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treatments-xml/data/A1/57/FD/A157FD05FFAF4B56FF2FFC37FEF7249D.xml
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<document id="874E45BA1B6FCF577E2876C4ED76F3C1" ID-CLB-Dataset="8167" ID-DOI="10.11646/zootaxa.4926.4.1" ID-GBIF-Dataset="f0b1c09e-298d-497d-b6de-cd74367c983e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4529512" ID-ZooBank="3ADD4A2B-A3F9-4379-A8FE-39DD867531F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612954843127" checkinUser="plazi" docAuthor="Fitzgerald, Scott J." docDate="2021" docId="A157FD05FFAF4B56FF2FFC37FEF7249D" docLanguage="en" docName="zootaxa.4926.4.1.pdf" docOrigin="Zootaxa 4926 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penthetria dolichopeza Fitzgerald 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="5D6E857DFFBD4B45FFB8FFBDFF98266C" masterDocTitle="Penthetria Meigen (Diptera: Bibionidae): Revision of the New World species and world catalog" masterLastPageNumber="500" masterPageNumber="451" pageNumber="469" updateTime="1698928650180" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="63476327C5CD775EC255B5C66F7912AE">Penthetria Meigen (Diptera: Bibionidae): Revision of the New World species and world catalog</mods:title>
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<mods:namePart id="F6078288B4D08E3C92D0E6BB8B13F058">Fitzgerald, Scott J.</mods:namePart>
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<mods:title id="C65A13196D47E0D94DC6B4FEC8495DFC">Zootaxa</mods:title>
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<mods:date id="A5BEC71FB3878491436DBB439191005E">2021</mods:date>
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<mods:number id="65B97C840C0EF682C659A1E3D0912B6B">2021-02-10</mods:number>
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<mods:number id="9A0334ECE7B8EC91D2A7E8CC304E5EFC">4926</mods:number>
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<mods:classification id="D013DB5053EAA58340B51008189C4222">journal article</mods:classification>
<mods:identifier id="B64C381B6783579C63A837F9CD9A3C3F" type="CLB-Dataset">8167</mods:identifier>
<mods:identifier id="5F73595D7C7E44A4BE9B4C858AC5472D" type="DOI">10.11646/zootaxa.4926.4.1</mods:identifier>
<mods:identifier id="72AEBB8B08C3E30BBD5ACCCBDF3827D5" type="GBIF-Dataset">f0b1c09e-298d-497d-b6de-cd74367c983e</mods:identifier>
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<treatment id="A157FD05FFAF4B56FF2FFC37FEF7249D" ID-DOI="http://doi.org/10.5281/zenodo.4529524" ID-GBIF-Taxon="177422184" ID-Zenodo-Dep="4529524" LSID="urn:lsid:plazi:treatment:A157FD05FFAF4B56FF2FFC37FEF7249D" httpUri="http://treatment.plazi.org/id/A157FD05FFAF4B56FF2FFC37FEF7249D" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<subSubSection id="61E41F98FFAF4B57FF2FFC37FDE725C9" box="[151,639,906,933]" pageId="18" pageNumber="469" type="nomenclature">
<paragraph id="29414C13FFAF4B57FF2FFC37FDE725C9" blockId="18.[151,639,906,969]" box="[151,639,906,933]" pageId="18" pageNumber="469">
<heading id="7209FB7FFFAF4B57FF2FFC37FDE725C9" bold="true" box="[151,639,906,933]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="1B8A9001FFAF4B57FF2FFC37FDE725C9" bold="true" box="[151,639,906,933]" pageId="18" pageNumber="469">
<taxonomicName id="EEFE3790FFAF4B57FF2FFC37FDAD25C8" ID-CoL="8GTBX" authority="Fitzgerald" authorityName="Fitzgerald" authorityYear="2021" box="[151,565,906,933]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dolichopeza" status="sp. nov.">
<emphasis id="1B8A9001FFAF4B57FF2FFC37FE3525C9" bold="true" box="[151,429,906,933]" italics="true" pageId="18" pageNumber="469">Penthetria dolichopeza</emphasis>
Fitzgerald
</taxonomicName>
<taxonomicNameLabel id="00B92D7AFFAF4B57FD84FC36FDE725C9" box="[572,639,907,933]" pageId="18" pageNumber="469" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAF4B57FF2FFC12FE3725A5" box="[151,431,942,969]" pageId="18" pageNumber="469" type="description">
<paragraph id="29414C13FFAF4B57FF2FFC12FE3725A5" blockId="18.[151,639,906,969]" box="[151,431,942,969]" pageId="18" pageNumber="469">
(
<figureCitation id="B1C55096FFAF4B57FF27FC12FEBF25A5" box="[159,295,942,969]" captionStart-0="FIGURES 12–13" captionStart-1="FIGURES 14–15" captionStartId-0="19.[151,264,1948,1972]" captionStartId-1="20.[151,264,1956,1980]" captionTargetBox-0="[193,1398,784,1919]" captionTargetBox-1="[389,1201,188,1928]" captionTargetId-0="figure-280@19.[187,1401,777,1924]" captionTargetId-1="figure-34@20.[382,1205,181,1932]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." captionText-1="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi-0="http://doi.org/10.5281/zenodo.4529528" figureDoi-1="http://doi.org/10.5281/zenodo.4529530" httpUri-0="https://zenodo.org/record/4529528/files/figure.png" httpUri-1="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Figs. 12–15</figureCitation>
,
<figureCitation id="B1C55096FFAF4B57FE8BFC12FEC925A5" box="[307,337,943,969]" captionStart="FIGURES 42–45" captionStartId="34.[151,264,1889,1913]" captionTargetBox="[167,1419,240,1859]" captionTargetId="figure-64@34.[177,1410,666,1865]" captionTargetPageId="34" captionText="FIGURES 42–45. Male terminalia, dorsal, tergite nine removed. 42. P. distincta. 43. P. dolichopeza n. sp., paratype. 44. P. appendicula. 45. P. neonigrita n. sp., paratype. Scale bar = ca. 0.5 mm. Abbreviations: dlp, divergent lobes of paramere; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; vml, ventromedian lobes of gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.4529558" httpUri="https://zenodo.org/record/4529558/files/figure.png" pageId="18" pageNumber="469">43</figureCitation>
,
<figureCitation id="B1C55096FFAF4B57FEE5FC12FE3E25A5" box="[349,422,943,969]" captionStart="MAP 2" captionStartId="39.[151,209,1832,1856]" captionTargetBox="[167,1424,945,1781]" captionTargetId="figure-37@39.[157,1431,936,1789]" captionText="MAP 2. Distribution of P. arizonensis (black circle) and P. dolichopeza (white circle)." figureDoi="http://doi.org/10.5281/zenodo.4546086" httpUri="https://zenodo.org/record/4546086/files/figure.png" pageId="18" pageNumber="469">Map 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAF4B57FF2FFC4AFC172211" pageId="18" pageNumber="469" type="materials_examined">
<paragraph id="29414C13FFAF4B57FF2FFC4AFC172211" blockId="18.[151,1437,1014,2013]" pageId="18" pageNumber="469">
<emphasis id="1B8A9001FFAF4B57FF2FFC4AFEDA227C" bold="true" box="[151,322,1014,1041]" pageId="18" pageNumber="469">Type Material</emphasis>
.
<materialsCitation id="9996464EFFAF4B57FEEAFC4AFCF42259" ID-GBIF-Occurrence="3037447332" collectingDate="1995-05-05" collectionCode="BYUC" collectorName="Baumann &amp; Houseman" country="Costa Rica" county="Male" location="La Abigail" municipality="Nubes de Coronado" pageId="18" pageNumber="469" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<emphasis id="1B8A9001FFAF4B57FEEAFC4AFE27227D" bold="true" box="[338,447,1015,1041]" pageId="18" pageNumber="469">
<typeStatus id="F645F2B1FFAF4B57FEEAFC4AFE27227D" box="[338,447,1015,1041]" pageId="18" pageNumber="469" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCounty id="C020349FFFAF4B57FE68FC4AFD92227D" box="[464,522,1015,1041]" pageId="18" pageNumber="469">Male</collectingCounty>
(
<figureCitation id="B1C55096FFAF4B57FDA2FC4AFD3D227D" box="[538,677,1014,1041]" captionStart-0="FIGURES 12–13" captionStart-1="FIGURES 14–15" captionStartId-0="19.[151,264,1948,1972]" captionStartId-1="20.[151,264,1956,1980]" captionTargetBox-0="[193,1398,784,1919]" captionTargetBox-1="[389,1201,188,1928]" captionTargetId-0="figure-280@19.[187,1401,777,1924]" captionTargetId-1="figure-34@20.[382,1205,181,1932]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." captionText-1="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi-0="http://doi.org/10.5281/zenodo.4529528" figureDoi-1="http://doi.org/10.5281/zenodo.4529530" httpUri-0="https://zenodo.org/record/4529528/files/figure.png" httpUri-1="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Figs. 12–15</figureCitation>
), point-pinned (
<collectionCode id="4FEFD4D6FFAF4B57FCE2FC4AFC35227D" box="[858,941,1015,1041]" pageId="18" pageNumber="469">BYUC</collectionCode>
),
<collectingCountry id="51E90C83FFAF4B57FC7AFC4AFBF0227D" box="[962,1128,1015,1041]" name="Costa Rica" pageId="18" pageNumber="469">COSTA RICA</collectingCountry>
,
<collectingRegion id="EB3A82F1FFAF4B57FBCDFC4AFB43227D" box="[1141,1243,1015,1041]" country="Costa Rica" name="San Jose" pageId="18" pageNumber="469">San Jose</collectingRegion>
,
<collectingMunicipality id="C925D669FFAF4B57FB50FC4AFF482259" pageId="18" pageNumber="469">Nubes de Coronado</collectingMunicipality>
,
<collectorName id="840B29C5FFAF4B57FF63FBA6FECC2259" box="[219,340,1051,1077]" pageId="18" pageNumber="469">
<location id="2C211AC8FFAF4B57FF63FBA6FECC2259" LSID="urn:lsid:plazi:treatment:A157FD05FFAF4B56FF2FFC37FEF7249D:2C211AC8FFAF4B57FF63FBA6FECC2259" box="[219,340,1051,1077]" country="Costa Rica" county="Male" municipality="Nubes de Coronado" name="La Abigail" pageId="18" pageNumber="469" stateProvince="San Jose">La Abigail</location>
</collectorName>
,
<date id="5D406AD3FFAF4B57FEE6FBA7FE532259" box="[350,459,1050,1077]" pageId="18" pageNumber="469" value="1995-05-05">
<collectingDate id="4D04933BFFAF4B57FEE6FBA7FE532259" box="[350,459,1050,1077]" pageId="18" pageNumber="469" value="1995-05-05">5-V-1995</collectingDate>
</date>
,
<collectorName id="840B29C5FFAF4B57FE6DFBA6FDD82259" box="[469,576,1051,1077]" pageId="18" pageNumber="469">Baumann</collectorName>
&amp;
<collectorName id="840B29C5FFAF4B57FDDAFBA6FD422259" box="[610,730,1051,1077]" pageId="18" pageNumber="469">Houseman</collectorName>
[white label]
</materialsCitation>
/
<typeStatus id="F645F2B1FFAF4B57FC38FBA6FB8F2259" box="[896,1047,1051,1077]" pageId="18" pageNumber="469" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="EEFE3790FFAF4B57FB99FBA6FA052259" authority="Fitzgerald" authorityName="Fitzgerald" authorityYear="2021" box="[1057,1437,1051,1077]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="469" phylum="Arthropoda" rank="species" species="dolichopeza">
<emphasis id="1B8A9001FFAF4B57FB99FBA6FABC2259" box="[1057,1316,1051,1077]" italics="true" pageId="18" pageNumber="469">Penthetria dolichopeza</emphasis>
Fitzgerald
</taxonomicName>
[red label], terminalia dissected. Missing right front, right middle, and left hind legs.
<materialsCitation id="9996464EFFAF4B57FBFDFB82FE762211" ID-GBIF-Occurrence="3037447349" collectionCode="VI" country="Mexico" location="Hwy" municipality="Yerba Buena" pageId="18" pageNumber="469" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="1B8A9001FFAF4B57FBFDFB82FB2A2235" bold="true" box="[1093,1202,1087,1113]" pageId="18" pageNumber="469">
<typeStatus id="F645F2B1FFAF4B57FBFDFB82FB2A2235" box="[1093,1202,1087,1113]" pageId="18" pageNumber="469" type="paratype">Paratype</typeStatus>
</emphasis>
:
<collectingCountry id="51E90C83FFAF4B57FB07FB82FAA82235" box="[1215,1328,1087,1113]" name="Mexico" pageId="18" pageNumber="469">MEXICO</collectingCountry>
,
<collectingRegion id="EB3A82F1FFAF4B57FA83FB82FA002235" box="[1339,1432,1087,1113]" country="Mexico" name="Chiapas" pageId="18" pageNumber="469">Chiapas</collectingRegion>
,
<collectingMunicipality id="C925D669FFAF4B57FF2FFBDEFEB22211" box="[151,298,1123,1149]" pageId="18" pageNumber="469">Yerba Buena</collectingMunicipality>
,
<location id="2C211AC8FFAF4B57FE8DFBDEFEF32211" LSID="urn:lsid:plazi:treatment:A157FD05FFAF4B56FF2FFC37FEF7249D:2C211AC8FFAF4B57FE8DFBDEFEF32211" box="[309,363,1123,1149]" country="Mexico" municipality="Yerba Buena" name="Hwy" pageId="18" pageNumber="469" stateProvince="Chiapas">Hwy</location>
195,
<date id="5D406AD3FFAF4B57FE13FBDEFE762211" box="[427,494,1123,1149]" pageId="18" pageNumber="469" value="1969-06-10">
10.
<collectionCode id="4FEFD4D6FFAF4B57FE76FBDEFE762211" box="[462,494,1123,1149]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="18" pageNumber="469">VI</collectionCode>
</date>
</materialsCitation>
.1969, B.
<collectionCode id="4FEFD4D6FFAF4B57FDEAFBDEFDFB2211" box="[594,611,1123,1149]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="18" pageNumber="469" type="Museum">V</collectionCode>
.
<materialsCitation id="9996464EFFAF4B57FDC9FBDEFC132211" ID-GBIF-Occurrence="3037447433" box="[625,907,1123,1149]" collectionCode="CNCI" country="Mexico" location="Peterson" pageId="18" pageNumber="469" specimenCount="1" specimenCount-male="1" stateProvince="Chiapas" typeStatus="paratype">
<location id="2C211AC8FFAF4B57FDC9FBDEFD4C2211" LSID="urn:lsid:plazi:treatment:A157FD05FFAF4B56FF2FFC37FEF7249D:2C211AC8FFAF4B57FDC9FBDEFD4C2211" box="[625,724,1123,1149]" country="Mexico" name="Peterson" pageId="18" pageNumber="469" stateProvince="Chiapas">Peterson</location>
,
<specimenCount id="3FF8879AFFAF4B57FD67FBDEFCB22211" box="[735,810,1123,1149]" pageId="18" pageNumber="469" type="male">1 male</specimenCount>
(
<collectionCode id="4FEFD4D6FFAF4B57FC81FBDEFC1A2211" box="[825,898,1123,1149]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="18" pageNumber="469">CNCI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAF4B57FF7FFB3BFE07211D" pageId="18" pageNumber="469" type="description">
<paragraph id="29414C13FFAF4B57FF7FFB3BFDFE2121" blockId="18.[151,1437,1014,2013]" pageId="18" pageNumber="469">
<emphasis id="1B8A9001FFAF4B57FF7FFB3BFEC922CC" bold="true" box="[199,337,1158,1184]" pageId="18" pageNumber="469">Description</emphasis>
. Male (
<figureCitation id="B1C55096FFAF4B57FE12FB3AFE6622CD" box="[426,510,1159,1185]" captionStart="FIGURES 12–13" captionStartId="19.[151,264,1948,1972]" captionTargetBox="[193,1398,784,1919]" captionTargetId="figure-280@19.[187,1401,777,1924]" captionTargetPageId="19" captionText="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529528" httpUri="https://zenodo.org/record/4529528/files/figure.png" pageId="18" pageNumber="469">Fig. 12</figureCitation>
). Body length: approx. [7.0] mm.
<emphasis id="1B8A9001FFAF4B57FC39FB3AFC2622CD" box="[897,958,1159,1185]" italics="true" pageId="18" pageNumber="469">Head</emphasis>
. Black. Compound eye holoptic, covering virtually all of dorsal and lateral surface of head, with lateral longitudinal step dividing upper and lower portion of eye. Eyes nearly bare, with some very minute, very sparse ommatrichia present.Antennae dark brown to black, eight flagellomeres with black setae. Flagellomeres wider than long except first flagellomere elongate, about three times as long as wide, apical flagellomere smaller. Three ocelli on well-developed tubercle. Thorax matte dark brown with light brown highlights on humeral ridge and posterior portions of pleura. Thorax largely bare except fine short dark hairs on scutellum and in broad dorsocentral rows.
<emphasis id="1B8A9001FFAF4B57FD5AFAE2FC862314" box="[738,798,1375,1400]" italics="true" pageId="18" pageNumber="469">Legs.</emphasis>
Slender elongate. Entirely dark brown with dense short dark appressed hairs. Hind femur gradually thickened (clavate) on apical half. Hind tibia slender elongate virtually parallel-sided. Hind basitarsus slender elongate, about twelve times as long as wide (
<figureCitation id="B1C55096FFAF4B57FBEBFA1AFB3E23AD" box="[1107,1190,1447,1473]" captionStart="FIGURES 12–13" captionStartId="19.[151,264,1948,1972]" captionTargetBox="[193,1398,784,1919]" captionTargetId="figure-280@19.[187,1401,777,1924]" captionTargetPageId="19" captionText="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529528" httpUri="https://zenodo.org/record/4529528/files/figure.png" pageId="18" pageNumber="469">Fig. 12</figureCitation>
, hb).
<emphasis id="1B8A9001FFAF4B57FB5FFA1AFAB323AC" box="[1255,1323,1447,1472]" italics="true" pageId="18" pageNumber="469">Wings</emphasis>
(
<figureCitation id="B1C55096FFAF4B57FA83FA1AFA1723AD" box="[1339,1423,1447,1473]" captionStart="FIGURES 12–13" captionStartId="19.[151,264,1948,1972]" captionTargetBox="[193,1398,784,1919]" captionTargetId="figure-280@19.[187,1401,777,1924]" captionTargetPageId="19" captionText="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529528" httpUri="https://zenodo.org/record/4529528/files/figure.png" pageId="18" pageNumber="469">Fig. 13</figureCitation>
). [8.0] mm, evenly brown fumose except slightly darker costally. Veins brown, pterostigma concolorous with membrane. Sc long, complete. R
<subScript id="B57A4E56FFAF4B57FE75F9BCFE7F2063" attach="left" box="[461,487,1537,1551]" fontSize="6" pageId="18" pageNumber="469">2+3</subScript>
elongate, subparallel to R
<subScript id="B57A4E56FFAF4B57FCB4F9BCFCBE2063" attach="left" box="[780,806,1537,1551]" fontSize="6" pageId="18" pageNumber="469">4+5</subScript>
, without basal appendix. CuA reaching wing edge, CuA and CuP parallel apically, not distinctly convergent or meeting.
<emphasis id="1B8A9001FFAF4B57FCEEF9AEFC502041" box="[854,968,1555,1581]" italics="true" pageId="18" pageNumber="469">Abdomen.</emphasis>
Dark brown. Terminalia (
<figureCitation id="B1C55096FFAF4B57FB5EF9AEFAF42041" box="[1254,1388,1554,1581]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Figs. 14–15</figureCitation>
,
<figureCitation id="B1C55096FFAF4B57FACFF9AEFA0C2041" box="[1399,1428,1555,1581]" captionStart="FIGURES 42–45" captionStartId="34.[151,264,1889,1913]" captionTargetBox="[167,1419,240,1859]" captionTargetId="figure-64@34.[177,1410,666,1865]" captionTargetPageId="34" captionText="FIGURES 42–45. Male terminalia, dorsal, tergite nine removed. 42. P. distincta. 43. P. dolichopeza n. sp., paratype. 44. P. appendicula. 45. P. neonigrita n. sp., paratype. Scale bar = ca. 0.5 mm. Abbreviations: dlp, divergent lobes of paramere; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; vml, ventromedian lobes of gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.4529558" httpUri="https://zenodo.org/record/4529558/files/figure.png" pageId="18" pageNumber="469">43</figureCitation>
) dark brown to black. Tergite nine broader than long, posterior margin with very shallow v-shaped emargination on posterior edge (
<figureCitation id="B1C55096FFAF4B57FEFDF9E6FE002019" box="[325,408,1627,1653]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Fig. 14</figureCitation>
). In dorsal view, gonocoxites dorsoapically with small apically rounded medially-projecting lobe just dorsal to gonostylar socket (
<figureCitation id="B1C55096FFAF4B57FD84F9C2FD1620F5" box="[572,654,1663,1689]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Fig. 14</figureCitation>
, dal). In ventral view, posterior edge of medially fused gonocoxites + hypandrium medially flat, without additional median cleft or development of lobes (
<figureCitation id="B1C55096FFAF4B57FB84F91EFB1720D1" box="[1084,1167,1698,1725]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Fig. 15</figureCitation>
). Gonostylus short stout gently curved, apically broadly rounded expect for small thorn-like nub anteroapically (
<figureCitation id="B1C55096FFAF4B57FBD5F97AFB6F208D" box="[1133,1271,1734,1761]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Figs. 14–15</figureCitation>
). In dorsal and ventral views, fused parameres projecting from out of genital capsule, shiny brown heavily sclerotized, posteriorly rounded dome-like, but roughly triangular in overall shape (much broader anteriorly than posteriorly) (
<figureCitation id="B1C55096FFAF4B57FA81F8B2FA082145" box="[1337,1424,1807,1833]" captionStart="FIGURES 42–45" captionStartId="34.[151,264,1889,1913]" captionTargetBox="[167,1419,240,1859]" captionTargetId="figure-64@34.[177,1410,666,1865]" captionTargetPageId="34" captionText="FIGURES 42–45. Male terminalia, dorsal, tergite nine removed. 42. P. distincta. 43. P. dolichopeza n. sp., paratype. 44. P. appendicula. 45. P. neonigrita n. sp., paratype. Scale bar = ca. 0.5 mm. Abbreviations: dlp, divergent lobes of paramere; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; vml, ventromedian lobes of gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.4529558" httpUri="https://zenodo.org/record/4529558/files/figure.png" pageId="18" pageNumber="469">Fig. 43</figureCitation>
). Cerci fleshy, apically rounded with setae.
</paragraph>
<paragraph id="29414C13FFAF4B57FF7FF8EAFE07211D" blockId="18.[151,1437,1014,2013]" box="[199,415,1879,1905]" pageId="18" pageNumber="469">
<emphasis id="1B8A9001FFAF4B57FF7FF8EAFEBB211D" box="[199,291,1879,1905]" italics="true" pageId="18" pageNumber="469">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAF4B56FF7FF8C7FC6A2695" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469" type="diagnosis">
<paragraph id="29414C13FFAF4B56FF7FF8C7FC6A2695" blockId="18.[151,1437,1014,2013]" lastBlockId="19.[151,1437,151,753]" lastPageId="19" lastPageNumber="470" pageId="18" pageNumber="469">
<emphasis id="1B8A9001FFAF4B57FF7FF8C7FEA221F8" bold="true" box="[199,314,1914,1940]" pageId="18" pageNumber="469">Diagnosis</emphasis>
. This species can be distinguished from all other New World species by the very long slender hind basitarsus (about 12 times as long as wide (
<figureCitation id="B1C55096FFAF4B57FD3FF822FD4321D5" box="[647,731,1951,1977]" captionStart="FIGURES 12–13" captionStartId="19.[151,264,1948,1972]" captionTargetBox="[193,1398,784,1919]" captionTargetId="figure-280@19.[187,1401,777,1924]" captionTargetPageId="19" captionText="FIGURES 12–13. P. dolichopeza n. sp., holotype male. 12. Habitus, lateral. 13. Wing. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529528" httpUri="https://zenodo.org/record/4529528/files/figure.png" pageId="18" pageNumber="469">Fig. 12</figureCitation>
, hb)), the shape of the gonostylus (short, stout, gently curved, apically broadly rounded with small thorn-like nub anteroapically (
<figureCitation id="B1C55096FFAF4B57FCC7F87EFB9D21B1" box="[895,1029,1986,2013]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="18" pageNumber="469">Figs. 14–15</figureCitation>
)), the paramere is roughly triangular (much broader anteriorly than posteriorly) in dorsal view (
<figureCitation id="B1C55096FFAE4B56FC95FF2AFC1A26DD" box="[813,898,151,177]" captionStart="FIGURES 42–45" captionStartId="34.[151,264,1889,1913]" captionTargetBox="[167,1419,240,1859]" captionTargetId="figure-64@34.[177,1410,666,1865]" captionTargetPageId="34" captionText="FIGURES 42–45. Male terminalia, dorsal, tergite nine removed. 42. P. distincta. 43. P. dolichopeza n. sp., paratype. 44. P. appendicula. 45. P. neonigrita n. sp., paratype. Scale bar = ca. 0.5 mm. Abbreviations: dlp, divergent lobes of paramere; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; vml, ventromedian lobes of gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.4529558" httpUri="https://zenodo.org/record/4529558/files/figure.png" pageId="19" pageNumber="470">Fig. 43</figureCitation>
), gonocoxites dorsoapically with small apically rounded medially-projecting lobe just dorsal to gonostylar socket (
<figureCitation id="B1C55096FFAE4B56FCC6FF06FC4826B9" box="[894,976,187,213]" captionStart="FIGURES 14–15" captionStartId="20.[151,264,1956,1980]" captionTargetBox="[389,1201,188,1928]" captionTargetId="figure-34@20.[382,1205,181,1932]" captionTargetPageId="20" captionText="FIGURES 14–15. P. dolichopeza n. sp., holotype, male terminalia. 14. Dorsal. 15. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; dal, dorsoapical lobe of gonocoxite; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529530" httpUri="https://zenodo.org/record/4529530/files/figure.png" pageId="19" pageNumber="470">Fig. 14</figureCitation>
, dal), ventromedian lobes of gonocoxites absent, and posterior edge of tergite with at most a very broad, shallow cleft.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAE4B56FF7FFEBEFD6C240D" pageId="19" pageNumber="470" type="discussion">
<paragraph id="29414C13FFAE4B56FF7FFEBEFAC827E5" blockId="19.[151,1437,151,753]" pageId="19" pageNumber="470">
<emphasis id="1B8A9001FFAE4B56FF7FFEBEFEAC2771" bold="true" box="[199,308,259,285]" pageId="19" pageNumber="470">Remarks</emphasis>
. The form of the gonostylus and paramere in
<taxonomicName id="EEFE3790FFAE4B56FC94FEBEFC552771" authorityName="Fitzgerald" authorityYear="2021" box="[812,973,259,285]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dolichopeza">
<emphasis id="1B8A9001FFAE4B56FC94FEBEFC552771" box="[812,973,259,285]" italics="true" pageId="19" pageNumber="470">P. dolichopeza</emphasis>
</taxonomicName>
are unlike any other species in the Nearctic and Neotropical regions, though this gonostylar form is known in some Palearctic species such as
<taxonomicName id="EEFE3790FFAE4B56FB4EFE9AFEA9270A" authority="(Gimmerthal)" baseAuthorityName="Gimmerthal" class="Insecta" family="Bibionidae" genus="Crapitula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="motschulskii">
<emphasis id="1B8A9001FFAE4B56FB4EFE9AFA04272D" box="[1270,1436,295,321]" italics="true" pageId="19" pageNumber="470">P. motschulskii</emphasis>
(Gimmerthal)
</taxonomicName>
from
<collectingCountry id="51E90C83FFAE4B56FEC0FEF6FE5B2709" box="[376,451,331,357]" name="Russia" pageId="19" pageNumber="470">Russia</collectingCountry>
&amp;
<collectingCountry id="51E90C83FFAE4B56FE50FEF6FDB32709" box="[488,555,331,357]" name="China" pageId="19" pageNumber="470">China</collectingCountry>
(e.g., see
<bibRefCitation id="4D6F31E2FFAE4B56FD18FEF6FC582709" author="Hardy, D. E. &amp; Takahashi, M." box="[672,960,331,357]" pageId="19" pageNumber="470" pagination="383 - 449" refId="ref34286" refString="Hardy, D. E. &amp; Takahashi, M. (1960) Revision of the Japanese Bibionidae (Diptera, Nematocera). Pacific Insects, 2 (4), 383 - 449." type="journal article" year="1960">Hardy &amp; Takahashi 1960</bibRefCitation>
,
<figureCitation id="B1C55096FFAE4B56FC74FEF6FB86270A" box="[972,1054,331,358]" captionStart="FIGURES 2–3" captionStartId="11.[151,264,1518,1542]" captionTargetBox="[218,1372,186,1488]" captionTargetId="figure-16@11.[212,1376,181,1494]" captionTargetPageId="11" captionText="FIGURES 2–3. P. appendicula. 2. Wing. 3. Female habitus, lateral view. Scale bars = ca. 1.0 mm. Abbreviations: ba, basal appendix of R2+3; aem, anepimeron; aes, anepisternum; c, coxa; f, femur; h, halter; hb, hind basitarsus; kem, katepimeron; kes, katepisternum; m, meron; mtaes, metanepisternum; mtg, mediotergite; mtkes, metakatepisternum; ltg, laterotergite; pltrch, pleu- rotrochantin; prn, pronotum; sct, scutum; t, trochanter." figureDoi="http://doi.org/10.5281/zenodo.4529518" httpUri="https://zenodo.org/record/4529518/files/figure.png" pageId="19" pageNumber="470">Fig. 3b</figureCitation>
) and
<taxonomicName id="EEFE3790FFAE4B56FBE6FEF6FB4F2709" box="[1118,1239,331,357]" class="Insecta" family="Bibionidae" genus="Amasia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="1B8A9001FFAE4B56FBE6FEF6FB4F2709" box="[1118,1239,331,357]" italics="true" pageId="19" pageNumber="470">P. funebris</emphasis>
</taxonomicName>
(Europe,
<collectingCountry id="51E90C83FFAE4B56FAF0FEF6FA0C270A" box="[1352,1428,331,358]" name="Russia" pageId="19" pageNumber="470">Russia</collectingCountry>
) and the roughly triangular paramere is very similar to that described for
<taxonomicName id="EEFE3790FFAE4B56FC78FED2FAD427E5" authority="(Krivosheina 1999)" baseAuthorityName="Krivosheina" baseAuthorityYear="1999" box="[960,1356,367,393]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="motschluskii">
<emphasis id="1B8A9001FFAE4B56FC78FED2FBFF27E5" box="[960,1127,367,393]" italics="true" pageId="19" pageNumber="470">P. motschluskii</emphasis>
(
<bibRefCitation id="4D6F31E2FFAE4B56FBCEFED2FADB27E5" author="Krivosheina, N. P." box="[1142,1347,367,393]" pageId="19" pageNumber="470" pagination="1 - 665" refId="ref34671" refString="Krivosheina, N. P. (1999) Pleciidae. In: Ler, P. A. (Ed.), Key to the insects of Russian Far East. Vol. VI. Diptera and Siphonaptera. Pt. 1. Dal'nauka, Vladivostok, pp. 1 - 665. [in Russian]" type="book chapter" year="1999">Krivosheina 1999</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="29414C13FFAE4B56FF7FFE2EFD6C240D" blockId="19.[151,1437,151,753]" pageId="19" pageNumber="470">
While the female of
<taxonomicName id="EEFE3790FFAE4B56FE0BFE2EFDCE27C1" authorityName="Fitzgerald" authorityYear="2021" box="[435,598,403,429]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dolichopeza">
<emphasis id="1B8A9001FFAE4B56FE0BFE2EFDCE27C1" box="[435,598,403,429]" italics="true" pageId="19" pageNumber="470">P. dolichopeza</emphasis>
</taxonomicName>
is currently unknown, the similarities of the male terminalia to the aforementioned Palearctic species might provide clues to the structure of the female terminalia of
<taxonomicName id="EEFE3790FFAE4B56FB2EFE0AFAAF27BD" authorityName="Fitzgerald" authorityYear="2021" box="[1174,1335,439,465]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="dolichopeza">
<emphasis id="1B8A9001FFAE4B56FB2EFE0AFAAF27BD" box="[1174,1335,439,465]" italics="true" pageId="19" pageNumber="470">P. dolichopeza</emphasis>
</taxonomicName>
; females of both Palearctic species have female tergite nine in dorsal view medially divided, but represented by a significant pair of sclerotized plates. In the New World, only females of
<taxonomicName id="EEFE3790FFAE4B56FCE9FE42FC7B2474" authorityName="Fitzgerald" authorityYear="2021" box="[849,995,511,536]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="neonigrita">
<emphasis id="1B8A9001FFAE4B56FCE9FE42FC7B2474" box="[849,995,511,536]" italics="true" pageId="19" pageNumber="470">P. neonigrita</emphasis>
</taxonomicName>
have tergite nine strongly represented, though it occurs to a lesser degree in
<taxonomicName id="EEFE3790FFAE4B56FD85FD9EFD7C2451" authorityName="Hardy" authorityYear="1945" box="[573,740,547,573]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="appendicula">
<emphasis id="1B8A9001FFAE4B56FD85FD9EFD7C2451" box="[573,740,547,573]" italics="true" pageId="19" pageNumber="470">P. appendicula</emphasis>
</taxonomicName>
and
<taxonomicName id="EEFE3790FFAE4B56FCA4FD9EFC002451" authorityName="Hardy" authorityYear="1945" box="[796,920,547,573]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="470" phylum="Arthropoda" rank="species" species="distincta">
<emphasis id="1B8A9001FFAE4B56FCA4FD9EFC002451" box="[796,920,547,573]" italics="true" pageId="19" pageNumber="470">P. distincta</emphasis>
</taxonomicName>
and is almost absent dorsally (represented by only a very thin strap) in the remainder of the species.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFAE4B56FF7FFDD6FEF7249D" pageId="19" pageNumber="470" type="etymology">
<paragraph id="29414C13FFAE4B56FF7FFDD6FE2B24C5" blockId="19.[151,1437,151,753]" pageId="19" pageNumber="470">
<emphasis id="1B8A9001FFAE4B56FF7FFDD6FED024E9" bold="true" box="[199,328,619,645]" pageId="19" pageNumber="470">Etymology</emphasis>
. The specific epithet is derived from the Greek
<emphasis id="1B8A9001FFAE4B56FCC9FDD6FC4824E9" box="[881,976,619,645]" italics="true" pageId="19" pageNumber="470">dolichos</emphasis>
(long) and
<emphasis id="1B8A9001FFAE4B56FBEFFDD6FB1224E8" box="[1111,1162,619,644]" italics="true" pageId="19" pageNumber="470">peza</emphasis>
(foot) for the long hind basitarsus of this species.
</paragraph>
<paragraph id="29414C13FFAE4B56FF7FFD0FFEF7249D" blockId="19.[151,1437,151,753]" pageId="19" pageNumber="470">
<emphasis id="1B8A9001FFAE4B56FF7FFD0FFDE324A0" bold="true" box="[199,635,690,716]" pageId="19" pageNumber="470">Geographic &amp; Seasonal Distribution</emphasis>
. Southern Mexico and
<collectingCountry id="51E90C83FFAE4B56FC3CFD0EFC6624A1" box="[900,1022,691,717]" name="Costa Rica" pageId="19" pageNumber="470">Costa Rica</collectingCountry>
(
<figureCitation id="B1C55096FFAE4B56FBB6FD0EFBC324A1" box="[1038,1115,691,717]" captionStart="MAP 2" captionStartId="39.[151,209,1832,1856]" captionTargetBox="[167,1424,945,1781]" captionTargetId="figure-37@39.[157,1431,936,1789]" captionText="MAP 2. Distribution of P. arizonensis (black circle) and P. dolichopeza (white circle)." figureDoi="http://doi.org/10.5281/zenodo.4546086" httpUri="https://zenodo.org/record/4546086/files/figure.png" pageId="19" pageNumber="470">Map 2</figureCitation>
). Seasonal distribution summarized in
<tableCitation id="647C79A8FFAE4B56FEADFD6AFEF0249D" box="[277,360,727,753]" captionStart="TABLE 1" captionStartId="34.[151,239,151,175]" captionText="TABLE 1. Summary of adult flight period (grey boxes indicate specimens collected during these months); Malaise trap data including more than one month have been excluded." pageId="19" pageNumber="470">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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