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<document id="06C0F836B62418DAEB7E7B7E4DD2D5E5" ID-CLB-Dataset="25807" ID-DOI="10.11646/zootaxa.4664.2.11" ID-GBIF-Dataset="192c8351-9f23-4350-ae38-75405264b53e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3384502" ID-ZooBank="ADA841C0-AF1F-4539-9C52-56C8347C3514" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567495487738" checkinUser="plazi" docAuthor="Balasubramanian, Chelliah, Muthukatturaja, Marimuthu &amp; Anbalagan, Sankarappan" docDate="2019" docId="F963592BFFB2FFEC7CEFFEB1E94A8655" docLanguage="en" docName="zootaxa.4664.2.11.pdf" docOrigin="Zootaxa 4664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhoenanthus (Rhoenanthus) tungaiensis Balasubramanian &amp; Muthukatturaja &amp; Anbalagan 2019, new species" docType="treatment" docVersion="7" lastPageNumber="299" masterDocId="055A2153FFB3FFEA7C78FFC0EC27850A" masterDocTitle="A new mayfly species of Rhoenanthus (Ephemeroptera: Potamanthidae) from Peninsular India" masterLastPageNumber="300" masterPageNumber="293" pageNumber="294" updateTime="1698754273683" updateUser="ExternalLinkService">
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<mods:title id="484C819388DE017709ECD472319C7BEE">A new mayfly species of Rhoenanthus (Ephemeroptera: Potamanthidae) from Peninsular India</mods:title>
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<mods:namePart id="9954C91E20E51D72082543A8F340EBD9">Balasubramanian, Chelliah</mods:namePart>
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<mods:namePart id="E1785C63B139C0C3E90F08F293719B9C">Muthukatturaja, Marimuthu</mods:namePart>
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<mods:namePart id="E292F22544134EDC76B1A368D8A011F2">Anbalagan, Sankarappan</mods:namePart>
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<mods:date id="01C2DD3C7773F2ACC6BF9DD4C4752368">2019</mods:date>
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<treatment id="F963592BFFB2FFEC7CEFFEB1E94A8655" ID-GBIF-Taxon="159359151" LSID="urn:lsid:plazi:treatment:F963592BFFB2FFEC7CEFFEB1E94A8655" httpUri="http://treatment.plazi.org/id/F963592BFFB2FFEC7CEFFEB1E94A8655" lastPageId="6" lastPageNumber="299" pageId="1" pageNumber="294">
<subSubSection id="39D0BBB6FFB2FFEB7CEFFEB1E93C8481" box="[151,1307,369,396]" pageId="1" pageNumber="294" type="nomenclature">
<paragraph id="7175E83DFFB2FFEB7CEFFEB1E93C8481" blockId="1.[151,1307,369,431]" box="[151,1307,369,396]" pageId="1" pageNumber="294">
<heading id="2A3D5F51FFB2FFEB7CEFFEB1E93C8481" bold="true" box="[151,1307,369,396]" fontSize="11" level="1" pageId="1" pageNumber="294" reason="1">
<emphasis id="43BE342FFFB2FFEB7CEFFEB1E93C8481" bold="true" box="[151,1307,369,396]" pageId="1" pageNumber="294">
<taxonomicName id="B6CA93BEFFB2FFEB7CEFFEB1E8598481" authority="Balasubramanian and Muthukatturaja" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[151,1150,369,396]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="tungaiensis" status="sp. nov." subGenus="Rhoenanthus">
<emphasis id="43BE342FFFB2FFEB7CEFFEB1ED1D8486" bold="true" box="[151,314,369,396]" italics="true" pageId="1" pageNumber="294">Rhoenanthus</emphasis>
(
<emphasis id="43BE342FFFB2FFEB7D33FEB1EDC98486" bold="true" box="[331,494,369,396]" italics="true" pageId="1" pageNumber="294">Rhoenanthus</emphasis>
)
<emphasis id="43BE342FFFB2FFEB7D87FEB1EEAD8481" bold="true" box="[511,650,369,395]" italics="true" pageId="1" pageNumber="294">tungaiensis</emphasis>
Balasubramanian and Muthukatturaja
</taxonomicName>
,
<taxonomicNameLabel id="588D8954FFB2FFEB78F3FEB1E93C8481" box="[1163,1307,369,395]" pageId="1" pageNumber="294" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="39D0BBB6FFB2FFEB7CEFFE55ED0684A5" box="[151,289,405,431]" pageId="1" pageNumber="294" type="description">
<paragraph id="7175E83DFFB2FFEB7CEFFE55ED0684A5" blockId="1.[151,1307,369,431]" box="[151,289,405,431]" pageId="1" pageNumber="294">
(
<figureCitation id="E9F1F4B8FFB2FFEB7CD8FE55ED3F84A5" box="[160,280,405,431]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 611" captionStart-2="FIGURE 12" captionStartId-0="1.[151,264,1965,1990]" captionStartId-1="2.[151,264,1783,1808]" captionStartId-2="3.[151,250,1945,1970]" captionTargetBox-0="[151,1436,453,1940]" captionTargetBox-1="[168,1413,182,1757]" captionTargetBox-2="[473,1103,181,1905]" captionTargetId-0="figure@1.[151,1436,453,1940]" captionTargetId-1="figure@2.[162,1426,181,1757]" captionTargetId-2="figure@3.[473,1114,181,1920]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." captionText-1="FIGURES 611. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 6, Foreleg; 7, Midleg; 8, Hindleg; 9, Gill I; 10, Gill III; 11, Gill VII." captionText-2="FIGURE 12. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov., dorsal view" httpUri-0="https://zenodo.org/record/3384504/files/figure.png" httpUri-1="https://zenodo.org/record/3384506/files/figure.png" httpUri-2="https://zenodo.org/record/3384508/files/figure.png" pageId="1" pageNumber="294">Figs.1 12</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="25B5B8B5FFB2FFEB7CEFF86DE8F982E0" httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="1" pageNumber="294" startId="1.[151,264,1965,1990]" targetBox="[151,1436,453,1940]" targetPageId="1">
<paragraph id="7175E83DFFB2FFEB7CEFF86DE8F982E0" blockId="1.[151,1436,1965,2026]" pageId="1" pageNumber="294">
<emphasis id="43BE342FFFB2FFEB7CEFF86DED1C82CC" bold="true" box="[151,315,1965,1990]" pageId="1" pageNumber="294">FIGURES 15.</emphasis>
Nymph of
<taxonomicName id="B6CA93BEFFB2FFEB7DC8F86EEF6D82CC" authority="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan, 2019" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[432,842,1966,1990]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="tungaiensis" status="sp. nov." subGenus="Rhoenanthus">
<emphasis id="43BE342FFFB2FFEB7DC8F86EEE1382CC" box="[432,564,1966,1990]" italics="true" pageId="1" pageNumber="294">Rhoenanthus</emphasis>
(
<emphasis id="43BE342FFFB2FFEB7E3BF86EEEE082CC" box="[579,711,1966,1990]" italics="true" pageId="1" pageNumber="294">Rhoenanthus</emphasis>
)
<emphasis id="43BE342FFFB2FFEB7EAFF86FEF6D82CC" box="[727,842,1967,1990]" italics="true" pageId="1" pageNumber="294">tungaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588D8954FFB2FFEB7F29F86EEF8782CF" box="[849,928,1966,1989]" pageId="1" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible.
</paragraph>
</caption>
<caption id="25B5B8B5FFB1FFE87CEFF937ED4F823E" httpUri="https://zenodo.org/record/3384506/files/figure.png" pageId="2" pageNumber="295" startId="2.[151,264,1783,1808]" targetBox="[168,1413,182,1757]" targetPageId="2">
<paragraph id="7175E83DFFB1FFE87CEFF937ED4F823E" blockId="2.[151,1436,1783,1844]" pageId="2" pageNumber="295">
<emphasis id="43BE342FFFB1FFE87CEFF937ED60821A" bold="true" box="[151,327,1783,1808]" pageId="2" pageNumber="295">FIGURES 611.</emphasis>
Nymph of
<taxonomicName id="B6CA93BEFFB1FFE87DC7F938EF7D821A" authority="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan, 2019" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[447,858,1784,1808]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="tungaiensis" status="sp. nov." subGenus="Rhoenanthus">
<emphasis id="43BE342FFFB1FFE87DC7F938EE64821A" box="[447,579,1784,1808]" italics="true" pageId="2" pageNumber="295">Rhoenanthus</emphasis>
(
<emphasis id="43BE342FFFB1FFE87E2BF938EEF0821A" box="[595,727,1784,1808]" italics="true" pageId="2" pageNumber="295">Rhoenanthus</emphasis>
)
<emphasis id="43BE342FFFB1FFE87E9FF939EF7D821A" box="[743,858,1785,1808]" italics="true" pageId="2" pageNumber="295">tungaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588D8954FFB1FFE87F1AF939EF95821A" box="[866,946,1785,1808]" pageId="2" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
6, Foreleg; 7, Midleg; 8, Hindleg; 9, Gill I; 10, Gill III; 11, Gill VII.
</paragraph>
</caption>
<subSubSection id="39D0BBB6FFB1FFEC7CEFF8A2E94A8655" lastPageId="6" lastPageNumber="299" pageId="2" pageNumber="295" type="materials_examined">
<paragraph id="7175E83DFFB1FFE87CEFF8A2E9B482E2" blockId="2.[151,1436,1890,2025]" pageId="2" pageNumber="295">
<emphasis id="43BE342FFFB1FFE87CEFF8A2ED5C8277" bold="true" box="[151,379,1890,1917]" pageId="2" pageNumber="295">Material examined</emphasis>
<materialsCitation id="C1A2E260FFB1FFE87DFBF8A2ED4182CE" ID-GBIF-Occurrence="2382361119" collectingDate="2018-04-01" collectorName="Reg. No. &amp; Balasubramanian &amp; Muthukatturaja" country="India" county="Tunga River" location="Chikkamagalore district" municipality="Sringeri" pageId="2" pageNumber="295" specimenCount="669" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="AE71569FFFB1FFE87DFBF8A2EDD78276" box="[387,496,1890,1916]" pageId="2" pageNumber="295" type="holotype">Holotype</typeStatus>
: male imago (
<collectorName id="DC3F8DEBFFB1FFE87EE9F8A3EED78277" box="[657,752,1891,1917]" pageId="2" pageNumber="295">Reg.No.</collectorName>
SRC/ ZSI-I/E 391),
<collectingCountry id="09DDA8ADFFB1FFE87FA6F8A3E8178277" box="[990,1072,1891,1917]" name="India" pageId="2" pageNumber="295">INDIA</collectingCountry>
,
<collectingCounty id="981490B1FFB1FFE87845F8A3E8EB8276" box="[1085,1228,1890,1917]" pageId="2" pageNumber="295">Tunga River</collectingCounty>
,
<collectingMunicipality id="91117247FFB1FFE878A0F8A2E9128276" box="[1240,1333,1890,1916]" pageId="2" pageNumber="295">Sringeri</collectingMunicipality>
,
<location id="7415BEE6FFB1FFE8793AF8A2ED7082AA" LSID="urn:lsid:plazi:treatment:F963592BFFB2FFEC7CEFFEB1E94A8655:7415BEE6FFB1FFE8793AF8A2ED7082AA" country="India" county="Tunga River" municipality="Sringeri" name="Chikkamagalore district" pageId="2" pageNumber="295" stateProvince="Karnataka">Chikkamagalore district</location>
,
<collectingRegion id="B30E26DFFFB1FFE87D19F846EE3782AB" box="[353,528,1926,1953]" country="India" name="Karnataka" pageId="2" pageNumber="295">Karnataka state</collectingRegion>
,
<date id="0574CEFDFFB1FFE87E62F847EEB882AB" box="[538,671,1927,1953]" pageId="2" pageNumber="295" value="2018-04-01">
<collectingDate id="15303715FFB1FFE87E62F847EEB882AB" box="[538,671,1927,1953]" pageId="2" pageNumber="295" value="2018-04-01">
01-IV-
<quantity id="B63245D8FFB1FFE87E1CF847EEB882AB" box="[612,671,1927,1953]" metricMagnitude="6" metricUnit="m" metricValue="2.018669" pageId="2" pageNumber="295" unit="m" value="2018669.0">2018</quantity>
</collectingDate>
</date>
,
<specimenCount id="67CC23B4FFB1FFE87ED1F846EED182AB" box="[681,758,1926,1953]" pageId="2" pageNumber="295" type="generic">669 m.</specimenCount>
(13º36.27″ N, 75º19.89″ E). Collected by
<collectorName id="DC3F8DEBFFB1FFE878A2F846E9BB82AA" box="[1242,1436,1926,1952]" pageId="2" pageNumber="295">Balasubramanian</collectorName>
&amp;
<collectorName id="DC3F8DEBFFB1FFE87CCEF86AED4182CE" box="[182,358,1962,1988]" pageId="2" pageNumber="295">Muthukatturaja</collectorName>
</materialsCitation>
.
<typeStatus id="AE71569FFFB1FFE87D0AF86BEDC482CF" box="[370,483,1963,1989]" pageId="2" pageNumber="295" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="67CC23B4FFB1FFE87D97F86BEE1B82CE" box="[495,572,1962,1989]" pageId="2" pageNumber="295" type="male">8 male</specimenCount>
imagoes,
<materialsCitation id="C1A2E260FFB1FFE87ECAF86BEDB182E2" ID-GBIF-Occurrence="2382361118" collectingDate="2018-04-01" collectorName="Reg. No." country="India" county="Tunga River" location="Chikkamagalore district" municipality="Sringeri" pageId="2" pageNumber="295" specimenCount="2" specimenCount-female="2" stateProvince="Karnataka" typeStatus="paratype">
<specimenCount id="67CC23B4FFB1FFE87ECAF86BEF3282CE" box="[690,789,1962,1989]" pageId="2" pageNumber="295" type="female">2 female</specimenCount>
imagoes and 35 mature nymphs (
<collectorName id="DC3F8DEBFFB1FFE878E2F86BE8DE82CF" box="[1178,1273,1963,1989]" pageId="2" pageNumber="295">Reg.No.</collectorName>
SRC/ ZSI-I/E 392 &amp; 393), same data
</materialsCitation>
<materialsCitation id="C1A2E260FFB1FFE87DE4F80EE9A882E2" ID-GBIF-Occurrence="2382361117" box="[412,1423,1998,2025]" country="India" county="Chennai" location="Zoological Survey of India" municipality="Southern Regional Centre" pageId="2" pageNumber="295" specimenCount="1" stateProvince="Karnataka" typeStatus="holotype">
<typeStatus id="AE71569FFFB1FFE87DE4F80EEE2382E2" box="[412,516,1998,2024]" pageId="2" pageNumber="295" type="holotype">Holotype</typeStatus>
is deposited in
<location id="7415BEE6FFB1FFE87ECCF80EEFC482E2" LSID="urn:lsid:plazi:treatment:F963592BFFB2FFEC7CEFFEB1E94A8655:7415BEE6FFB1FFE87ECCF80EEFC482E2" box="[692,995,1998,2025]" country="India" county="Chennai" municipality="Southern Regional Centre" name="Zoological Survey of India" pageId="2" pageNumber="295" stateProvince="Karnataka">Zoological Survey of India</location>
(
<collectingMunicipality id="91117247FFB1FFE87F89F80EE93C82E3" box="[1009,1307,1998,2025]" pageId="2" pageNumber="295">Southern Regional Centre</collectingMunicipality>
),
<collectingCounty id="981490B1FFB1FFE87956F80EE9A882E2" box="[1326,1423,1998,2024]" pageId="2" pageNumber="295">Chennai</collectingCounty>
</materialsCitation>
.
</paragraph>
<caption id="25B5B8B5FFB0FFE97CEFF859E82482B8" box="[151,1027,1945,1970]" httpUri="https://zenodo.org/record/3384508/files/figure.png" pageId="3" pageNumber="296" startId="3.[151,250,1945,1970]" targetBox="[473,1103,181,1905]" targetPageId="3">
<paragraph id="7175E83DFFB0FFE97CEFF859E82482B8" blockId="3.[151,1027,1945,1970]" box="[151,1027,1945,1970]" pageId="3" pageNumber="296">
<emphasis id="43BE342FFFB0FFE97CEFF859ED0782B8" bold="true" box="[151,288,1945,1970]" pageId="3" pageNumber="296">FIGURE 12.</emphasis>
Nymph of
<taxonomicName id="B6CA93BEFFB0FFE97DECF85AEF0A82B8" authority="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan, 2019" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[404,813,1946,1970]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="tungaiensis" status="sp. nov." subGenus="Rhoenanthus">
<emphasis id="43BE342FFFB0FFE97DECF85AEE3F82B8" box="[404,536,1946,1970]" italics="true" pageId="3" pageNumber="296">Rhoenanthus</emphasis>
(
<emphasis id="43BE342FFFB0FFE97E5FF85AEE8C82B8" box="[551,683,1946,1970]" italics="true" pageId="3" pageNumber="296">Rhoenanthus</emphasis>
)
<emphasis id="43BE342FFFB0FFE97EC2F85BEF0A82B8" box="[698,813,1947,1970]" italics="true" pageId="3" pageNumber="296">tungaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588D8954FFB0FFE97F4CF85AEFA282BB" box="[820,901,1946,1969]" pageId="3" pageNumber="296" rank="species">sp. nov.</taxonomicNameLabel>
, dorsal view
</paragraph>
</caption>
<paragraph id="7175E83DFFB7FFEE7CBFFF58E94F85F0" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFFF58ED9485B9" bold="true" box="[199,435,152,179]" pageId="4" pageNumber="297">
<collectionCode id="17DB70F8FFB7FFEE7CBFFF58ED9485B9" box="[199,435,152,179]" pageId="4" pageNumber="297">MATURE NYMPH</collectionCode>
</emphasis>
(
<figureCitation id="E9F1F4B8FFB7FFEE7DBBFF58EE3685B9" box="[451,529,152,179]" captionStart="FIGURE 12" captionStartId="3.[151,250,1945,1970]" captionTargetBox="[473,1103,181,1905]" captionTargetId="figure@3.[473,1114,181,1920]" captionTargetPageId="3" captionText="FIGURE 12. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov., dorsal view" httpUri="https://zenodo.org/record/3384508/files/figure.png" pageId="4" pageNumber="297">Fig.12</figureCitation>
): Body length (excluding cerci):
<quantity id="B63245D8FFB7FFEE7FFCFF59EFCF85B8" box="[900,1000,152,179]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" metricValueMax="5.334" metricValueMin="4.826" pageId="4" pageNumber="297" unit="in" value="20.0" valueMax="21.0" valueMin="19.0">1921 in</quantity>
male and female nymphs; cerci length half times longer than body length. General body coloration yellow to dark brown, yellowish brown to dark brown dorsally and pale to yellowish brown ventrally, young specimens generally paler; legs light to medium brown.
</paragraph>
<paragraph id="7175E83DFFB7FFEE7CBFFEC5ED2686E4" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFFEC5ED378415" bold="true" box="[199,272,261,287]" pageId="4" pageNumber="297">Head:</emphasis>
Brown with cranial and epicranial sutures pale. Three crescent shaped ocelli present; dorsal diameter of male compound eyes
<quantity id="B63245D8FFB7FFEE7DF6FEE8EDD28449" box="[398,501,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="297" unit="mm" value="1.05">1.05 mm</quantity>
, female compound eyes 0.59 mm
<quantity id="B63245D8FFB7FFEE7FFBFEE8EFCF8449" box="[899,1000,296,323]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="4" pageNumber="297" unit="mm" value="0.61">0.61 mm</quantity>
, distance between compound eye
<quantity id="B63245D8FFB7FFEE7913FEE8ECE1846D" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="4" pageNumber="297" unit="mm" value="1.86">1.86 mm</quantity>
; Antenna 1.3 times longer than head width, flagellum medium to light brown. Labrum (
<figureCitation id="E9F1F4B8FFB7FFEE78E4FE8CE8FA846D" box="[1180,1245,332,359]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig. 1</figureCitation>
) 2.5 times wider than long, with smooth anterior emargination; dorsal face covered medially by 2 rows of long feathered setae; dorsal setae with feather structure; anterior margin with a row of small thin setae along with a row of mono pectinate (feathered on one side) setae and bipectinate (feathered on both side) setae at emargination. Mandibles slender with few thin setae in outer margin, mandibular tusks 0.8 times longer than head length (
<figureCitation id="E9F1F4B8FFB7FFEE7832FE1CE8BA84FD" box="[1098,1181,476,503]" captionStart="FIGURE 12" captionStartId="3.[151,250,1945,1970]" captionTargetBox="[473,1103,181,1905]" captionTargetId="figure@3.[473,1114,181,1920]" captionTargetPageId="3" captionText="FIGURE 12. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov., dorsal view" httpUri="https://zenodo.org/record/3384508/files/figure.png" pageId="4" pageNumber="297">Fig. 12</figureCitation>
), outer incisor of right mandible (
<figureCitation id="E9F1F4B8FFB7FFEE7D68FDC0ED448711" box="[272,355,512,539]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig. 5b</figureCitation>
) with 3 large and 3 small teeth; inner incisor with 2 teeth and line of setae laterally; prostheca bifid with sharp subapical spines, comprising a tufts of hairs (
<figureCitation id="E9F1F4B8FFB7FFEE7F31FDE4EFBA8735" box="[841,925,548,575]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig. 5b</figureCitation>
); a tuft of hairy setae between mola and prostheca; ten long setae below the molar; mandibular body with row of stout yellowish brown setae at mesal margin; outer margin with 2022 scattered blunt setae, 1315 spine-like setae and 1012 hair-like long setae; apex of left mandibular tusk (
<figureCitation id="E9F1F4B8FFB7FFEE7D24FD50ED8B87A1" box="[348,428,656,683]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig. 5a</figureCitation>
) with two thin setae; incisors with three teeth; no thin setae, between mola and prostheca, no tuft of hairy setae. Hypopharynx (
<figureCitation id="E9F1F4B8FFB7FFEE7E4CFD74EE5687C5" box="[564,625,692,719]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig.2</figureCitation>
) with lingua apically truncate and covered with thin setae; super-linguae with two basal rows of sharp setae and covered apically with long simple setae. Maxillae (
<figureCitation id="E9F1F4B8FFB7FFEE78F0FD18E90587F9" box="[1160,1314,728,755]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Figs. 4a &amp; 4b</figureCitation>
) slender, 2 indented dentisetae with 710 serrations, and 3 long setae on inner apical part and a small row of long thin setae at crown; inner margin from apex to base of lacinia with ca 22 long monopectinate setae ventrally and 3 long simple setae dorsally; segment I of maxillary palp 1.7 times longer than segment II; maxillary palp with 58 simple setae; cardo and stipes of the maxilla with numerous thin setae dorsally. Labium (
<figureCitation id="E9F1F4B8FFB7FFEE7FA9FCA8E8338689" box="[977,1044,872,899]" captionStart="FIGURES 15" captionStartId="1.[151,264,1965,1990]" captionTargetBox="[151,1436,453,1940]" captionTargetId="figure@1.[151,1436,453,1940]" captionTargetPageId="1" captionText="FIGURES 15. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 1, Labrum (dorsal view); 2, Hypopharynx (dorsal view); 3, Labium; 4a, Left Maxilla; 4b, Right maxilla; 5a, Right mandible (dorsal view); 5b, Left Mandible." httpUri="https://zenodo.org/record/3384504/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
): labial palp of segment III convex, inner margin covered with simple stout setae, 1.53 times longer than segment II, lengths of segments IIII (mm) = 0.54 /0.35 /0.57; paraglossae densely covered with spine like setae on ventral surface; glossa with similar setae ventrally.
</paragraph>
<paragraph id="7175E83DFFB7FFEE7CBFFC39EF4B8070" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFFC39ED0E8119" bold="true" box="[199,297,1017,1043]" pageId="4" pageNumber="297">Thorax:</emphasis>
medium brown to dark brown; prothorax with two anterolateral dorsal tubercles, meso and metathorax each with two anteromedial tubercles. Legs: yellow to brown, with a row of 69 thin setae on distal end of coxa; a row of simple setae over the trochanter; three medium to dark brown stripes at femora; length ratio range of foreleg femur: tibia: tarsus: claw (mm) = 3.03.2 / 3.03.2 / 1.31.5 / 0.020.03; midleg femur: tibia: tarsus: claw (mm) = 2.22.3 / 1.8 2.0 / 0.70.8 / 0.02; hindleg femur: tibia: tarsus: claw (mm) = 3.03.2 / 2.32.5 / 0.91.0 / 0.020.03; fore femur (
<figureCitation id="E9F1F4B8FFB7FFEE7D5AFB6CED4F81CD" box="[290,360,1196,1223]" captionStart="FIGURES 611" captionStartId="2.[151,264,1783,1808]" captionTargetBox="[168,1413,182,1757]" captionTargetId="figure@2.[162,1426,181,1757]" captionTargetPageId="2" captionText="FIGURES 611. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 6, Foreleg; 7, Midleg; 8, Hindleg; 9, Gill I; 10, Gill III; 11, Gill VII." httpUri="https://zenodo.org/record/3384506/files/figure.png" pageId="4" pageNumber="297">Fig. 6</figureCitation>
) 2.9 times longer than its greatest width; outer margin covered by stout and long setae, inner margin with a short row of stout and long setae; mid femora similar to that of fore femur with few stout yellowish brown setae dorsally; apex of hind femur with clump of prominent bipectinate setae ventrally; mid and hind femora 1.61.8 times longer than the greatest width (
<figureCitation id="E9F1F4B8FFB7FFEE7EF5FAD8EF2C8039" box="[653,779,1304,1331]" captionStart="FIGURES 611" captionStartId="2.[151,264,1783,1808]" captionTargetBox="[168,1413,182,1757]" captionTargetId="figure@2.[162,1426,181,1757]" captionTargetPageId="2" captionText="FIGURES 611. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 6, Foreleg; 7, Midleg; 8, Hindleg; 9, Gill I; 10, Gill III; 11, Gill VII." httpUri="https://zenodo.org/record/3384506/files/figure.png" pageId="4" pageNumber="297">Figs. 7 &amp; 8</figureCitation>
); dorsal and inner margins of femur with a row of long and stout setae; tibia with a row of long and stout setae on dorsal and lateral margin. Tarsal claw simple, hooked with tufts of setae at the base, 2.5 times longer than its greatest width.
</paragraph>
<paragraph id="7175E83DFFB7FFEE7CBFFA44EED283B4" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFFA44ED628095" bold="true" box="[199,325,1412,1439]" pageId="4" pageNumber="297">Abdomen:</emphasis>
Yellow to brown, markings with dark brown; terga with no scattered bifurcate plumose setae on surfaces, with setae slightly distributed along margins; well-developed apically pale posterolateral projections present on terga IIIIX, with largest projection on tergum IX; terga IX each with inverted bell shaped marking, two pale anteromedial spot; sterna pale yellow, with no distinct pale spots anteromedially on segments IIVIII. Terminal filament is subequal or slightly longer than cerci, with long setae at junction of each segment. Gills (
<figureCitation id="E9F1F4B8FFB7FFEE78A3F9D4E9698325" box="[1243,1358,1556,1583]" captionStart="FIGURES 611" captionStartId="2.[151,264,1783,1808]" captionTargetBox="[168,1413,182,1757]" captionTargetId="figure@2.[162,1426,181,1757]" captionTargetPageId="2" captionText="FIGURES 611. Nymph of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 6, Foreleg; 7, Midleg; 8, Hindleg; 9, Gill I; 10, Gill III; 11, Gill VII." httpUri="https://zenodo.org/record/3384506/files/figure.png" pageId="4" pageNumber="297">Figs. 9-11</figureCitation>
) white with medium to dark brown tracheae, gills IVII attached laterally, Gill 1 single, segmented, basal segment
<quantity id="B63245D8FFB7FFEE7913F9F8ECE4837D" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="4" pageNumber="297" unit="mm" value="0.67">0.67 mm</quantity>
; apical segment
<quantity id="B63245D8FFB7FFEE7DFEF99DEDCC837D" box="[390,491,1629,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="297" unit="mm" value="1.03">1.03 mm</quantity>
, with numerous hair like setae. Gills II- VII composed of well developed bipectinate fibrillae with lamella, well marked tracheation and a tuft of dark brown fibrillae, gill III with 4648 marginal fibrillae on dorsal lamellae, and 3537 on ventral lamellae.
</paragraph>
<paragraph id="7175E83DFFB7FFEE7CBFF908EDAF8221" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFF908EDAF83E9" bold="true" box="[199,392,1736,1763]" pageId="4" pageNumber="297">
<collectionCode id="17DB70F8FFB7FFEE7CBFF908EDA483E9" box="[199,387,1736,1763]" pageId="4" pageNumber="297">MALE ADULT</collectionCode>
:
</emphasis>
Length of body
<quantity id="B63245D8FFB7FFEE7E3CF909EEB183E9" box="[580,662,1737,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="297" unit="mm" value="14.0">14 mm</quantity>
, length of fore wing
<quantity id="B63245D8FFB7FFEE7FFAF908EFCF83E9" box="[898,1000,1736,1763]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="297" unit="mm" value="12.5">12.5 mm</quantity>
, width of forewing 6.0 mm, hind wing length 6.0 mm, width of hind wing 3.0 mm; dorsal diameter of compound eyes
<quantity id="B63245D8FFB7FFEE785CF92DE8AB820D" box="[1060,1164,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="4" pageNumber="297" unit="mm" value="0.78">0.78 mm</quantity>
; distance between compound eyes
<quantity id="B63245D8FFB7FFEE7D65F8D1EDA38221" box="[285,388,1809,1835]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="297" unit="mm" value="0.1">0.10 mm</quantity>
.
</paragraph>
<paragraph id="7175E83DFFB7FFEE7CBFF8F5EDE082B1" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFF8F5ED208245" bold="true" box="[199,263,1845,1871]" pageId="4" pageNumber="297">Head</emphasis>
(
<figureCitation id="E9F1F4B8FFB7FFEE7D6FF8F4ED4B8245" box="[279,364,1844,1871]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="4" pageNumber="297">Fig. 17</figureCitation>
). Color yellowish brown. Compound eyes blackish yellow in alcohol, large, nearly contiguous (ES=0.14). Relative size of compound eyes of mature larvae was measured by the formula ES= B/D. where ES: compound eye size, B: shortest distance between compound eye, D: longest dorsal diameter of a compound eye (
<bibRefCitation id="155B95CCFFB7FFEE7CE7F861ED9D82B1" author="Bae, Y. J. &amp; McCafferty, W. P." box="[159,442,1952,1979]" pageId="4" pageNumber="297" pagination="1 - 143" refId="ref3676" refString="Bae, Y. J. &amp; McCafferty, W. P. (1991) Phylogenetic systematics of the Potamanthidae (Ephemeroptera), Transactions of the American Entomological Society, 117 (3 - 4), 1 - 143." type="journal article" year="1991">Bae &amp; McCafferty, 1991</bibRefCitation>
).
</paragraph>
<paragraph id="7175E83DFFB7FFEF7CBFF805EF3B84A5" blockId="4.[151,1437,152,2015]" lastBlockId="5.[151,1437,153,431]" lastPageId="5" lastPageNumber="298" pageId="4" pageNumber="297">
<emphasis id="43BE342FFFB7FFEE7CBFF805ED0E82D5" bold="true" box="[199,297,1989,2015]" pageId="4" pageNumber="297">Thorax:</emphasis>
Color whitish brown to brown: nota somewhat darker than pleura and sterna. Forewings (
<figureCitation id="E9F1F4B8FFB7FFEE7938F804E9B482D5" box="[1344,1427,1988,2015]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="4" pageNumber="297">Fig. 13</figureCitation>
) transparent, with pinkish brown to brown stained marks variously developed at base of forks; longitudinal veins yellowish brown to brown; costal cross veins reddish brown and infuscated, other cross veins purplish brown and infuscate, numbering ca 30 between Sc and R
<subScript id="ED4EEA78FFB6FFEF7EECFF32EE86840A" attach="left" box="[660,673,242,256]" fontSize="6" pageId="5" pageNumber="298">1;</subScript>
21 between R
<subScript id="ED4EEA78FFB6FFEF7F39FF32EF6E840A" attach="left" box="[833,841,242,256]" fontSize="6" pageId="5" pageNumber="298">1</subScript>
and R
<subScript id="ED4EEA78FFB6FFEF7FF7FF32EFBA840A" attach="left" box="[911,925,242,256]" fontSize="6" pageId="5" pageNumber="298">2;</subScript>
MAs 0.75 x length of MA
<subScript id="ED4EEA78FFB6FFEF78BDFF32E8F5840A" attach="left" box="[1221,1234,242,256]" fontSize="6" pageId="5" pageNumber="298">1;</subScript>
between
<collectionCode id="17DB70F8FFB6FFEF7944FF21E94385F1" box="[1340,1380,225,251]" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="298">MP</collectionCode>
<subScript id="ED4EEA78FFB6FFEF791DFF32E94A840A" attach="left" box="[1381,1389,242,256]" fontSize="6" pageId="5" pageNumber="298">2</subScript>
and CuA a large cell present; Cu regions with
<specimenCount id="67CC23B4FFB6FFEF7E15FEC5EF2C8415" box="[621,779,261,287]" pageId="5" pageNumber="298" type="intercalary">4 intercalaries</specimenCount>
(3 forked): A
<subScript id="ED4EEA78FFB6FFEF7FDDFED6EF8A842E" attach="left" box="[933,941,278,292]" fontSize="6" pageId="5" pageNumber="298">1</subScript>
forked once. Hind wings (
<figureCitation id="E9F1F4B8FFB6FFEF78A0FEC5E90D8415" box="[1240,1322,261,287]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="5" pageNumber="298">Fig. 14</figureCitation>
) transparent, with brown stained marks variously at base of forks: longitudinal veins brown; Cross veins light purplish brown and infuscated in central area: intercalaries same as forewing; costal projection acute (&gt;90°): base of R
<subScript id="ED4EEA78FFB6FFEF7962FE9EE9058466" attach="left" box="[1306,1314,350,364]" fontSize="6" pageId="5" pageNumber="298">1</subScript>
not arched (ca.110°) toward Sc. Forelegs yellowish; middle region of the fore tibiae and fore tarsal segment 4 and 5 brown; middle legs and hind legs pale yellow; claws dark brown.
</paragraph>
<caption id="25B5B8B5FFB6FFEF7CEFF8CDEF468240" httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="5" pageNumber="298" startId="5.[151,264,1805,1830]" targetBox="[256,1341,431,1764]" targetPageId="5">
<paragraph id="7175E83DFFB6FFEF7CEFF8CDEF468240" blockId="5.[151,1436,1805,1866]" pageId="5" pageNumber="298">
<emphasis id="43BE342FFFB6FFEF7CEFF8CDED70822C" bold="true" box="[151,343,1805,1830]" pageId="5" pageNumber="298">FIGURES 1318.</emphasis>
Adult of
<taxonomicName id="B6CA93BEFFB6FFEF7DBBF8CDEF44822F" authority="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan, 2019" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[451,867,1805,1829]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="tungaiensis" status="sp. nov." subGenus="Rhoenanthus">
<emphasis id="43BE342FFFB6FFEF7DBBF8CDEE60822F" box="[451,583,1805,1829]" italics="true" pageId="5" pageNumber="298">Rhoenanthus</emphasis>
(
<emphasis id="43BE342FFFB6FFEF7E21F8CDEEFA822F" box="[601,733,1805,1829]" italics="true" pageId="5" pageNumber="298">Rhoenanthus</emphasis>
)
<emphasis id="43BE342FFFB6FFEF7E88F8CEEF44822F" box="[752,867,1806,1829]" italics="true" pageId="5" pageNumber="298">tungaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588D8954FFB6FFEF7F15F8CEEF98822F" box="[877,959,1806,1829]" pageId="5" pageNumber="298" rank="species">sp. nov.</taxonomicNameLabel>
13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia.
</paragraph>
</caption>
<paragraph id="7175E83DFFB6FFEC7CBFF8B7EEC18449" blockId="5.[151,1436,1911,2010]" lastBlockId="6.[151,1437,152,863]" lastPageId="6" lastPageNumber="299" pageId="5" pageNumber="298">
<emphasis id="43BE342FFFB6FFEF7CBFF8B7ED628298" bold="true" box="[199,325,1911,1938]" pageId="5" pageNumber="298">Abdomen:</emphasis>
terga (
<figureCitation id="E9F1F4B8FFB6FFEF7DE2F8B8EDC88298" box="[410,495,1911,1938]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="5" pageNumber="298">Fig. 16</figureCitation>
) I-X each pinkish yellow, with dark brown, broad mid longitudinal inverted bell shaped markings containing a pairs of pale trabeculae (rod-like marking) outwards, with pair dark brown, broad stripes running from anterolateral corner to median posterior margin and with single narrow, elongated, spots ar- ranged on lateral margin. Sterna (
<figureCitation id="E9F1F4B8FFB5FFEC7E6CFF59EE4085B9" box="[532,615,152,179]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="6" pageNumber="299">Fig. 15</figureCitation>
) IX each yellow, with 2 pairs of dark, median tear drop spots, sub median spots (anterior pair larger and more widely separated) and pair of narrow, longitudinal, lateral stripes. Genital forceps (
<figureCitation id="E9F1F4B8FFB5FFEC7C83FF21ED6885F1" box="[251,335,225,251]" captionStart="FIGURES 1318" captionStartId="5.[151,264,1805,1830]" captionTargetBox="[256,1341,431,1764]" captionTargetId="figure@5.[237,1349,453,1780]" captionTargetPageId="5" captionText="FIGURES 1318. Adult of Rhoenanthus (Rhoenanthus) tungaiensis sp. nov. 13, Forewing; 14, Hindwing; 15, Abdominal sterna IV-IX; 16, Abdominal terga IV-X; 17, Head; 18, Male genitalia." httpUri="https://zenodo.org/record/3384510/files/figure.png" pageId="6" pageNumber="299">Fig. 18</figureCitation>
) pale yellow; penes with ventral surface pale yellow, dorsal surface light brown, apex of penes bifurcated dorsoventrally and rounded apically. Cerci pale yellow to brown relatively long (1.92 x length of body); terminal filament rudimentary 0.04 x length of cerci.
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFE8CE84B84A5" blockId="6.[151,1437,152,863]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFE8CED8B846D" bold="true" box="[199,428,332,359]" pageId="6" pageNumber="299">
<collectionCode id="17DB70F8FFB5FFEC7CBFFE8CED80846D" box="[199,423,332,359]" pageId="6" pageNumber="299">FEMALE ADULT</collectionCode>
:
</emphasis>
Length of body 15.0 mm, length of fore wing 21.0 mm, width of forewing 5.0 mm, Hind wing length 4.0 mm, width of hind wing 3.0 mm; forelegs
<quantity id="B63245D8FFB5FFEC7F5CFEB1EF5C8481" box="[804,891,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="299" unit="mm" value="2.1">2.1 mm</quantity>
, cerci
<quantity id="B63245D8FFB5FFEC7FBBFEB1E8338481" box="[963,1044,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="299" unit="mm" value="17.0">17 mm</quantity>
, median terminal filament 1.0 mm; dorsal diameter of compound eyes
<quantity id="B63245D8FFB5FFEC7E58FE55EEA084A5" box="[544,647,405,431]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="299" unit="mm" value="0.78">0.78 mm</quantity>
; distance between compound eyes
<quantity id="B63245D8FFB5FFEC7877FE55E84F84A5" box="[1039,1128,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="299" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFE79E91084DE" blockId="6.[151,1437,152,863]" box="[199,1335,441,468]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFE79ED3784D9" bold="true" box="[199,272,441,467]" pageId="6" pageNumber="299">Head:</emphasis>
Color brownish yellow. Compound eyes black in alcohol, medium, widely separated (ES=0.6)
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFE1DE9BA878D" blockId="6.[151,1437,152,863]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFE1DED0E84FD" bold="true" box="[199,297,477,503]" pageId="6" pageNumber="299">Thorax:</emphasis>
prothorax with brownish yellow to brown, longitudinal, lateral stripes on notum and pleura. Forewings with brownish brown, infuscated, costal cross veins reddish brown and with 4 cubital intercalleries 4 (3foreked); other venation as in male. Hind wings with acute costal projections (&lt;90 degree). Fore legs pinkish brown to yellow, fore tibiae pinkish brown to yellow, tarsal segment 5 reddish brown, claw bifid, dark brown. Fore tarsal segments 1-4 pale yellow, mid and hind legs pale yellow; fore tibiae 0.9 x length of foreleg, 1.05 x length of fore tarsi.
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFD50EEFB87C5" blockId="6.[151,1437,152,863]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFD50ED6287A1" bold="true" box="[199,325,656,683]" pageId="6" pageNumber="299">Abdomen:</emphasis>
Color pattern as in male. Cerci pale yellow to brown, with dark band at each suture: median terminal filament brown, rudimentary, 0.07 x length of cerci.
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFD19ED23861D" blockId="6.[151,1437,152,863]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFD19ED7587F9" bold="true" box="[199,338,729,755]" pageId="6" pageNumber="299">Etymology:</emphasis>
The species name
<taxonomicName id="B6CA93BEFFB5FFEC7E5CFD1AEE8387F9" authorityName="Balasubramanian &amp; Muthukatturaja &amp; Anbalagan" authorityYear="2019" box="[548,676,730,755]" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="299" phylum="Arthropoda" rank="species" species="tungaiensis">
<emphasis id="43BE342FFFB5FFEC7E5CFD1AEE8387F9" box="[548,676,730,755]" italics="true" pageId="6" pageNumber="299">tungaiensis</emphasis>
</taxonomicName>
refers to the locality name, Tunga River, where the new species was collected.
</paragraph>
<paragraph id="7175E83DFFB5FFEC7CBFFCE1E94A8655" blockId="6.[151,1437,152,863]" pageId="6" pageNumber="299">
<emphasis id="43BE342FFFB5FFEC7CBFFCE1ED088631" bold="true" box="[199,303,801,827]" pageId="6" pageNumber="299">Ecology:</emphasis>
This species was found in the loose soil places at boundary of the stream in middle to fast current, associated with
<taxonomicName id="B6CA93BEFFB5FFEC7D32FC85ED9A8655" box="[330,445,837,863]" class="Insecta" family="Ephemeridae" genus="Ephemera" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="43BE342FFFB5FFEC7D32FC85ED9A8655" box="[330,445,837,863]" italics="true" pageId="6" pageNumber="299">Ephemera</emphasis>
</taxonomicName>
sp. possibly to feed on fine particulate organic matter available in the water current.
</paragraph>
</subSubSection>
</treatment>
</document>