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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1058.68628" ID-GBIF-Dataset="4477866e-bbd0-4938-86ed-bd708b00559c" ID-PMC="PMC8421324" ID-Pensoft-Pub="1313-2970-1058-1" ID-Pensoft-UUID="61FB9E86BEE45968AAC868B21AF7ED86" ID-PubMed="34557053" ID-ZooBank="654932353DDB4E1B8848EAB69F2C20FD" ModsDocID="1313-2970-1058-1" checkinTime="1630349293316" checkinUser="pensoft" docAuthor="Vagalinski, Boyan &amp; Golovatch, Sergei I." docDate="2021" docId="FA33AAF820C65286AECF2C84C63A72A7" docLanguage="en" docName="ZooKeys 1058: 1-127" docOrigin="ZooKeys 1058" docPubDate="2021-08-30" docSource="http://dx.doi.org/10.3897/zookeys.1058.68628" docTitle="Omobrachyiulus ponticus Vagalinski 2021, sp. nov." docType="treatment" docUuid="5F4E33FA-A9FF-445F-B33A-27BC59971AB5" docUuidSource="ZooBank" docVersion="5" id="61FB9E86BEE45968AAC868B21AF7ED86" lastPageNumber="1" masterDocId="61FB9E86BEE45968AAC868B21AF7ED86" masterDocTitle="The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)" masterLastPageNumber="127" masterPageNumber="1" pageNumber="1" updateTime="1668150880892" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vagalinski, Boyan</mods:namePart>
<mods:affiliation>Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier type="email">boyan_vagalinski@excite.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1058</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>127</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1058.68628</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1058.68628</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1058-1</mods:identifier>
<mods:identifier type="ZooBank">654932353DDB4E1B8848EAB69F2C20FD</mods:identifier>
<mods:identifier type="Pensoft-UUID">61FB9E86BEE45968AAC868B21AF7ED86</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7019454" ID-GBIF-Taxon="185371665" ID-Zenodo-Dep="7019454" LSID="urn:lsid:zoobank.org:act:5F4E33FA-A9FF-445F-B33A-27BC59971AB5" httpUri="http://treatment.plazi.org/id/FA33AAF820C65286AECF2C84C63A72A7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/5F4E33FA-A9FF-445F-B33A-27BC59971AB5" authority="Vagalinski" authorityName="Vagalinski" authorityYear="2021" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus" status="sp. nov.">Omobrachyiulus ponticus Vagalinski</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Omobrachyiulus ponticus sp. nov. A habitus of ♀ paratype, lateral, slightly dorsal view B telson of ♂ holotype, lateral view. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure41" httpUri="https://binary.pensoft.net/fig/581976" pageId="0" pageNumber="1">Figs 41</figureCitation>
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Omobrachyiulus ponticus sp. nov., ♂ holotype (A-C) and ♀ paratype (D, E) A leg pair 1, caudal view B left flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right vulva, lateral view (receptaculum seminis seen by transparency, but drawn with solid lines for emphasis; setae omitted) E same, caudal view. Scale bars: 0.2 mm (D), 0.1 mm (A-C, E). Abbreviations: ct central tube, op operculum, pa posterior ampulla, pt posterior tube." figureDoi="10.3897/zookeys.1058.68628.figure42" httpUri="https://binary.pensoft.net/fig/581977" pageId="0" pageNumber="1">, 42</figureCitation>
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">, 43</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(ZMUM).
<materialsCitation collectingDate="1983-04-08" collectingDateMax="1983-04-10" collectingDateMin="1983-04-08" country="Georgia" elevation="75" location="Myussera" municipality="Pitsunda-Myussera Nature Reserve" specimenCount="♂" stateProvince="Abkhazia" typeStatus="Holotype">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<specimenCount>♂</specimenCount>
(in head, collum to ring 3, ring 4 to ring 6, and rest of body; leg pair 1, penis, right leg 3, gonopods, left flange of pleurotergum 7, and a mid-body leg dissected) (ZMUM),
<collectingCountry name="Georgia">Georgia</collectingCountry>
, AR
<collectingRegion country="Georgia" name="Abkhazia">Abkhazia</collectingRegion>
,
<collectingMunicipality>Pitsunda-Myussera Nature Reserve</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:FA33AAF820C65286AECF2C84C63A72A7:791520E6A6A946E3A2E7CE569F686A9A" country="Georgia" municipality="Pitsunda-Myussera Nature Reserve" name="Myussera" stateProvince="Abkhazia">Myussera</location>
part,
<quantity metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="13.0" metricValueMin="2.0" unit="m" value="75.0" valueMax="130.0" valueMin="20.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="13.0" metricValueMin="2.0" unit="m" value="75.0" valueMax="130.0" valueMin="20.0">20-130 m</elevation>
</quantity>
, mixed deciduous forest (
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Castanea" kingdom="Plantae" lsidName="Castanea" order="Fagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Castanea</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" lsidName="Alnus" order="Fagales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Alnus</emphasis>
</taxonomicName>
, etc.), litter, under bark and stones,
<collectingDate value="1983-04-08" valueMax="1983-04-10" valueMin="1983-04-08">8-10.IV.1983</collectingDate>
, SIG leg
</materialsCitation>
</emphasis>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<specimenCount type="female">2 ♀♀</specimenCount>
(one unbroken, the other in head to ring 3 and rest of body, right vulva dissected), same collecting data as of the
<typeStatus>holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A species of
<taxonomicName authorityName="Lohmander" authorityYear="1936" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus</emphasis>
</taxonomicName>
most similar to
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. sevangensis" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sevangensis">
<emphasis italics="true" pageId="0" pageNumber="1">O. sevangensis</emphasis>
</taxonomicName>
(Lohmander, 1936) comb. nov.,
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. kvavadzei" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kvavadzei">
<emphasis italics="true" pageId="0" pageNumber="1">O. kvavadzei</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. trochiloides" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="trochiloides">
<emphasis italics="true" pageId="0" pageNumber="1">O. trochiloides</emphasis>
</taxonomicName>
sp. nov.; sharing with O.
<taxonomicName authorityName="Vagalinski &amp; Golovatch" authorityYear="2021" baseAuthorityName="Lohmander" baseAuthorityYear="1932" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus sevangensis" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sevangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus sevangensis</emphasis>
</taxonomicName>
and
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. trochiloides" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="trochiloides">
<emphasis italics="true" pageId="0" pageNumber="1">O. trochiloides</emphasis>
</taxonomicName>
sp. nov. the presence of a lamellar ridge-like anterior process of the opisthomere. Differs from the former species by the presence of a distinct, rounded, apicolateral part of the mesomeroidal lobe of the opisthomere, as well as by the solenomere considerably outreaching both anterior process and apical outgrowth of the basoposterior process, and from the latter species by the much more obscure, poorly differentiated basoposterior process ending in an unciform, rather than a fine rod-like, apical outgrowth. Resembling
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. kvavadzei" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kvavadzei">
<emphasis italics="true" pageId="0" pageNumber="1">O. kvavadzei</emphasis>
</taxonomicName>
sp. nov. mostly by the promere having a distomesal lobe and by the mesomeroidal lobe of the opisthomere bearing an apicolateral part, but differing in other gonopod characters included in the tabular key to the
<taxonomicName authorityName="Vagalinski &amp; Golovatch" authorityYear="2021" baseAuthorityName="Lohmander" baseAuthorityYear="1932" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus sevangensis" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sevangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus sevangensis</emphasis>
</taxonomicName>
group (Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Key to the species of the Omobrachyiulus sevangensis group." httpUri="http://table.plazi.org/id/5DDD582B315D587F9E11FAC806897695" pageId="0" pageNumber="1" tableUuid="5DDD582B315D587F9E11FAC806897695">2</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="5DDD582B315D587F9E11FAC806897695" httpUri="http://table.plazi.org/id/5DDD582B315D587F9E11FAC806897695" pageId="0" pageNumber="1" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 2.</emphasis>
Key to the species of the
<taxonomicName authorityName="Vagalinski &amp; Golovatch" authorityYear="2021" baseAuthorityName="Lohmander" baseAuthorityYear="1932" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus sevangensis" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sevangensis">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus sevangensis</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Character</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. sevangensis" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="sevangensis">
<emphasis italics="true" pageId="0" pageNumber="1">O. sevangensis</emphasis>
</taxonomicName>
comb. nov.
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. kvavadzei" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kvavadzei">
<emphasis italics="true" pageId="0" pageNumber="1">O. kvavadzei</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis italics="true" pageId="0" pageNumber="1">O. ponticus</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. trochiloides" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="trochiloides">
<emphasis italics="true" pageId="0" pageNumber="1">O. trochiloides</emphasis>
</taxonomicName>
sp. nov.
</th>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">apical outgrowth of basoposterior process</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">rather slender, distally tapering, pointing disto-frontad</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
stout, distally tapering, apex abruptly bent frontad (Figs
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Omobrachyiulus kvavadzei sp. nov., ♂ (A-F) and ♀ (H) paratypes from near mouth of Khekpara River, AR Ajara, Georgia, and ♂ paratype from Zeraboseli, AR Ajara, Georgia (ZMUM) (G) A hypoproct, ventral view B leg 7 C right flange of pleurotergum 7, ventro-lateral view D penis, caudal view E right gonopods, mesal view F right opisthomere, caudo-lateral view G distal part of right opisthomere, mesal view H right vulva, caudo-lateral view. Scale bars: 0.1 mm (A), 0.2 mm (B-H). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ct central tube, dt distal tip of anterior process f flagellum, op operculum, pa posterior ampulla, pt posterior tube s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure39" httpUri="https://binary.pensoft.net/fig/581974" pageId="0" pageNumber="1">39G-F</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Omobrachyiulus kvavadzei sp. nov., gonopods of ♂ paratype from near Chakhati, AR Ajara, Georgia (ZMUM) A left promere, caudal view (flagellum broken off near base) B right opisthomere (with flagellum in its channel), mesal view C left opisthomere, latero-caudal view D distal part of the same aspect E distal part of right opisthomere (with flagellum tip protruding out of its channel), mesal view. Scale bars: 0.1 mm (A-C), 0.05 mm (D, E). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, dt distal tip of anterior process, f flagellum, fc flagellum channel, g (supposed) gonocoxal gland, mg median groove, ml mesomeroidal lobe, mr median ridge, s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure40" httpUri="https://binary.pensoft.net/fig/581975" pageId="0" pageNumber="1">40B-E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
stout, distally tapering, pointing disto-frontad (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43A, C-E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
very fine and slender, pointing disto-frontad (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Omobrachyiulus trochiloides sp. nov., gonopods of ♂ paratypes from Baniskhevi Valley, Georgia (ZMUM) (A-D) and from near Nabeglavi, Georgia (NMNHS) (E) A left promere (flagellum broken off), caudal view B right opisthomere, mesal view C distal part of the same aspect D right opisthomere, lateral, somewhat caudal view E distal part of right opisthomere, lateral view. Scale bars: 0.2 mm (D), 0.1 mm (A, B, E), 0.05 mm (C). Abbreviations: ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure46" httpUri="https://binary.pensoft.net/fig/581981" pageId="0" pageNumber="1">46B-E</figureCitation>
)
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">mesomeroidal lobe</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">weakly pronounced, without apicolateral part</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
rather weakly pronounced, with an apicolateral part pointing frontad (Figs
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Omobrachyiulus kvavadzei sp. nov., ♂ (A-F) and ♀ (H) paratypes from near mouth of Khekpara River, AR Ajara, Georgia, and ♂ paratype from Zeraboseli, AR Ajara, Georgia (ZMUM) (G) A hypoproct, ventral view B leg 7 C right flange of pleurotergum 7, ventro-lateral view D penis, caudal view E right gonopods, mesal view F right opisthomere, caudo-lateral view G distal part of right opisthomere, mesal view H right vulva, caudo-lateral view. Scale bars: 0.1 mm (A), 0.2 mm (B-H). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ct central tube, dt distal tip of anterior process f flagellum, op operculum, pa posterior ampulla, pt posterior tube s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure39" httpUri="https://binary.pensoft.net/fig/581974" pageId="0" pageNumber="1">39G</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Omobrachyiulus kvavadzei sp. nov., gonopods of ♂ paratype from near Chakhati, AR Ajara, Georgia (ZMUM) A left promere, caudal view (flagellum broken off near base) B right opisthomere (with flagellum in its channel), mesal view C left opisthomere, latero-caudal view D distal part of the same aspect E distal part of right opisthomere (with flagellum tip protruding out of its channel), mesal view. Scale bars: 0.1 mm (A-C), 0.05 mm (D, E). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, dt distal tip of anterior process, f flagellum, fc flagellum channel, g (supposed) gonocoxal gland, mg median groove, ml mesomeroidal lobe, mr median ridge, s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure40" httpUri="https://binary.pensoft.net/fig/581975" pageId="0" pageNumber="1">40B-E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
moderately pronounced, with an apicolateral part pointing frontad (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43C-E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
strongly pronounced, without apicolateral part (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Omobrachyiulus trochiloides sp. nov., gonopods of ♂ paratypes from Baniskhevi Valley, Georgia (ZMUM) (A-D) and from near Nabeglavi, Georgia (NMNHS) (E) A left promere (flagellum broken off), caudal view B right opisthomere, mesal view C distal part of the same aspect D right opisthomere, lateral, somewhat caudal view E distal part of right opisthomere, lateral view. Scale bars: 0.2 mm (D), 0.1 mm (A, B, E), 0.05 mm (C). Abbreviations: ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure46" httpUri="https://binary.pensoft.net/fig/581981" pageId="0" pageNumber="1">46B</figureCitation>
)
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">anterior process</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">well-developed lamellar ridge without free apical part</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
distally deeply bifurcated, with only a vestigial lamella (Figs
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Omobrachyiulus kvavadzei sp. nov., ♂ (A-F) and ♀ (H) paratypes from near mouth of Khekpara River, AR Ajara, Georgia, and ♂ paratype from Zeraboseli, AR Ajara, Georgia (ZMUM) (G) A hypoproct, ventral view B leg 7 C right flange of pleurotergum 7, ventro-lateral view D penis, caudal view E right gonopods, mesal view F right opisthomere, caudo-lateral view G distal part of right opisthomere, mesal view H right vulva, caudo-lateral view. Scale bars: 0.1 mm (A), 0.2 mm (B-H). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ct central tube, dt distal tip of anterior process f flagellum, op operculum, pa posterior ampulla, pt posterior tube s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure39" httpUri="https://binary.pensoft.net/fig/581974" pageId="0" pageNumber="1">39G</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Omobrachyiulus kvavadzei sp. nov., gonopods of ♂ paratype from near Chakhati, AR Ajara, Georgia (ZMUM) A left promere, caudal view (flagellum broken off near base) B right opisthomere (with flagellum in its channel), mesal view C left opisthomere, latero-caudal view D distal part of the same aspect E distal part of right opisthomere (with flagellum tip protruding out of its channel), mesal view. Scale bars: 0.1 mm (A-C), 0.05 mm (D, E). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, dt distal tip of anterior process, f flagellum, fc flagellum channel, g (supposed) gonocoxal gland, mg median groove, ml mesomeroidal lobe, mr median ridge, s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure40" httpUri="https://binary.pensoft.net/fig/581975" pageId="0" pageNumber="1">40B, E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
well-developed lamellar ridge with free apical part (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43C-E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
well-developed lamellar ridge without free apical part (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Omobrachyiulus trochiloides sp. nov., gonopods of ♂ paratypes from Baniskhevi Valley, Georgia (ZMUM) (A-D) and from near Nabeglavi, Georgia (NMNHS) (E) A left promere (flagellum broken off), caudal view B right opisthomere, mesal view C distal part of the same aspect D right opisthomere, lateral, somewhat caudal view E distal part of right opisthomere, lateral view. Scale bars: 0.2 mm (D), 0.1 mm (A, B, E), 0.05 mm (C). Abbreviations: ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure46" httpUri="https://binary.pensoft.net/fig/581981" pageId="0" pageNumber="1">46B-E</figureCitation>
)
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">solenomere</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">rather stout, unipartite, apically shortly bifurcate</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
rather stout, unipartite, apically hollow (Figs
<figureCitation captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. Omobrachyiulus kvavadzei sp. nov., ♂ (A-F) and ♀ (H) paratypes from near mouth of Khekpara River, AR Ajara, Georgia, and ♂ paratype from Zeraboseli, AR Ajara, Georgia (ZMUM) (G) A hypoproct, ventral view B leg 7 C right flange of pleurotergum 7, ventro-lateral view D penis, caudal view E right gonopods, mesal view F right opisthomere, caudo-lateral view G distal part of right opisthomere, mesal view H right vulva, caudo-lateral view. Scale bars: 0.1 mm (A), 0.2 mm (B-H). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ct central tube, dt distal tip of anterior process f flagellum, op operculum, pa posterior ampulla, pt posterior tube s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure39" httpUri="https://binary.pensoft.net/fig/581974" pageId="0" pageNumber="1">39G, F</figureCitation>
,
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Omobrachyiulus kvavadzei sp. nov., gonopods of ♂ paratype from near Chakhati, AR Ajara, Georgia (ZMUM) A left promere, caudal view (flagellum broken off near base) B right opisthomere (with flagellum in its channel), mesal view C left opisthomere, latero-caudal view D distal part of the same aspect E distal part of right opisthomere (with flagellum tip protruding out of its channel), mesal view. Scale bars: 0.1 mm (A-C), 0.05 mm (D, E). Abbreviations: al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, dt distal tip of anterior process, f flagellum, fc flagellum channel, g (supposed) gonocoxal gland, mg median groove, ml mesomeroidal lobe, mr median ridge, s solenomere, sf spiniform filaments, si anteromesal sinus, tp proximal tip of anterior process." figureDoi="10.3897/zookeys.1058.68628.figure40" httpUri="https://binary.pensoft.net/fig/581975" pageId="0" pageNumber="1">40C, D</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
slender, bipartite (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43C, E</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
slender, unipartite (Fig.
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Omobrachyiulus trochiloides sp. nov., gonopods of ♂ paratypes from Baniskhevi Valley, Georgia (ZMUM) (A-D) and from near Nabeglavi, Georgia (NMNHS) (E) A left promere (flagellum broken off), caudal view B right opisthomere, mesal view C distal part of the same aspect D right opisthomere, lateral, somewhat caudal view E distal part of right opisthomere, lateral view. Scale bars: 0.2 mm (D), 0.1 mm (A, B, E), 0.05 mm (C). Abbreviations: ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure46" httpUri="https://binary.pensoft.net/fig/581981" pageId="0" pageNumber="1">46B-E</figureCitation>
)
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="name">
<paragraph pageId="0" pageNumber="1">Name.</paragraph>
<paragraph pageId="0" pageNumber="1">After the Greek name of the Black Sea, Pontos Euxeinos, to emphasise the type locality of the new species at the eastern coast of that sea. Adjective.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
:
</emphasis>
holotype ♂ in S VII, 35+1+T, L = 10 mm, H = 0.9 mm; paratype ♀♀ in S VII-VIII, 34-35+2+T, L = 11.5-12 mm, H = 1.2-1.35 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
(Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Omobrachyiulus ponticus sp. nov. A habitus of ♀ paratype, lateral, slightly dorsal view B telson of ♂ holotype, lateral view. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure41" httpUri="https://binary.pensoft.net/fig/581976" pageId="0" pageNumber="1">41A</figureCitation>
): Head and collum with the usual colour pattern; body gradually lighter from dorsal to ventral side; prozonae grey, dorsally with a transverse dark brown stripe near pro-metazonal suture; metazonae light brown-beige; dorsum with a continuous whitish axial line.
</paragraph>
<caption doi="10.3897/zookeys.1058.68628.figure41" httpUri="https://binary.pensoft.net/fig/581976" pageId="0" pageNumber="1" start="Figure 41" startId="F41">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 41.</emphasis>
<taxonomicName authorityName="Vagalinski" authorityYear="2021" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus ponticus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus of ♀ paratype, lateral, slightly dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
telson of ♂ holotype, lateral view. Not to scale.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">External structures</emphasis>
:
</emphasis>
Eye patches in adults consisting of 20-23 relatively large, clearly convex ommatidia arranged in mostly easily recognisable vertical rows. Vertigial, supralabral, and labral setae: two, four, and 17, respectively. Antennae ca. 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as head in males and 1.1-1.15
<normalizedToken originalValue="×">x</normalizedToken>
in females; antennomere 2&gt; 5&gt; 4 ≥ 3&gt; 6. Gnathochilarium with a conspicuously large promentum separating lamellae linguales&gt; halfway, the latter with three or four setae in a longitudinal row; stipites in males parabasally each with a longitudinal row of several setae, non-setose in females. Collum smooth, with a single deep groove running along entire lateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Body rings somewhat vaulted. Prozonae with scattered, very short and shallow, longitudinal grooves. Metazonae rather deeply striated, n
<emphasis italics="true" pageId="0" pageNumber="1">Schub</emphasis>
= 6 or 7; metazonal setae relatively short, ca. 2/5 of (in more anterior rings), to equal to (in more posterior rings), metazonal length, arranged in very dense whorls. Ozopores placed right in pro-metazonal suture in more anterior rings, and tightly behind suture in more posterior ones; sutures not being sinuous in front of ozopores. Tarsus of mid-body legs 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as tibia and 3.8
<normalizedToken originalValue="×">x</normalizedToken>
as long as apical claw.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Telson</emphasis>
(Fig.
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Omobrachyiulus ponticus sp. nov. A habitus of ♀ paratype, lateral, slightly dorsal view B telson of ♂ holotype, lateral view. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure41" httpUri="https://binary.pensoft.net/fig/581976" pageId="0" pageNumber="1">41B</figureCitation>
): Epiproct very long (considerably surpassing the longest paraproctal setae in males, somewhat less pronounced in females), straight, distally bent ventrad, ending with a pointed hyaline tip; with a few setae. Hypoproct in males narrowly trapezoidal, tridentate, the teeth blunt, protruding behind rear contour of paraprocts; the same being narrowly rounded in females. Paraprocts rather sparsely setose, without distinct row of shorter setae along caudal margins.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Male sexual characters</emphasis>
:
</emphasis>
Mandibular stipites rather weakly expanded, forming a rounded ventro-anterior corner. Leg pair 1 (Fig.
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Omobrachyiulus ponticus sp. nov., ♂ holotype (A-C) and ♀ paratype (D, E) A leg pair 1, caudal view B left flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right vulva, lateral view (receptaculum seminis seen by transparency, but drawn with solid lines for emphasis; setae omitted) E same, caudal view. Scale bars: 0.2 mm (D), 0.1 mm (A-C, E). Abbreviations: ct central tube, op operculum, pa posterior ampulla, pt posterior tube." figureDoi="10.3897/zookeys.1058.68628.figure42" httpUri="https://binary.pensoft.net/fig/581977" pageId="0" pageNumber="1">42A</figureCitation>
) considerably converging (turned against one another) hooks with slender tibial outgrowths. Leg pair 2 and following pairs with crested adhesive pads, both tibial and postfemoral ones gradually reduced towards telson, but still visible after mid-body; femora of pairs 3-7 with an oval groove in the centre of a bulge. Pleurotergum 7 ventrally forming broadly rounded lobes (Fig.
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Omobrachyiulus ponticus sp. nov., ♂ holotype (A-C) and ♀ paratype (D, E) A leg pair 1, caudal view B left flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right vulva, lateral view (receptaculum seminis seen by transparency, but drawn with solid lines for emphasis; setae omitted) E same, caudal view. Scale bars: 0.2 mm (D), 0.1 mm (A-C, E). Abbreviations: ct central tube, op operculum, pa posterior ampulla, pt posterior tube." figureDoi="10.3897/zookeys.1058.68628.figure42" httpUri="https://binary.pensoft.net/fig/581977" pageId="0" pageNumber="1">42B</figureCitation>
) originating from the zone around pro-metazonal suture, protruding mesoventrad behind gonopods. Penis (42C) very small, subquadrate, without apical lobes and with very short, rounded, terminal lamellae directed distolaterad.
</paragraph>
<caption doi="10.3897/zookeys.1058.68628.figure42" httpUri="https://binary.pensoft.net/fig/581977" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
<taxonomicName authorityName="Vagalinski" authorityYear="2021" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus ponticus</emphasis>
</taxonomicName>
sp. nov., ♂ holotype (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
) and ♀ paratype (
<emphasis bold="true" pageId="0" pageNumber="1">D, E</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
leg pair 1, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
left flange of pleurotergum 7, ventro-lateral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
penis, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
right vulva, lateral view (receptaculum seminis seen by transparency, but drawn with solid lines for emphasis; setae omitted)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
same, caudal view. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
), 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A-C, E</emphasis>
). Abbreviations:
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ct</emphasis>
central tube,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">op</emphasis>
operculum,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">pa</emphasis>
posterior ampulla,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">pt</emphasis>
posterior tube.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43</figureCitation>
): Promere (Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43B, p</figureCitation>
in Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Omobrachyiulus ponticus sp. nov., gonopods of ♂ holotype A left gonopods, mesal view B right promere, caudal view C right opisthomere, mesal view D same, caudo-lateral view E distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations: abs anterior branch of solenomere, al apicolateral part of mesomeroidal lobe, ao apical outgrowth of basoposterior process, ap anterior process, bpp basoposterior process, dml distomesal lobe, f flagellum, fc flagellum channel, ml mesomeroidal lobe, mr median ridge, p promere, pbs posterior branch of solenomere, s solenomere." figureDoi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1">43A</figureCitation>
) slightly exceeded in height by solenomere, slender, with nearly symmetrical mesal and lateral margins joining in a narrow, blunt apex; median ridge short and strongly pronounced; a well-developed distomesal lobe. Opisthomere slender; basoposterior process weakly pronounced, ending in a tapering apical outgrowth turned slightly anteriad; anterior process shaped as a micro-serrate lamellar ridge with a freely protruding, tapering, apical part; mesomeroidal lobe with a moderately pronounced basal part, flattened distally and ending in an apicolateral part being very similar in shape to that in
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. kvavadzei" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kvavadzei">
<emphasis italics="true" pageId="0" pageNumber="1">O. kvavadzei</emphasis>
</taxonomicName>
sp. nov.; solenomere deeply divided into two fine, slender, parallel branches: an anterior and a posterior one, the latter being slightly longer than the former; spiniform filaments along flagellum channel either absent or too small and slanting to be detected by light microscopy.
</paragraph>
<caption doi="10.3897/zookeys.1058.68628.figure43" httpUri="https://binary.pensoft.net/fig/581978" pageId="0" pageNumber="1" start="Figure 43" startId="F43">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 43.</emphasis>
<taxonomicName authorityName="Vagalinski" authorityYear="2021" class="Diplopoda" family="Julidae" genus="Omobrachyiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omobrachyiulus ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis italics="true" pageId="0" pageNumber="1">Omobrachyiulus ponticus</emphasis>
</taxonomicName>
sp. nov., gonopods of ♂ holotype
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
left gonopods, mesal view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
right promere, caudal view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
right opisthomere, mesal view
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
same, caudo-lateral view
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
distal part of the same aspect. Scale bars: 0.1 mm. Abbreviations:
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">abs</emphasis>
anterior branch of solenomere,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">al</emphasis>
apicolateral part of mesomeroidal lobe,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ao</emphasis>
apical outgrowth of basoposterior process,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ap</emphasis>
anterior process,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">bpp</emphasis>
basoposterior process,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">dml</emphasis>
distomesal lobe,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">f</emphasis>
flagellum,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">fc</emphasis>
flagellum channel,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ml</emphasis>
mesomeroidal lobe,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">mr</emphasis>
median ridge,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">p</emphasis>
promere,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">pbs</emphasis>
posterior branch of solenomere,
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">s</emphasis>
solenomere.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Female sexual characters</emphasis>
:
</emphasis>
Leg pairs 1 and 2 somewhat shorter, first also thicker, than following legs. Vulva (Fig.
<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Omobrachyiulus ponticus sp. nov., ♂ holotype (A-C) and ♀ paratype (D, E) A leg pair 1, caudal view B left flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right vulva, lateral view (receptaculum seminis seen by transparency, but drawn with solid lines for emphasis; setae omitted) E same, caudal view. Scale bars: 0.2 mm (D), 0.1 mm (A-C, E). Abbreviations: ct central tube, op operculum, pa posterior ampulla, pt posterior tube." figureDoi="10.3897/zookeys.1058.68628.figure42" httpUri="https://binary.pensoft.net/fig/581977" pageId="0" pageNumber="1">42D, E</figureCitation>
) rather slender, similarly shaped to that of
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. kvavadzei" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kvavadzei">
<emphasis italics="true" pageId="0" pageNumber="1">O. kvavadzei</emphasis>
</taxonomicName>
sp. nov., the main difference being the less strongly slanting apex of bursa, and the smaller, apically positioned opening in
<taxonomicName class="Diplopoda" kingdom="Animalia" lsidName="O. ponticus" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ponticus">
<emphasis italics="true" pageId="0" pageNumber="1">O. ponticus</emphasis>
</taxonomicName>
sp. nov.; both bursa and operculum moderately densely covered with long setae. Receptaculum seminis consisting of a short finger-shaped central tube, and a long, spirally twisted, posterior tube ending in an ovoid posterior ampulla with a minute pocket at bottom.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">General distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">SWGC.</paragraph>
</subSubSection>
</treatment>
</document>
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