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<document id="610E681A838A483D0B72FB2C9035734F" ID-DOI="10.11646/zootaxa.3974.1.7" ID-GBIF-Dataset="e34081ab-9685-4aa0-b9aa-60d903b10448" ID-ISSN="1175-5326" ID-Zenodo-Dep="232028" ID-ZooBank="6EA64894-DAC1-4C9B-A3A7-D075AF5B87DF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461216788271" checkinUser="plazi" docAuthor="Celis, Liliana V. &amp; Rangel-Ch, Orlando" docDate="2015" docId="EE301571FFE5602C79C4FC6DFB7EFB57" docLanguage="en" docName="zt03974p114.pdf" docOrigin="Zootaxa 3974 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Righiodrilus lebrijae Celis &amp; Rangel-Ch, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="113" masterDocId="12096D09FFE0602B7953FFCAFF8FFFEB" masterDocTitle="Two new earthworm species (Oligochaeta: Annelida) of the Caribbean region of Colombia" masterLastPageNumber="114" masterPageNumber="106" pageNumber="111" updateTime="1698602636948" updateUser="plazi">
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<mods:title id="8C6F03B2178D9E6EA08032A5C394B798">Two new earthworm species (Oligochaeta: Annelida) of the Caribbean region of Colombia</mods:title>
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<mods:namePart id="7D5E9A8EB7F2A5E932665FFC67611CB8">Celis, Liliana V.</mods:namePart>
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<mods:namePart id="A4CDF7D19EA2E5E629AE39BB91FB7CB3">Rangel-Ch, Orlando</mods:namePart>
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<mods:date id="0374802D67AC3C2BE9A6E3D6851210E0">2015</mods:date>
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<treatment id="EE301571FFE5602C79C4FC6DFB7EFB57" ID-DOI="http://doi.org/10.5281/zenodo.5658073" ID-GBIF-Taxon="119630847" ID-Zenodo-Dep="5658073" LSID="urn:lsid:plazi:treatment:EE301571FFE5602C79C4FC6DFB7EFB57" httpUri="http://treatment.plazi.org/id/EE301571FFE5602C79C4FC6DFB7EFB57" lastPageId="7" lastPageNumber="113" pageId="5" pageNumber="111">
<subSubSection id="2E83F7ECFFE5602E79C4FC6DFCACFBC3" pageId="5" pageNumber="111" type="nomenclature">
<paragraph id="6626A467FFE5602E79C4FC6DFE79FC2A" blockId="5.[151,502,935,995]" box="[151,502,935,962]" pageId="5" pageNumber="111">
<heading id="3D6E130BFFE5602E79C4FC6DFE79FC2A" bold="true" box="[151,502,935,962]" fontSize="11" level="1" pageId="5" pageNumber="111" reason="1">
<emphasis id="54ED7875FFE5602E79C4FC6DFE79FC2A" bold="true" box="[151,502,935,962]" pageId="5" pageNumber="111">
<taxonomicName id="A199DFE4FFE5602E79C4FC6DFE1FFC2A" box="[151,400,935,961]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="111" phylum="Annelida" rank="species" species="lebrijae" status="sp. nov.">
<emphasis id="54ED7875FFE5602E79C4FC6DFE1FFC2A" bold="true" box="[151,400,935,961]" italics="true" pageId="5" pageNumber="111">Righiodrilus lebrijae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE5602E78CBFC62FE79FC2A" box="[408,502,935,962]" pageId="5" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6626A467FFE5602E79C4FC00FED9FC08" blockId="5.[151,502,935,995]" box="[151,342,970,995]" pageId="5" pageNumber="111">
<figureCitation id="FEA2B8E2FFE5602E79C4FC00FF79FC08" box="[151,246,970,995]" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Figure 3</figureCitation>
,
<tableCitation id="2B1B91DCFFE5602E7850FC00FED9FC08" box="[259,342,970,995]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionTargetBox="[159,1375,202,687]" captionTargetPageId="7" captionText="TABLE 2. Comparison of species of the genus Righiodrilus with two pairs of spermathecae in segments 8 / 9 9 / 10." httpUri="http://table.plazi.org/id/32E6F4EFFFE7602C79C4FF52FAD1FF44" pageId="5" pageNumber="111" tableUuid="32E6F4EFFFE7602C79C4FF52FAD1FF44">Table 2</tableCitation>
</paragraph>
<paragraph id="6626A467FFE5602E79C4FBD8FCACFBC3" blockId="5.[151,803,1042,1064]" box="[151,803,1042,1064]" pageId="5" pageNumber="111">
<taxonomicName id="A199DFE4FFE5602E79C4FBD8FE99FBC3" box="[151,278,1042,1064]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="111" phylum="Annelida" rank="genus" status="sp. nov.">
<emphasis id="54ED7875FFE5602E79C4FBD8FE99FBC3" box="[151,278,1042,1064]" italics="true" pageId="5" pageNumber="111">Righiodrilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE5602E784FFBD9FEE1FBC3" box="[284,366,1043,1064]" pageId="5" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
: Celis &amp; Rangel.-Ch. 2013:
<specimenCount id="709F6FEEFFE5602E7BDFFBD8FD4BFBC3" box="[652,708,1042,1064]" pageId="5" pageNumber="111" type="generic">488f.</specimenCount>
, fig. 268.
</paragraph>
</subSubSection>
<subSubSection id="2E83F7ECFFE5602E79C4FB9FFE7AFAC9" pageId="5" pageNumber="111" type="materials_examined">
<paragraph id="6626A467FFE5602E79C4FB9FFE7AFAC9" blockId="5.[151,1437,1109,1998]" pageId="5" pageNumber="111">
<emphasis id="54ED7875FFE5602E79C4FB9FFE09FB85" bold="true" box="[151,390,1109,1134]" pageId="5" pageNumber="111">Material examined.</emphasis>
<typeStatus id="B9221AC5FFE5602E78C1FB9FFE70FB85" box="[402,511,1109,1134]" pageId="5" pageNumber="111" type="holotype">Holotype</typeStatus>
: LVCelis007, clitellate specimen;
<collectingCountry id="1E8EE4F7FFE5602E7AC7FB9FFB8AFB85" box="[916,1029,1109,1134]" name="Colombia" pageId="5" pageNumber="111">Colombia</collectingCountry>
, Department of Cesar, Aguachica, Barranca Lebrija, Doña María wetland (
<geoCoordinate id="03ADC2A0FFE5602E7B3DFBB3FC81FB79" box="[622,782,1145,1170]" direction="west" orientation="longitude" pageId="5" pageNumber="111" precision="1" value="-73.730446">73°43'49.6''W</geoCoordinate>
<geoCoordinate id="03ADC2A0FFE5602E7A4AFBB0FC16FB79" box="[793,921,1146,1170]" direction="north" orientation="latitude" pageId="5" pageNumber="111" precision="1" value="8.079972">8°4'47.9''N</geoCoordinate>
),
<quantity id="A1610982FFE5602E7AE2FBB3FC60FB7A" box="[945,1007,1145,1170]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="111" unit="m" value="50.0">50 m</quantity>
asl; L.V. Celis,
<date id="122782A7FFE5602E7DE7FBB0FA17FB79" box="[1204,1432,1145,1170]" pageId="5" pageNumber="111" value="2011-11-13">13 November 2011</date>
.
<typeStatus id="B9221AC5FFE5602E79C4FB54FE86FB5D" box="[151,265,1182,1206]" pageId="5" pageNumber="111" type="paratype">Paratypes</typeStatus>
: LVCelis006,
<specimenCount id="709F6FEEFFE5602E78E0FB54FDDEFB5D" box="[435,593,1181,1206]" pageId="5" pageNumber="111" type="generic">42 specimens</specimenCount>
(
<specimenCount id="709F6FEEFFE5602E7B35FB54FD59FB5D" box="[614,726,1181,1206]" pageId="5" pageNumber="111" type="adult">27 adults</specimenCount>
,
<specimenCount id="709F6FEEFFE5602E7BB5FB57FCF5FB5D" box="[742,890,1181,1206]" pageId="5" pageNumber="111" type="juvenile">15 juveniles</specimenCount>
);
<collectingCountry id="1E8EE4F7FFE5602E7AC0FB57FB8BFB5D" box="[915,1028,1181,1206]" name="Colombia" pageId="5" pageNumber="111">Colombia</collectingCountry>
, Department of Cesar, Aguachica, Barranca Lebrija, Doña María wetland,
<quantity id="A1610982FFE5602E7B0EFB0BFD4DFB32" box="[605,706,1217,1242]" metricMagnitude="1" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="5" pageNumber="111" unit="m" value="49.0" valueMax="53.0" valueMin="45.0">4553 m</quantity>
asl; L.V. Celis,
<date id="122782A7FFE5602E7A29FB0BFBBCFB31" box="[890,1075,1217,1242]" pageId="5" pageNumber="111" value="2011-11">November 2011</date>
. LVCelis003,
<specimenCount id="709F6FEEFFE5602E7D86FB0BFAE0FB31" box="[1237,1391,1217,1242]" pageId="5" pageNumber="111" type="generic">36 specimens</specimenCount>
(
<specimenCount id="709F6FEEFFE5602E7C2CFB08FF52FB15" pageId="5" pageNumber="111" type="adult">28 adults</specimenCount>
,
<specimenCount id="709F6FEEFFE5602E79B4FB2CFEE8FB15" box="[231,359,1253,1278]" pageId="5" pageNumber="111" type="juvenile">8 juveniles</specimenCount>
);
<collectingCountry id="1E8EE4F7FFE5602E782AFB2FFE65FB15" box="[377,490,1253,1278]" name="Colombia" pageId="5" pageNumber="111">Colombia</collectingCountry>
, Cesar, San Martin, Terraplen,
<collectingCountry id="1E8EE4F7FFE5602E7A15FB2FFC3FFB15" box="[838,944,1253,1278]" name="Republic of the Congo" pageId="5" pageNumber="111">El Congo</collectingCountry>
wetland (
<geoCoordinate id="03ADC2A0FFE5602E7D48FB2FFB3AFB15" box="[1051,1205,1253,1278]" direction="west" orientation="longitude" pageId="5" pageNumber="111" precision="1" value="-73.645164">73°38'42.6''W</geoCoordinate>
<geoCoordinate id="03ADC2A0FFE5602E7DE8FB2FFACCFB15" box="[1211,1347,1253,1278]" direction="north" orientation="latitude" pageId="5" pageNumber="111" precision="1" value="7.8801947">7°52'48.7''N</geoCoordinate>
),
<quantity id="A1610982FFE5602E7C05FB2FFF22FACA" metricMagnitude="1" metricUnit="m" metricValue="5.85" metricValueMax="5.9" metricValueMin="5.8" pageId="5" pageNumber="111" unit="m" value="58.5" valueMax="59.0" valueMin="58.0">5859 m</quantity>
asl. L.V. Celis,
<date id="122782A7FFE5602E780CFAC0FE7DFAC9" box="[351,498,1290,1314]" pageId="5" pageNumber="111" value="2011-08">August 2011</date>
.
</paragraph>
</subSubSection>
<subSubSection id="2E83F7ECFFE5602E7994FAE7FA87F825" pageId="5" pageNumber="111" type="description">
<paragraph id="6626A467FFE5602E7994FAE7FBF9F9AA" blockId="5.[151,1437,1109,1998]" pageId="5" pageNumber="111">
<emphasis id="54ED7875FFE5602E7994FAE7FED6FAAD" bold="true" box="[199,345,1325,1350]" pageId="5" pageNumber="111">Description.</emphasis>
Body cylindrical, without pigment. Body dimensions,
<typeStatus id="B9221AC5FFE5602E7A84FAE7FBB1FAAD" box="[983,1086,1325,1350]" pageId="5" pageNumber="111" type="holotype">holotype</typeStatus>
: length
<quantity id="A1610982FFE5602E7DCFFAE7FB7FFAAE" box="[1180,1264,1325,1350]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="111" unit="mm" value="65.0">65 mm</quantity>
, diameter
<quantity id="A1610982FFE5602E7C38FAE7FF4CFA82" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="111" unit="mm" value="1.95">1.95 mm</quantity>
in preclitellar region,
<quantity id="A1610982FFE5602E7885FA98FDBCFA82" box="[470,563,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="111" unit="mm" value="3.1">3.1 mm</quantity>
at clitellum,
<quantity id="A1610982FFE5602E7B89FA98FCB8FA82" box="[730,823,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="111" unit="mm" value="1.9">1.9 mm</quantity>
in postclitellar region (segment
<emphasis id="54ED7875FFE5602E7D91FA9BFB59FA82" box="[1218,1238,1361,1385]" italics="true" pageId="5" pageNumber="111">xl</emphasis>
), 133 segments;
<typeStatus id="B9221AC5FFE5602E79C4FABCFE88FA65" box="[151,263,1398,1422]" pageId="5" pageNumber="111" type="paratype">paratypes</typeStatus>
: length
<quantity id="A1610982FFE5602E7831FABFFE7CFA66" box="[354,499,1397,1422]" metricMagnitude="-2" metricUnit="m" metricValue="6.534999999999999" metricValueMax="7.0" metricValueMin="6.07" pageId="5" pageNumber="111" unit="mm" value="65.35" valueMax="70.0" valueMin="60.7">60.770 mm</quantity>
, diameter
<quantity id="A1610982FFE5602E7B3FFABFFD78FA66" box="[620,759,1397,1422]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.5" metricValueMin="1.8" pageId="5" pageNumber="111" unit="mm" value="2.65" valueMax="3.5" valueMin="1.8">1.83.5 mm</quantity>
, 112135 segments. Setae visible from segment
<emphasis id="54ED7875FFE5602E7C77FABCFAB4FA66" box="[1316,1339,1398,1421]" italics="true" pageId="5" pageNumber="111">iii</emphasis>
; closely paired, 8 per segment, in two ventro-lateral and two latero-dorsal rows. Setae slightly sigmoid without node. Genital setae not found. Distance between setae in mid-body region (segments
<emphasis id="54ED7875FFE5602E7D44FA77FBCDFA3E" box="[1047,1090,1469,1493]" italics="true" pageId="5" pageNumber="111">xll</emphasis>
) aa: ab: bc: cd: dd = 25: 1.0: 5.0: 1.0: 32.5 (
<typeStatus id="B9221AC5FFE5602E7809FA2BFE4CFA11" box="[346,451,1505,1530]" pageId="5" pageNumber="111" type="holotype">holotype</typeStatus>
). Clitellum saddle-shaped with barely defined margins in segments
<emphasis id="54ED7875FFE5602E7D89FA28FAAEFA12" box="[1242,1313,1506,1529]" italics="true" pageId="5" pageNumber="111">xvxxi</emphasis>
; tubercula pubertatis in ½
<emphasis id="54ED7875FFE5602E7814F9CCFEE8F9F6" box="[327,359,1542,1565]" italics="true" pageId="5" pageNumber="111">xvi</emphasis>
–½
<emphasis id="54ED7875FFE5602E78C1F9CCFE3CF9F6" box="[402,435,1542,1565]" italics="true" pageId="5" pageNumber="111">xix</emphasis>
(
<figureCitation id="FEA2B8E2FFE5602E7890F9CFFD89F9F5" box="[451,518,1541,1566]" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Fig. 3</figureCitation>
A). Male pores in intersegment 17/18, female pores in segment
<emphasis id="54ED7875FFE5602E7D85F9CCFB7AF9F6" box="[1238,1269,1542,1565]" italics="true" pageId="5" pageNumber="111">xiv</emphasis>
. Spermathecal pores visible in
<date id="122782A7FFE5602E781AF9E0FE57F9A9" box="[329,472,1577,1602]" pageId="5" pageNumber="111" value="2010-08-09" valueMax="2010-08-09" valueMin="2010-08-09">8/9 and 9/10</date>
. Nephridiopores above setae
<emphasis id="54ED7875FFE5602E7A72F9E3FCA0F9AA" box="[801,815,1577,1601]" italics="true" pageId="5" pageNumber="111">b</emphasis>
visible from intersegment
<emphasis id="54ED7875FFE5602E7D0FF9E0FBE1F9AA" box="[1116,1134,1578,1601]" italics="true" pageId="5" pageNumber="111">iv</emphasis>
.
</paragraph>
<paragraph id="6626A467FFE5602E7994F987FA87F825" blockId="5.[151,1437,1109,1998]" pageId="5" pageNumber="111">
<emphasis id="54ED7875FFE5602E7994F987FE2BF98D" bold="true" box="[199,420,1613,1638]" pageId="5" pageNumber="111">Internal anatomy.</emphasis>
<collectingCountry id="1E8EE4F7FFE5602E78E2F984FE60F98D" box="[433,495,1614,1638]" name="Spain" pageId="5" pageNumber="111">Septa</collectingCountry>
<date id="122782A7FFE5602E78AFF984FDE2F98D" box="[508,621,1614,1638]" pageId="5" pageNumber="111" value="2011-08-07" valueMax="2011-10-07" valueMin="2011-08-07">7/810/11</date>
thick and muscularized. Gizzard strongly muscularized, rounded, in segment
<emphasis id="54ED7875FFE5602E79ADF9B8FE9DF962" box="[254,274,1650,1673]" italics="true" pageId="5" pageNumber="111">vi</emphasis>
. One pair of extramural, pyriform calciferous glands with tubular structure (
<figureCitation id="FEA2B8E2FFE5602E7DDBF9BBFB43F961" box="[1160,1228,1649,1674]" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Fig. 3</figureCitation>
B) emerging from postero-lateral region of esophagus of segment
<emphasis id="54ED7875FFE5602E7BFCF95CFD4CF946" box="[687,707,1686,1709]" italics="true" pageId="5" pageNumber="111">xi</emphasis>
and extending to segment
<emphasis id="54ED7875FFE5602E7AA7F95CFB9AF946" box="[1012,1045,1686,1709]" italics="true" pageId="5" pageNumber="111">xiv</emphasis>
; glandular part in segments
<emphasis id="54ED7875FFE5602E7C05F95CFAE5F946" box="[1366,1386,1686,1709]" italics="true" pageId="5" pageNumber="111">xi</emphasis>
,
<emphasis id="54ED7875FFE5602E7C2AF95CFA1AF946" box="[1401,1429,1686,1709]" italics="true" pageId="5" pageNumber="111">xii</emphasis>
, membranous part in segments
<emphasis id="54ED7875FFE5602E78AFF970FDAFF93A" box="[508,544,1722,1745]" italics="true" pageId="5" pageNumber="111">xiii</emphasis>
,
<emphasis id="54ED7875FFE5602E7B61F970FDDCF93A" box="[562,595,1722,1745]" italics="true" pageId="5" pageNumber="111">xiv</emphasis>
(
<figureCitation id="FEA2B8E2FFE5602E7B35F973FD23F939" box="[614,684,1721,1746]" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Fig. 3</figureCitation>
C). Transition of esophagus-intestine in segment
<emphasis id="54ED7875FFE5602E7DB9F970FA81F93A" box="[1258,1294,1722,1745]" italics="true" pageId="5" pageNumber="111">xiii</emphasis>
. Typhlosole simple, from segment
<emphasis id="54ED7875FFE5602E78C7F914FE3CF91E" box="[404,435,1758,1781]" italics="true" pageId="5" pageNumber="111">xiv</emphasis>
, occupying 1/3 of gut diameter, folded between segments
<emphasis id="54ED7875FFE5602E7D1FF914FB15F91E" box="[1100,1178,1758,1781]" italics="true" pageId="5" pageNumber="111">xivxxi</emphasis>
, with folds decreasing in number and size from segment
<emphasis id="54ED7875FFE5602E7B4FF8C8FDCBF8F2" box="[540,580,1794,1817]" italics="true" pageId="5" pageNumber="111">xxii</emphasis>
to
<emphasis id="54ED7875FFE5602E7B3FF8CBFDFBF8F2" box="[620,628,1793,1817]" italics="true" pageId="5" pageNumber="111">l</emphasis>
. Caeca absent. Three pairs of lateral hearts in segments
<emphasis id="54ED7875FFE5602E7DABF8C8FA9BF8F2" box="[1272,1300,1794,1817]" italics="true" pageId="5" pageNumber="111">vii</emphasis>
,
<emphasis id="54ED7875FFE5602E7C77F8C8FAC7F8F2" box="[1316,1352,1794,1817]" italics="true" pageId="5" pageNumber="111">viii</emphasis>
and
<emphasis id="54ED7875FFE5602E7CD2F8C8FA1AF8F2" box="[1409,1429,1794,1817]" italics="true" pageId="5" pageNumber="111">ix</emphasis>
, and two pairs of voluminous intestinal hearts in segments
<emphasis id="54ED7875FFE5602E7A66F8ECFCCEF8D6" box="[821,833,1830,1853]" italics="true" pageId="5" pageNumber="111">x</emphasis>
and
<emphasis id="54ED7875FFE5602E7A2FF8ECFC1FF8D6" box="[892,912,1830,1853]" italics="true" pageId="5" pageNumber="111">xi</emphasis>
. One pair of holonephridia per segment (
<figureCitation id="FEA2B8E2FFE5602E7C3CF8EFFF2AF889" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Fig. 3</figureCitation>
D). Male funnels free in segment
<emphasis id="54ED7875FFE5602E7B77F880FDB7F88A" box="[548,568,1866,1889]" italics="true" pageId="5" pageNumber="111">xi</emphasis>
and connected to deferent ducts. One pair of seminal vesicles from segment
<emphasis id="54ED7875FFE5602E79C4F8A4FF3CF86E" box="[151,179,1902,1925]" italics="true" pageId="5" pageNumber="111">xii</emphasis>
, extending backwards along the lateral body region to segments
<emphasis id="54ED7875FFE5602E7AD7F8A4FC3EF86E" box="[900,945,1902,1925]" italics="true" pageId="5" pageNumber="111">xxix</emphasis>
,
<emphasis id="54ED7875FFE5602E7AEDF8A4FC64F86E" box="[958,1003,1902,1925]" italics="true" pageId="5" pageNumber="111">xxx,</emphasis>
or
<emphasis id="54ED7875FFE5602E7D5CF8A4FBB3F86E" box="[1039,1084,1902,1925]" italics="true" pageId="5" pageNumber="111">xxxi</emphasis>
. Male ducts run along the body wall down to 17/18. Ovaries in segment
<emphasis id="54ED7875FFE5602E7BD3F858FD2BF842" box="[640,676,1938,1961]" italics="true" pageId="5" pageNumber="111">xiii</emphasis>
. Two pairs of spermathecae in intersegments 8/9, 9/10; anterior spermathcae with long duct and rounded ampulla, posterior spermatheca with oval ampulla (
<figureCitation id="FEA2B8E2FFE5602E7DF7F87FFB69F825" box="[1188,1254,1973,1998]" captionStart="FIGURE 3" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[199,1393,219,1630]" captionTargetId="figure@6.[192,1407,207,1652]" captionTargetPageId="6" captionText="FIGURE 3. Righiodrilus lebrijae sp. nov. A. Ventral view. B. Calciferous gland. C. Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle). D. Nephridium. E. Spermathecae." httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="5" pageNumber="111">Fig. 3</figureCitation>
E).
</paragraph>
</subSubSection>
<caption id="32E6F4EFFFE6602D79C4F957FAF9F939" httpUri="https://zenodo.org/record/232031/files/figure.png" pageId="6" pageNumber="112" targetBox="[199,1393,219,1630]" targetPageId="6">
<paragraph id="6626A467FFE6602D79C4F957FAF9F939" blockId="6.[151,1436,1693,1746]" pageId="6" pageNumber="112">
<emphasis id="54ED7875FFE6602D79C4F957FDEAF958" bold="true" box="[151,613,1693,1715]" pageId="6" pageNumber="112">
FIGURE 3.
<taxonomicName id="A199DFE4FFE6602D784BF957FE7FF958" box="[280,496,1693,1715]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="112" phylum="Annelida" rank="species" species="lebrijae" status="sp. nov.">
<emphasis id="54ED7875FFE6602D784BF957FE7FF958" bold="true" box="[280,496,1693,1715]" italics="true" pageId="6" pageNumber="112">Righiodrilus lebrijae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE6602D78A6F957FDC9F958" box="[501,582,1693,1715]" pageId="6" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Ventral view.
<emphasis id="54ED7875FFE6602D7BA5F957FC82F958" bold="true" box="[758,781,1693,1715]" pageId="6" pageNumber="112">B.</emphasis>
Calciferous gland.
<emphasis id="54ED7875FFE6602D7A81F957FC64F958" bold="true" box="[978,1003,1693,1715]" pageId="6" pageNumber="112">C.</emphasis>
Internal dorsal view (Gp, glandular part of the calciferous glands; Mp, membranous part of calciferous glands; Sv, seminal vesicle).
<emphasis id="54ED7875FFE6602D7D4DF976FBB9F939" bold="true" box="[1054,1078,1724,1746]" pageId="6" pageNumber="112">D.</emphasis>
Nephridium.
<emphasis id="54ED7875FFE6602D7D97F976FB54F939" bold="true" box="[1220,1243,1724,1746]" pageId="6" pageNumber="112">E.</emphasis>
Spermathecae.
</paragraph>
</caption>
<subSubSection id="2E83F7ECFFE6602D7994F934FBA7F84C" pageId="6" pageNumber="112" type="etymology">
<paragraph id="6626A467FFE6602D7994F934FE82F8D0" blockId="6.[151,1436,1790,1959]" pageId="6" pageNumber="112">
<emphasis id="54ED7875FFE6602D7994F934FEC1F8FC" bold="true" box="[199,334,1790,1815]" pageId="6" pageNumber="112">Etymology.</emphasis>
The species is named after the Lebrija River, which flows into several wetlands in southern Cesar,
<collectingCountry id="1E8EE4F7FFE6602D79C4F8E8FE85F8D0" box="[151,266,1826,1851]" name="Colombia" pageId="6" pageNumber="112">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="6626A467FFE6602D7994F88CFBA7F84C" blockId="6.[151,1436,1790,1959]" pageId="6" pageNumber="112">
<emphasis id="54ED7875FFE6602D7994F88CFE67F8B4" bold="true" box="[199,488,1862,1887]" pageId="6" pageNumber="112">Habitat characteristics.</emphasis>
<taxonomicName id="A199DFE4FFE6602D78A4F88DFD64F8B4" box="[503,747,1863,1887]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="112" phylum="Annelida" rank="species" species="lebrijae" status="sp. nov.">
<emphasis id="54ED7875FFE6602D78A4F88DFD64F8B4" box="[503,747,1863,1887]" italics="true" pageId="6" pageNumber="112">Righiodrilus lebrijae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE6602D7BA8F88CFCD2F8B4" box="[763,861,1862,1887]" pageId="6" pageNumber="112" rank="species">
<emphasis id="54ED7875FFE6602D7BA8F88CFCD2F8B4" bold="true" box="[763,861,1862,1887]" pageId="6" pageNumber="112">sp. nov.</emphasis>
</taxonomicNameLabel>
was collected in three physiognomic
<typeStatus id="B9221AC5FFE6602D7C69F882FAFAF88B" box="[1338,1397,1864,1888]" pageId="6" pageNumber="112">types</typeStatus>
of vegetation (relicts of tropical dry forests and savannas and herbaceous plant formation with trees) in
<collectingCountry id="1E8EE4F7FFE6602D7C53F8A0FAE4F868" box="[1280,1387,1898,1923]" name="Republic of the Congo" pageId="6" pageNumber="112">El Congo</collectingCountry>
and Doña Maria wetlands, but not in highly transformed areas (
<bibRefCitation id="0208D996FFE6602D7A79F844FB92F84C" author="Celis" box="[810,1053,1934,1959]" pageId="6" pageNumber="112" refString="Celis, L. V. &amp; Rangel-Ch., J. O. (2013) Lombrices de tierra: riqueza y abundancia en las zonas aledanas a complejos cenagosos del Sur del departamento del Cesar. In: Rangel-Ch., J. O. (Ed.), Colombia Diversidad Biotica XIII. Complejo cenagoso Zapatosa y cienagas del Sur del Cesar. Biodiversidad, conservacion y manejo: Universidad Nacional de Colombia - Instituto de Ciencias Naturales. Bogota, D. C., pp. 485 - 498." type="book chapter" year="2013">Celis &amp; Rangel 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="32E6F4EFFFE7602C79C4FF52FAD1FF44" ID-Table-UUID="32E6F4EFFFE7602C79C4FF52FAD1FF44" box="[151,1374,151,175]" httpUri="http://table.plazi.org/id/32E6F4EFFFE7602C79C4FF52FAD1FF44" pageId="7" pageNumber="113" targetBox="[159,1375,202,687]" targetIsTable="true" targetPageId="7">
<paragraph id="6626A467FFE7602C79C4FF52FAD1FF44" blockId="7.[151,1374,151,175]" box="[151,1374,151,175]" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C79C4FF52FE86FF44" bold="true" box="[151,265,152,175]" pageId="7" pageNumber="113">TABLE 2.</emphasis>
Comparison of species of the genus
<taxonomicName id="A199DFE4FFE7602C7BC0FF5DFC97FF44" box="[659,792,151,175]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7BC0FF5DFC97FF44" box="[659,792,151,175]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
with two pairs of spermathecae in segments 8/99/10.
</paragraph>
</caption>
<paragraph id="6626A467FFE7602C79CCFF00FAD1FD7B" pageId="7" pageNumber="113">
<table id="149956C7FFE79FD479CCFF00FAD0FD44" box="[159,1375,202,687]" gridcols="6" gridrows="7" pageId="7" pageNumber="113">
<tr id="D8A9A625FFE79FD479CCFF00FAD0FF14" box="[159,1375,202,255]" gridrow="0" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFF00FEEFFF14" box="[159,352,202,255]" gridcol="0" gridrow="0" pageId="7" pageNumber="113">Species</th>
<th id="9B78CF59FFE79FD478C7FF00FE62FF14" box="[404,493,202,255]" gridcol="1" gridrow="0" pageId="7" pageNumber="113">Length (mm)</th>
<th id="9B78CF59FFE79FD47B4AFF00FD28FF14" box="[537,679,202,255]" gridcol="2" gridrow="0" pageId="7" pageNumber="113">Clitellum</th>
<th id="9B78CF59FFE79FD47BBFFF00FC33FF14" box="[748,956,202,255]" gridcol="3" gridrow="0" pageId="7" pageNumber="113">Tubercula pubertatis</th>
<th id="9B78CF59FFE79FD47ABAFF00FB62FF14" box="[1001,1261,202,255]" gridcol="4" gridrow="0" pageId="7" pageNumber="113">Posterior extension of seminal vesicles</th>
<th id="9B78CF59FFE79FD47C76FF00FAD0FF14" box="[1317,1375,202,255]" gridcol="5" gridrow="0" pageId="7" pageNumber="113">Male pores</th>
</tr>
<tr id="D8A9A625FFE79FD479CCFED8FAD0FEAC" box="[159,1375,274,327]" gridrow="1" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFED8FEEFFEAC" box="[159,352,274,327]" gridcol="0" gridrow="1" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFED9FE99FEAC" authority="Righi, 1982" authorityName="Righi" authorityYear="1982" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="arapaco">
<emphasis id="54ED7875FFE7602C79CCFED9FE80FEC3" box="[159,271,275,296]" italics="true" pageId="7" pageNumber="113">R. arapaco</emphasis>
Righi, 1982
</taxonomicName>
</th>
<td id="9B78CF59FFE79FD478C7FED8FE62FEAC" box="[404,493,274,327]" gridcol="1" gridrow="1" pageId="7" pageNumber="113">4560</td>
<td id="9B78CF59FFE79FD47B4AFED8FD28FEAC" box="[537,679,274,327]" gridcol="2" gridrow="1" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7B4AFED8FD15FEC3" box="[537,666,274,296]" italics="true" pageId="7" pageNumber="113">½ xvi½ xxii</emphasis>
(6 segments)
</td>
<td id="9B78CF59FFE79FD47BBFFED8FC33FEAC" box="[748,956,274,327]" gridcol="3" gridrow="1" pageId="7" pageNumber="113">½ xviii½ xx (2 segments)</td>
<td id="9B78CF59FFE79FD47ABAFED8FB62FEAC" box="[1001,1261,274,327]" gridcol="4" gridrow="1" pageId="7" pageNumber="113">
to segment
<emphasis id="54ED7875FFE7602C7D37FED9FB0EFEC3" box="[1124,1153,275,296]" italics="true" pageId="7" pageNumber="113">xiv</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FED8FAD0FEAC" box="[1317,1375,274,327]" gridcol="5" gridrow="1" pageId="7" pageNumber="113">19/20</td>
</tr>
<tr id="D8A9A625FFE79FD479CCFE90FAD0FE64" box="[159,1375,346,399]" gridrow="2" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFE90FEEFFE64" box="[159,352,346,399]" gridcol="0" gridrow="2" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFE91FE99FE64" authority="Righi, 1971" authorityName="Righi" authorityYear="1971" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="itajo">
<emphasis id="54ED7875FFE7602C79CCFE91FF65FE9B" box="[159,234,347,368]" italics="true" pageId="7" pageNumber="113">R. itajo</emphasis>
Righi, 1971
</taxonomicName>
</th>
<td id="9B78CF59FFE79FD478C7FE90FE62FE64" box="[404,493,346,399]" gridcol="1" gridrow="2" pageId="7" pageNumber="113">3874</td>
<td id="9B78CF59FFE79FD47B4AFE90FD28FE64" box="[537,679,346,399]" gridcol="2" gridrow="2" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7B4AFE91FDEFFE9B" box="[537,608,347,368]" italics="true" pageId="7" pageNumber="113">xvxxii</emphasis>
(8 segments)
</td>
<td id="9B78CF59FFE79FD47BBFFE90FC33FE64" box="[748,956,346,399]" gridcol="3" gridrow="2" pageId="7" pageNumber="113">½ xvi½ xviii (2 segments)</td>
<td id="9B78CF59FFE79FD47ABAFE90FB62FE64" box="[1001,1261,346,399]" gridcol="4" gridrow="2" pageId="7" pageNumber="113">
to segments
<emphasis id="54ED7875FFE7602C7D34FE90FB2CFE9B" box="[1127,1187,346,368]" italics="true" pageId="7" pageNumber="113">llviii</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FE90FAD0FE64" box="[1317,1375,346,399]" gridcol="5" gridrow="2" pageId="7" pageNumber="113">16/17</td>
</tr>
<tr id="D8A9A625FFE79FD479CCFE68FAD0FE3C" box="[159,1375,418,471]" gridrow="3" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFE68FEEFFE3C" box="[159,352,418,471]" gridcol="0" gridrow="3" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFE69FE99FE3C" authority="Righi, 1982" authorityName="Righi" authorityYear="1982" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="mairaro">
<emphasis id="54ED7875FFE7602C79CCFE69FE82FE53" box="[159,269,419,440]" italics="true" pageId="7" pageNumber="113">R. mairaro</emphasis>
Righi, 1982
</taxonomicName>
</th>
<td id="9B78CF59FFE79FD478C7FE68FE62FE3C" box="[404,493,418,471]" gridcol="1" gridrow="3" pageId="7" pageNumber="113">60115</td>
<td id="9B78CF59FFE79FD47B4AFE68FD28FE3C" box="[537,679,418,471]" gridcol="2" gridrow="3" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7B4AFE68FD15FE53" box="[537,666,418,440]" italics="true" pageId="7" pageNumber="113">½ xvi½ xxii</emphasis>
(6 segments)
</td>
<td id="9B78CF59FFE79FD47BBFFE68FC33FE3C" box="[748,956,418,471]" gridcol="3" gridrow="3" pageId="7" pageNumber="113">½ xviii½ xxxx (2,2½ segments)</td>
<td id="9B78CF59FFE79FD47ABAFE68FB62FE3C" box="[1001,1261,418,471]" gridcol="4" gridrow="3" pageId="7" pageNumber="113">
to segments
<emphasis id="54ED7875FFE7602C7D34FE69FB5EFE53" box="[1127,1233,419,440]" italics="true" pageId="7" pageNumber="113">xxivxxxvi</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FE68FAD0FE3C" box="[1317,1375,418,471]" gridcol="5" gridrow="3" pageId="7" pageNumber="113">18/19</td>
</tr>
<tr id="D8A9A625FFE79FD479CCFE20FAD0FDF4" box="[159,1375,490,543]" gridrow="4" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFE20FEEFFDF4" box="[159,352,490,543]" gridcol="0" gridrow="4" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFE21FE99FDF4" authority="Righi, 1982" authorityName="Righi" authorityYear="1982" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="oliveirae">
<emphasis id="54ED7875FFE7602C79CCFE21FE9AFDEB" box="[159,277,490,512]" italics="true" pageId="7" pageNumber="113">R. oliveirae</emphasis>
Righi, 1982
</taxonomicName>
</th>
<td id="9B78CF59FFE79FD478C7FE20FE62FDF4" box="[404,493,490,543]" gridcol="1" gridrow="4" pageId="7" pageNumber="113">180250</td>
<td id="9B78CF59FFE79FD47B4AFE20FD28FDF4" box="[537,679,490,543]" gridcol="2" gridrow="4" pageId="7" pageNumber="113">xvxxiv (10 segments)</td>
<td id="9B78CF59FFE79FD47BBFFE20FC33FDF4" box="[748,956,490,543]" gridcol="3" gridrow="4" pageId="7" pageNumber="113">⅔xviiixx (2⅔ segments)</td>
<td id="9B78CF59FFE79FD47ABAFE20FB62FDF4" box="[1001,1261,490,543]" gridcol="4" gridrow="4" pageId="7" pageNumber="113">
to segment
<emphasis id="54ED7875FFE7602C7D0EFE21FBF5FDEB" box="[1117,1146,491,512]" italics="true" pageId="7" pageNumber="113">xiv</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FE20FAD0FDF4" box="[1317,1375,490,543]" gridcol="5" gridrow="4" pageId="7" pageNumber="113">⅔19</td>
</tr>
<tr id="D8A9A625FFE79FD479CCFDF8FAD0FD8C" box="[159,1375,562,615]" gridrow="5" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFDF8FEEFFD8C" box="[159,352,562,615]" gridcol="0" gridrow="5" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFDF9FE99FD8C" authority="Righi, 1984" authorityName="Righi" authorityYear="1984" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C79CCFDF9FE80FDA3" box="[159,271,562,584]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
Righi, 1984
</taxonomicName>
a
</th>
<td id="9B78CF59FFE79FD478C7FDF8FE62FD8C" box="[404,493,562,615]" gridcol="1" gridrow="5" pageId="7" pageNumber="113">30</td>
<td id="9B78CF59FFE79FD47B4AFDF8FD28FD8C" box="[537,679,562,615]" gridcol="2" gridrow="5" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7B4AFDF9FDD6FDA3" box="[537,601,563,584]" italics="true" pageId="7" pageNumber="113">xvxxi</emphasis>
(7 segments)
</td>
<td id="9B78CF59FFE79FD47BBFFDF8FC33FD8C" box="[748,956,562,615]" gridcol="3" gridrow="5" pageId="7" pageNumber="113">xvii½ xix (2 ½ segments)</td>
<td id="9B78CF59FFE79FD47ABAFDF8FB62FD8C" box="[1001,1261,562,615]" gridcol="4" gridrow="5" pageId="7" pageNumber="113">
to segment
<emphasis id="54ED7875FFE7602C7D0EFDF9FB09FDA3" box="[1117,1158,563,584]" italics="true" pageId="7" pageNumber="113">xxvi</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FDF8FAD0FD8C" box="[1317,1375,562,615]" gridcol="5" gridrow="5" pageId="7" pageNumber="113">17</td>
</tr>
<tr id="D8A9A625FFE79FD479CCFDB0FAD0FD44" box="[159,1375,634,687]" gridrow="6" pageId="7" pageNumber="113">
<th id="9B78CF59FFE79FD479CCFDB0FEEFFD44" box="[159,352,634,687]" gridcol="0" gridrow="6" pageId="7" pageNumber="113">
<taxonomicName id="A199DFE4FFE7602C79CCFDB1FE85FD7B" box="[159,266,634,656]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="lebrijae" status="sp. nov.">
<emphasis id="54ED7875FFE7602C79CCFDB1FE85FD7B" box="[159,266,634,656]" italics="true" pageId="7" pageNumber="113">R. lebrijae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE7602C7843FDB0FEEFFD7B" box="[272,352,634,656]" pageId="7" pageNumber="113" rank="species">
<emphasis id="54ED7875FFE7602C7843FDB0FEEFFD7B" bold="true" box="[272,352,634,656]" pageId="7" pageNumber="113">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<td id="9B78CF59FFE79FD478C7FDB0FE62FD44" box="[404,493,634,687]" gridcol="1" gridrow="6" pageId="7" pageNumber="113">60.770</td>
<td id="9B78CF59FFE79FD47B4AFDB0FD28FD44" box="[537,679,634,687]" gridcol="2" gridrow="6" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7B4AFDB1FDD6FD7B" box="[537,601,635,656]" italics="true" pageId="7" pageNumber="113">xvxxi</emphasis>
(7 segments)
</td>
<td id="9B78CF59FFE79FD47BBFFDB0FC33FD44" box="[748,956,634,687]" gridcol="3" gridrow="6" pageId="7" pageNumber="113">½ xvi½ xix (3 segments)</td>
<td id="9B78CF59FFE79FD47ABAFDB0FB62FD44" box="[1001,1261,634,687]" gridcol="4" gridrow="6" pageId="7" pageNumber="113">
to segments
<emphasis id="54ED7875FFE7602C7D34FDB1FB62FD7B" box="[1127,1261,635,656]" italics="true" pageId="7" pageNumber="113">xxix,xxxxxxi</emphasis>
</td>
<td id="9B78CF59FFE79FD47C76FDB0FAD0FD44" box="[1317,1375,634,687]" gridcol="5" gridrow="6" pageId="7" pageNumber="113">17/18</td>
</tr>
</table>
</paragraph>
<subSubSection id="2E83F7ECFFE7602C7994FD3EFB7EFB57" pageId="7" pageNumber="113" type="discussion">
<paragraph id="6626A467FFE7602C7994FD3EFB7EFB57" blockId="7.[151,1436,756,1213]" pageId="7" pageNumber="113">
<emphasis id="54ED7875FFE7602C7994FD3EFEC0FCE6" bold="true" box="[199,335,756,781]" pageId="7" pageNumber="113">Comments.</emphasis>
<bibRefCitation id="0208D996FFE7602C780AFD3EFE7CFCE6" author="Zicsi" box="[345,499,756,781]" pageId="7" pageNumber="113" refString="Zicsi, A. (1995 b) Revision der Gattung Glossodrilus Cognetti, 1905 auf Grund der Arten aus dem Andengebiet (Oligochaeta: Glossoscolecidae). Regenwurmer aus Sudamerika, 25. Opuscula Zoologica, Budapest, 27 / 28, 79 - 116." type="journal article" year="1995" yearSuffix="b">Zicsi (1995b)</bibRefCitation>
separated the genus
<taxonomicName id="A199DFE4FFE7602C7BB4FD3EFCFCFCE7" box="[743,883,756,780]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7BB4FD3EFCFCFCE7" box="[743,883,756,780]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
from the genus
<taxonomicName id="A199DFE4FFE7602C7D67FD3EFAF9FCE6" authority="Cognetti, 1905" authorityName="Cognetti" authorityYear="1905" box="[1076,1398,756,781]" class="Clitellata" family="Glossoscolecidae" genus="Glossodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7D67FD3EFB4CFCE7" box="[1076,1219,756,780]" italics="true" pageId="7" pageNumber="113">Glossodrilus</emphasis>
Cognetti, 1905
</taxonomicName>
by two major criteria, one being taxonomic (the genus
<taxonomicName id="A199DFE4FFE7602C7BB6FCD2FCFEFCDB" box="[741,881,792,816]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7BB6FCD2FCFEFCDB" box="[741,881,792,816]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
has a membranous part in the calciferous gland) and the other one referring to its geographical distribution: species of
<taxonomicName id="A199DFE4FFE7602C7AFDFCF6FBB5FCBF" box="[942,1082,828,852]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7AFDFCF6FBB5FCBF" box="[942,1082,828,852]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
are found in
<collectingCountry id="1E8EE4F7FFE7602C7D80FCF6FA95FCBE" box="[1235,1306,828,853]" name="Brazil" pageId="7" pageNumber="113">Brazil</collectingCountry>
,
<collectingCountry id="1E8EE4F7FFE7602C7C75FCF6FA18FCBE" box="[1318,1431,828,853]" name="Colombia" pageId="7" pageNumber="113">Colombia</collectingCountry>
,
<collectingCountry id="1E8EE4F7FFE7602C79C4FC95FF7BFC93" box="[151,244,863,888]" name="Ecuador" pageId="7" pageNumber="113">Ecuador</collectingCountry>
and
<collectingCountry id="1E8EE4F7FFE7602C787FFC95FE10FC93" box="[300,415,863,888]" name="Venezuela" pageId="7" pageNumber="113">Venezuela</collectingCountry>
in rainforests below
<quantity id="A1610982FFE7602C7BDDFCAAFD6CFC9C" box="[654,739,864,888]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="113" unit="m" value="1000.0">1000 m</quantity>
asl, whereas species of
<taxonomicName id="A199DFE4FFE7602C7AA0FCAAFB0DFC93" box="[1011,1154,864,888]" class="Clitellata" family="Glossoscolecidae" genus="Glossodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7AA0FCAAFB0DFC93" box="[1011,1154,864,888]" italics="true" pageId="7" pageNumber="113">Glossodrilus</emphasis>
</taxonomicName>
are found above
<quantity id="A1610982FFE7602C7C15FCAAFA13FC9C" box="[1350,1436,864,888]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="113" unit="m" value="1000.0">1000 m</quantity>
in altitude (
<bibRefCitation id="0208D996FFE7602C7844FC4EFD8BFC77" author="Feijoo" box="[279,516,900,925]" pageId="7" pageNumber="113" refString="Feijoo, A. &amp; Celis, L. V. (2010) Tres nuevas especies de Righiodrilus Zicsi, 1995 (Annelida, Oligochaeta: Glossoscolecidae) de la Amazonia colombiana. Acta Amazonica, 40, 231 - 240. http: // dx. doi. org / 10.1590 / S 0044 - 59672010000100030" type="journal article" year="2010">Feijoo &amp; Celis 2010</bibRefCitation>
). The new species belongs to a group of species of
<taxonomicName id="A199DFE4FFE7602C7D1BFC4EFB5BFC77" box="[1096,1236,900,924]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C7D1BFC4EFB5BFC77" box="[1096,1236,900,924]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
with two pairs of spermathecae in intersegments
<date id="122782A7FFE7602C78ACFC62FDE8FC2B" box="[511,615,936,960]" pageId="7" pageNumber="113" value="2010-08-09" valueMax="2010-08-09" valueMin="2010-08-09">8/99/10</date>
.
<taxonomicName id="A199DFE4FFE7602C7B27FC62FCEEFC2B" box="[628,865,936,960]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="lebrijae" status="sp. nov.">
<emphasis id="54ED7875FFE7602C7B27FC62FCEEFC2B" box="[628,865,936,960]" italics="true" pageId="7" pageNumber="113">Righiodrilus lebrijae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FDEC50EFFE7602C7A3FFC6DFC46FC2B" box="[876,969,935,960]" pageId="7" pageNumber="113" rank="species">
<emphasis id="54ED7875FFE7602C7A3FFC6DFC46FC2B" bold="true" box="[876,969,935,960]" pageId="7" pageNumber="113">sp. nov.</emphasis>
</taxonomicNameLabel>
is close to
<taxonomicName id="A199DFE4FFE7602C7D07FC63FAFAFC2B" authority="Righi, 1984" authorityName="Righi" authorityYear="1984" box="[1108,1397,935,960]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7D07FC63FB5BFC2B" box="[1108,1236,936,960]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
<bibRefCitation id="0208D996FFE7602C7D8DFC6DFAFAFC2B" author="Righi" box="[1246,1397,935,960]" pageId="7" pageNumber="113" refString="Righi, G. (1984 a) Nova contribuicao ao conhecimento dos Oligochaeta da Venezuela. Papeis Avulsos de Zoologia, Sao Paulo, 35, 243 - 256." type="journal article" year="1984" yearSuffix="a">Righi, 1984a</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="A199DFE4FFE7602C7CD7FC63FE0CFC0F" authority="Righi, 1982" authorityName="Righi" authorityYear="1982" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="arapaco">
<emphasis id="54ED7875FFE7602C7CD7FC63FF7BFC0F" italics="true" pageId="7" pageNumber="113">R. arapaco</emphasis>
<bibRefCitation id="0208D996FFE7602C79AFFC06FE0CFC0F" author="Righi" box="[252,387,972,997]" pageId="7" pageNumber="113" refString="Righi, G. (1982) Oligochaeta, Glossoscolecidae, do Parque Nacional da Amazonia, Tapajos. Revista Brasileira de Biologia, 42, 107 - 116." type="journal article" year="1982">Righi, 1982</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="A199DFE4FFE7602C7893FC07FD2CFC0F" authority="Righi, 1971" authorityName="Righi" authorityYear="1971" box="[448,675,972,997]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="itajo">
<emphasis id="54ED7875FFE7602C7893FC07FD9BFC0F" box="[448,532,973,996]" italics="true" pageId="7" pageNumber="113">R. itajo</emphasis>
<bibRefCitation id="0208D996FFE7602C7B4FFC06FD2CFC0F" author="Righi" box="[540,675,972,997]" pageId="7" pageNumber="113" refString="Righi, G. (1971) Sobre a familia Glossoscolecidae (Oligochaeta) no Brasil. Arquivos de Zoologia, 20, 1 - 96. http: // dx. doi. org / 10.11606 / issn. 2176 - 7793. v 20 i 1 p 1 - 95" type="journal article" year="1971">Righi, 1971</bibRefCitation>
</taxonomicName>
, in the position of the clitellum. Within this group, the new species has the same extension of the clitellum as
<taxonomicName id="A199DFE4FFE7602C7BC9FC3BFC94FBE3" box="[666,795,1008,1032]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7BC9FC3BFC94FBE3" box="[666,795,1008,1032]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
</taxonomicName>
(
<emphasis id="54ED7875FFE7602C7A62FC3AFCF7FBEC" box="[817,888,1008,1031]" italics="true" pageId="7" pageNumber="113">xvxxi</emphasis>
), but differs in the extension of the tubercula pubertatis (3 segments
<emphasis id="54ED7875FFE7602C78C9FBDFFE3EFBC7" box="[410,433,1045,1068]" italics="true" pageId="7" pageNumber="113">vs</emphasis>
. 2 ½ in
<taxonomicName id="A199DFE4FFE7602C7B5EFBDFFD05FBC7" box="[525,650,1044,1068]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7B5EFBDFFD05FBC7" box="[525,650,1044,1068]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
</taxonomicName>
), the extension of the seminal vesicles (
<emphasis id="54ED7875FFE7602C7D1FFBDFFB3CFBC7" box="[1100,1203,1045,1068]" italics="true" pageId="7" pageNumber="113">xxix, xxx,</emphasis>
or
<emphasis id="54ED7875FFE7602C7D8AFBDFFAABFBC7" box="[1241,1316,1045,1068]" italics="true" pageId="7" pageNumber="113">xxxi vs</emphasis>
.
<emphasis id="54ED7875FFE7602C7C60FBDFFAEFFBC7" box="[1331,1376,1045,1068]" italics="true" pageId="7" pageNumber="113">xxvi</emphasis>
in
<taxonomicName id="A199DFE4FFE7602C7CD7FBDFFF7BFBBB" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7CD7FBDFFF7BFBBB" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
</taxonomicName>
), the position of the male pores (17/18
<emphasis id="54ED7875FFE7602C7BF8FBF3FD4DFBBB" box="[683,706,1081,1104]" italics="true" pageId="7" pageNumber="113">vs</emphasis>
. anterior part of segment
<emphasis id="54ED7875FFE7602C7AB2FBF2FB86FBA4" box="[993,1033,1080,1103]" italics="true" pageId="7" pageNumber="113">xvii</emphasis>
in
<taxonomicName id="A199DFE4FFE7602C7D7FFBF3FB28FBBB" box="[1068,1191,1080,1104]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7D7FFBF3FB28FBBB" box="[1068,1191,1080,1104]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
</taxonomicName>
), the arrangements of setae (closely paired
<emphasis id="54ED7875FFE7602C78D7FB97FE14FB9F" box="[388,411,1117,1140]" italics="true" pageId="7" pageNumber="113">vs</emphasis>
. widely paired in
<taxonomicName id="A199DFE4FFE7602C7B3BFB97FD6BFB9F" box="[616,740,1116,1140]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="species" species="paolettii">
<emphasis id="54ED7875FFE7602C7B3BFB97FD6BFB9F" box="[616,740,1116,1140]" italics="true" pageId="7" pageNumber="113">R. paolettii</emphasis>
</taxonomicName>
) and in the shape of the spermathecae (anterior spermathecae with long duct and rounded ampulla, posterior spermatheca with oval ampulla
<emphasis id="54ED7875FFE7602C7D63FB4BFBC0FB73" box="[1072,1103,1153,1176]" italics="true" pageId="7" pageNumber="113">vs.</emphasis>
long duct and arrow-shaped ampulla). A comparison of
<taxonomicName id="A199DFE4FFE7602C789FFB6EFDD7FB57" box="[460,600,1188,1212]" class="Clitellata" family="Glossoscolecidae" genus="Righiodrilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="113" phylum="Annelida" rank="genus">
<emphasis id="54ED7875FFE7602C789FFB6EFDD7FB57" box="[460,600,1188,1212]" italics="true" pageId="7" pageNumber="113">Righiodrilus</emphasis>
</taxonomicName>
species with two pairs of spermathecae is given in
<tableCitation id="2B1B91DCFFE7602C7DC4FB6EFB62FB57" box="[1175,1261,1188,1213]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionTargetBox="[159,1375,202,687]" captionTargetPageId="7" captionText="TABLE 2. Comparison of species of the genus Righiodrilus with two pairs of spermathecae in segments 8 / 9 9 / 10." httpUri="http://table.plazi.org/id/32E6F4EFFFE7602C79C4FF52FAD1FF44" pageId="7" pageNumber="113" tableUuid="32E6F4EFFFE7602C79C4FF52FAD1FF44">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>