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<document id="6626D0CD9001DEB9045677412B3346C5" ID-CLB-Dataset="8167" ID-DOI="10.11646/zootaxa.4926.4.1" ID-GBIF-Dataset="f0b1c09e-298d-497d-b6de-cd74367c983e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4529512" ID-ZooBank="3ADD4A2B-A3F9-4379-A8FE-39DD867531F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612954843127" checkinUser="plazi" docAuthor="Fitzgerald, Scott J." docDate="2021" docId="A157FD05FFA24B64FF2FFD3EFD1625ED" docLanguage="en" docName="zootaxa.4926.4.1.pdf" docOrigin="Zootaxa 4926 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penthetria yakima Fitzgerald 2021, n. sp." docType="treatment" docVersion="5" lastPageNumber="484" masterDocId="5D6E857DFFBD4B45FFB8FFBDFF98266C" masterDocTitle="Penthetria Meigen (Diptera: Bibionidae): Revision of the New World species and world catalog" masterLastPageNumber="500" masterPageNumber="451" pageNumber="482" updateTime="1698928650180" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B7413FB6B262FB98EB51A115F1949B3C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4D9CEB0707AFDBE69526E8ACA9E7B2DF">
<mods:title id="C24FAFC808A7EA8845323D8B66EC6D56">Penthetria Meigen (Diptera: Bibionidae): Revision of the New World species and world catalog</mods:title>
</mods:titleInfo>
<mods:name id="38617E66647609E58BAFFCC5152EA6CB" type="personal">
<mods:role id="81CDA8429E5343F05F1E0F50E18706FB">
<mods:roleTerm id="26D37DBEFFC931380925DDC44870CB88">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8CFFD42A31E20718AE620382CF999ED">Fitzgerald, Scott J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="792621FBAA551FDFEFD503077BB2C1C8">text</mods:typeOfResource>
<mods:relatedItem id="E411B75CF0BAEA9C019793B96B03FC6D" type="host">
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<mods:title id="3C1D89C44BA9B948DFB0160BD9068860">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="96B6B5E17D1D82E07EC349478F9420EC">
<mods:date id="8DA374A6C7662CB0E6D995AC1A0D658D">2021</mods:date>
<mods:detail id="9F65CF7A927ECC012744E1F806FAECC3" type="pubDate">
<mods:number id="4D0DE49ACAE5074E49FFA84A526507CC">2021-02-10</mods:number>
</mods:detail>
<mods:detail id="9B394508899B16017306A06486634EB3" type="volume">
<mods:number id="0A3CE0467269FB7E08F13C12A1D0EA46">4926</mods:number>
</mods:detail>
<mods:detail id="0E855322E1A3099F36929D1A15FA170A" type="issue">
<mods:number id="698115184BA466E7A066765DC946D0B5">4</mods:number>
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<mods:start id="86C5D42DA939F4A31383E0C9095217A5">451</mods:start>
<mods:end id="65A690EA32FE2AE73F9FC1066DE06612">500</mods:end>
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<mods:classification id="B7EA505B0A865F2F93C645B085F8DCBC">journal article</mods:classification>
<mods:identifier id="BD0F0F08C5628D5503E87CA31CA38941" type="CLB-Dataset">8167</mods:identifier>
<mods:identifier id="E35DAE5854C7CB1C4C77DD06BD360809" type="DOI">10.11646/zootaxa.4926.4.1</mods:identifier>
<mods:identifier id="3D7DD729DC5966898597F152C8BCB0A8" type="GBIF-Dataset">f0b1c09e-298d-497d-b6de-cd74367c983e</mods:identifier>
<mods:identifier id="DFD9B80EDA9273881679AA97DD1B7F4E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1F5EDAEF6056DF77D11652DA4B3468FA" type="Zenodo-Dep">4529512</mods:identifier>
<mods:identifier id="9265440E6E99A15819A32EB2066D0C59" type="ZooBank">3ADD4A2B-A3F9-4379-A8FE-39DD867531F7</mods:identifier>
</mods:mods>
<treatment id="A157FD05FFA24B64FF2FFD3EFD1625ED" ID-DOI="http://doi.org/10.5281/zenodo.4546104" ID-GBIF-Taxon="177422188" ID-Zenodo-Dep="4546104" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED" httpUri="http://treatment.plazi.org/id/A157FD05FFA24B64FF2FFD3EFD1625ED" lastPageId="33" lastPageNumber="484" pageId="31" pageNumber="482">
<subSubSection id="61E41F98FFA24B5AFF2FFD3EFDDF24F2" box="[151,583,643,670]" pageId="31" pageNumber="482" type="nomenclature">
<paragraph id="29414C13FFA24B5AFF2FFD3EFDDF24F2" blockId="31.[151,583,643,707]" box="[151,583,643,670]" pageId="31" pageNumber="482">
<heading id="7209FB7FFFA24B5AFF2FFD3EFDDF24F2" bold="true" box="[151,583,643,670]" fontSize="11" level="1" pageId="31" pageNumber="482" reason="1">
<emphasis id="1B8A9001FFA24B5AFF2FFD3EFDDF24F2" bold="true" box="[151,583,643,670]" pageId="31" pageNumber="482">
<taxonomicName id="EEFE3790FFA24B5AFF2FFD3EFE6624F2" ID-CoL="8GTZS" authority="Fitzgerald" authorityName="Fitzgerald" authorityYear="2021" box="[151,510,643,670]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="yakima" status="sp. nov.">
<emphasis id="1B8A9001FFA24B5AFF2FFD3EFEEE24F2" bold="true" box="[151,374,643,670]" italics="true" pageId="31" pageNumber="482">Penthetria yakima</emphasis>
Fitzgerald
</taxonomicName>
<taxonomicNameLabel id="00B92D7AFFA24B5AFDBDFD39FDDF24F2" box="[517,583,644,670]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFA24B5AFF2FFD15FDB424AE" box="[151,556,680,707]" pageId="31" pageNumber="482" type="description">
<paragraph id="29414C13FFA24B5AFF2FFD15FDB424AE" blockId="31.[151,583,643,707]" box="[151,556,680,707]" pageId="31" pageNumber="482">
(
<figureCitation id="B1C55096FFA24B5AFF27FD15FF6524AE" box="[159,253,680,706]" captionStart="FIGURES 30–32" captionStartId="29.[151,264,1324,1348]" captionTargetBox="[248,1339,186,1295]" captionTargetId="figure-16@29.[243,1343,181,1299]" captionTargetPageId="29" captionText="FIGURES 30–32. Holotype males, lateral. 30. P. neonigrita n. sp.. 31. P. yakima n. sp.. 32. P. arizonensis n. sp.. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529546" httpUri="https://zenodo.org/record/4529546/files/figure.png" pageId="31" pageNumber="482">Figs. 31</figureCitation>
, 36–37, 39, 41, 56,
<figureCitation id="B1C55096FFA24B5AFE63FD14FDBB24AE" box="[475,547,680,707]" captionStart="MAP 3" captionStartId="40.[151,209,1161,1185]" captionTargetBox="[170,1420,191,1128]" captionTargetId="figure-34@40.[160,1427,181,1137]" captionText="MAP 3. Distribution of P. heteroptera (black circle) and P. yakima (star)." figureDoi="http://doi.org/10.5281/zenodo.4546088" httpUri="https://zenodo.org/record/4546088/files/figure.png" pageId="31" pageNumber="482">Map 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFA24B5AFF2FFD4DFCD82246" pageId="31" pageNumber="482" type="materials_examined">
<paragraph id="29414C13FFA24B5AFF2FFD4DFAC9251A" blockId="31.[151,1437,752,2039]" pageId="31" pageNumber="482">
<emphasis id="1B8A9001FFA24B5AFF2FFD4DFEDB2566" bold="true" box="[151,323,752,778]" pageId="31" pageNumber="482">Type Material</emphasis>
.
<materialsCitation id="9996464EFFA24B5AFEECFD4DFE8A253E" ID-GBIF-Occurrence="3037447407" collectingDate="1903-09-25" collectionCode="USNM" collectorName="N. Yakima" country="United States of America" location="Male" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="1B8A9001FFA24B5AFEECFD4DFE592566" bold="true" box="[340,449,752,778]" pageId="31" pageNumber="482">
<typeStatus id="F645F2B1FFA24B5AFEECFD4DFE592566" box="[340,449,752,778]" pageId="31" pageNumber="482" type="holotype">Holotype</typeStatus>
</emphasis>
:
<location id="2C211AC8FFA24B5AFE6AFD4DFD942566" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED:2C211AC8FFA24B5AFE6AFD4DFD942566" box="[466,524,752,778]" country="United States of America" name="Male" pageId="31" pageNumber="482" stateProvince="Washington">Male</location>
(
<figureCitation id="B1C55096FFA24B5AFDA6FD4DFDEB2566" box="[542,627,752,778]" captionStart="FIGURES 30–32" captionStartId="29.[151,264,1324,1348]" captionTargetBox="[248,1339,186,1295]" captionTargetId="figure-16@29.[243,1343,181,1299]" captionTargetPageId="29" captionText="FIGURES 30–32. Holotype males, lateral. 30. P. neonigrita n. sp.. 31. P. yakima n. sp.. 32. P. arizonensis n. sp.. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529546" httpUri="https://zenodo.org/record/4529546/files/figure.png" pageId="31" pageNumber="482">Fig. 31</figureCitation>
), point-pinned (
<collectionCode id="4FEFD4D6FFA24B5AFC93FD4DFC1A2566" box="[811,898,752,778]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
),
<collectingCountry id="51E90C83FFA24B5AFC2FFD4DFC4A2566" box="[919,978,752,778]" name="United States of America" pageId="31" pageNumber="482">USA</collectingCountry>
,
<collectingRegion id="EB3A82F1FFA24B5AFC67FD4DFBFF2566" box="[991,1127,752,778]" country="United States of America" name="Washington" pageId="31" pageNumber="482">Washington</collectingRegion>
,
<collectorName id="840B29C5FFA24B5AFBCDFD4CFB6A2566" box="[1141,1266,752,779]" pageId="31" pageNumber="482">N. Yakima</collectorName>
,
<date id="5D406AD3FFA24B5AFB47FD4DFF4A2542" pageId="31" pageNumber="482" value="1903-09-25">
<collectingDate id="4D04933BFFA24B5AFB47FD4DFF4A2542" pageId="31" pageNumber="482" value="1903-09-25">25 September 1903</collectingDate>
</date>
,
<collectorName id="840B29C5FFA24B5AFF64FCA9FEF52543" box="[220,365,788,815]" pageId="31" pageNumber="482">Eldred Jenne</collectorName>
[white label] /
<collectorName id="840B29C5FFA24B5AFDABFCA9FCAE2542" box="[531,822,788,814]" pageId="31" pageNumber="482">A.L. Melander Collection</collectorName>
, 1961 [white label] / USNMENT 01556096 [white barcode label]
</materialsCitation>
/
<typeStatus id="F645F2B1FFA24B5AFE90FC85FE27253E" box="[296,447,824,850]" pageId="31" pageNumber="482" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="EEFE3790FFA24B5AFE72FC85FC88253E" authority="Fitzgerald" authorityName="Fitzgerald" box="[458,784,824,850]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FFA24B5AFE72FC85FD00253E" box="[458,664,824,850]" italics="true" pageId="31" pageNumber="482">Penthetria yakima</emphasis>
Fitzgerald
</taxonomicName>
[red label]; terminalia dissected, missing middle and right front legs.
<materialsCitation id="9996464EFFA24B5AFEA8FCE1FAD5251A" ID-GBIF-Occurrence="3037447554" box="[272,1357,860,887]" collectingDate="1903-09-25" collectionCode="USNM" collectorName="N. Yakima" country="United States of America" location="Male" pageId="31" pageNumber="482" specimenCount="2" specimenCount-male="2" stateProvince="Washington" typeStatus="paratype">
<emphasis id="1B8A9001FFA24B5AFEA8FCE1FE10251A" bold="true" box="[272,392,860,886]" pageId="31" pageNumber="482">
<typeStatus id="F645F2B1FFA24B5AFEA8FCE1FE10251A" box="[272,392,860,886]" pageId="31" pageNumber="482" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as holotype except USNMENT 01556099 &amp; 01556098 (
<specimenCount id="3FF8879AFFA24B5AFB32FCE1FB7B251A" box="[1162,1251,860,886]" pageId="31" pageNumber="482" type="male">2 males</specimenCount>
,
<collectionCode id="4FEFD4D6FFA24B5AFB57FCE1FADD251A" box="[1263,1349,860,886]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="29414C13FFA24B5AFF7FFC3DFCD82246" blockId="31.[151,1437,752,2039]" pageId="31" pageNumber="482">
<emphasis id="1B8A9001FFA24B5AFF7FFC3DFDA825F6" bold="true" box="[199,560,896,922]" pageId="31" pageNumber="482">Additional material examined</emphasis>
:
<materialsCitation id="9996464EFFA24B5AFDF8FC3DFE3C25D3" ID-GBIF-Occurrence="3037447390" collectingDate="1957-08-19" collectionCode="USNM, R" collectorName="Coquillett" country="Canada" location="Robson" pageId="31" pageNumber="482" specimenCount="1" stateProvince="British Columbia">
<emphasis id="1B8A9001FFA24B5AFDF8FC3DFD2125F6" bold="true" box="[576,697,896,922]" pageId="31" pageNumber="482">
<collectingCountry id="51E90C83FFA24B5AFDF8FC3DFD2125F6" box="[576,697,896,922]" name="Canada" pageId="31" pageNumber="482">CANADA</collectingCountry>
</emphasis>
: “Can.,”
<collectorName id="840B29C5FFA24B5AFC9EFC3DFC0125F6" box="[806,921,896,922]" pageId="31" pageNumber="482">Coquillett</collectorName>
, 1M (
<collectionCode id="4FEFD4D6FFA24B5AFC65FC3DFBAC25F6" box="[989,1076,896,922]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
);
<collectingRegion id="EB3A82F1FFA24B5AFBF1FC3DFAC125F6" box="[1097,1369,896,922]" country="Canada" name="British Columbia" pageId="31" pageNumber="482">BRITISH COLUMBIA</collectingRegion>
:
<location id="2C211AC8FFA24B5AFADCFC3DFF5825D3" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED:2C211AC8FFA24B5AFADCFC3DFF5825D3" country="Canada" name="Robson" pageId="31" pageNumber="482" stateProvince="British Columbia">Robson</location>
,
<date id="5D406AD3FFA24B5AFF76FC19FEFE25D3" box="[206,358,932,959]" pageId="31" pageNumber="482" value="1957-08-19">
<collectingDate id="4D04933BFFA24B5AFF76FC19FEFE25D3" box="[206,358,932,959]" pageId="31" pageNumber="482" value="1957-08-19">19 VIII 1957</collectingDate>
</date>
, H.
<collectionCode id="4FEFD4D6FFA24B5AFE36FC18FE3C25D3" box="[398,420,933,959]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="482">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9996464EFFA24B5AFE09FC19FD13258F" ID-GBIF-Occurrence="3037447515" collectingDate="1949-09-11" collectingDateMax="1989-09-07" collectingDateMin="1949-09-11" collectionCode="CNCI, SFC, R" collectorName="Foxlee &amp; A. Borkent" country="Canada" location="Moose Factory" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Ontario">
<collectorName id="840B29C5FFA24B5AFE09FC19FD9825D2" box="[433,512,932,958]" pageId="31" pageNumber="482">Foxlee</collectorName>
, 1M (
<collectionCode id="4FEFD4D6FFA24B5AFDFCFC19FD1525D2" box="[580,653,932,958]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="482">CNCI</collectionCode>
);
<locationDeviation id="38F35531FFA24B5AFD1BFC19FC0B25D2" box="[675,915,932,959]" location="Salmon Arm" pageId="31" pageNumber="482">
<quantity id="EE06E1F6FFA24B5AFD1BFC19FD4525D2" box="[675,733,932,958]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="482" unit="km" value="6.0">6 km</quantity>
E Salmon Arm
</locationDeviation>
,
<date id="5D406AD3FFA24B5AFC18FC18FB8025D2" box="[928,1048,932,959]" pageId="31" pageNumber="482" value="1989-09-07">
<collectingDate id="4D04933BFFA24B5AFC18FC18FB8025D2" box="[928,1048,932,959]" pageId="31" pageNumber="482" value="1989-09-07">7 IX 1989</collectingDate>
</date>
,
<collectorName id="840B29C5FFA24B5AFB9BFC19FB3C25D2" box="[1059,1188,932,958]" pageId="31" pageNumber="482">A. Borkent</collectorName>
, CD1071, 1M (
<collectionCode id="4FEFD4D6FFA24B5AFAE0FC19FA1725D2" box="[1368,1423,932,958]" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="31" pageNumber="482">SFC</collectionCode>
);
<collectingRegion id="EB3A82F1FFA24B5AFF2FFC75FE8F258E" box="[151,279,968,994]" country="Canada" name="Ontario" pageId="31" pageNumber="482">ONTARIO</collectingRegion>
:
<location id="2C211AC8FFA24B5AFE9AFC74FE54258F" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED:2C211AC8FFA24B5AFE9AFC74FE54258F" box="[290,460,969,995]" country="Canada" name="Moose Factory" pageId="31" pageNumber="482" stateProvince="Ontario">Moose Factory</location>
,
<date id="5D406AD3FFA24B5AFE6FFC75FDC0258E" box="[471,600,968,995]" pageId="31" pageNumber="482" value="1949-09-11">
<collectingDate id="4D04933BFFA24B5AFE6FFC75FDC0258E" box="[471,600,968,995]" pageId="31" pageNumber="482" value="1949-09-11">11 IX 1949</collectingDate>
</date>
, J.
<collectionCode id="4FEFD4D6FFA24B5AFDCCFC74FD13258F" box="[628,651,969,995]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="482">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9996464EFFA24B5AFD2DFC75FE88226A" ID-GBIF-Occurrence="3037447542" collectingDate="1948-08-26" collectionCode="CNCI, R" collectorName="Vockeroth" country="Canada" location="Christopher" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Saskatchewan">
<collectorName id="840B29C5FFA24B5AFD2DFC75FC93258E" box="[661,779,968,994]" pageId="31" pageNumber="482">Vockeroth</collectorName>
, 1M (
<collectionCode id="4FEFD4D6FFA24B5AFCF4FC75FC0D258E" box="[844,917,968,994]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="482">CNCI</collectionCode>
);
<collectingRegion id="EB3A82F1FFA24B5AFC10FC75FB16258E" box="[936,1166,968,994]" country="Canada" name="Saskatchewan" pageId="31" pageNumber="482">SASKATCHEWAN</collectingRegion>
:
<location id="2C211AC8FFA24B5AFB20FC75FA84258E" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED:2C211AC8FFA24B5AFB20FC75FA84258E" box="[1176,1308,968,994]" country="Canada" name="Christopher" pageId="31" pageNumber="482" stateProvince="Saskatchewan">Christopher</location>
L.,
<date id="5D406AD3FFA24B5AFAF2FC75FF4A226A" pageId="31" pageNumber="482" value="1948-08-26">
<collectingDate id="4D04933BFFA24B5AFAF2FC75FF4A226A" pageId="31" pageNumber="482" value="1948-08-26">26 VIII 1948</collectingDate>
</date>
, A.
<collectionCode id="4FEFD4D6FFA24B5AFF42FC51FE88226A" box="[250,272,1004,1030]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="482">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9996464EFFA24B5AFEA5FC51FDA0226A" ID-GBIF-Occurrence="3037447344" box="[285,568,1004,1030]" collectionCode="CNCI" collectorName="Brooks" country="Canada" pageId="31" pageNumber="482" specimenCount="1">
<collectorName id="840B29C5FFA24B5AFEA5FC51FEE9226A" box="[285,369,1004,1030]" pageId="31" pageNumber="482">Brooks</collectorName>
, 1M, 1F (
<collectionCode id="4FEFD4D6FFA24B5AFE5FFC51FDA8226A" box="[487,560,1004,1030]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="31" pageNumber="482">CNCI</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9996464EFFA24B5AFDFEFC51FCA42246" ID-GBIF-Occurrence="3037447435" collectingDate="1996-09-13" collectionCode="BYUC, SFC" collectorName="B. O. Huntsman &amp; C. Wallace" country="United States of America" county="Uinta Co." location="Bear River State Park" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Wyoming">
<emphasis id="1B8A9001FFA24B5AFDFEFC51FDE6226A" bold="true" box="[582,638,1004,1030]" pageId="31" pageNumber="482">
<collectingCountry id="51E90C83FFA24B5AFDFEFC51FDE6226A" box="[582,638,1004,1030]" name="United States of America" pageId="31" pageNumber="482">USA</collectingCountry>
</emphasis>
:
<collectingRegion id="EB3A82F1FFA24B5AFD37FC51FCB9226A" box="[655,801,1004,1030]" country="United States of America" name="Wyoming" pageId="31" pageNumber="482">WYOMING</collectingRegion>
:
<collectingCounty id="C020349FFFA24B5AFC97FC51FC39226A" box="[815,929,1004,1030]" pageId="31" pageNumber="482">Uinta Co.</collectingCounty>
,
<location id="2C211AC8FFA24B5AFC17FC50FB36226A" LSID="urn:lsid:plazi:treatment:A157FD05FFA24B64FF2FFD3EFD1625ED:2C211AC8FFA24B5AFC17FC50FB36226A" box="[943,1198,1004,1031]" country="United States of America" county="Uinta Co." name="Bear River State Park" pageId="31" pageNumber="482" stateProvince="Wyoming">Bear River State Park</location>
,
<date id="5D406AD3FFA24B5AFB05FC51FAC1226A" box="[1213,1369,1004,1030]" pageId="31" pageNumber="482" value="1996-09-13">
<collectingDate id="4D04933BFFA24B5AFB05FC51FAC1226A" box="[1213,1369,1004,1030]" pageId="31" pageNumber="482" value="1996-09-13">13 Sept 1996</collectingDate>
</date>
,
<collectorName id="840B29C5FFA24B5AFADFFC51FE952247" pageId="31" pageNumber="482">B.O. Huntsman</collectorName>
, 1M (
<collectionCode id="4FEFD4D6FFA24B5AFEF6FBADFE392246" box="[334,417,1040,1066]" pageId="31" pageNumber="482">BYUC</collectionCode>
), 1F (
<collectionCode id="4FEFD4D6FFA24B5AFE59FBADFD802246" box="[481,536,1040,1066]" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="31" pageNumber="482">SFC</collectionCode>
),
<collectorName id="840B29C5FFA24B5AFD94FBADFD302246" box="[556,680,1040,1066]" pageId="31" pageNumber="482">C. Wallace</collectorName>
, 1F (
<collectionCode id="4FEFD4D6FFA24B5AFD58FBADFCAC2246" box="[736,820,1040,1066]" pageId="31" pageNumber="482">BYUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FFA24B64FF7FFB89FC8D2695" lastPageId="33" lastPageNumber="484" pageId="31" pageNumber="482" type="description">
<paragraph id="29414C13FFA24B5AFF7FFB89FAC5210B" blockId="31.[151,1437,752,2039]" pageId="31" pageNumber="482">
<emphasis id="1B8A9001FFA24B5AFF7FFB89FEC92222" bold="true" box="[199,337,1076,1102]" pageId="31" pageNumber="482">Description</emphasis>
. Male (
<figureCitation id="B1C55096FFA24B5AFE12FB89FE662222" box="[426,510,1076,1102]" captionStart="FIGURES 30–32" captionStartId="29.[151,264,1324,1348]" captionTargetBox="[248,1339,186,1295]" captionTargetId="figure-16@29.[243,1343,181,1299]" captionTargetPageId="29" captionText="FIGURES 30–32. Holotype males, lateral. 30. P. neonigrita n. sp.. 31. P. yakima n. sp.. 32. P. arizonensis n. sp.. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529546" httpUri="https://zenodo.org/record/4529546/files/figure.png" pageId="31" pageNumber="482">Fig. 31</figureCitation>
). Body length: approx. [6.0] mm.
<emphasis id="1B8A9001FFA24B5AFC39FB89FC262222" box="[897,958,1076,1102]" italics="true" pageId="31" pageNumber="482">Head</emphasis>
. Black. Compound eye holoptic, covering virtually all of dorsal and lateral surface of head, with lateral longitudinal step dividing upper and lower portion of eye. Eyes nearly bare, with some very minute, very sparse ommatrichia present. Antennae dark brown to black, nine flagellomeres with black setae. Flagellomeres wider than long except first flagellomere more elongate, about twice as long as wide, and apical flagellomere which is slighter longer than wide and tapers to a rounded point. Three ocelli on well-developed tubercle. Thorax matte dark brown to black with light brown highlights on humeral ridge and posterior portions of pleurae. Thorax largely bare except fine short dark hairs on scutellum and in broad dorsocentral rows.
<emphasis id="1B8A9001FFA24B5AFED6FA8CFE322326" box="[366,426,1329,1354]" italics="true" pageId="31" pageNumber="482">Legs.</emphasis>
Slender elongate. Entirely dark brown with dense short dark appressed hairs. Hind femur gradually slightly thickened (clavate) on apical half. Hind tibia slender elongate virtually parallel-sided. Hind basitarsus slender elongate to slightly robust, but not swollen, about five times as long as wide (
<figureCitation id="B1C55096FFA24B5AFB0AFAC5FA9E23FE" box="[1202,1286,1400,1426]" captionStart="FIGURES 30–32" captionStartId="29.[151,264,1324,1348]" captionTargetBox="[248,1339,186,1295]" captionTargetId="figure-16@29.[243,1343,181,1299]" captionTargetPageId="29" captionText="FIGURES 30–32. Holotype males, lateral. 30. P. neonigrita n. sp.. 31. P. yakima n. sp.. 32. P. arizonensis n. sp.. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529546" httpUri="https://zenodo.org/record/4529546/files/figure.png" pageId="31" pageNumber="482">Fig. 31</figureCitation>
).
<emphasis id="1B8A9001FFA24B5AFAA3FAC4FAC723FE" box="[1307,1375,1401,1426]" italics="true" pageId="31" pageNumber="482">Wings</emphasis>
(
<figureCitation id="B1C55096FFA24B5AFAD7FAC5FF2E23DA" captionStart="FIGURES 30–32" captionStartId="29.[151,264,1324,1348]" captionTargetBox="[248,1339,186,1295]" captionTargetId="figure-16@29.[243,1343,181,1299]" captionTargetPageId="29" captionText="FIGURES 30–32. Holotype males, lateral. 30. P. neonigrita n. sp.. 31. P. yakima n. sp.. 32. P. arizonensis n. sp.. Scale bars = ca. 1.0 mm. Abbreviations: hb, hind basitarsus." figureDoi="http://doi.org/10.5281/zenodo.4529546" httpUri="https://zenodo.org/record/4529546/files/figure.png" pageId="31" pageNumber="482">Fig. 31</figureCitation>
). 6.0–[6.5] mm (n=4), evenly brown fumose except slightly darker costally and cell sc which is slightly more pale. Veins dark brown, pterostigma brown, slightly contrasting to concolorous with membrane. Sc long, complete. R
<subScript id="B57A4E56FFA24B5AFF12FA4BFF5B2068" attach="left" box="[170,195,1526,1540]" fontSize="6" pageId="31" pageNumber="482">2+3</subScript>
elongate, subparallel to R
<subScript id="B57A4E56FFA24B5AFE50FA4BFD9A2068" attach="left" box="[488,514,1526,1540]" fontSize="6" pageId="31" pageNumber="482">4+5</subScript>
, without basal appendix. CuA reaching wing edge, CuA and CuP parallel apically, not distinctly convergent or meeting.
<emphasis id="1B8A9001FFA24B5AFD87F9B5FD29204E" box="[575,689,1544,1570]" italics="true" pageId="31" pageNumber="482">Abdomen.</emphasis>
Dark brown with blackish areas. Terminalia (Figs. 36–37, 39, 41) dark brown to black. Tergite nine broader than long, posterior margin with v-shaped emargination on posterior edge that is about one-third to slightly over one-half the length of the tergite; lobes resulting from emargination broadly rounded posteriorly. In ventral view, posterior edge of medially fused gonocoxites + hypandrium with a pair of small, rounded, weakly to moderately projecting, ventromedian lobes, separated by a shallow, broad ventromedian cleft. In dorsal and ventral views, gonostylus elongate, relatively narrow, gently curved, apically minutely truncate or with small notch (Figs. 36–37). In posterior view, gonostylus relatively slender, tapering apically (
<figureCitation id="B1C55096FFA24B5AFAA2F95DFAF72096" box="[1306,1391,1760,1786]" captionStart="FIGURES 38–41" captionStartId="33.[151,264,1888,1912]" captionTargetBox="[168,1417,924,1848]" captionTargetId="figure-337@33.[164,1424,921,1864]" captionTargetPageId="33" captionText="FIGURES 38–41. Male terminalia. 38. P. heteroptera, dorsal, tergite nine removed, topotypic (Maryland, USA). 39. P. yakima n. sp., holotype, dorsal, tergite nine removed. 40. P. heteroptera, posterior, topotypic (Maryland, USA). 41. P. yakima n. sp., holotype, posterior. Scale bar = ca. 0.5 mm.Abbreviations: c, cerci; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529554" httpUri="https://zenodo.org/record/4529554/files/figure.png" pageId="31" pageNumber="482">Fig. 41</figureCitation>
). In dorsal and ventral views fused parameres (dorsal sclerite) typically projecting slightly beyond genital capsule, shiny, brown, heavily-sclerotized, dome-like, roughly diamond-shaped (broadest medially), apically coming to a broadly rounded point, without divergent horn-like lobes apically (
<figureCitation id="B1C55096FFA24B5AFC9EF8F1FCE2210A" box="[806,890,1868,1894]" captionStart="FIGURES 38–41" captionStartId="33.[151,264,1888,1912]" captionTargetBox="[168,1417,924,1848]" captionTargetId="figure-337@33.[164,1424,921,1864]" captionTargetPageId="33" captionText="FIGURES 38–41. Male terminalia. 38. P. heteroptera, dorsal, tergite nine removed, topotypic (Maryland, USA). 39. P. yakima n. sp., holotype, dorsal, tergite nine removed. 40. P. heteroptera, posterior, topotypic (Maryland, USA). 41. P. yakima n. sp., holotype, posterior. Scale bar = ca. 0.5 mm.Abbreviations: c, cerci; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529554" httpUri="https://zenodo.org/record/4529554/files/figure.png" pageId="31" pageNumber="482">Fig. 39</figureCitation>
). Cerci fleshy, apically rounded with setae.
</paragraph>
<paragraph id="29414C13FFA24B64FF7FF8CDFC8D2695" blockId="31.[151,1437,752,2039]" lastBlockId="33.[151,1437,151,897]" lastPageId="33" lastPageNumber="484" pageId="31" pageNumber="482">
<emphasis id="1B8A9001FFA24B5AFF7FF8CDFEBB21E6" box="[199,291,1904,1930]" italics="true" pageId="31" pageNumber="482">Female.</emphasis>
Similar to male. Body about 5.0 mm and wing 6.0 mm (n=1). Eyes dichoptic. Antennae with 7–8 flagellomeres (apically flagellomeres somewhat fused and difficult to enumerate). Female hind basitarsus slender elongate. Abdomen stout. Terminalia (
<figureCitation id="B1C55096FFA24B5AFDACF805FDF121BF" box="[532,617,1976,2003]" captionStart="FIGURES 54–57" captionStartId="37.[151,264,1708,1732]" captionTargetBox="[168,1421,209,1673]" captionTargetId="figure-16@37.[164,1424,181,1684]" captionTargetPageId="37" captionText="FIGURES 54–57. Female terminalia and eggs. 54–56. Female terminalia. 54. P. neonigrita n. sp., dorsal. 55. P. neonigrita n. sp., ventral. 56. P. yakima n. sp., ventral. 57. P. heteroptera, eggs (dissected from abdomen). Scale bars = ca. 0.25 mm. Abbreviations: c1, cercus 1; c2, cercus 2; s7, sternite seven; s8, sternite eight; s10, sternite ten; sp, spermatheca; t7, tergite seven; t8, tergite eight; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529566" httpUri="https://zenodo.org/record/4529566/files/figure.png" pageId="31" pageNumber="482">Fig. 56</figureCitation>
): Tergite nine present dorsally as a very narrow transverse strap medially subdivided; tergite more produced ventrolaterally. Tergite ten minute, longitudinally elongate. Cerci two-segmented, ultimate segment relatively short. Subgenital plate (= sternite eight) large, longitudinally subdivided, posterior margin with a pair of broad lobes medially.Y-shaped genital fork present. Sternite 10 broadly rounded posteriorly. Three rounded, sclerotized, capsule-like, spermathecae present.
</paragraph>
</subSubSection>
<caption id="7D811C9BFF9D4B65FF2FF820FB7021B5" ID-DOI="http://doi.org/10.5281/zenodo.4529552" ID-Zenodo-Dep="4529552" httpUri="https://zenodo.org/record/4529552/files/figure.png" pageId="32" pageNumber="483" startId="32.[151,264,1949,1973]" targetBox="[334,1258,216,1918]" targetPageId="32">
<paragraph id="29414C13FF9D4B65FF2FF820FB7021B5" blockId="32.[151,1437,1949,2009]" pageId="32" pageNumber="483">
<emphasis id="1B8A9001FF9D4B65FF2FF820FE9021D9" bold="true" box="[151,264,1949,1973]" pageId="32" pageNumber="483">FIGURES</emphasis>
3
<emphasis id="1B8A9001FF9D4B65FEA3F820FECD21D9" bold="true" box="[283,341,1949,1973]" pageId="32" pageNumber="483">6–37.</emphasis>
<taxonomicName id="EEFE3790FF9D4B65FEE3F823FE2521D9" authority="Fitzgerald, 2021" authorityName="Fitzgerald" authorityYear="2021" box="[347,445,1949,1973]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="483" phylum="Arthropoda" rank="species" species="yakima" status="sp. nov.">
<emphasis id="1B8A9001FF9D4B65FEE3F823FE2521D9" box="[347,445,1949,1973]" italics="true" pageId="32" pageNumber="483">P. yakima</emphasis>
</taxonomicName>
<emphasis id="1B8A9001FF9D4B65FE7CF820FE6621D9" bold="true" box="[452,510,1949,1973]" pageId="32" pageNumber="483">
<taxonomicNameLabel id="00B92D7AFF9D4B65FE7CF820FE6621D9" box="[452,510,1949,1973]" pageId="32" pageNumber="483" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male terminalia. 36. Dorsal. 37. Ventral. Scale bar = ca. 0.5 mm. Abbreviations: c, cerci; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine; vml, ventromedian lobes of gonocoxite.
</paragraph>
</caption>
<subSubSection id="61E41F98FF9C4B64FF7FFEBFFD6227C1" pageId="33" pageNumber="484" type="diagnosis">
<paragraph id="29414C13FF9C4B64FF7FFEBFFD6227C1" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="484">
<emphasis id="1B8A9001FF9C4B64FF7FFEBFFEA22770" bold="true" box="[199,314,258,284]" pageId="33" pageNumber="484">Diagnosis</emphasis>
. Males of
<taxonomicName id="EEFE3790FF9C4B64FE0AFEBEFD872771" box="[434,543,259,285]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FE0AFEBEFD872771" box="[434,543,259,285]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
can be distinguished from all New World congeners by the following combination of characters: gonostylus relatively slender, apically truncate and sometimes notched (Figs. 36–37, 39, 41), paramere diamond-shaped (
<figureCitation id="B1C55096FF9C4B64FE68FEF6FDBC2709" box="[464,548,331,357]" captionStart="FIGURES 38–41" captionStartId="33.[151,264,1888,1912]" captionTargetBox="[168,1417,924,1848]" captionTargetId="figure-337@33.[164,1424,921,1864]" captionTargetPageId="33" captionText="FIGURES 38–41. Male terminalia. 38. P. heteroptera, dorsal, tergite nine removed, topotypic (Maryland, USA). 39. P. yakima n. sp., holotype, dorsal, tergite nine removed. 40. P. heteroptera, posterior, topotypic (Maryland, USA). 41. P. yakima n. sp., holotype, posterior. Scale bar = ca. 0.5 mm.Abbreviations: c, cerci; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529554" httpUri="https://zenodo.org/record/4529554/files/figure.png" pageId="33" pageNumber="484">Fig. 39</figureCitation>
), tergite nine posteromedially cleft about one-third to slightly over one-half its length, without shelf-like development or a field of densely-set, short, black, spine-like setae, and ventromedian lobes of gonocoxites weakly to moderately developed.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FF9C4B64FF7FFE0AFC80240D" pageId="33" pageNumber="484" type="discussion">
<paragraph id="29414C13FF9C4B64FF7FFE0AFC242799" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="484">
<emphasis id="1B8A9001FF9C4B64FF7FFE0AFEAC27BD" bold="true" box="[199,308,439,465]" pageId="33" pageNumber="484">Remarks</emphasis>
. Males of
<taxonomicName id="EEFE3790FF9C4B64FE0CFE0AFDBC27BD" box="[436,548,439,465]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FE0CFE0AFDBC27BD" box="[436,548,439,465]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
are most similar to
<taxonomicName id="EEFE3790FF9C4B64FCA9FE0AFC2A27BD" box="[785,946,439,465]" class="Insecta" family="Phoridae" genus="Penthetira" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="heteroptera">
<emphasis id="1B8A9001FF9C4B64FCA9FE0AFC2A27BD" box="[785,946,439,465]" italics="true" pageId="33" pageNumber="484">P. heteroptera</emphasis>
</taxonomicName>
, but can be differentiated by the narrower gonostylus and more diamond-shaped paramere (compare
<figureCitation id="B1C55096FF9C4B64FC9FFE66FC282799" box="[807,944,475,501]" captionStart="FIGURES 38–41" captionStartId="33.[151,264,1888,1912]" captionTargetBox="[168,1417,924,1848]" captionTargetId="figure-337@33.[164,1424,921,1864]" captionTargetPageId="33" captionText="FIGURES 38–41. Male terminalia. 38. P. heteroptera, dorsal, tergite nine removed, topotypic (Maryland, USA). 39. P. yakima n. sp., holotype, dorsal, tergite nine removed. 40. P. heteroptera, posterior, topotypic (Maryland, USA). 41. P. yakima n. sp., holotype, posterior. Scale bar = ca. 0.5 mm.Abbreviations: c, cerci; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gc, gonocoxite; gs, gonostylus; p, paramere; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529554" httpUri="https://zenodo.org/record/4529554/files/figure.png" pageId="33" pageNumber="484">Figs. 38–41</figureCitation>
).
</paragraph>
<paragraph id="29414C13FF9C4B64FF7FFE42FC80240D" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="484">
Females of
<taxonomicName id="EEFE3790FF9C4B64FEF4FE42FE212475" box="[332,441,511,537]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FEF4FE42FE212475" box="[332,441,511,537]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
(
<figureCitation id="B1C55096FF9C4B64FE72FE42FD842475" box="[458,540,510,537]" captionStart="FIGURES 54–57" captionStartId="37.[151,264,1708,1732]" captionTargetBox="[168,1421,209,1673]" captionTargetId="figure-16@37.[164,1424,181,1684]" captionTargetPageId="37" captionText="FIGURES 54–57. Female terminalia and eggs. 54–56. Female terminalia. 54. P. neonigrita n. sp., dorsal. 55. P. neonigrita n. sp., ventral. 56. P. yakima n. sp., ventral. 57. P. heteroptera, eggs (dissected from abdomen). Scale bars = ca. 0.25 mm. Abbreviations: c1, cercus 1; c2, cercus 2; s7, sternite seven; s8, sternite eight; s10, sternite ten; sp, spermatheca; t7, tergite seven; t8, tergite eight; t9, tergite nine." figureDoi="http://doi.org/10.5281/zenodo.4529566" httpUri="https://zenodo.org/record/4529566/files/figure.png" pageId="33" pageNumber="484">Fig. 56</figureCitation>
) are indistinguishable from females of
<taxonomicName id="EEFE3790FF9C4B64FC63FE42FBE12475" box="[987,1145,511,537]" class="Insecta" family="Phoridae" genus="Penthetira" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="heteroptera">
<emphasis id="1B8A9001FF9C4B64FC63FE42FBE12475" box="[987,1145,511,537]" italics="true" pageId="33" pageNumber="484">P. heteroptera</emphasis>
</taxonomicName>
and
<taxonomicName id="EEFE3790FF9C4B64FB09FE42FAD62474" authorityName="Fitzgerald" authorityYear="2021" box="[1201,1358,511,536]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="arizonensis">
<emphasis id="1B8A9001FF9C4B64FB09FE42FAD62474" box="[1201,1358,511,536]" italics="true" pageId="33" pageNumber="484">P. arizonensis</emphasis>
</taxonomicName>
. However, unassociated females from the northwestern
<collectingCountry id="51E90C83FF9C4B64FD7FFD9EFD652451" box="[711,765,547,573]" name="United States of America" pageId="33" pageNumber="484">USA</collectingCountry>
(west of the Rocky Mountains) likely belong to this species as it is the only species currently recorded from this area.
</paragraph>
</subSubSection>
<subSubSection id="61E41F98FF9C4B64FF7FFDD6FD1625ED" pageId="33" pageNumber="484" type="etymology">
<paragraph id="29414C13FF9C4B64FF7FFDD6FA1E24E9" blockId="33.[151,1437,151,897]" box="[199,1414,619,645]" pageId="33" pageNumber="484">
<emphasis id="1B8A9001FF9C4B64FF7FFDD6FED024E9" bold="true" box="[199,328,619,645]" pageId="33" pageNumber="484">Etymology</emphasis>
. The specific epithet
<taxonomicName id="EEFE3790FF9C4B64FD82FDD6FD1324E9" box="[570,651,619,645]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FD82FDD6FD1324E9" box="[570,651,619,645]" italics="true" pageId="33" pageNumber="484">yakima</emphasis>
</taxonomicName>
is based on the name of the
<typeStatus id="F645F2B1FF9C4B64FC71FDD6FC6124E9" box="[969,1017,619,645]" pageId="33" pageNumber="484">type</typeStatus>
locality “N. Yakima,”
<collectingRegion id="EB3A82F1FF9C4B64FB41FDD6FA1A24E9" box="[1273,1410,619,645]" country="United States of America" name="Washington" pageId="33" pageNumber="484">Washington</collectingRegion>
.
</paragraph>
<paragraph id="29414C13FF9C4B64FF7FFD33FD1625ED" blockId="33.[151,1437,151,897]" pageId="33" pageNumber="484">
<emphasis id="1B8A9001FF9C4B64FF7FFD33FDE424C4" bold="true" box="[199,636,654,680]" pageId="33" pageNumber="484">Geographic &amp; Seasonal Distribution</emphasis>
. Northwestern
<collectingCountry id="51E90C83FF9C4B64FC93FD32FCF924C5" box="[811,865,655,681]" name="United States of America" pageId="33" pageNumber="484">USA</collectingCountry>
(
<collectingRegion id="EB3A82F1FF9C4B64FCC9FD32FC6024C5" box="[881,1016,655,681]" country="United States of America" name="Washington" pageId="33" pageNumber="484">Washington</collectingRegion>
&amp; southwestern
<collectingRegion id="EB3A82F1FF9C4B64FB00FD32FAB024C6" box="[1208,1320,655,682]" country="United States of America" name="Wyoming" pageId="33" pageNumber="484">Wyoming</collectingRegion>
) and
<collectingRegion id="EB3A82F1FF9C4B64FADFFD32FEB424A1" country="Canada" name="British Columbia" pageId="33" pageNumber="484">British Columbia</collectingRegion>
east to
<collectingRegion id="EB3A82F1FF9C4B64FE3AFD0EFE4024A1" box="[386,472,691,717]" country="Canada" name="Ontario" pageId="33" pageNumber="484">Ontario</collectingRegion>
in
<collectingCountry id="51E90C83FF9C4B64FE43FD0EFDD724A1" box="[507,591,691,717]" name="Canada" pageId="33" pageNumber="484">Canada</collectingCountry>
(
<figureCitation id="B1C55096FF9C4B64FDE5FD0EFD3124A1" box="[605,681,691,717]" captionStart="MAP 3" captionStartId="40.[151,209,1161,1185]" captionTargetBox="[170,1420,191,1128]" captionTargetId="figure-34@40.[160,1427,181,1137]" captionText="MAP 3. Distribution of P. heteroptera (black circle) and P. yakima (star)." figureDoi="http://doi.org/10.5281/zenodo.4546088" httpUri="https://zenodo.org/record/4546088/files/figure.png" pageId="33" pageNumber="484">Map 3</figureCitation>
). Based on current distribution records,
<taxonomicName id="EEFE3790FF9C4B64FBD1FD0EFB4D24A1" box="[1129,1237,691,717]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FBD1FD0EFB4D24A1" box="[1129,1237,691,717]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
and the similar
<taxonomicName id="EEFE3790FF9C4B64FA30FD0EFE81249D" class="Insecta" family="Phoridae" genus="Penthetira" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="heteroptera">
<emphasis id="1B8A9001FF9C4B64FA30FD0EFE81249D" italics="true" pageId="33" pageNumber="484">P. heteroptera</emphasis>
</taxonomicName>
are allopatric in the
<collectingCountry id="51E90C83FF9C4B64FDB9FD6AFDAF249D" box="[513,567,727,753]" name="United States of America" pageId="33" pageNumber="484">USA</collectingCountry>
and sympatric across most of
<collectingCountry id="51E90C83FF9C4B64FC36FD6AFC7E249D" box="[910,998,727,753]" name="Canada" pageId="33" pageNumber="484">Canada</collectingCountry>
. The
<taxonomicName id="EEFE3790FF9C4B64FB9AFD6AFB17249D" box="[1058,1167,727,753]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FB9AFD6AFB17249D" box="[1058,1167,727,753]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
specimens known from western
<collectingRegion id="EB3A82F1FF9C4B64FF40FD46FEF12579" box="[248,361,763,789]" country="United States of America" name="Wyoming" pageId="33" pageNumber="484">Wyoming</collectingRegion>
,
<collectingCountry id="51E90C83FF9C4B64FECFFD46FE352579" box="[375,429,763,789]" name="United States of America" pageId="33" pageNumber="484">USA</collectingCountry>
occur at the southwestern edge of the
<collectingRegion id="EB3A82F1FF9C4B64FCCAFD46FC782579" box="[882,992,763,789]" country="United States of America" name="Wyoming" pageId="33" pageNumber="484">Wyoming</collectingRegion>
Basin which is essentially high desert habitat along the Bear River in the low areas between the
<collectingRegion id="EB3A82F1FF9C4B64FC93FCA2FC012555" box="[811,921,799,825]" country="United States of America" name="Wyoming" pageId="33" pageNumber="484">Wyoming</collectingRegion>
Range to the North and the Uinta Mountains of
<collectingRegion id="EB3A82F1FF9C4B64FF0DFCFEFF732531" box="[181,235,835,861]" country="United States of America" name="Utah" pageId="33" pageNumber="484">Utah</collectingRegion>
to the South; this is the farthest eastern and southern record for this species in the
<collectingCountry id="51E90C83FF9C4B64FB3BFCFEFB272531" box="[1155,1215,835,861]" name="United States of America" pageId="33" pageNumber="484">USA</collectingCountry>
.
<taxonomicName id="EEFE3790FF9C4B64FB70FCFEFAAD2531" box="[1224,1333,835,861]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="yakima">
<emphasis id="1B8A9001FF9C4B64FB70FCFEFAAD2531" box="[1224,1333,835,861]" italics="true" pageId="33" pageNumber="484">P. yakima</emphasis>
</taxonomicName>
has been recorded in August and September (
<tableCitation id="647C79A8FF9C4B64FD91FCDAFDE525ED" box="[553,637,871,897]" captionStart="TABLE 1" captionStartId="34.[151,239,151,175]" captionText="TABLE 1. Summary of adult flight period (grey boxes indicate specimens collected during these months); Malaise trap data including more than one month have been excluded." pageId="33" pageNumber="484">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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