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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt &amp; Liew, Thor-Seng" docDate="2017" docId="A71DE5D2E9B438DD2BFC57C6A576C002" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus jousseaumei Morgan 1885" docType="treatment" docVersion="5" lastPageNumber="30" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="27" updateTime="1668164728433" updateUser="ExternalLinkService">
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<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Foon, Junn Kitt</mods:namePart>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>692</mods:number>
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<mods:start>1</mods:start>
<mods:end>81</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.692.14706</mods:url>
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<mods:classification>journal article</mods:classification>
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<treatment ID-GBIF-Taxon="132786249" LSID="urn:lsid:plazi:treatment:A71DE5D2E9B438DD2BFC57C6A576C002" httpUri="http://treatment.plazi.org/id/A71DE5D2E9B438DD2BFC57C6A576C002" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27">
<subSubSection pageId="26" pageNumber="27" type="nomenclature">
<paragraph pageId="26" pageNumber="27">
<taxonomicName authority="Morgan, 1885 a" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei Morgan, 1885a</taxonomicName>
Figures 7
<normalizedToken originalValue="HI">H-I</normalizedToken>
, 19, 31F
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="reference_group">
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="jousseaumei">
<pageBreakToken pageId="27" pageNumber="28" start="start">Alycaeus</pageBreakToken>
jousseaumei
</taxonomicName>
Morgan, 1885a: 70;
<bibRefCitation author="Morgan, de J" journalOrPublisher="Bulletin de la Societe zoologique de France" pageId="52" pageNumber="53" pagination="353 - 429" title="Mollusques terrestes et fiuviatiles de Royaume de Perak et des pais voisins (presque'Ile Malaise)." url="https://doi.org/10.5962/bhl.part.14300" volume="10" year="1885 b">Morgan 1885b</bibRefCitation>
: 402, plate 8, figure 4;
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="52" pageNumber="53" pagination="299 - 316" title="The landshells of Perak." url="http://www.biodiversitylibrary.org/part/145559/summary" volume="55" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
: 312;
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
: 343;
<bibRefCitation author="Tarruella, A" journalOrPublisher="Spira" pageId="53" pageNumber="54" pagination="71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." url="http://www.molluscat.com/assets/spira_4_12_7.pdf" volume="4" year="2011">
Tarruella and
<normalizedToken originalValue="Domènech">Domenech</normalizedToken>
2011
</bibRefCitation>
: 72, figure 2B;
<bibRefCitation author="Foon, JK" journalOrPublisher="ZooKeys" pageId="50" pageNumber="51" pagination="1 - 94" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
: 12, figure 4D.
</paragraph>
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) jousseaumi" order="Architaenioglossa" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="jousseaumi" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) jousseaumi</taxonomicName>
[sic]: Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
<bibRefCitation author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="51" pageNumber="52" pagination="73 - 192" title="Catalog der gegenwaertig lebend bekannten Pneumonopomen." url="https://doi.org/10.5962/bhl.title.14877" volume="1" year="1897">1897</bibRefCitation>
: 151.
</paragraph>
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) jousseaumei" order="Architaenioglossa" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="jousseaumei" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) jousseaumei</taxonomicName>
:
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="51" pageNumber="52" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In: Kobelt W (Ed.) In Verbindung mit der deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://doi.org/10.5962/bhl.title.1227" year="1902">Kobelt 1902</bibRefCitation>
: 357;
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 36;
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="4 - 5" title="A brief collecting trip to Perak, West Malaysia: Part One." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
: 4;
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="1 - 2" title="A brief collecting trip to Perak, West Malaysia: Part Two." volume="12" year="1998 b">Chan 1998b</bibRefCitation>
: 1.
</paragraph>
<paragraph pageId="27" pageNumber="28">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus jousseaumei" order="Architaenioglossa" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="jousseaumei">Chamalycaeus jousseaumei</taxonomicName>
:
<bibRefCitation author="Benthem Jutting, van WSS" journalOrPublisher="Basteria" pageId="49" pageNumber="50" pagination="10 - 20" title="Some notes on land and freshwater mollusca of Malaya." url="http://natuurtijdschriften.nl/record/596402" volume="24" year="1960 a">Benthem Jutting 1960a</bibRefCitation>
: 13;
<bibRefCitation author="Berry, AJ" journalOrPublisher="Malayan Nature Journal" pageId="49" pageNumber="50" pagination="1 - 17" title="An introduction to the non-marine molluscs of Malaya." volume="17" year="1963">Berry 1963</bibRefCitation>
: plate 4;
<bibRefCitation pageId="27" pageNumber="28">Maassen 2001</bibRefCitation>
: 24;
<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="49" pageNumber="50" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
: 2760.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="type locality">
<paragraph pageId="27" pageNumber="28">Type locality.</paragraph>
<paragraph pageId="27" pageNumber="28">
Mont Lano (= PRK 18 Gunung Lanno), Perak (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.5252776">4°31'31&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.14667">101°08'48&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="type material">
<paragraph pageId="28" pageNumber="29">
<pageBreakToken pageId="28" pageNumber="29" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="28" pageNumber="29">Syntypes. Mont Lano (= PRK 18 Gunung Lanno), Perak: MNHN IM-2000-31800/1 (Not seen), MNHN IM-2000-31801/1 (Seen).</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="materials_examined">
<paragraph pageId="28" pageNumber="29">Other examined materials.</paragraph>
<paragraph pageId="28" pageNumber="29">PRK 01 Gunung Tempurung, Perak: BOR/MOL 6847/1, BOR/MOL 8346/3, BOR/MOL 11136/11, BOR/MOL 11211/37, BOR/MOL 11408/10. PRK 15 Gunung Keroh, Perak: ZRC 1975.2.21.247-248/2. PRK 18 Gunung Lanno, Perak: BOR/MOL 6841/1, BOR/MOL 6916/3, BOR/MOL 8337/2, BOR/MOL 8338/2. PRK 23 Gunung Rapat, Perak: BOR/MOL 293/5, BOR/MOL 6255/1, BOR/MOL 6844/1, BOR/MOL 6846/1, BOR/MOL 6848/1, BOR/MOL 7133/6, BOR/MOL 8341/14, BOR/MOL 8342/2, BOR/MOL 8345/22, BOR/MOL 8347/25, BOR/MOL 8348/4, BOR/MOL 10046/8, BOR/MOL 10202/3, BOR/MOL 10228/9, BOR/MOL 10253/10, BOR/MOL 12963/1, BOR/MOL 12998/2, ZRC 1997.18/2. PRK 27 Gunung Datok, Perak: BOR/MOL 10412/9, BOR/MOL 10441/8, BOR/MOL 10474/30, BOR/MOL 10494/8. PRK 34 Gunung Tasek, Perak: BOR/MOL 6839/1, BOR/MOL 8336/6, BOR/MOL 10788/7, BOR/MOL 11054/1. PRK 42 Gunung Bercham, Perak: BOR/MOL 10593/4. PRK 47 Gunung Kanthan South, Perak: BOR/MOL 9101/2, ZRC 1975.2.21.2/2. PRK 53 Hill KF, Perak: BOR/MOL 10658/12, BOR/MOL 10689/13, BOR/MOL 10717/5. Mykarst-189, Perak: BOR/MOL 6842/1, BOR/MOL 8339/8. Mykarst-068, Perak: BOR/MOL 6843/1, BOR/MOL 8340/1, RMNH 153708/3. Mykarst-025, Perak: BOR/MOL 9383/1. Mykarst-027 Gunung Kanthan North, Perak: BOR/MOL 9053/3, BOR/MOL 9374/1. Mykarst-070 Kramat Pulai, Perak: ZRC 1975.2.21.238-245/8. Perak: RMNH 153477/1.</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph pageId="28" pageNumber="29">Description.</paragraph>
<paragraph pageId="28" pageNumber="29">Protoconch. Smooth.</paragraph>
<paragraph pageId="28" pageNumber="29">Shell shape. Flat. Shell height: 7.50-8.76 mm. Shell width: 12.42-15.03 mm.</paragraph>
<paragraph pageId="28" pageNumber="29">Spire. Spire height: 1.77-2.27 mm. Spire width: 2.94-3.45 mm. Number of whorls: up to 3. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open.</paragraph>
<paragraph pageId="28" pageNumber="29">
Whorl constriction. At about 2
<normalizedToken originalValue="¾">3/4</normalizedToken>
whorls posterior of protoconch.
</paragraph>
<paragraph pageId="28" pageNumber="29">Breathing tube. Length: 5.44-5.97 mm.</paragraph>
<paragraph pageId="28" pageNumber="29">Aperture and peristome. Aperture circular, very expanded. Aperture height: 4.88-7.30 mm. Aperture width: 4.92-6.59 mm. Peristome triple. The first (most anterior) peristome is moderately expanded but not thickened. The second (outer) peristome is very expanded but not thickened. The second peristome is impressed on the thickened third (inner), non-expanded peristome. Interspace between first and second peristome wide. Interspace between second and third peristome narrow. Peristome orientation is 44-55° oblique with respect to the coiling axis.</paragraph>
<paragraph pageId="28" pageNumber="29">Spiral lines. Absent.</paragraph>
<paragraph pageId="28" pageNumber="29">Radial ribs running anterior of breathing tube. Radial ribs indistinct, evenly spaced. Approximately 10-18 ribs in 1 mm.</paragraph>
<paragraph pageId="28" pageNumber="29">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced and thick, evenly spaced. Approximately 8-15 ribs in 1 mm.</paragraph>
<paragraph pageId="28" pageNumber="29">Radial ribs running posterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 10-25 ribs per 1 mm.</paragraph>
<paragraph pageId="28" pageNumber="29">Operculum. Flat, conical. Exterior calcareous, smooth but occasionally with calcareous deposits. Interior covered by proteinaceous layer, smooth.</paragraph>
<paragraph pageId="29" pageNumber="30">
<pageBreakToken pageId="29" pageNumber="30" start="start">Shell</pageBreakToken>
colour. First 3 whorls usually red, orange, purple or white. Ultimate whorl white. Whorl constriction generally white, occasionally red.
</paragraph>
<paragraph pageId="29" pageNumber="30">Living animal. Body light grey to light brown. Head pinkish-brown to dark grey. Tentacles brown.</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="habitat">
<paragraph pageId="29" pageNumber="30">Habitat and ecology.</paragraph>
<paragraph pageId="29" pageNumber="30">Lives on limestone rock surfaces and crevices. In wet, shady forests on limestone hills.</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="distribution">
<paragraph pageId="29" pageNumber="30">Distribution range.</paragraph>
<paragraph pageId="29" pageNumber="30">Restricted to the karsts of Kinta Valley, Perak. Its known northern limit is at Gunung Tchehel and southern limit at Gunung Tempurung.</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="differential diagnosis">
<paragraph pageId="29" pageNumber="30">Differential diagnosis.</paragraph>
<paragraph pageId="29" pageNumber="30">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
is distinguished from all other
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
species in Peninsular Malaysia by its very low spire, very expanded, obtuse ultimate whorl and very expanded, double peristome with wide interspace.
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="discussion">
<paragraph pageId="29" pageNumber="30">Discussion.</paragraph>
<paragraph pageId="29" pageNumber="30">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
is morphologically unique and unmistakable among Peninsular Malaysian
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
has been subjected to attempts of supraspecific reclassification.
<bibRefCitation author="Morgan, de J" journalOrPublisher="Le Naturaliste" pageId="52" pageNumber="53" pagination="68 - 70" title="Note sur quelques especes nouvelles de mollusques terrestres recueillis dans la Peninsule Malaise." url="http://www.biodiversitylibrary.org/item/105819page/83" volume="7" year="1885 a">Morgan (1885a)</bibRefCitation>
initially placed
<taxonomicName lsidName="A. jousseaumei" pageId="29" pageNumber="30" rank="species" species="jousseaumei">A. jousseaumei</taxonomicName>
in the genus
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
but Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
(
<bibRefCitation author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="51" pageNumber="52" pagination="73 - 192" title="Catalog der gegenwaertig lebend bekannten Pneumonopomen." url="https://doi.org/10.5962/bhl.title.14877" volume="1" year="1897">1897</bibRefCitation>
) reassigned
<taxonomicName lsidName="A. jousseaumei" pageId="29" pageNumber="30" rank="species" species="jousseaumei">A. jousseaumei</taxonomicName>
in
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Chamalycaeus</taxonomicName>
without any justification. The genus
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Chamalycaeus</taxonomicName>
in Peninsular Malaysia is characterised by its small to minute shell (shell width &lt;6.0 mm), discoid shell shape, strong radial ribs and a breathing tube of variable length (
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="51" pageNumber="52" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In: Kobelt W (Ed.) In Verbindung mit der deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://doi.org/10.5962/bhl.title.1227" year="1902">Kobelt 1902</bibRefCitation>
,
<bibRefCitation pageId="29" pageNumber="30">Maassen 2001</bibRefCitation>
,
<bibRefCitation author="Egorov, RV" journalOrPublisher="Colus-Doverie, Moscow" pageId="50" pageNumber="51" title="Treasure of Russian Shells; Supplement 3, A review of the genera of the recent terrestrial pectinibranch molluscs (synopsis mainly based on published data); Part III, Littoriniformes: Liareidae, Pupinidae, Diplommatinidae, Alycaeidae, Cochlostomidae." year="2013">Egorov 2013</bibRefCitation>
,
<bibRefCitation author="Foon, JK" journalOrPublisher="ZooKeys" pageId="50" pageNumber="51" pagination="1 - 94" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
).
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
is clearly not a
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Chamalycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chamalycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Chamalycaeus</taxonomicName>
species. Thus,
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
is herein reassigned back to
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
is remarkably similar to
<taxonomicName lsidName="A. kapayanensis" pageId="29" pageNumber="30" rank="species" species="kapayanensis">A. kapayanensis</taxonomicName>
in being range-restricted to the karsts of Kinta Valley (
<bibRefCitation author="Morgan, de J" journalOrPublisher="Le Naturaliste" pageId="52" pageNumber="53" pagination="68 - 70" title="Note sur quelques especes nouvelles de mollusques terrestres recueillis dans la Peninsule Malaise." url="http://www.biodiversitylibrary.org/item/105819page/83" volume="7" year="1885 a">Morgan 1885a</bibRefCitation>
,
<bibRefCitation author="Morgan, de J" journalOrPublisher="Bulletin de la Societe zoologique de France" pageId="52" pageNumber="53" pagination="353 - 429" title="Mollusques terrestes et fiuviatiles de Royaume de Perak et des pais voisins (presque'Ile Malaise)." url="https://doi.org/10.5962/bhl.part.14300" volume="10" year="1885 b">1885b</bibRefCitation>
,
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="52" pageNumber="53" pagination="299 - 316" title="The landshells of Perak." url="http://www.biodiversitylibrary.org/part/145559/summary" volume="55" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
,
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
,
<bibRefCitation author="Benthem Jutting, van WSS" journalOrPublisher="Basteria" pageId="49" pageNumber="50" pagination="10 - 20" title="Some notes on land and freshwater mollusca of Malaya." url="http://natuurtijdschriften.nl/record/596402" volume="24" year="1960 a">Benthem Jutting 1960a</bibRefCitation>
,
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="4 - 5" title="A brief collecting trip to Perak, West Malaysia: Part One." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
,
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="1 - 2" title="A brief collecting trip to Perak, West Malaysia: Part Two." volume="12" year="1998 b">1998b</bibRefCitation>
).
</paragraph>
<caption pageId="29" pageNumber="30">
<paragraph pageId="29" pageNumber="30">
Figure 19.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus jousseaumei" order="Architaenioglossa" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="jousseaumei">Alycaeus jousseaumei</taxonomicName>
Morgan, 1885a.
<normalizedToken originalValue="AE">A-E</normalizedToken>
Shell of syntype, MNHN-IM-2000-31801
<normalizedToken originalValue="FJ">F-J</normalizedToken>
Shell of BOR/MOL 11211
<normalizedToken originalValue="KM">K-M</normalizedToken>
Close up of shell of BOR/MOL 11211
<normalizedToken originalValue="NQ">N-Q</normalizedToken>
Operculum of BOR/MOL 11211. Scale bars:
<normalizedToken originalValue="AQ">A-Q</normalizedToken>
1 mm. Photographs
<normalizedToken originalValue="AE">A-E</normalizedToken>
by Manuel Caballer (reproduced herein under CC-BY 4.0 license, Project RECOLNAT, MNHN). Photographs
<normalizedToken originalValue="FV">F-V</normalizedToken>
by Junn Kitt Foon.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>