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<document id="FC65BA6E22B3F036928FB9A8E4FAD7AA" ID-CLB-Dataset="26038" ID-DOI="10.11646/zootaxa.4653.1.1" ID-GBIF-Dataset="2429a921-404f-482c-8f90-741676225442" ID-ISSN="1175-5326" ID-Zenodo-Dep="3366542" ID-ZooBank="FCDFD71B-D7AD-480D-A8CA-18297FD8D5EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565679017730" checkinUser="plazi" docAuthor="Meshram, Naresh M." docDate="2019" docId="CD4987FE4A7DE00CFF0CFA5DFB9CF8F4" docLanguage="en" docName="zootaxa.4653.1.1.pdf" docOrigin="Zootaxa 4653 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mahellus ungulatus Viraktamath &amp; Meshram. 2017" docType="treatment" docVersion="19" lastPageNumber="18" masterDocId="3170FF864A6CE01DFF9BFF9CFFA0FFEE" masterDocTitle="Leafhopper tribe Coelidiini (Hemiptera: Cicadellidae: Coelidiinae) of the Indian subcontinent" masterLastPageNumber="91" masterPageNumber="1" pageNumber="18" updateTime="1698688541026" updateUser="plazi">
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<mods:title id="2263ED5D31CBC8184954084D1052AA99">Leafhopper tribe Coelidiini (Hemiptera: Cicadellidae: Coelidiinae) of the Indian subcontinent</mods:title>
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<mods:namePart id="3C0C02A64E59C7EA30FECAAD768C0CC0">Meshram, Naresh M.</mods:namePart>
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<mods:date id="ABE60E745DE41EE80555A4F46AF7AF02">2019</mods:date>
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<mods:number id="81057EB4FB1F9AE85E3FDF62D36652B9">2019-08-12</mods:number>
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<treatment id="CD4987FE4A7DE00CFF0CFA5DFB9CF8F4" ID-DOI="http://doi.org/10.5281/zenodo.5191080" ID-GBIF-Taxon="159125923" ID-Zenodo-Dep="5191080" LSID="urn:lsid:plazi:treatment:CD4987FE4A7DE00CFF0CFA5DFB9CF8F4" httpUri="http://treatment.plazi.org/id/CD4987FE4A7DE00CFF0CFA5DFB9CF8F4" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="0DFA65634A7DE00CFF0CFA5DFD74FA32" box="[151,724,1473,1500]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="455F36E84A7DE00CFF0CFA5DFD74FA32" blockId="17.[151,724,1473,1535]" box="[151,724,1473,1500]" pageId="17" pageNumber="18">
<heading id="1E1781844A7DE00CFF0CFA5DFD74FA32" bold="true" box="[151,724,1473,1500]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="82E04D6B4A7DE00CFF0CFA5DFD74FA32" ID-CoL="8JWDT" authority="Viraktamath &amp; Meshram" authorityName="Viraktamath &amp; Meshram." authorityYear="2017" box="[151,724,1473,1500]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ungulatus">
<emphasis id="7794EAFA4A7DE00CFF0CFA5DFD74FA32" bold="true" box="[151,724,1473,1500]" pageId="17" pageNumber="18">
<emphasis id="7794EAFA4A7DE00CFF0CFA5DFE2BFA32" bold="true" box="[151,395,1473,1500]" italics="true" pageId="17" pageNumber="18">Mahellus ungulatus</emphasis>
Viraktamath &amp; Meshram
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DFA65634A7DE00CFF0CFA79FE9BFA11" box="[151,315,1509,1535]" pageId="17" pageNumber="18" type="description">
<paragraph id="455F36E84A7DE00CFF0CFA79FE9BFA11" blockId="17.[151,724,1473,1535]" box="[151,315,1509,1535]" pageId="17" pageNumber="18">
<figureCitation id="DDDB2A6D4A7DE00CFF0CFA79FF51FA11" box="[151,241,1509,1535]" captionStart="FIGURES 28" captionStartId="63.[151,264,1851,1876]" captionTargetBox="[176,1379,207,1822]" captionTargetId="figure@63.[166,1421,181,1831]" captionTargetPageId="63" captionText="FIGURES 28A–M. Species of Mahellus Nielson, male genitalia. M. distanti Nielson: A. Pygofer, lateral view; B. Subgenital plate, ventral view; C. Style, dorsal view; D. Aedeagus, lateral view; E.Aedeagus dorsal view; F. Apex of aedeagal shaft, dorsal view. M. ungulatus Viraktamath &amp; Meshram: G. Pygofer, lateral view, H. Pygofer process, lateral view; J. Style dorsal view; K. Aedeagus, lateral view; L. Aedeagus, dorsal view; M. Apex of aedeagal shaft, dorsal view. Modified and reproduced from Viraktamath &amp; Meshram (2017)." httpUri="https://zenodo.org/record/3366600/files/figure.png" pageId="17" pageNumber="18">Figs. 28</figureCitation>
G–M.
</paragraph>
</subSubSection>
<subSubSection id="0DFA65634A7DE00CFF0CF9B6FBB1F9AC" box="[151,1041,1577,1602]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="455F36E84A7DE00CFF0CF9B6FBB1F9AC" blockId="17.[151,1041,1577,1602]" box="[151,1041,1577,1602]" pageId="17" pageNumber="18">
<taxonomicName id="82E04D6B4A7DE00CFF0CF9B6FCA6F9AC" ID-CoL="8JWDT" authority="Viraktamath &amp; Meshram 2017: 279 - 280" authorityName="Viraktamath &amp; Meshram." authorityPageNumber="279" authorityYear="2017" box="[151,774,1577,1602]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ungulatus">
<emphasis id="7794EAFA4A7DE00CFF0CF9B6FEC0F9AC" box="[151,352,1578,1602]" italics="true" pageId="17" pageNumber="18">Mahellus ungulatus</emphasis>
<bibRefCitation id="21714B194A7DE00CFEFDF9B5FCA6F9AC" author="Viraktamath, C. A. &amp; Meshram, N. M." box="[358,774,1577,1602]" pageId="17" pageNumber="18" pagination="271 - 280" refId="ref39134" refString="Viraktamath, C. A. &amp; Meshram, N. M. (2017) A review of the coelidiine leafhopper genus Mahellus (Hemiptera: Cicadellidae: Coelidiinae) with description of two new species from the Oriental region. Zootaxa, 4258 (3), 271 - 280 https: // doi. org / 10.11646 / zootaxa. 4258.3.4" type="journal article" year="2017">Viraktamath &amp; Meshram 2017: 279−280</bibRefCitation>
</taxonomicName>
,
<figureCitation id="DDDB2A6D4A7DE00CFC8BF9B5FCD3F9AC" box="[784,883,1577,1602]" captionStart-0="FIGURES 8" captionStart-1="FIGURES 9" captionStartId-0="43.[151,264,1859,1884]" captionStartId-1="44.[151,264,1879,1904]" captionTargetBox-0="[166,1420,295,1729]" captionTargetBox-1="[171,1415,283,1778]" captionTargetId-0="figure@43.[151,1435,254,1729]" captionTargetId-1="figure@44.[151,1435,237,1793]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="FIGURES 8A–P. Faces of Coelidiini species.A &amp; B. Olidiana brevis (Walker), male and female; C. O. fletcheri sp. nov., male; D. O. indica (Walker), male; E &amp; F. O. kirkaldyi (Nielson), male and female; G. O. lanceolata sp. nov., male; H. O. perculta (Distant), male; I. O. serra (Neilson), male; J &amp; K, O. umroensis sp. nov., male and female; L. O. unidenta sp. nov., male; M. Singillatus parapectitus sp. nov., male; N &amp; O. S. serratispatulatus sp. nov., male and female; P. Trinoridia dialata sp. nov., male." captionText-1="FIGURES 9 A–M. Faces of Coelidiini species. A. Trinoridia dialata sp. nov., female; B. T. ochrocephala sp. nov., male; C &amp; D. T. piperica sp. nov., male and female; E &amp; F. T. ramamurthyi sp. nov., male and female; G. T. saraikela sp. nov., male; H. T. timlivana sp. nov., male; I. T. tripectinata (Nielson), male; J &amp; K. Webbolidia andamana sp. nov., male and female; L. W. burmanica sp. nov., male; M. Zhangolidia weicongi sp. nov., male." httpUri-0="https://zenodo.org/record/3366560/files/figure.png" httpUri-1="https://zenodo.org/record/3366562/files/figure.png" pageId="17" pageNumber="18">Figs. 8−9</figureCitation>
,
<figureCitation id="DDDB2A6D4A7DE00CFCE6F9B6FC38F9AC" box="[893,920,1578,1602]" captionStart="FIGURES 17" captionStartId="52.[151,264,1965,1990]" captionTargetBox="[181,1370,194,1945]" captionTargetId="figure@52.[164,1423,166,1945]" captionTargetPageId="52" captionText="FIGURES 17 A–H. Calodia kodikanelensis Nielson, male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366578/files/figure.png" pageId="17" pageNumber="18">17</figureCitation>
,
<figureCitation id="DDDB2A6D4A7DE00CFC38F9B6FC46F9AC" box="[931,998,1578,1602]" captionStart-0="FIGURES 40" captionStart-1="FIGURES 41" captionStart-2="FIGURES 42" captionStart-3="FIGURES 43" captionStart-4="FIGURES 44" captionStart-5="FIGURES 45" captionStart-6="FIGURES 46" captionStart-7="FIGURES 47" captionStartId-0="75.[151,264,1929,1954]" captionStartId-1="76.[151,264,1970,1995]" captionStartId-2="77.[151,264,1934,1959]" captionStartId-3="78.[151,264,1924,1949]" captionStartId-4="79.[151,264,1964,1989]" captionStartId-5="80.[151,264,1964,1989]" captionStartId-6="81.[151,264,1968,1993]" captionStartId-7="82.[151,264,1983,2008]" captionTargetBox-0="[206,1398,207,1885]" captionTargetBox-1="[187,1380,191,1938]" captionTargetBox-2="[174,1356,188,1874]" captionTargetBox-3="[156,1394,203,1876]" captionTargetBox-4="[209,1363,176,1922]" captionTargetBox-5="[189,1378,191,1932]" captionTargetBox-6="[169,1395,181,1926]" captionTargetBox-7="[165,1373,199,1944]" captionTargetId-0="figure@75.[157,1429,181,1909]" captionTargetId-1="figure@76.[162,1425,166,1950]" captionTargetId-2="figure@77.[174,1412,166,1915]" captionTargetId-3="figure@78.[148,1419,181,1905]" captionTargetId-4="figure@79.[164,1423,166,1944]" captionTargetId-5="figure@80.[164,1423,166,1944]" captionTargetId-6="figure@81.[163,1425,166,1948]" captionTargetId-7="figure@82.[165,1423,179,1955]" captionTargetPageId-0="75" captionTargetPageId-1="76" captionTargetPageId-2="77" captionTargetPageId-3="78" captionTargetPageId-4="79" captionTargetPageId-5="80" captionTargetPageId-6="81" captionTargetPageId-7="82" captionText-0="FIGURES 40 A–P. Species of Orbisolidia Nielson and Singillatus Nielson, male genitalia. A–D. Orbisolidia paracava (Nielson); E–H. O. spinocava (Nielson); I–L. Singillatus curtus (Nielson); M–P. S. furcatus (Nielson): Modified and reproduced from Nielson 1982 (Figs. A–L) and Nielson 1991 (Figs. M–P)." captionText-1="FIGURES 41 A–G. Singillatus parapectitus sp. nov., male genitalia.A &amp; B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view." captionText-2="FIGURES 42 A–G. Singillatus serratispatulatus sp. nov., male genitalia. A. Aedeagus, dorsal view; B. Aedeagus, lateral view; C. Subgenital plate, dorsal view; D. Style, dorsal view; E. Connective, dorsal view; F. Pygofer ventral process, lateral view. G. Pygofer, lateral view." captionText-3="FIGURES 43 A–Q. Species of Singillatus Nielson and Trinoridia Nielson. A–D. S. singularis (Nielson); E–I. S. ventrospinatus Nielson; J–M. Trinoridia calcaris Nielson. N–Q. T. rama (Kirkaldy). Modified and reproduced from Nielson 1991 (Figs.A–D), Nielson 2015 (Figs. I–M) and Nielson 1982 (Figs. N–Q)." captionText-4="FIGURES 44 A–G. Trinoridia dialata sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view." captionText-5="FIGURES 45 A–G. Trinoridia ochrocephala sp. nov., male genitalia.A.Aedeagus, dorsal view; B &amp; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view." captionText-6="FIGURES 46 A–H. Trinoridia piperica sp. nov., male genitalia.A &amp; B. Aedeagus, dorsal view; C &amp; D.Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." captionText-7="FIGURES 47 A–G. Trinoridia ramamuthyi sp. nov., male genitalia.A &amp; B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view." httpUri-0="https://zenodo.org/record/3366624/files/figure.png" httpUri-1="https://zenodo.org/record/3366626/files/figure.png" httpUri-2="https://zenodo.org/record/3366628/files/figure.png" httpUri-3="https://zenodo.org/record/3366630/files/figure.png" httpUri-4="https://zenodo.org/record/3366632/files/figure.png" httpUri-5="https://zenodo.org/record/3366634/files/figure.png" httpUri-6="https://zenodo.org/record/3366636/files/figure.png" httpUri-7="https://zenodo.org/record/3366638/files/figure.png" pageId="17" pageNumber="18">40−47</figureCitation>
,
<figureCitation id="DDDB2A6D4A7DE00CFC6BF9B5FBACF9AC" box="[1008,1036,1577,1602]" captionStart="FIGURES 52" captionStartId="87.[151,264,1939,1964]" captionTargetBox="[195,1387,191,1911]" captionTargetId="figure@87.[173,1415,166,1920]" captionTargetPageId="87" captionText="FIGURES 52 A–J. Webbolidia burmanica sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view; I &amp; J. Dorsal pygofer process, lateral views." httpUri="https://zenodo.org/record/3366648/files/figure.png" pageId="17" pageNumber="18">52</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DFA65634A7DE00CFF0CF9ECFB9CF8F4" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="455F36E84A7DE00CFF0CF9ECFAFCF964" blockId="17.[151,1436,1648,1819]" box="[151,1372,1648,1675]" pageId="17" pageNumber="18">
<emphasis id="7794EAFA4A7DE00CFF0CF9ECFED3F964" box="[151,371,1648,1674]" italics="true" pageId="17" pageNumber="18">Material examined.</emphasis>
<materialsCitation id="F5883CB54A7DE00CFEE2F9ECFAFCF964" ID-GBIF-Occurrence="2332229223" box="[377,1372,1648,1675]" collectingDate="1981-11-10" collectionCode="USAB" collectorName="S. Viraktamath Coll." country="India" location="Sukna" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="holotype">
<typeStatus id="9A5B884A4A7DE00CFEE2F9ECFDADF964" box="[377,525,1648,1674]" pageId="17" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="53E6FD614A7DE00CFD8FF9EDFD8BF965" box="[532,555,1649,1675]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
,
<collectingCountry id="3DF776784A7DE00CFDACF9ECFD2BF964" box="[567,651,1648,1674]" name="India" pageId="17" pageNumber="18">INDIA</collectingCountry>
:
<collectingRegion id="8724F80A4A7DE00CFD0EF9EDFC87F964" box="[661,807,1648,1675]" country="India" name="West Bengal" pageId="17" pageNumber="18">West Bengal</collectingRegion>
:
<location id="403F60334A7DE00CFCAAF9ECFCDAF964" LSID="urn:lsid:plazi:treatment:CD4987FE4A7DE00CFF0CFA5DFB9CF8F4:403F60334A7DE00CFCAAF9ECFCDAF964" box="[817,890,1648,1674]" country="India" name="Sukna" pageId="17" pageNumber="18" stateProvince="West Bengal">Sukna</location>
,
<date id="315E10284A7DE00CFC1EF9ECFBA0F964" box="[901,1024,1648,1674]" pageId="17" pageNumber="18" value="1981-11-10">
<collectingDate id="211AE9C04A7DE00CFC1EF9ECFBA0F964" box="[901,1024,1648,1674]" pageId="17" pageNumber="18" value="1981-11-10">10.xi.1981</collectingDate>
</date>
,
<collectorName id="E815533E4A7DE00CFB90F9ECFB51F964" box="[1035,1265,1648,1674]" pageId="17" pageNumber="18">S. Viraktamath Coll.</collectorName>
(
<collectionCode id="23F1AE2D4A7DE00CFA9BF9ECFAF1F964" box="[1280,1361,1648,1674]" pageId="17" pageNumber="18">USAB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="455F36E84A7DE00CFF5CF908FB9CF8F4" blockId="17.[151,1436,1648,1819]" pageId="17" pageNumber="18">
<emphasis id="7794EAFA4A7DE00CFF5CF908FE94F940" bold="true" box="[199,308,1684,1710]" pageId="17" pageNumber="18">Remarks</emphasis>
.
<bibRefCitation id="21714B194A7DE00CFEDFF908FD16F940" author="Viraktamath, C. A. &amp; Meshram, N. M." box="[324,694,1684,1710]" pageId="17" pageNumber="18" pagination="271 - 280" refId="ref39134" refString="Viraktamath, C. A. &amp; Meshram, N. M. (2017) A review of the coelidiine leafhopper genus Mahellus (Hemiptera: Cicadellidae: Coelidiinae) with description of two new species from the Oriental region. Zootaxa, 4258 (3), 271 - 280 https: // doi. org / 10.11646 / zootaxa. 4258.3.4" type="journal article" year="2017">Viraktamath &amp; Meshram (2017)</bibRefCitation>
adequately described and illustrated the species based on single male collected from Sukna (
<collectingRegion id="8724F80A4A7DE00CFE48F924FDC8F93D" box="[467,616,1720,1747]" country="India" name="West Bengal" pageId="17" pageNumber="18">West Bengal</collectingRegion>
). It is so far known from northeast
<collectingCountry id="3DF776784A7DE00CFC68F924FB8DF93C" box="[1011,1069,1720,1746]" name="India" pageId="17" pageNumber="18">India</collectingCountry>
only. It differs from
<taxonomicName id="82E04D6B4A7DE00CFA83F925FEE0F919" authority="(Distant)" baseAuthorityName="Distant" class="Insecta" family="Cicadellidae" genus="Jassus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="determinatus">
<emphasis id="7794EAFA4A7DE00CFA83F925FF74F918" italics="true" pageId="17" pageNumber="18">M. determinatus</emphasis>
(Distant)
</taxonomicName>
in having a much narrower apophysis of the style (
<figureCitation id="DDDB2A6D4A7DE00CFC11F940FC50F919" box="[906,1008,1756,1783]" captionStart="FIGURES 28" captionStartId="63.[151,264,1851,1876]" captionTargetBox="[176,1379,207,1822]" captionTargetId="figure@63.[166,1421,181,1831]" captionTargetPageId="63" captionText="FIGURES 28A–M. Species of Mahellus Nielson, male genitalia. M. distanti Nielson: A. Pygofer, lateral view; B. Subgenital plate, ventral view; C. Style, dorsal view; D. Aedeagus, lateral view; E.Aedeagus dorsal view; F. Apex of aedeagal shaft, dorsal view. M. ungulatus Viraktamath &amp; Meshram: G. Pygofer, lateral view, H. Pygofer process, lateral view; J. Style dorsal view; K. Aedeagus, lateral view; L. Aedeagus, dorsal view; M. Apex of aedeagal shaft, dorsal view. Modified and reproduced from Viraktamath &amp; Meshram (2017)." httpUri="https://zenodo.org/record/3366600/files/figure.png" pageId="17" pageNumber="18">Fig. 28 J</figureCitation>
) and from
<taxonomicName id="82E04D6B4A7DE00CFBEBF941FEA1F8F4" authority="Viraktamath &amp; Meshram" authorityName="Viraktamath &amp; Meshram" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis id="7794EAFA4A7DE00CFBEBF941FB4CF918" box="[1136,1260,1756,1782]" italics="true" pageId="17" pageNumber="18">M. cardoni</emphasis>
Viraktamath &amp; Meshram
</taxonomicName>
in the curvature of the dorsal marginal sclerotized process of the pygofer.
</paragraph>
</subSubSection>
</treatment>
</document>
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