231 lines
24 KiB
XML
231 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1189.107506" ID-Pensoft-Pub="1313-2970-1189-83" ID-Pensoft-UUID="9CB018B9E4D3586DB83ED2D956E86983" ID-ZooBank="FF5E2B3943DB497EB546587BD91F794B" ModsDocID="1313-2970-1189-83" checkinTime="1705458508311" checkinUser="pensoft" docAuthor="Griebenow, Zachary" docDate="2024" docId="FF818E27A13E582C9003A6BC077EF6BC" docLanguage="en" docName="ZooKeys 1189: 83-184" docOrigin="ZooKeys 1189" docPubDate="2024-01-16" docSource="http://dx.doi.org/10.3897/zookeys.1189.107506" docTitle="Leptanilla Emery 1870" docType="treatment" docVersion="1" id="9CB018B9E4D3586DB83ED2D956E86983" lastPageNumber="83" masterDocId="9CB018B9E4D3586DB83ED2D956E86983" masterDocTitle="Systematic revision of the ant subfamily Leptanillinae (Hymenoptera, Formicidae)" masterLastPageNumber="184" masterPageNumber="83" pageNumber="83" updateTime="1705458508311" updateUser="pensoft">
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<mods:mods id="33C17AB83BD528C88B959398F0E264FC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="937F9692000CE68FB422234FB3FADEE5">
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<mods:title id="3B087ACF08547B4307ED99FD4E925023">Systematic revision of the ant subfamily Leptanillinae (Hymenoptera, Formicidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="4859F37FE17BFC2AF758B2D73B9FF913" type="personal">
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<mods:roleTerm id="9B7FEB61DCF18EF4049F52FA018E72DD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FD7C20F5B6ADD348E35BD898B0D57ECA">Griebenow, Zachary</mods:namePart>
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<mods:nameIdentifier id="9BBFD31BF9FC7518074DC66C60EA6FD1" type="ORCID">https://orcid.org/0000-0003-3385-8479</mods:nameIdentifier>
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<mods:affiliation id="C867E5C801151A6B6412A88AC03A5172">Department of Entomology & Nematology, University of California, Davis, CA USA & Department of Agricultural Biology, Colorado State University, Fort Collins, CO USA</mods:affiliation>
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<mods:nameIdentifier id="DE39C4A3DFA6728271E2A13A30D62101" type="email">zachary.griebenow@colostate.edu</mods:nameIdentifier>
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<mods:typeOfResource id="7B195E29C99EE44E672EE5B54A5A3C98">text</mods:typeOfResource>
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<mods:title id="2A07782DAA38B78FD39DB1FDCEBBF69D">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="BB04B3F57025A95C010D931824AFA7F6">
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<mods:date id="FEA0380A5C51A4990668D1323093BDE7">2024</mods:date>
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<mods:detail id="89E2E68A06BDF839C1E3C30433AA7999" type="pubDate">
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<mods:number id="C734CDC7AD7A1CC61DCD9DE969468590">2024-01-16</mods:number>
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</mods:detail>
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<mods:detail id="E48D389533D7909BB86FA3A9F2D9948C" type="volume">
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<mods:number id="7D84186439F716C38526F250D3330CA0">1189</mods:number>
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<mods:start id="DAC000237BC4418CB14E5D3BA0543B1D">83</mods:start>
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<mods:end id="433BE936CA3A90C214FEE38251BB473D">184</mods:end>
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<mods:url id="1D61F86D081286FD50A0FDA02F8F4A64">http://dx.doi.org/10.3897/zookeys.1189.107506</mods:url>
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</mods:location>
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<mods:classification id="08F862BE28570843EFD2C7538917DE62">journal article</mods:classification>
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<mods:identifier id="9F9BAB6DB7568855040A5AC20C009ADA" type="DOI">http://dx.doi.org/10.3897/zookeys.1189.107506</mods:identifier>
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<mods:identifier id="155E31C3EDC133FB3C311D6F9F2A0EC0" type="Pensoft-Pub">1313-2970-1189-83</mods:identifier>
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<mods:identifier id="FB7C4CEEFDAA04DDEDB18A2268C54F47" type="ZooBank">FF5E2B3943DB497EB546587BD91F794B</mods:identifier>
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</mods:mods>
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<treatment id="FF818E27A13E582C9003A6BC077EF6BC" LSID="urn:lsid:plazi:treatment:FF818E27A13E582C9003A6BC077EF6BC" httpUri="http://treatment.plazi.org/id/FF818E27A13E582C9003A6BC077EF6BC" lastPageNumber="83" pageId="0" pageNumber="83">
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<subSubSection id="91C943A00576B4817B3CD03806E89B90" pageId="0" pageNumber="83" type="nomenclature">
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<paragraph id="0395019F68D7FEC5FC9189D6DC61DD47" pageId="0" pageNumber="83">
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<taxonomicName id="109CC6B620C06C3F70B2B4DA59913A41" ID-CoL="JT8" ID-ENA="213868" LSID="FF818E27-A13E-582C-9003-A6BC077EF6BC" authority="Emery, 1910" authorityName="Emery" authorityYear="1870" family="Formicidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="83" rank="subFamily" subFamily="Leptanillinae">Leptanillinae Emery, 1910</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="84C6F3A7C28ECA6F539C87A94900EF76" pageId="0" pageNumber="83" type="type genus">
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<paragraph id="DB7BF13D7AFAB68733B14EEF4D3957CB" pageId="0" pageNumber="83">Type genus.</paragraph>
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<paragraph id="584B16B0644BF8F048F56CFF7AAD63FA" pageId="0" pageNumber="83">
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<taxonomicName id="2289501E1F6DB627DF6ED39BC0A6E2C8" authorityName="Emery" authorityYear="1870" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="00DA9F1CD314ED790CB6B36BC403349A" italics="true" pageId="0" pageNumber="83">Leptanilla</emphasis>
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</taxonomicName>
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Emery, 1870: 196.
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</paragraph>
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</subSubSection>
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<subSubSection id="B2C35027B3EE5F944BF47AFBDCA0EA7A" pageId="0" pageNumber="83" type="worker diagnosis">
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<paragraph id="D942847E07BE2515219D80FC2401B46A" pageId="0" pageNumber="83">Worker diagnosis</paragraph>
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<paragraph id="ACF6FAE1D6B5E294BDF2C66FFD415B94" pageId="0" pageNumber="83">
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(modified from
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<bibRefCitation id="C637C1687422BEEBEDD2FE1F1B8EA612" author="Bolton, B" journalOrPublisher="The American Entomological Institute, Gainesville" pageId="0" pageNumber="83" refId="B14" refString="Bolton, B, 2003. 71 Synopsis and Classification of the Formicidae. The American Entomological Institute, Gainesville" title="71 Synopsis and Classification of the Formicidae." year="2003">Bolton 2003</bibRefCitation>
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):
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</paragraph>
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<paragraph id="E6B034F08D3A5A86FB5677A3EED94853" pageId="0" pageNumber="83">Mandibles without differentiated basal and masticatory margins.</paragraph>
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<paragraph id="D30489BF4A0BC6BF26BB43E6A0BA8F93" pageId="0" pageNumber="83">At least one preapical tooth or lobe present on mandible.</paragraph>
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<paragraph id="E1097084E4DBD97747F55AB52014A239" pageId="0" pageNumber="83">Frontal lobes absent.</paragraph>
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<paragraph id="AB43265F8B931E1013BF1F82761D3FE9" pageId="0" pageNumber="83">Antennal sockets dorsal, fully exposed.</paragraph>
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<paragraph id="F0BBB7AA7F03E033395D70EC83BB9A47" pageId="0" pageNumber="83">
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Compound eyes absent, if present (
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<taxonomicName id="E56830F3AB144AD5580366F8986DC8ED" class="Insecta" family="Formicidae" genus="Protanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanilla izanagi" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="izanagi">
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<emphasis id="C65C9EBD4C0E3009505AA3AB727836C2" italics="true" pageId="0" pageNumber="83">Protanilla izanagi</emphasis>
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</taxonomicName>
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Terayama, 2013) then reduced to two ommatidia (Fig.
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<figureCitation id="85A76369257610B1031830C3683EB759" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Aspects of Protanilla izanagi, worker A profile view of posterior half of cranium B ventral view of the mandibles. Abbreviation: com = compound eye. Scale bars: 0.1 mm (A); 0.2 mm (B)." figureDoi="10.3897/zookeys.1189.107506.figure17" httpUri="https://binary.pensoft.net/fig/968317" pageId="0" pageNumber="83">17A</figureCitation>
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).
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</paragraph>
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<paragraph id="50227F04DFF1A723AA4F7D034B527E7F" pageId="0" pageNumber="83">Ocelli absent.</paragraph>
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<paragraph id="2AB283ECC7DF08921A612F7BB1BD19E8" pageId="0" pageNumber="83">Antenna 12-merous.</paragraph>
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<paragraph id="B5200DFECB11ECC1E79B5DEC6FFAF2C6" pageId="0" pageNumber="83">Promesonotal suture fully articulated.</paragraph>
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<paragraph id="1DE36B1CDCDA8E8B5A8843937436F3F5" pageId="0" pageNumber="83">Propodeal lobes weakly present (Opamyrmini) or absent (Leptanillini).</paragraph>
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<paragraph id="3573F51830E8166BD590EE137C370E27" pageId="0" pageNumber="83">Propodeal spiracle situated low on propodeum.</paragraph>
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<paragraph id="A775A74BEE806DD150256DDBD08146AD" pageId="0" pageNumber="83">
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Metacoxal foramen small, fully closed (Fig.
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<figureCitation id="DBDF16BEC2E7EEC347BA046EC6AD50F4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Metacoxal foramen of Leptanilla havilandi (CASENT 0010809), ventral view, worker. Scale bar: 0.05 mm." figureDoi="10.3897/zookeys.1189.107506.figure18" httpUri="https://binary.pensoft.net/fig/968318" pageId="0" pageNumber="83">18</figureCitation>
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).
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</paragraph>
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<paragraph id="803734F30213A66CA8D4E0695924E9FB" pageId="0" pageNumber="83">Suture absent from annulus surrounding metacoxal foramen.</paragraph>
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<paragraph id="BF826EB014EB4C7C5CE67418F6AF0C51" pageId="0" pageNumber="83">Metapleural gland present.</paragraph>
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<paragraph id="BD9EC960CCB9A4965BCC58FCC8909A62" pageId="0" pageNumber="83">Orifice of metapleural gland covered by dorsal cuticular flange.</paragraph>
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<paragraph id="010E9AEFD384BA195A12FF9E12154C18" pageId="0" pageNumber="83">Helcial sternite reduced and partly covered by corresponding tergite.</paragraph>
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<paragraph id="146F592AD9874A0E6F612077F8D9841F" pageId="0" pageNumber="83">Spiracle of abdominal segment III large and placed far forward.</paragraph>
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<paragraph id="CF2AC70B4FF8BA105ABCFFE7F91E3061" pageId="0" pageNumber="83">Spiracles of abdominal segments IV-VII concealed by posterior margins of preceding tergites.</paragraph>
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<paragraph id="6310778879C52E5B5C651C6F85FC0069" pageId="0" pageNumber="83">
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Petiole sessile, rarely subsessile (
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<taxonomicName id="B4816B72987FBB0550709FCCC3A479F4" authorityName="Parker, 1932: 342 (Non Teratolepis Taylori Parker" authorityYear="1942" class="Insecta" family="Formicidae" genus="Protanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanilla taylori" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="taylori">
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<emphasis id="82F636B75B11AC72F798CBABD9520BD0" italics="true" pageId="0" pageNumber="83">Protanilla taylori</emphasis>
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</taxonomicName>
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species group).
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</paragraph>
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<paragraph id="F0932B3F2E02708AC7BC6E69B83EEDC5" pageId="0" pageNumber="83">Abdominal postsclerites II with (Leptanillini) or without (Opamyrmini) complete tergosternal fusion.</paragraph>
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<paragraph id="910E1250327AA29D79BD704BC8C4BC6D" pageId="0" pageNumber="83">Abdominal postsclerites III with (Leptanillini) or without (Opamyrmini) tergosternal fusion.</paragraph>
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<paragraph id="979FCA4B48D06142484ED72F94AFCC6F" pageId="0" pageNumber="83">Abdominal segment III petiolate (Leptanillini) or not (Opamyrmini).</paragraph>
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<paragraph id="58C489E9163E9402EB0592A8F71BC37B" pageId="0" pageNumber="83">Abdominal segment IV without tergosternal fusion.</paragraph>
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<paragraph id="70D6A1CC852387D299CE9203DCA1BFF7" pageId="0" pageNumber="83">Stridulitrum absent from abdominal segment IV.</paragraph>
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<paragraph id="7EE6D371377966CE082711ADE7DBEF71" pageId="0" pageNumber="83">Abdominal tergite VII large, with simple posterior margin.</paragraph>
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<paragraph id="C86083C25DF9FD6CC1435CEEB6ED803B" pageId="0" pageNumber="83">Sting present.</paragraph>
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<paragraph id="7E45BFD84F84F64B27EF8A190CAD3FF2" pageId="0" pageNumber="83">Pretarsal claws edentate.</paragraph>
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</subSubSection>
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<subSubSection id="F2E983DB5CAAF118176084BA0463E800" pageId="0" pageNumber="83" type="gyne diagnosis">
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<paragraph id="2D85150D43593C20E44CEE70A35E705B" pageId="0" pageNumber="83">Gyne diagnosis.</paragraph>
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<paragraph id="EF2389555709838347EEFC7A8FC1C01D" pageId="0" pageNumber="83">As above, but alate or dichthadiiform (rarely ergatoid). If alate then with ocelli and pterostigma; hindwing with R + Rs and 1A tubular, not intersecting distal wing margin. If dichthadiiform then compound eyes reduced to one or two ommatidia, or absent; ocelli absent; mandibles sometimes edentate.</paragraph>
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<caption id="A76B683CC44F6C364FA79D9D7CBEF765" doi="10.3897/zookeys.1189.107506.figure17" httpUri="https://binary.pensoft.net/fig/968317" pageId="0" pageNumber="83" start="Figure 17" startId="F17">
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<paragraph id="6A8EC7F7E9FA7C3690D57D25703FBAA6" pageId="0" pageNumber="83">
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<emphasis id="C69ED4367E1F7495982FDDDB140A698D" bold="true" pageId="0" pageNumber="83">Figure 17.</emphasis>
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Aspects of
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<taxonomicName id="A408CEBD1427762D415952C1E64C6EDB" class="Insecta" family="Formicidae" genus="Protanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanilla izanagi" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="izanagi">
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<emphasis id="248EFEF561CAAD54D96ADD9FBF0FDC3A" italics="true" pageId="0" pageNumber="83">Protanilla izanagi</emphasis>
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</taxonomicName>
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, worker
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<emphasis id="B984F8EA9380B99598673D6FACB9136F" bold="true" pageId="0" pageNumber="83">A</emphasis>
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profile view of posterior half of cranium
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<emphasis id="17709466C11912D3A1D1E2FF4C9F128A" bold="true" pageId="0" pageNumber="83">B</emphasis>
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ventral view of the mandibles. Abbreviation: com = compound eye. Scale bars: 0.1 mm (
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<emphasis id="882B175A5BA03D865C9FE98862ED629E" bold="true" pageId="0" pageNumber="83">A</emphasis>
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); 0.2 mm (
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<emphasis id="5DF7EB00562E6EE2A6F8A07302D73F93" bold="true" pageId="0" pageNumber="83">B</emphasis>
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).
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</paragraph>
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</caption>
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<caption id="6DDBD7CF0337B671DB70F5869046FA65" doi="10.3897/zookeys.1189.107506.figure18" httpUri="https://binary.pensoft.net/fig/968318" pageId="0" pageNumber="83" start="Figure 18" startId="F18">
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<paragraph id="16E40D6F4378280FA70E3F79BA51A817" pageId="0" pageNumber="83">
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<emphasis id="9E5AEE340ACCCF92105C073E3E8EE99F" bold="true" pageId="0" pageNumber="83">Figure 18.</emphasis>
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Metacoxal foramen of
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<taxonomicName id="A64FBB4B169DD2C399E4E40835C821DE" authorityName="Forel" authorityYear="1901" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla havilandi" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="havilandi">
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<emphasis id="6C217E51713784655D8819CDFED2D810" italics="true" pageId="0" pageNumber="83">Leptanilla havilandi</emphasis>
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</taxonomicName>
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(CASENT0010809), ventral view, worker. Scale bar: 0.05 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="A29766462E46ED371D6C191D798CE23C" pageId="0" pageNumber="83" type="male diagnosis">
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<paragraph id="43E5256A3C07E41D03A8A3588D233C80" pageId="0" pageNumber="83">Male diagnosis</paragraph>
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<paragraph id="B8C200968526276C8AE747D81AC6F658" pageId="0" pageNumber="83">
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(modified from
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<bibRefCitation id="4012F007D11CFA44D4A313A4BB5F1FE8" DOI="https://doi.org/10.5852/ejt.2015.120" author="Boudinot, BE" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="83" pagination="1 - 62" refId="B20" refString="Boudinot, BE, 2015. Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): A new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages. European Journal of Taxonomy 120 (120): 1 - 62, DOI: https://doi.org/10.5852/ejt.2015.120" title="Contributions to the knowledge of Formicidae (Hymenoptera, Aculeata): A new diagnosis of the family, the first global male-based key to subfamilies, and a treatment of early branching lineages." url="https://doi.org/10.5852/ejt.2015.120" volume="120" year="2015">Boudinot 2015</bibRefCitation>
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):
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</paragraph>
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<paragraph id="419E00984A02857E2718E22841CF759B" pageId="0" pageNumber="83">
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Mandible edentate, nub-like or spatulate (
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<taxonomicName id="F128847B25D6B24B4A8E7D287B9047DC" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla anomala" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="anomala">
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<emphasis id="B90EA3795A8F2A169A362DD5432B039D" italics="true" pageId="0" pageNumber="83">Leptanilla anomala</emphasis>
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</taxonomicName>
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(Brues, 1925), comb. nov.).
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</paragraph>
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<paragraph id="C035A780AF614663746ED14F251F9B7B" pageId="0" pageNumber="83">Frontal carinae absent.</paragraph>
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<paragraph id="DF684BE30A14D8FDEE3BFFC1806FBB7B" pageId="0" pageNumber="83">Cuticular pegs absent from anterior clypeal margin.</paragraph>
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<paragraph id="165540D5DDC2D31B6942967AA8447F08" pageId="0" pageNumber="83">Antenna 13-merous.</paragraph>
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<paragraph id="5E26FA7746D486B90F25ED97D042A4CD" pageId="0" pageNumber="83">Funiculus filiform to submoniliform.</paragraph>
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<paragraph id="118C7879DD8D3B9DE10E033B416A36D5" pageId="0" pageNumber="83">Oblique mesopleural sulcus present or absent.</paragraph>
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<paragraph id="FF63D09546464236FC589D5F0EF36328" pageId="0" pageNumber="83">Metapleural spiracular plate absent.</paragraph>
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<paragraph id="0CBA4642EDC944E8435E8F52BE513452" pageId="0" pageNumber="83">Propodeal lobes inconspicuous or absent.</paragraph>
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<paragraph id="E46BFDC64EEBA4744AE0B891BE88D70D" pageId="0" pageNumber="83">Metacoxal foramen small, fully closed.</paragraph>
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<paragraph id="F8D241EEEE9AD76A569B8FE61F4DB9A4" pageId="0" pageNumber="83">Mesotibia with one or two spurs or none.</paragraph>
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<paragraph id="6A5E2512D41317CF21F3B32CD49618D3" pageId="0" pageNumber="83">Metatibia with one or two spurs.</paragraph>
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<paragraph id="3C04FF44A2F5255C2BDF0104F7987D32" pageId="0" pageNumber="83">Metatarsus lacking posterolateral line of dense differentiated setae.</paragraph>
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<paragraph id="9CCDECACA55486053F60375FE268436E" pageId="0" pageNumber="83">Pretarsal claws edentate.</paragraph>
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<paragraph id="F40FF88F041779A9FC3EEEB5B79B88E7" pageId="0" pageNumber="83">Pterostigma present or absent.</paragraph>
|
|
<paragraph id="CC001275243CC92835E20F42E3B6BD69" pageId="0" pageNumber="83">Rs+M absent (Leptanillini) or present, nebulous (Opamyrmini).</paragraph>
|
|
<paragraph id="ACB844689FC0B43218678AA46A44AA25" pageId="0" pageNumber="83">
|
|
<emphasis id="77D8E24F9956848A265839AE5CD9F514" italics="true" pageId="0" pageNumber="83">1m-cu</emphasis>
|
|
absent (Leptanillini) or present, nebulous (Opamyrmini).
|
|
</paragraph>
|
|
<paragraph id="DB5C93D13A2EB89B4BAB3B5BAE61076F" pageId="0" pageNumber="83">Jugal lobe absent.</paragraph>
|
|
<paragraph id="005A2B4701654A9874E1171EB183C4E7" pageId="0" pageNumber="83">Hindwing venation reduced, at most R+Rs and 1A tubular.</paragraph>
|
|
<paragraph id="B0AF35AC2CE7547C53713212F4F6B0C2" pageId="0" pageNumber="83">
|
|
Metapleural gland absent (Fig.
|
|
<figureCitation id="CCDE1092EBE934AC3AD8DBADDD00F515" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Metapleuron in male Leptanillinae A Leptanilla nr. indica (CASENT 0106381) B Leptanilla zhg-th 02 (CASENT 0842615). Abbreviation: mpl = metapleural gland orifice. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1189.107506.figure19" httpUri="https://binary.pensoft.net/fig/968319" pageId="0" pageNumber="83">19A</figureCitation>
|
|
) or rarely present (Fig.
|
|
<figureCitation id="119E1C823B36620F4BFE54134DA03FA9" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Metapleuron in male Leptanillinae A Leptanilla nr. indica (CASENT 0106381) B Leptanilla zhg-th 02 (CASENT 0842615). Abbreviation: mpl = metapleural gland orifice. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1189.107506.figure19" httpUri="https://binary.pensoft.net/fig/968319" pageId="0" pageNumber="83">19B</figureCitation>
|
|
) (e.g.,
|
|
<taxonomicName id="C8502398D57D2C4D33B384B449F7DBFE" authorityName="Emery" authorityYear="1870" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="DAE2535C43EB51D476BAB31FF4F48F5C" italics="true" pageId="0" pageNumber="83">Leptanilla</emphasis>
|
|
</taxonomicName>
|
|
zhg-th02).
|
|
</paragraph>
|
|
<paragraph id="3E4F106704425987E59F9CDFC0F93A67" pageId="0" pageNumber="83">
|
|
Petiole present or reduced to absent (
|
|
<taxonomicName id="50D60D5773FEE1642C809D337B70697B" authorityName="Baroni Urbani" authorityYear="1977" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla thai" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="thai">
|
|
<emphasis id="96793CAFD6BA0F595C617EFCE6B60EFA" italics="true" pageId="0" pageNumber="83">Leptanilla thai</emphasis>
|
|
</taxonomicName>
|
|
species group,
|
|
<taxonomicName id="7B9033CE305EF2D5B72C8E2F889593CD" authorityName="Forel" authorityYear="1901" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla havilandi" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="havilandi">
|
|
<emphasis id="6E6F59358D15E4D9B719BF40621E3B1C" italics="true" pageId="0" pageNumber="83">Leptanilla havilandi</emphasis>
|
|
</taxonomicName>
|
|
species group).
|
|
</paragraph>
|
|
<paragraph id="E080C1C0497498A0A8979BC5671FD051" pageId="0" pageNumber="83">Helcium axial or infra-axial.</paragraph>
|
|
<paragraph id="0E2441BEEF1C2FB1956AB1FF6BD989C8" pageId="0" pageNumber="83">
|
|
Abdominal segment III not petiolate, or rarely petiolate (
|
|
<taxonomicName id="532CAB1A4778A24C2F6C65E402280472" class="Insecta" family="Formicidae" genus="Protanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Protanilla bicolor" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="bicolor">
|
|
<emphasis id="C4F6D14B1434B582B7FC64C0B2A3445D" italics="true" pageId="0" pageNumber="83">Protanilla bicolor</emphasis>
|
|
</taxonomicName>
|
|
species group).
|
|
</paragraph>
|
|
<paragraph id="C48F7F92AF97CB633F8C4C2913BFE405" pageId="0" pageNumber="83">Abdominal segment IV not vaulted, as long as, or distinctly longer than posterad abdominal segments.</paragraph>
|
|
<paragraph id="9D3E8ED122401EF4CC3493297C56F700" pageId="0" pageNumber="83">Abdominal spiracles IV-VIII obscured by preceding tergites.</paragraph>
|
|
<paragraph id="0117387AC9E74AC2F8B57B5877659D6F" pageId="0" pageNumber="83">Posterior margin of abdominal sternite IX with posteromedian process, or entire, or emarginate, or with mulceators.</paragraph>
|
|
<paragraph id="67B6D3CD65D98AF2BA9B835262FE9FFA" pageId="0" pageNumber="83">Cerci absent.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F2E0F10EDAB9024FE4C6AFC62603D179" pageId="0" pageNumber="83" type="larval diagnosis">
|
|
<paragraph id="032E0912D5956E9D0DB189E0CCABEBBD" pageId="0" pageNumber="83">Larval diagnosis.</paragraph>
|
|
<paragraph id="87513E2F6A327182AC2312E819181ED3" pageId="0" pageNumber="83">Stenocephalous, with post-cranial soma moderately (i.e., habitus pogonomyrmecoid) to extremely (i.e., habitus leptanilloid) elongate. Mandibles typhlomyrmecoid or leptanilloid.</paragraph>
|
|
<caption id="B0E910AA8358994A91E5A2B0A43F127B" doi="10.3897/zookeys.1189.107506.figure19" httpUri="https://binary.pensoft.net/fig/968319" pageId="0" pageNumber="83" start="Figure 19" startId="F19">
|
|
<paragraph id="60044D86AB349972224E9F91E1C88AAB" pageId="0" pageNumber="83">
|
|
<emphasis id="FCCB190826FD88837D9FEC5E8636D86C" bold="true" pageId="0" pageNumber="83">Figure 19.</emphasis>
|
|
Metapleuron in male
|
|
<taxonomicName id="7A0FFFA74D9399256F35C7B01C233756" authorityName="Emery" authorityYear="1910" lsidName="" pageId="0" pageNumber="83" rank="subFamily" subFamily="Leptanillinae">Leptanillinae</taxonomicName>
|
|
<emphasis id="9E7D3EDF47740E337B8C00566F039310" bold="true" pageId="0" pageNumber="83">A</emphasis>
|
|
<taxonomicName id="56C2380BEAB9B8BC8E334553DB84FCBF" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla indica" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="indica">Leptanilla nr. indica</taxonomicName>
|
|
(CASENT0106381)
|
|
<emphasis id="3463653A505B6666B5FFF301E255A11B" bold="true" pageId="0" pageNumber="83">B</emphasis>
|
|
<taxonomicName id="40B36346A5ADFDD09B6982D88A1E9A58" authorityName="Emery" authorityYear="1870" class="Insecta" family="Formicidae" genus="Leptanilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptanilla" order="Hymenoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="5C9D6218996D2376A61C4626C8273CC8" italics="true" pageId="0" pageNumber="83">Leptanilla</emphasis>
|
|
</taxonomicName>
|
|
zhg-th02 (CASENT0842615). Abbreviation: mpl = metapleural gland orifice. Scale bars: 0.05 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |