shepard W. D. & Sites R. W." box="[531,836,649,676]" pageId="83" pageNumber="84" pagination="89 - 126" refId="ref100782" refString="Shepard W. D. & Sites R. W. 2016. Aquatic beetles of the families Dryopidae and Elmidae (Insecta: Coleoptera: Byrrhoidea) of Thailand: Annotated list and illustrated key to genera. Natural History Bulletin of the Siam Society 61 (2): 89 - 126." type="journal article" year="2016">Shepard & Sites 2016: 93</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C338B29CDA5BE71CFF23FD72FDB927F1" pageId="83" pageNumber="84" type="diagnosis">
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<paragraph id="8B9DE117DA5BE71CFF23FD72FE2A26A2" blockId="83.[189,457,712,738]" box="[189,457,712,738]" pageId="83" pageNumber="84">
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<heading id="D0D5567BDA5BE71CFF23FD72FE2A26A2" bold="true" box="[189,457,712,738]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
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<emphasis id="B9563D05DA5BE71CFF23FD72FE2A26A2" bold="true" box="[189,457,712,738]" pageId="83" pageNumber="84">Differential diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9DE117DA5BE71CFF23FD4EFDB927F1" blockId="83.[188,1400,756,945]" pageId="83" pageNumber="84">
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<taxonomicName id="4C229A94DA5BE71CFF23FD4EFE1B274E" authorityName="Hinton" authorityYear="1935" box="[189,504,756,782]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
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<emphasis id="B9563D05DA5BE71CFF23FD4EFE1B274E" box="[189,504,756,782]" italics="true" pageId="83" pageNumber="84">Elmomorphus prosternalis</emphasis>
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</taxonomicName>
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is characterised by the presence of a plastron on the dorsal cranial surface (except for a small area on the vertex), on the anterolateral portions of the pronotum, and the lateral sides of the elytra. Elytral punctures are scattered over the entire surface, striae absent. The species most closely resembles
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<taxonomicName id="4C229A94DA5BE71CFE08FCECFDCB272F" baseAuthorityName="Grouvelle" baseAuthorityYear="1913" box="[406,552,854,879]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="montanus">
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<emphasis id="B9563D05DA5BE71CFE08FCECFDCB272F" box="[406,552,854,879]" italics="true" pageId="83" pageNumber="84">E. montanus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C229A94DA5BE71CFDFDFCECFCCE272F" authorityName="Selnekovič & Jäch & Kodada" authorityYear="2024" box="[611,813,854,879]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="paramontanus" status="sp. nov.">
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||
<emphasis id="B9563D05DA5BE71CFDFDFCECFCCE272F" box="[611,813,854,879]" italics="true" pageId="83" pageNumber="84">E. paramontanus</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A265807EDA5BE71CFCABFCECFC732730" box="[821,912,854,880]" pageId="83" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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but differs in the prominent and strongly arched median keel of the prosternal process and the presence of a short median keel along the posterior part of the metaventrite (
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<figureCitation id="1319FD92DA5BE71CFE41FC2DFDAE27F1" box="[479,589,919,945]" captionStart="Fig" captionStartId="84.[189,232,1811,1837]" captionTargetBox="[196,1393,285,1746]" captionTargetId="figure-62@84.[189,1399,265,1759]" captionTargetPageId="84" captionText="Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="83" pageNumber="84">Fig. 39A</figureCitation>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C338B29CDA5BE71CFF23FC6CFD6521E2" pageId="83" pageNumber="84" type="materials_examined">
|
||
<paragraph id="8B9DE117DA5BE71CFF23FC6CFE8F27AF" blockId="83.[189,364,981,1008]" box="[189,364,981,1008]" pageId="83" pageNumber="84">
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<heading id="D0D5567BDA5BE71CFF23FC6CFE8F27AF" bold="true" box="[189,364,981,1008]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
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<emphasis id="B9563D05DA5BE71CFF23FC6CFE8F27AF" bold="true" box="[189,364,981,1008]" pageId="83" pageNumber="84">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9DE117DA5BE71CFF74FBB8FEBF205C" blockId="83.[189,1399,1026,1442]" box="[234,348,1026,1052]" pageId="83" pageNumber="84">
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<emphasis id="B9563D05DA5BE71CFF74FBB8FEBF205C" bold="true" box="[234,348,1026,1052]" pageId="83" pageNumber="84">
|
||
<typeStatus id="54995FB5DA5BE71CFF74FBB8FEBF205C" box="[234,348,1026,1052]" pageId="83" pageNumber="84" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
</paragraph>
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||
<paragraph id="8B9DE117DA5BE71CFF23FB98FD6521E2" blockId="83.[189,1399,1026,1442]" pageId="83" pageNumber="84">
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<collectingCountry id="F335A187DA5BE71CFF23FB98FEB6207C" box="[189,341,1058,1084]" name="Thailand" pageId="83" pageNumber="84">THAILAND</collectingCountry>
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||
–
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<emphasis id="B9563D05DA5BE71CFEE1FB98FCC9207C" bold="true" box="[383,810,1058,1084]" pageId="83" pageNumber="84">
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||
<collectingRegion id="49E62FF5DA5BE71CFEE1FB98FCC9207C" box="[383,810,1058,1084]" country="Thailand" name="Nakhon Si Thammarat" pageId="83" pageNumber="84">Nakhon Si Thammarat Province</collectingRegion>
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||
</emphasis>
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•
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||
<specimenCount id="9D242A9EDA5BE71CFCCFFB98FC8B207C" box="[849,872,1058,1084]" count="1" pageId="83" pageNumber="84" type="female">♀</specimenCount>
|
||
; “PENINSULAR
|
||
<collectingCountry id="F335A187DA5BE71CFBD1FB98FB7B207C" box="[1103,1176,1058,1084]" name="Thailand" pageId="83" pageNumber="84">SIAM</collectingCountry>
|
||
[last third of the “M” cut off] NAKON SRI TAMARA [last “A” slightly cut off at the end] [modern spelling:
|
||
<collectingRegion id="49E62FF5DA5BE71CFA84FBF8FE8E203D" country="Thailand" name="Nakhon Si Thammarat" pageId="83" pageNumber="84">Nakhon Si Thammarat</collectingRegion>
|
||
] KHAO RAM [all printed] at light 750 [handwritten, with printed underlining] FT. [printed] March 2 sd [handwritten, with printed underlining] 1922 H. M. PENDLEBURY [printed] | Type [printed on cirular label with red frame] | comp.type [printed]
|
||
<taxonomicName id="4C229A94DA5BE71CFB74FB1EFA9420FE" authorityName="Hinton" authorityYear="1935" box="[1258,1399,1188,1214]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
|
||
<emphasis id="B9563D05DA5BE71CFB74FB1EFA9420FE" box="[1258,1399,1188,1214]" italics="true" pageId="83" pageNumber="84">prosternalis</emphasis>
|
||
</taxonomicName>
|
||
[handwritten] H.E. Hinton [printed] [pink rectangular label] | H. E. Hinton collection. B.M. 1977—566 [printed on white label] |
|
||
<taxonomicName id="4C229A94DA5BE71CFD3AFB5FFB8520BF" authority="HINTON" authorityName="Hinton" authorityYear="1935" box="[676,1126,1253,1279]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="prosternalis">
|
||
<emphasis id="B9563D05DA5BE71CFD3AFB5FFC0520BF" box="[676,998,1253,1279]" italics="true" pageId="83" pageNumber="84">Elmomorphus prosternalis</emphasis>
|
||
HINTON
|
||
</taxonomicName>
|
||
det. Kodada & Jäch [printed on white label]”;
|
||
<collectionCode id="ED3379D2DA5BE71CFD97FABCFD9F2160" box="[521,636,1286,1312]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="83" pageNumber="84" type="Museum">NHMUK</collectionCode>
|
||
. Photographs of the
|
||
<typeStatus id="54995FB5DA5BE71CFC18FABCFC0E2160" box="[902,1005,1286,1312]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
and its labels are available at: https://www.flickr.com/photos/nhm_beetle_id/52809944410/in/album-72177720297734809 (
|
||
<typeStatus id="54995FB5DA5BE71CFA99FA9CFA912100" box="[1287,1394,1318,1344]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
, labels); https://www.flickr.com/photos/nhm_beetle_id/52809739549/in/album-72177720297734809/ (
|
||
<typeStatus id="54995FB5DA5BE71CFF5BFADDFED321C1" box="[197,304,1383,1409]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
, ventral view); https://www.flickr.com/photos/nhm_beetle_id/52809538071/in/album-721777 20297734809/ (
|
||
<typeStatus id="54995FB5DA5BE71CFEE8FA32FE0221E2" box="[374,481,1416,1442]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
, dorsal view).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338B29CDA5BE71CFF23FA7DFA94224C" box="[189,1399,1479,1549]" pageId="83" pageNumber="84" type="discussion">
|
||
<paragraph id="8B9DE117DA5BE71CFF23FA7DFECC21A1" blockId="83.[189,303,1479,1505]" box="[189,303,1479,1505]" pageId="83" pageNumber="84">
|
||
<heading id="D0D5567BDA5BE71CFF23FA7DFECC21A1" bold="true" box="[189,303,1479,1505]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
|
||
<emphasis id="B9563D05DA5BE71CFF23FA7DFECC21A1" bold="true" box="[189,303,1479,1505]" pageId="83" pageNumber="84">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5BE71CFF23FA48FA94224C" blockId="83.[189,1399,1522,1549]" box="[189,1399,1522,1549]" pageId="83" pageNumber="84">
|
||
The
|
||
<typeStatus id="54995FB5DA5BE71CFF6FFA48FEBB224C" box="[241,344,1522,1548]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
is an immature female of light brown colour; femora, tibiae and tarsi of hind legs missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338B29CDA5BE71CFF23F98BFC452299" pageId="83" pageNumber="84" type="materials_examined">
|
||
<paragraph id="8B9DE117DA5BE71CFF23F98BFEBE220B" blockId="83.[189,349,1585,1611]" box="[189,349,1585,1611]" pageId="83" pageNumber="84">
|
||
<heading id="D0D5567BDA5BE71CFF23F98BFEBE220B" bold="true" box="[189,349,1585,1611]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
|
||
<emphasis id="B9563D05DA5BE71CFF23F98BFEBE220B" bold="true" box="[189,349,1585,1611]" pageId="83" pageNumber="84">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5BE71CFF23F9E7FC452299" blockId="83.[189,1399,1629,1753]" pageId="83" pageNumber="84">
|
||
South
|
||
<collectingCountry id="F335A187DA5BE71CFE90F9E7FE9A2237" box="[270,377,1629,1655]" name="Thailand" pageId="83" pageNumber="84">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E62FF5DA5BE71CFE14F9E7FCF42237" box="[394,791,1629,1655]" country="Thailand" name="Nakhon Si Thammarat" pageId="83" pageNumber="84">Nakhon Si Thammarat Province</collectingRegion>
|
||
, ca
|
||
<quantity id="4CDA4CF2DA5BE71CFCD0F9E7FC7D2238" box="[846,926,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="83" pageNumber="84" unit="m" value="230.0">230 m</quantity>
|
||
a.s.l. According to the label data, the
|
||
<typeStatus id="54995FB5DA5BE71CFF23F9C4FEC722D8" box="[189,292,1662,1688]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
was collected at a place called “Khao Ram”, which was spelled “Khao Rom” in the original description (
|
||
<bibRefCitation id="EFB39CE6DA5BE71CFECEF924FE0F22F9" author="Hinton H. E." box="[336,492,1694,1721]" pageId="83" pageNumber="84" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton 1935</bibRefCitation>
|
||
). It probably refers to Khao Ram Rome [mountain], which lies inside Khao Luang National Park, east of the city of
|
||
<collectingRegion id="49E62FF5DA5BE71CFD0DF905FC422299" box="[659,929,1727,1753]" country="Thailand" name="Nakhon Si Thammarat" pageId="83" pageNumber="84">Nakhon Si Thammarat</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338B29CDA5BE71AFF23F947FE952739" lastPageId="85" lastPageNumber="86" pageId="83" pageNumber="84" type="description">
|
||
<paragraph id="8B9DE117DA5BE71CFF23F947FE1D2358" blockId="83.[189,510,1789,1816]" box="[189,510,1789,1816]" pageId="83" pageNumber="84">
|
||
<heading id="D0D5567BDA5BE71CFF23F947FE1D2358" bold="true" box="[189,510,1789,1816]" fontSize="11" level="3" pageId="83" pageNumber="84" reason="3">
|
||
<emphasis id="B9563D05DA5BE71CFF23F947FE1D2358" bold="true" box="[189,510,1789,1816]" pageId="83" pageNumber="84">
|
||
Redescription of
|
||
<typeStatus id="54995FB5DA5BE71CFE0CF944FE1D2358" box="[402,510,1790,1816]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5BE71CFF23F890FBF72303" blockId="83.[189,1044,1833,1860]" box="[189,1044,1833,1860]" pageId="83" pageNumber="84">Measurements (mm): TL: 3.20; PL: 0.70; PW: 1.20; EL: 2.50; EW: 1.40.</paragraph>
|
||
<paragraph id="8B9DE117DA5BE71CFF23F8D2FCD023A6" blockId="83.[189,1401,1896,2023]" pageId="83" pageNumber="84">Body elongate oval, moderately convex dorsally, widest close before midlength of elytra; brown, legs slightly paler. Dorsal plastron present on frontoclypeus, anterolateral portions of vertex, anterolateral portions of pronotum and lateral bands on elytra. Ventral plastron absent on prosternal process, median part of metaventrite, and middle of first two ventrites.</paragraph>
|
||
<caption id="DF5DB19FDA5CE71BFF23F8A9FB0323D7" ID-DOI="http://doi.org/10.5281/zenodo.13773490" ID-Zenodo-Dep="13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="84" pageNumber="85" startId="84.[189,232,1811,1837]" targetBox="[196,1393,285,1746]" targetPageId="84">
|
||
<paragraph id="8B9DE117DA5CE71BFF23F8A9FB0323D7" blockId="84.[189,1398,1811,1943]" pageId="84" pageNumber="85">
|
||
<emphasis id="B9563D05DA5CE71BFF23F8A9FED6236D" bold="true" box="[189,309,1811,1837]" pageId="84" pageNumber="85">Fig. 39. A</emphasis>
|
||
.
|
||
<taxonomicName id="4C229A94DA5CE71BFEDDF8A9FCC3236E" authority="Hinton, 1935" authorityName="Hinton" authorityYear="1935" box="[323,800,1811,1838]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="prosternalis">
|
||
<emphasis id="B9563D05DA5CE71BFEDDF8A9FD9F236D" box="[323,636,1811,1837]" italics="true" pageId="84" pageNumber="85">Elmomorphus prosternalis</emphasis>
|
||
<bibRefCitation id="EFB39CE6DA5CE71BFD1DF8A9FCC3236E" author="Hinton H. E." box="[643,800,1811,1838]" pageId="84" pageNumber="85" pagination="169 - 179" refId="ref99677" refString="Hinton H. E. 1935. Notes on the Dryopoidea (Col.). Stylops 4 (8): 169 - 179. https: // doi. org / 10.1111 / j. 1365 - 3113.1935. tb 00660. x" type="journal article" year="1935">Hinton, 1935</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype (BMNH), prosternal process and median part of metaventrite.
|
||
<emphasis id="B9563D05DA5CE71BFE20F88CFE312310" bold="true" box="[446,466,1846,1872]" pageId="84" pageNumber="85">B</emphasis>
|
||
.
|
||
<taxonomicName id="4C229A94DA5CE71BFE7AF88DFCBD2310" authority="(Grouvelle, 1913)" baseAuthorityName="Grouvelle" baseAuthorityYear="1913" box="[484,862,1846,1872]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9563D05DA5CE71BFE7AF88DFD992310" box="[484,634,1847,1872]" italics="true" pageId="84" pageNumber="85">E. montanus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB39CE6DA5CE71BFD13F88CFCB72310" author="Grouvelle A." box="[653,852,1846,1872]" pageId="84" pageNumber="85" pagination="99 - 117" refId="ref99512" refString="Grouvelle A. 1913. [Zoological results of the Abor expedition, 1911 - 1912] VIII [.] Coleoptera, I: Rhysodidae, Nitidulidae, Colydiidae, Cucujidae, Passandridae, Discolomidae, Cryptophagidae, Mycetophagidae, Dryopidae. Records of the Indian Museum VIII (2): 99 - 117. https: // doi. org / 10.26515 / rzsi / v 8 / i 2 / 1913 / 163160" type="journal article" year="1913">Grouvelle, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, syntype, male (MNHN), prosternal process and median part of metaventrite.
|
||
<emphasis id="B9563D05DA5CE71BFDDDF8E3FDBB2333" bold="true" box="[579,600,1881,1907]" pageId="84" pageNumber="85">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C229A94DA5CE71BFDF9F8E0FD1A2333" baseAuthorityName="Grouvelle" baseAuthorityYear="1913" box="[615,761,1882,1907]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B9563D05DA5CE71BFDF9F8E0FD1A2333" box="[615,761,1882,1907]" italics="true" pageId="84" pageNumber="85">E. montanus</emphasis>
|
||
</taxonomicName>
|
||
, syntype (MNHN), aedeagus, lateral aspect.
|
||
<emphasis id="B9563D05DA5CE71BFA94F8E3FAFC2333" bold="true" box="[1290,1311,1881,1907]" pageId="84" pageNumber="85">D</emphasis>
|
||
. Same, ventral aspect, detail of parameres and penis.
|
||
<emphasis id="B9563D05DA5CE71BFD4AF8C7FD0B23D7" bold="true" box="[724,744,1917,1943]" pageId="84" pageNumber="85">E</emphasis>
|
||
. Same, ventral aspect. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FEB3FBAF25CD" blockId="85.[189,1398,265,398]" pageId="85" pageNumber="86">Cranial surface with round setiferous punctures, each about as wide as an eye facet, separated by approximately a puncture diameter. Plastron present on frontoclypeus and anterolateral portions of vertex, posteromedian portion without plastron about as wide as eye (dorsal aspect). Labrum transverse, exposed portion microreticulate, with decumbent setae along anterior margin.</paragraph>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FE0FFAD82687" blockId="85.[189,1399,437,711]" pageId="85" pageNumber="86">
|
||
Pronotum moderately convex, widest at base, PW/PL: 1.71; round setiferous punctures slightly larger than those on head; plastron confined to anterolateral angles; anterior margin slightly arcuate; anterolateral angles strongly deflexed, acute. Prosternal process with lateral edges rounded; lateral margins posteriorly expanded and distinctly separated from conspicuously strongly arched median keel (
|
||
<figureCitation id="1319FD92DA5DE71AFF58FDF9FED5261D" box="[198,310,579,605]" captionStart="Fig" captionStartId="84.[189,232,1811,1837]" captionTargetBox="[196,1393,285,1746]" captionTargetId="figure-62@84.[189,1399,265,1759]" captionTargetPageId="84" captionText="Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="85" pageNumber="86">Fig. 39A</figureCitation>
|
||
). Metaventrite convex, median portion without plastron; discrimen distinct, with short keel and adjacent groove posterior of metakatepisternal suture (
|
||
<figureCitation id="1319FD92DA5DE71AFCE1FDDCFC1326C0" box="[895,1008,614,640]" captionStart="Fig" captionStartId="84.[189,232,1811,1837]" captionTargetBox="[196,1393,285,1746]" captionTargetId="figure-62@84.[189,1399,265,1759]" captionTargetPageId="84" captionText="Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="85" pageNumber="86">Fig. 39A</figureCitation>
|
||
). Elytra moderately convex, EL/ EW: 1.79; weakly microreticulate; punctures scattered on entire surface; plastron forming two lateral bands, almost reaching elytral base, slightly widening posteriorly and connecting at elytral apices.
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FD55FC2C2749" blockId="85.[189,975,751,777]" box="[189,975,751,777]" pageId="85" pageNumber="86">Ventrites covered with plastron, except for middle of ventrites 1–2.</paragraph>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FC8AFDD4270A" blockId="85.[189,567,816,842]" box="[189,567,816,842]" pageId="85" pageNumber="86">
|
||
<heading id="D0D5567BDA5DE71AFF23FC8AFDD4270A" bold="true" box="[189,567,816,842]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
|
||
<emphasis id="B9563D05DA5DE71AFF23FC8AFDD4270A" bold="true" box="[189,567,816,842]" pageId="85" pageNumber="86">Secondary sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FCE5FE952739" blockId="85.[189,374,863,889]" box="[189,374,863,889]" pageId="85" pageNumber="86">Male unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C338B29CDA5DE71AFF23FC1AFD2D204D" pageId="85" pageNumber="86" type="distribution">
|
||
<paragraph id="8B9DE117DA5DE71AFF23FC1AFEB527FA" blockId="85.[189,342,928,954]" box="[189,342,928,954]" pageId="85" pageNumber="86">
|
||
<heading id="D0D5567BDA5DE71AFF23FC1AFEB527FA" bold="true" box="[189,342,928,954]" fontSize="11" level="3" pageId="85" pageNumber="86" reason="3">
|
||
<emphasis id="B9563D05DA5DE71AFF23FC1AFEB527FA" bold="true" box="[189,342,928,954]" pageId="85" pageNumber="86">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9DE117DA5DE71AFF23FC75FD2D204D" blockId="85.[189,1398,975,1037]" pageId="85" pageNumber="86">
|
||
Southern
|
||
<collectingCountry id="F335A187DA5DE71AFEB3FC75FE7627A9" box="[301,405,975,1001]" name="Thailand" pageId="85" pageNumber="86">Thailand</collectingCountry>
|
||
(
|
||
<figureCitation id="1319FD92DA5DE71AFE3BFC75FDFE27AA" box="[421,541,975,1002]" captionStart="Fig" captionStartId="218.[189,232,1871,1897]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-40@218.[189,1399,265,1830]" captionTargetPageId="218" captionText="Fig. 110. Distribution maps.A. Elmomorphus corpulentus sp. nov.B. E. ellipticus sp. nov.C. E. mazzoldii sp. nov. D. E. bryanti Hinton, 1935. E. E. prosternalis Hinton, 1935. F. E. montanus (Grouvelle, 1913)." figureDoi="http://doi.org/10.5281/zenodo.13773714" httpUri="https://zenodo.org/record/13773714/files/figure.png" pageId="85" pageNumber="86">Fig. 110E</figureCitation>
|
||
). This species (and the genus
|
||
<taxonomicName id="4C229A94DA5DE71AFCEBFC75FBF927A9" authorityName="Sharp" authorityYear="1888" box="[885,1050,975,1001]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9563D05DA5DE71AFCEBFC75FBF927A9" box="[885,1050,975,1001]" italics="true" pageId="85" pageNumber="86">Elmomorphus</emphasis>
|
||
</taxonomicName>
|
||
) was erroneously recorded as new for
|
||
<collectingCountry id="F335A187DA5DE71AFE80FC49FE65204D" box="[286,390,1011,1037]" name="Thailand" pageId="85" pageNumber="86">Thailand</collectingCountry>
|
||
by
|
||
<bibRefCitation id="EFB39CE6DA5DE71AFE2CFC49FD28204D" author="Shepard W. D. & Sites R. W." box="[434,715,1011,1037]" pageId="85" pageNumber="86" pagination="89 - 126" refId="ref100782" refString="Shepard W. D. & Sites R. W. 2016. Aquatic beetles of the families Dryopidae and Elmidae (Insecta: Coleoptera: Byrrhoidea) of Thailand: Annotated list and illustrated key to genera. Natural History Bulletin of the Siam Society 61 (2): 89 - 126." type="journal article" year="2016">Shepard & Sites (2016)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |