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<document id="71C9DA8DC1B4A6C1F0BFFED1758A066B" ID-CLB-Dataset="21099" ID-DOI="10.1080/00222933.2014.909072" ID-GBIF-Dataset="3bad6762-1788-4f94-9cfd-1281b8619c55" ID-ISSN="1464-5262" ID-Zenodo-Dep="4002597" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598467980037" checkinUser="carolina" docAuthor="Frolov, Andrey V." docDate="2014" docId="F91EE008102EFFAAFE9BDE049DB96E5C" docLanguage="en" docName="JNATHIST.49.3-4.129-154.pdf" docOrigin="Journal of Natural History 49 (3)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Metacatharsius janssensi Frolov 2014, comb. nov." docType="treatment" docVersion="11" lastPageNumber="153" masterDocId="052798701039FFB2FFECDA539E466F14" masterDocTitle="Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa" masterLastPageNumber="154" masterPageNumber="129" pageNumber="152" updateTime="1698845524048" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A282ADA996B918BDA207C4406DEE6B49">Revision of the genus Delopleurus Boheman (Coleoptera: Scarabaeidae: Scarabaeinae) with description of new species from Africa</mods:title>
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<mods:namePart id="37261FB70574454AC5520B9899162003">Frolov, Andrey V.</mods:namePart>
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<mods:title id="FE487FB2703D38F0479D322FC0331C94">Journal of Natural History</mods:title>
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<mods:date id="CC6D381707C08862933E86DE7D70258A">2014</mods:date>
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<mods:number id="DCCB0D6D582BB55575363DD4E24D5737">2014-06-30</mods:number>
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<mods:number id="2483143055B0D3826716D2BEC17B1584">49</mods:number>
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<mods:classification id="A932275E06A27E547DEE5D62D866B637">journal article</mods:classification>
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<mods:identifier id="B7E8F640B921896063B6D93CB34A9816" type="DOI">10.1080/00222933.2014.909072</mods:identifier>
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<treatment id="F91EE008102EFFAAFE9BDE049DB96E5C" ID-DOI="http://doi.org/10.5281/zenodo.4331163" ID-GBIF-Taxon="167368581" ID-Zenodo-Dep="4331163" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C" httpUri="http://treatment.plazi.org/id/F91EE008102EFFAAFE9BDE049DB96E5C" lastPageId="24" lastPageNumber="153" pageId="23" pageNumber="152">
<subSubSection id="39AD0295102EFFA5FE9BDE049D2F6B64" box="[375,873,1111,1136]" pageId="23" pageNumber="152" type="nomenclature">
<paragraph id="7108511E102EFFA5FE9BDE049D2F6B64" blockId="23.[375,873,1111,1168]" box="[375,873,1111,1136]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FE9BDE049D2F6B64" box="[375,873,1111,1136]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="152" reason="2">
<taxonomicName id="B6B72A9D102EFFA5FE9BDE049C9D6B64" ID-CoL="428Y3" authority="(Frey)" authorityName="Frolov" authorityYear="2014" baseAuthorityName="Frolov" baseAuthorityYear="1963" box="[375,731,1111,1136]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi" status="comb. nov.">
<emphasis id="43C38D0C102EFFA5FE9BDE049CCC6B7B" bold="true" box="[375,650,1111,1135]" italics="true" pageId="23" pageNumber="152">Metacatharsius janssensi</emphasis>
(Frey)
</taxonomicName>
,
<taxonomicNameLabel id="58F03077102EFFA5FD05DE0B9D2F6B64" box="[745,873,1112,1136]" pageId="23" pageNumber="152" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FDC2DE2B9CF46B84" box="[558,690,1144,1168]" pageId="23" pageNumber="152" type="description">
<paragraph id="7108511E102EFFA5FDC2DE2B9CF46B84" blockId="23.[375,873,1111,1168]" box="[558,690,1144,1168]" pageId="23" pageNumber="152">
(
<figureCitation id="E98C4D9B102EFFA5FDDBDE2B9CEF6B84" box="[567,681,1144,1168]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF7CDEF69D966AE8" pageId="23" pageNumber="152" type="reference_group">
<paragraph id="7108511E102EFFA5FF7CDEF69FB96BA9" blockId="23.[144,511,1189,1214]" box="[144,511,1189,1214]" pageId="23" pageNumber="152">
<heading id="2A40E672102EFFA5FF7CDEF69FB96BA9" box="[144,511,1189,1214]" fontSize="10" level="3" pageId="23" pageNumber="152" reason="5">
<taxonomicName id="B6B72A9D102EFFA5FF7CDEF69FB96BA9" ID-CoL="34H58" authority="Frey, 1963" authorityName="Frey" authorityYear="1963" box="[144,511,1189,1214]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FF7CDEF69F3A6BA9" box="[144,380,1189,1213]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
<bibRefCitation id="15262CEF102EFFA5FE69DEF59FB96BA9" author="Frey G" box="[389,511,1189,1214]" pageId="23" pageNumber="152" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey, 1963</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="7108511E102EFFA5FF7CDEB69D966AE8" blockId="23.[144,1104,1253,1691]" pageId="23" pageNumber="152">
<taxonomicName id="B6B72A9D102EFFA5FF7CDEB69F3D6BE9" ID-CoL="34H58" authorityName="Frey" authorityYear="1963" box="[144,379,1253,1277]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FF7CDEB69F3D6BE9" box="[144,379,1253,1277]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
</taxonomicName>
was described from two specimens from Warder,
<collectingRegion id="B3739FFC102EFFA5FC5EDEB69A166BE9" box="[946,1104,1253,1277]" country="Ethiopia" name="Sumale" pageId="23" pageNumber="152">Somali region</collectingRegion>
of
<collectingCountry id="09A0118E102EFFA5FF5FDF579F5E6A08" box="[179,280,1284,1308]" name="Ethiopia" pageId="23" pageNumber="152">Ethiopia</collectingCountry>
. The
<typeStatus id="AE0CEFBC102EFFA5FE8DDF579F806A08" box="[353,454,1284,1308]" pageId="23" pageNumber="152" type="holotype">holotype</typeStatus>
, female, has all the diagnostic characters of the genus
<taxonomicName id="B6B72A9D102EFFA5FF7CDF769FDD6A28" ID-CoL="5Q6D" authority="Paulian" authorityName="Paulian" authorityYear="1939" box="[144,411,1316,1341]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102EFFA5FF7CDF769F7D6A29" box="[144,315,1317,1341]" italics="true" pageId="23" pageNumber="152">Metacatharsius</emphasis>
Paulian
</taxonomicName>
(large dorsal eye parts can be seen on the habitual photo,
<figureCitation id="E98C4D9B102EFFA5FF7CDF179F446A48" box="[144,258,1348,1372]" captionStart="Figure 9" captionStartId="11.[144,211,939,961]" captionTargetBox="[147,1104,168,906]" captionTargetId="figure@11.[144,1104,168,907]" captionTargetPageId="11" captionText="Figure 9. Delopleurus spp. and Metacatharsius janssensi. (A, E) habitus (A, D. striatus, E, M. janssensi); (B) D. striatus, head and pronotum in dorsal view; (C) D. striatus, pygidium; (D) D. striatus, locality map; (F) D. fossatus sp. nov., ventral view of head showing gula with distinct longitudinal groove." figureDoi="http://doi.org/10.5281/zenodo.4002615" httpUri="https://zenodo.org/record/4002615/files/figure.png" pageId="23" pageNumber="152">Figure 9E</figureCitation>
). The
<typeStatus id="AE0CEFBC102EFFA5FEA3DF169FF46A49" box="[335,434,1349,1373]" pageId="23" pageNumber="152" type="paratype">paratype</typeStatus>
is similar to and apparently conspecific with the
<typeStatus id="AE0CEFBC102EFFA5FC0ADF179A0A6A48" box="[998,1100,1348,1372]" pageId="23" pageNumber="152" type="holotype">holotype</typeStatus>
. In the original description, the author compared this species with
<taxonomicName id="B6B72A9D102EFFA5FC63DF369DB36A68" ID-CoL="6CHFF" authorityName="Boheman" authorityYear="1857" box="[911,1013,1380,1404]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="pullus">
<emphasis id="43C38D0C102EFFA5FC63DF369DB36A68" box="[911,1013,1380,1404]" italics="true" pageId="23" pageNumber="152">D. pullus</emphasis>
</taxonomicName>
and
<taxonomicName id="B6B72A9D102EFFA5FBD8DF369E886A88" authorityName="Frolov" authorityYear="2014" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="gilleti">
<emphasis id="43C38D0C102EFFA5FBD8DF369E886A88" italics="true" pageId="23" pageNumber="152">D. gilleti</emphasis>
</taxonomicName>
but did not provide characters specific for the genus
<emphasis id="43C38D0C102EFFA5FCCADFD79DF76A88" box="[806,945,1412,1436]" italics="true" pageId="23" pageNumber="152">
<taxonomicName id="B6B72A9D102EFFA5FCCADFD79DEB6A88" ID-CoL="42PQ" authorityName="Erichson" authorityYear="1847" box="[806,941,1412,1436]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">Delopleurus</taxonomicName>
.
</emphasis>
However, the original description and locality data agree well with the type specimens. There is no doubt that
<taxonomicName id="B6B72A9D102EFFA5FEF5DF979C4F6AC8" ID-CoL="34H58" authorityName="Frey" authorityYear="1963" box="[281,521,1476,1500]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FEF5DF979C4F6AC8" box="[281,521,1476,1500]" italics="true" pageId="23" pageNumber="152">Delopleurus janssensi</emphasis>
</taxonomicName>
was described from misidentified
<taxonomicName id="B6B72A9D102EFFA5FC49DF979A166AC8" ID-CoL="5Q6D" authorityName="Paulian" authorityYear="1939" box="[933,1104,1476,1500]" class="Insecta" family="Scarabaeidae" genus="Metacatharsius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="genus">
<emphasis id="43C38D0C102EFFA5FC49DF979A166AC8" box="[933,1104,1476,1500]" italics="true" pageId="23" pageNumber="152">Metacatharsius</emphasis>
</taxonomicName>
specimens and the new generic combination is therefore established here.
</paragraph>
</subSubSection>
<subSubSection id="39AD0295102EFFA5FF56DC579CE0698F" pageId="23" pageNumber="152" type="description">
<paragraph id="7108511E102EFFA5FF56DC579CE0698F" blockId="23.[144,1104,1253,1691]" pageId="23" pageNumber="152">
It is possible that
<taxonomicName id="B6B72A9D102EFFA5FE68DC569C4B6908" authorityName="Frey" authorityYear="1963" box="[388,525,1541,1564]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FE68DC569C4B6908" box="[388,525,1541,1564]" italics="true" pageId="23" pageNumber="152">M. janssensi</emphasis>
</taxonomicName>
is conspecific with another described species of the genus. There are up to 18 species of this genus known from adjacent
<collectingCountry id="09A0118E102EFFA5FC18DC709A09692F" box="[1012,1103,1571,1595]" name="Somalia" pageId="23" pageNumber="152">Somalia</collectingCountry>
(
<bibRefCitation id="15262CEF102EFFA5FF7BDC109F74694F" author="Ferreira M" box="[151,306,1603,1627]" pageId="23" pageNumber="152" pagination="5 - 1088" refId="ref9860" refString="Ferreira M. 1972. Os escarabideos de Africa (Sul do Saara), I. Rev Ent Mocambique. 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972</bibRefCitation>
) and the genus is currently a subject of taxonomic revision (François Génier, pers. comm.). It is out of the scope of the present work to specify the taxonomic status of
<taxonomicName id="B6B72A9D102EFFA5FE95DCD79C45698F" authorityName="Frey" authorityYear="1963" box="[377,515,1668,1691]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="152" phylum="Arthropoda" rank="species" species="janssensi">
<emphasis id="43C38D0C102EFFA5FE95DCD79C45698F" box="[377,515,1668,1691]" italics="true" pageId="23" pageNumber="152">M. janssensi</emphasis>
</taxonomicName>
more exactly.
</paragraph>
</subSubSection>
<subSubSection id="39AD02951021FFAAFF38DAFA9DB96E5C" pageId="24" pageNumber="153" type="materials_examined">
<paragraph id="7108511E1021FFAAFF38DAFA9F366FD4" blockId="24.[212,1172,168,328]" box="[212,368,168,192]" pageId="24" pageNumber="153">
<heading id="2A40E6721021FFAAFF38DAFA9F366FD4" box="[212,368,168,192]" fontSize="10" level="3" pageId="24" pageNumber="153" reason="5">
<emphasis id="43C38D0C1021FFAAFF38DAFA9F366FD4" box="[212,368,168,192]" italics="true" pageId="24" pageNumber="153">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="7108511E1021FFAAFF38DA839DB96E5C" blockId="24.[212,1172,168,328]" pageId="24" pageNumber="153">
<materialsCitation id="C1DF5B431021FFAAFF38DA839A1D6E1C" ID-GBIF-Occurrence="2849122365" collectionCode="BMNH" determinerName="G. Frey" location="Wardere Somalia'" municipality="Wardere" pageId="24" pageNumber="153" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="AE0CEFBC1021FFAAFF38DA839F076FFC" box="[212,321,208,232]" pageId="24" pageNumber="153" type="holotype">Holotype</typeStatus>
,
<specimenCount id="67B19A971021FFAAFEA0DA839FD06FFC" box="[332,406,208,232]" pageId="24" pageNumber="153" type="female">female</specimenCount>
with the labels: ‘
<location id="746807C51021FFAAFDB4DA839D626FFC" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C:746807C51021FFAAFDB4DA839D626FFC" box="[600,804,208,232]" municipality="Wardere" name="Wardere Somalia'" pageId="24" pageNumber="153">Wardere Somalia’</location>
, ‘Holotype’, ‘TYPE’, ‘Brit. Mus 1963-277’, ‘
<taxonomicName id="B6B72A9D1021FFAAFEB1DAA39C106E1C" authorityName="Frey" authorityYear="1963" box="[349,598,240,264]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="janssensi">Delopleurus janssensi</taxonomicName>
det.
<determinerName id="84D4BDDA1021FFAAFD70DAA39CA86E1C" box="[668,750,240,264]" pageId="24" pageNumber="153">
G.
<bibRefCitation id="15262CEF1021FFAAFD59DAA39CA86E1C" author="Frey G" box="[693,750,240,264]" pageId="24" pageNumber="153" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey</bibRefCitation>
</determinerName>
1963, n. sp. Typus’ (
<collectionCode id="17A6C9DB1021FFAAFC16DAA39A156E1C" box="[1018,1107,240,264]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="24" pageNumber="153">BMNH</collectionCode>
)
</materialsCitation>
and
<materialsCitation id="C1DF5B431021FFAAFF38DB429DB96E5C" ID-GBIF-Occurrence="3757906302" collectingDate="1943-10" collectionCode="NHMB" country="Somalia" determinerName="G. Frey" location="Delopleurus" pageId="24" pageNumber="153" specimenCount="1" typeStatus="paratype">
<typeStatus id="AE0CEFBC1021FFAAFF38DB429F716E3D" box="[212,311,273,297]" pageId="24" pageNumber="153" type="paratype">paratype</typeStatus>
with the labels: ‘Wardere
<collectingCountry id="09A0118E1021FFAAFD2EDB439D5B6E3C" box="[706,797,272,296]" name="Somalia" pageId="24" pageNumber="153">Somalia</collectingCountry>
<date id="050977DE1021FFAAFCD5DB439DFB6E3C" box="[825,957,272,296]" pageId="24" pageNumber="153" value="1943-10">
<collectingDate id="154D8E361021FFAAFCD5DB439DFB6E3C" box="[825,957,272,296]" pageId="24" pageNumber="153" value="1943-10">Oct. 1943</collectingDate>
</date>
, T.H.E.Jackson’, ‘
<taxonomicName id="B6B72A9D1021FFAAFF37DB639F896E5C" authorityName="Frey" authorityYear="1963" box="[219,463,304,328]" class="Insecta" family="Scarabaeidae" genus="Delopleurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="153" phylum="Arthropoda" rank="species" species="janssensi">
<location id="746807C51021FFAAFF37DB639F226E5C" LSID="urn:lsid:plazi:treatment:F91EE008102EFFAAFE9BDE049DB96E5C:746807C51021FFAAFF37DB639F226E5C" box="[219,356,304,328]" country="Somalia" name="Delopleurus" pageId="24" pageNumber="153">Delopleurus</location>
janssensi
</taxonomicName>
det.
<determinerName id="84D4BDDA1021FFAAFDE6DB639C1A6E5C" box="[522,604,304,328]" pageId="24" pageNumber="153">
G.
<bibRefCitation id="15262CEF1021FFAAFDCFDB639C1A6E5C" author="Frey G" box="[547,604,304,328]" pageId="24" pageNumber="153" pagination="538 - 549" refId="ref9887" refString="Frey G. 1963. Neue Onthophagen (Col. Scarab.) aus Australien (Queenslandausbeute von H. Demarz) und aus Afrika. Ent Arb Mus Frey. 14: 538 - 549." type="journal article" year="1963">Frey</bibRefCitation>
</determinerName>
1963, p. Typus’, ‘Cotype’ (
<collectionCode id="17A6C9DB1021FFAAFC7BDB639DB56E5C" box="[919,1011,304,328]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="24" pageNumber="153">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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