381 lines
43 KiB
XML
381 lines
43 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1084.78744" ID-GBIF-Dataset="fa64803a-85ce-4f7d-8341-f6649b5b6eee" ID-PMC="PMC8825426" ID-Pensoft-Pub="1313-2970-1084-183" ID-Pensoft-UUID="72D7F87358C157C0A26EA3E21B731456" ID-PubMed="35233169" ID-ZooBank="42DE4083042B4F1BACDDD126C67ACC87" ModsDocID="1313-2970-1084-183" checkinTime="1643754958652" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2022" docId="D613257E4D9C5D2A92E2134C97B764E6" docLanguage="en" docName="ZooKeys 1084: 183-199" docOrigin="ZooKeys 1084" docPubDate="2022-02-01" docSource="http://dx.doi.org/10.3897/zookeys.1084.78744" docTitle="Nepalella siamensis Likhitrakarn & Golovatch & Panha 2022, sp. nov." docType="treatment" docUuid="3768467C-2FB4-4E2F-88A8-3977AE5ADDFF" docUuidSource="ZooBank" docVersion="4" id="72D7F87358C157C0A26EA3E21B731456" lastPageNumber="183" masterDocId="72D7F87358C157C0A26EA3E21B731456" masterDocTitle="The Oriental millipede genus Nepalella Shear, 1979, with the description of a new species from Thailand and an updated key (Diplopoda, Chordeumatida, Megalotylidae)" masterLastPageNumber="199" masterPageNumber="183" pageNumber="183" updateTime="1668151447016" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Oriental millipede genus Nepalella Shear, 1979, with the description of a new species from Thailand and an updated key (Diplopoda, Chordeumatida, Megalotylidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation>Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1084</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>183</mods:start>
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<mods:end>199</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1084.78744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1084.78744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1084-183</mods:identifier>
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<mods:identifier type="ZooBank">42DE4083042B4F1BACDDD126C67ACC87</mods:identifier>
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<mods:identifier type="Pensoft-UUID">72D7F87358C157C0A26EA3E21B731456</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="193244743" LSID="urn:lsid:zoobank.org:act:3768467C-2FB4-4E2F-88A8-3977AE5ADDFF" httpUri="http://treatment.plazi.org/id/D613257E4D9C5D2A92E2134C97B764E6" lastPageNumber="183" pageId="0" pageNumber="183">
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<subSubSection pageId="0" pageNumber="183" type="nomenclature">
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<paragraph pageId="0" pageNumber="183">
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<taxonomicName LSID="http://zoobank.org/3768467C-2FB4-4E2F-88A8-3977AE5ADDFF" authority="Likhitrakarn & Golovatch & Panha, 2022" authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">Nepalella siamensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="183">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="183" type="description">
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<paragraph pageId="0" pageNumber="183">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A, B anterior gonopods, front and caudal views, respectively C, D posterior gonopods, caudal and front views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nepalella siamensis sp. nov., SEM ♂ holotype (CUMZ) A, B anterior gonopods, caudal and superior views, respectively C, E posterior gonopods, front and caudal views, respectively D, F left gonopod, front and caudal views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="183" type="holotype">
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<paragraph pageId="0" pageNumber="183">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="183">
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♂ (CUMZ), Thailand, Sa Kaeo Province, Ta Phraya District, Ta Phraya National Park, 183 m a.s.l.,
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<geoCoordinate degrees="14" direction="north" minutes="08" orientation="latitude" precision="15" seconds="22" value="14.139444">14°08'22"N</geoCoordinate>
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,
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<geoCoordinate degrees="102" direction="east" minutes="40" orientation="longitude" precision="15" seconds="11" value="102.669716">102°40'11"E</geoCoordinate>
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, 27.10.2010, leg. N. Likhitrakarn and S.I. Golovatch. The holotype of
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="183">Nepalella siamensis</emphasis>
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</taxonomicName>
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sp. nov. is housed in the Museum of Zoology, Chulalongkorn University (CUMZ), Bangkok, Thailand.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="183" type="etymology">
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<paragraph pageId="0" pageNumber="183">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="183">
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To emphasize
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<normalizedToken originalValue="“Siam”">"Siam"</normalizedToken>
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, referring to the former name of Thailand as the
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<emphasis italics="true" pageId="0" pageNumber="183">terra typica</emphasis>
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; adjective.
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</paragraph>
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<caption doi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="183">
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<emphasis bold="true" pageId="0" pageNumber="183">Figure 2.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="183">Nepalella siamensis</emphasis>
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</taxonomicName>
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sp. nov., ♂ holotype (CUMZ)
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<emphasis bold="true" pageId="0" pageNumber="183">A-C</emphasis>
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anterior part of body, lateral, dorsal and ventral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="183">D-F</emphasis>
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body segments 8-10, sublateral, dorsal and ventral views, respectively
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<emphasis bold="true" pageId="0" pageNumber="183">G-I</emphasis>
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posterior part of body, lateral, dorsal and ventral views, respectively.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="183" type="diagnosis">
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<paragraph pageId="0" pageNumber="183">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="183">
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Differs from the congeners by ♂ femora 3 and 4 each with a small mushroom-like protuberance (
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<emphasis bold="true" pageId="0" pageNumber="183">mp</emphasis>
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) ventrally (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3C</figureCitation>
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); ♂ coxa 10 with a conspicuous horn-shaped process (
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<emphasis bold="true" pageId="0" pageNumber="183">h</emphasis>
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) dorsally (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3E, F</figureCitation>
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); ♂ coxa 11 with a small, medial, digitiform process (
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<emphasis bold="true" pageId="0" pageNumber="183">m</emphasis>
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) and a high, basal, funnel-shaped process (
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<emphasis bold="true" pageId="0" pageNumber="183">b</emphasis>
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) (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3G, H</figureCitation>
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); anterior gonopod sternum carrying a median lobe and two small lateral lobules (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A, B anterior gonopods, front and caudal views, respectively C, D posterior gonopods, caudal and front views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183">4A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nepalella siamensis sp. nov., SEM ♂ holotype (CUMZ) A, B anterior gonopods, caudal and superior views, respectively C, E posterior gonopods, front and caudal views, respectively D, F left gonopod, front and caudal views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183">5A</figureCitation>
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), coupled with posterior gonopod equipped with a foot-shaped colpocoxite (
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<emphasis bold="true" pageId="0" pageNumber="183">c</emphasis>
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) and a rounded bulge (
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<emphasis bold="true" pageId="0" pageNumber="183">r</emphasis>
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) at base in frontal view (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A, B anterior gonopods, front and caudal views, respectively C, D posterior gonopods, caudal and front views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183">4C, D</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nepalella siamensis sp. nov., SEM ♂ holotype (CUMZ) A, B anterior gonopods, caudal and superior views, respectively C, E posterior gonopods, front and caudal views, respectively D, F left gonopod, front and caudal views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183">5C, D</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="183">
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<emphasis bold="true" pageId="0" pageNumber="183">Figure 3.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="183">Nepalella siamensis</emphasis>
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</taxonomicName>
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sp. nov., ♂ holotype (CUMZ)
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<emphasis bold="true" pageId="0" pageNumber="183">A</emphasis>
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gnathochilarium, ventral view
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<emphasis bold="true" pageId="0" pageNumber="183">B</emphasis>
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antenna
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<emphasis bold="true" pageId="0" pageNumber="183">C</emphasis>
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leg 4, caudal view
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<emphasis bold="true" pageId="0" pageNumber="183">D</emphasis>
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leg 7, caudal view
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<emphasis bold="true" pageId="0" pageNumber="183">E</emphasis>
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leg 10, front view
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<emphasis bold="true" pageId="0" pageNumber="183">F</emphasis>
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coxa 10, subcaudal view
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<emphasis bold="true" pageId="0" pageNumber="183">G</emphasis>
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leg 11, front view
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<emphasis bold="true" pageId="0" pageNumber="183">H</emphasis>
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coxa 11, caudal view
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<emphasis bold="true" pageId="0" pageNumber="183">I</emphasis>
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leg 12, caudal view. Abbreviations:
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<emphasis bold="true" pageId="0" pageNumber="183">b</emphasis>
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basal process,
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<emphasis bold="true" pageId="0" pageNumber="183">cg</emphasis>
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coxal gland,
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<emphasis bold="true" pageId="0" pageNumber="183">m</emphasis>
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medial process,
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<emphasis bold="true" pageId="0" pageNumber="183">mp</emphasis>
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mushroom-shaped protuberance,
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<emphasis bold="true" pageId="0" pageNumber="183">p</emphasis>
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parabasal process,
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<emphasis bold="true" pageId="0" pageNumber="183">h</emphasis>
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horn-shaped process. Scale bars: 0.25 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="183" type="description">
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<paragraph pageId="0" pageNumber="183">Description.</paragraph>
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<paragraph pageId="0" pageNumber="183">
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Length of holotype ca 33 mm, maximum width 3.2 mm. Coloration light brown (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A, B, D, E, G, H</figureCitation>
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); head light brown, venter and legs light yellowish to pallid (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2C, F, I</figureCitation>
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). Eye patches and antennae brownish black (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A, C</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="183">
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<emphasis bold="true" pageId="0" pageNumber="183">Figure 4.</emphasis>
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<taxonomicName authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis">
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<emphasis italics="true" pageId="0" pageNumber="183">Nepalella siamensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., ♂ holotype (CUMZ)
|
|
<emphasis bold="true" pageId="0" pageNumber="183">A, B</emphasis>
|
|
anterior gonopods, front and caudal views, respectively
|
|
<emphasis bold="true" pageId="0" pageNumber="183">C, D</emphasis>
|
|
posterior gonopods, caudal and front views, respectively. Abbreviations:
|
|
<emphasis bold="true" pageId="0" pageNumber="183">c</emphasis>
|
|
colpocoxite,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cl</emphasis>
|
|
lateral lobe,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cxi</emphasis>
|
|
coxites,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ll</emphasis>
|
|
lateral lobules,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ml</emphasis>
|
|
median lobe,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">r</emphasis>
|
|
rounded bulge,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t1</emphasis>
|
|
telopoditomere 1,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t2</emphasis>
|
|
telopoditomere 2. Scale bars: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="183">In width, collum <segment 2 <3 <head with genae = segment 4 <5 <6 <7 = 20; thereafter, body very gradually tapering towards telson.</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Body with 30 segments (29 pleurotergites with free sternites, plus telson, or
|
|
<normalizedToken originalValue="“rings”">"rings"</normalizedToken>
|
|
, in terms of
|
|
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1993.tb00305.x" author="Enghoff, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="183" pagination="103 - 234" refId="B2" refString="Enghoff, H, Dohle, W, Blower, JG, 1993. Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations. Zoological Journal of the Linnean Society 109: 103 - 234, DOI: https://doi.org/10.1111/j.1096-3642.1993.tb00305.x" title="Anamorphosis in millipedes (Diplopoda) - the present state of knowledge with some developmental and phylogenetic considerations." url="https://doi.org/10.1111/j.1096-3642.1993.tb00305.x" volume="109" year="1993">Enghoff et al. (1993</bibRefCitation>
|
|
,
|
|
<bibRefCitation DOI="https://doi.org/10.1163/9789004188273_017" author="Enghoff, H" editor="Minelli, A" journalOrPublisher="The Myriapoda 2 (16)" pageId="0" pageNumber="183" pagination="363 - 453" refId="B3" refString="Enghoff, H, Golovatch, SI, Short, M, Stoev, P, Wesener, T, 2015. Diplopoda. In: Minelli, A, Ed., Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda 2(16): 363 - 453, DOI: https://doi.org/10.1163/9789004188273_017" title="Diplopoda." url="https://doi.org/10.1163/9789004188273_017" volumeTitle="Treatise on Zoology-Anatomy, Taxonomy, Biology." year="2015">2015</bibRefCitation>
|
|
)).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Head densely setose, clypeolabral region slightly convex. Eye patches triangular, each composed of 27 and 28 convex ommatidia (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A, C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Antennae very long and slender (Figs
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3A</figureCitation>
|
|
), reaching past body segment 6 when stretched posteriorly; antennomere 7 with four apical cones.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Gnathochilarium without promentum (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Collum as usual (for heterochordeumatoideans), obcordate in shape, with rudimentary paraterga (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A</figureCitation>
|
|
). Tegument smooth, shining, only prozonae distinctly and densely striolate transversely (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2D, E</figureCitation>
|
|
). Metatergal setation 3 + 3, typical of
|
|
<taxonomicName authorityName="Shear" authorityYear="1979" class="Diplopoda" kingdom="Animalia" lsidName="" order="Chordeumatida" pageId="0" pageNumber="183" rank="order">Chordeumatida</taxonomicName>
|
|
; macrochaetae long, rather thick, pointed, placed on clear knobs (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A, B, D, E, G, H</figureCitation>
|
|
); stricture between pro- and metazona shallow, inconspicuous (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2A, D, E, G, H</figureCitation>
|
|
). Paraterga poorly developed, with small dorsolateral bulges in anterior part of body (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2B</figureCitation>
|
|
), following segments rather regularly rounded in dorsal view (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2D, E, H</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
CIX (ring 15) = 0.62; MIX (ring 15) = 0.87; MA (ring 15) ≈ 145°; PIX impossible to evaluate due to insufficiently developed paraterga. Axial suture distinct, pallid, as usual (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A-C anterior part of body, lateral, dorsal and ventral views, respectively D-F body segments 8 - 10, sublateral, dorsal and ventral views, respectively G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.1084.78744.figure2" httpUri="https://binary.pensoft.net/fig/640691" pageId="0" pageNumber="183">2B, D, E, H</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
♂ legs long and slender, ca 1.5 times as long as midbody height. Legs 1 and 2 slightly reduced, tarsi with usual ventral brushes, but without papillae; ♂ coxa 2 with a distal mediocaudal cone perforated by gonopore orifice. All following legs conspicuously papillate on ventral face of tarsi (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3C, D, E, G, I</figureCitation>
|
|
). ♂ legs 3-7 distinctly and increasingly crassate, pairs 3 and 4 particularly so. Femora 3 and 4 each with a small, but evident mushroom-shaped protuberance (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">mp</emphasis>
|
|
) at midway ventrally (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3C</figureCitation>
|
|
). Coxa 7 with a small, but evident distoventral digitiform outgrowth (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
♂ legs 10 and 11 each with a small coxal gland (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cg</emphasis>
|
|
) (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3E-H</figureCitation>
|
|
); each coxa 10 dorsally with a large horn-shaped process (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">h</emphasis>
|
|
) conspicuously enlarged at base (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3E, F</figureCitation>
|
|
); each coxa 11 with a small, medial, digitiform process (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">m</emphasis>
|
|
) and a high, basal, funnel-shaped process (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">b</emphasis>
|
|
) (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3G, H</figureCitation>
|
|
); prefemur 11 with a small parabasal process (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">p</emphasis>
|
|
) ventrally (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A gnathochilarium, ventral view B antenna C leg 4, caudal view D leg 7, caudal view E leg 10, front view F coxa 10, subcaudal view G leg 11, front view H coxa 11, caudal view I leg 12, caudal view. Abbreviations: b basal process, cg coxal gland, m medial process, mp mushroom-shaped protuberance, p parabasal process, h horn-shaped process. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1084.78744.figure3" httpUri="https://binary.pensoft.net/fig/640692" pageId="0" pageNumber="183">3G, H</figureCitation>
|
|
). Claws simple, rather long.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Anterior gonopods (♂ leg-pair 8) very strongly reduced, sternum with a median lobe (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ml</emphasis>
|
|
) distally in oral view and with two small lateral lobules (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ll</emphasis>
|
|
); coxites (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cxi</emphasis>
|
|
) long, slender and horn-shaped (Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A, B anterior gonopods, front and caudal views, respectively C, D posterior gonopods, caudal and front views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183">4A, B</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nepalella siamensis sp. nov., SEM ♂ holotype (CUMZ) A, B anterior gonopods, caudal and superior views, respectively C, E posterior gonopods, front and caudal views, respectively D, F left gonopod, front and caudal views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183">5A, B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
|
|
Posterior gonopods (♂ leg-pair 9) (Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Nepalella siamensis sp. nov., ♂ holotype (CUMZ) A, B anterior gonopods, front and caudal views, respectively C, D posterior gonopods, caudal and front views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure4" httpUri="https://binary.pensoft.net/fig/640693" pageId="0" pageNumber="183">4C, D</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Nepalella siamensis sp. nov., SEM ♂ holotype (CUMZ) A, B anterior gonopods, caudal and superior views, respectively C, E posterior gonopods, front and caudal views, respectively D, F left gonopod, front and caudal views, respectively. Abbreviations: c colpocoxite, cl lateral lobe, cxi coxites, ll lateral lobules, ml median lobe, r rounded bulge, t 1 telopoditomere 1, t 2 telopoditomere 2. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183">5C, D, E, F</figureCitation>
|
|
) hypertrophied, each with a prominent, foot-shaped colpocoxite (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">c</emphasis>
|
|
), this being higher than telopodite, and with three evident longitudinal lamellae in caudal view; a rather conspicuous lateral lobe (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cl</emphasis>
|
|
) at midway in caudal view; with a rounded bulge (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">r</emphasis>
|
|
) at base in frontal view; telopoditomere 1 (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t1</emphasis>
|
|
) particularly strongly setose on posterior face, expanded apically, telopoditomere 2 (
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t2</emphasis>
|
|
) subpyriform, likewise voluminous, only slightly setose laterally in basal half.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1084.78744.figure5" httpUri="https://binary.pensoft.net/fig/640694" pageId="0" pageNumber="183" start="Figure 5" startId="F5">
|
|
<paragraph pageId="0" pageNumber="183">
|
|
<emphasis bold="true" pageId="0" pageNumber="183">Figure 5.</emphasis>
|
|
<taxonomicName authorityName="Likhitrakarn & Golovatch & Panha" authorityYear="2022" class="Diplopoda" family="Megalotylidae" genus="Nepalella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nepalella siamensis" order="Chordeumatida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="siamensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="183">Nepalella siamensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov., SEM ♂ holotype (CUMZ)
|
|
<emphasis bold="true" pageId="0" pageNumber="183">A, B</emphasis>
|
|
anterior gonopods, caudal and superior views, respectively
|
|
<emphasis bold="true" pageId="0" pageNumber="183">C, E</emphasis>
|
|
posterior gonopods, front and caudal views, respectively
|
|
<emphasis bold="true" pageId="0" pageNumber="183">D, F</emphasis>
|
|
left gonopod, front and caudal views, respectively. Abbreviations:
|
|
<emphasis bold="true" pageId="0" pageNumber="183">c</emphasis>
|
|
colpocoxite,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cl</emphasis>
|
|
lateral lobe,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">cxi</emphasis>
|
|
coxites,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ll</emphasis>
|
|
lateral lobules,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">ml</emphasis>
|
|
median lobe,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">r</emphasis>
|
|
rounded bulge,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t1</emphasis>
|
|
telopoditomere 1,
|
|
<emphasis bold="true" pageId="0" pageNumber="183">t2</emphasis>
|
|
telopoditomere 2. Scale bars: 0.2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="183" type="remarks">
|
|
<paragraph pageId="0" pageNumber="183">Remark.</paragraph>
|
|
<paragraph pageId="0" pageNumber="183">
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The specimen was collected by hand while it was moving very fast on the leaf litter surface. The type locality is situated in a dipterocarp forest on the side of a road near the Ta Phraya Waterfall. The species was found syntopically together with
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<taxonomicName baseAuthorityName="Attems" baseAuthorityYear="1931" class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha uncinata" order="Spirostreptida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="uncinata">
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<emphasis italics="true" pageId="0" pageNumber="183">Antheromorpha uncinata</emphasis>
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</taxonomicName>
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(Attems, 1931) (
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<taxonomicName authorityName="Daday" authorityYear="1889" class="Diplopoda" family="Paradoxosomatidae" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="family">Paradoxosomatidae</taxonomicName>
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,
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<taxonomicName authorityName="Jeekel" authorityYear="1968" class="Diplopoda" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="183" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
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) (
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.571.7566" author="Likhitrakarn, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="183" pagination="21 - 57" refId="B11" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2016. Review of the Southeast Asian millipede genus Antheromorpha Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 571: 21 - 57, DOI: https://doi.org/10.3897/zookeys.571.7566" title="Review of the Southeast Asian millipede genus Antheromorpha Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.571.7566" volume="571" year="2016">Likhitrakarn et al. 2016</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |